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<document id="C16D3868A9DA8E7E643872360B2FFD49" ID="10.11646/zootaxa.4193.3.6" ID-DOI="10.11646/zootaxa.4193.3.6" ID-GBIF-Dataset="3425fed3-d6a1-4b69-a730-4293fb3f073f" ID-ISSN="1175-5326" ID-Zenodo-Dep="167241" ID-ZooBank="3F565C0C-A00E-4B4C-B59A-1FA5144DBB29" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1479369947288" checkinUser="plazi" docAuthor="Colmenares, Pío A. &amp; Tourinho, Ana Lúcia" docDate="2016" docId="07009602FFC1BB3BF18AF9B1FD6CE520" docLanguage="en" docName="zootaxa.4193.3.6.pdf" docOrigin="Zootaxa 4193 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Maracaynatum isadorae Colmenares &amp; Tourinho, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="569" masterDocId="FB39EE7AFFC0BB3EF11DFF8DFFE7E43D" masterDocTitle="First Amazonian species of Maracaynatum, with comments on the genus (Opiliones: Laniatores: Samoidae)" masterLastPageNumber="572" masterPageNumber="565" pageNumber="566" updateTime="1698393891430" updateUser="plazi">
<mods:mods id="4CC9357286EAE55F0088C8B175318564" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0EBDF2413816EEB2502107031A4FDA8C">
<mods:title id="FFFC21C0013B084037E6E6F106E42540">First Amazonian species of Maracaynatum, with comments on the genus (Opiliones: Laniatores: Samoidae)</mods:title>
</mods:titleInfo>
<mods:name id="E5A89F4CB9E989893C71991F8D7510F9" type="personal">
<mods:role id="046D9A06725CBBF4B760B00BD6E5D92C">
<mods:roleTerm id="93221F178C337D894D5B3DE3AA1A58AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB8C58B34B034271B0D2A83265AA91D4">Colmenares, Pío A.</mods:namePart>
</mods:name>
<mods:name id="0CDCE42E97FAAFC266E544CE117FCFF0" type="personal">
<mods:role id="80F9F58013331854D4FB2B8581A23528">
<mods:roleTerm id="1AC99AF3946DBAAC56F9C867BE8941A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="90EF57ECEC42BD2D7F17A2B94AA3A381">Tourinho, Ana Lúcia</mods:namePart>
</mods:name>
<mods:typeOfResource id="2DBA79FD0E1D8C27E9CC7CE46804694B">text</mods:typeOfResource>
<mods:relatedItem id="EA48FFBACA826B71415069FA345182A1" type="host">
<mods:titleInfo id="5007A4F9E864C39E259DB5934C6C555F">
<mods:title id="091BAC76BBA8F10FAF068320AC750F99">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DD584B0F4E0621512ABDDA08AC1848FC">
<mods:date id="16DF2CFBCEE719AC81E36C174A4B7897">2016</mods:date>
<mods:detail id="EE7469CB460197BE74F7C41576E6BE07" type="volume">
<mods:number id="AD31600073B85B8276CBB29CABC67F92">4193</mods:number>
</mods:detail>
<mods:detail id="A62D5D8E6195D0347B45BF79F25768AB" type="issue">
<mods:number id="0827AEA5645CB7DE7F9C1E2E65C266DF">3</mods:number>
</mods:detail>
<mods:extent id="CC61A87C387F3387EADA24615D195D40" unit="page">
<mods:start id="B5358A0B7110004B26C48951AC3068AA">565</mods:start>
<mods:end id="640798F2A863C7E3FA83C5A2D510B995">572</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0BABB4E9C0F7F15641A7C309972F3A4A">journal article</mods:classification>
<mods:identifier id="2958B8820C7474D19359ABBD04CB729B" type="DOI">10.11646/zootaxa.4193.3.6</mods:identifier>
<mods:identifier id="FBC562255DB5D4024EDF11A32A245C89" type="GBIF-Dataset">3425fed3-d6a1-4b69-a730-4293fb3f073f</mods:identifier>
<mods:identifier id="25072C6EE2467E504BC5D5A75D6F2776" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="014D71781EBF96D81D492FBA81B839C0" type="Zenodo-Dep">167241</mods:identifier>
<mods:identifier id="C19170AB90DD481A71DFF853E2161AF3" type="ZooBank">3F565C0C-A00E-4B4C-B59A-1FA5144DBB29</mods:identifier>
</mods:mods>
<treatment id="07009602FFC1BB3BF18AF9B1FD6CE520" ID-DOI="http://doi.org/10.5281/zenodo.6063892" ID-GBIF-Taxon="124770314" ID-Zenodo-Dep="6063892" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520" httpUri="http://treatment.plazi.org/id/07009602FFC1BB3BF18AF9B1FD6CE520" lastPageId="5" lastPageNumber="569" pageId="1" pageNumber="566">
<subSubSection id="C7B3749FFFC1BB3FF18AF9B1FEE7E24A" pageId="1" pageNumber="566" type="nomenclature">
<paragraph id="8F162714FFC1BB3FF18AF9B1FDC4E26B" blockId="1.[151,547,1596,1655]" box="[151,547,1596,1622]" pageId="1" pageNumber="566">
<heading id="D45E9078FFC1BB3FF18AF9B1FDC4E26B" bold="true" box="[151,547,1596,1622]" fontSize="11" level="1" pageId="1" pageNumber="566" reason="1">
<emphasis id="BDDDFB06FFC1BB3FF18AF9B1FDC4E26B" bold="true" box="[151,547,1596,1622]" pageId="1" pageNumber="566">
<taxonomicName id="48A95C97FFC1BB3FF18AF9B1FE59E26B" ID-CoL="7X55M" box="[151,446,1596,1622]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="566" phylum="Arthropoda" rank="species" species="isadorae" status="sp. nov.">
<emphasis id="BDDDFB06FFC1BB3FF18AF9B1FE59E26B" bold="true" box="[151,446,1596,1622]" italics="true" pageId="1" pageNumber="566">Maracaynatum isadorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE467DFFC1BB3FF0D8F9B1FDC4E26B" box="[453,547,1596,1622]" pageId="1" pageNumber="566" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8F162714FFC1BB3FF18AF9D3FEE7E24A" blockId="1.[151,547,1596,1655]" box="[151,256,1630,1655]" pageId="1" pageNumber="566">
<figureCitation id="17923B91FFC1BB3FF18AF9D3FEE7E24A" box="[151,256,1630,1655]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1829,1851]" captionStartId-1="3.[151,250,1385,1407]" captionStartId-2="5.[151,250,1246,1268]" captionTargetBox-0="[163,1423,563,1793]" captionTargetBox-1="[228,1354,191,1346]" captionTargetBox-2="[163,1427,343,1218]" captionTargetId-0="figure@2.[151,1436,553,1807]" captionTargetId-1="figure@3.[220,1366,179,1364]" captionTargetId-2="figure@5.[151,1436,339,1224]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionText-0="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." captionText-1="FIGURE 2. Maracaynatum isadorae sp. nov. Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm." captionText-2="FIGURE 3. Distribution of Maracaynatum spp." httpUri-0="https://zenodo.org/record/167242/files/figure.png" httpUri-1="https://zenodo.org/record/261023/files/figure.png" httpUri-2="https://zenodo.org/record/261024/files/figure.png" pageId="1" pageNumber="566">Figs. 1–3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C7B3749FFFC1BB3CF18AF92BFCF5E5C9" lastPageId="2" lastPageNumber="567" pageId="1" pageNumber="566" type="materials_examined">
<paragraph id="8F162714FFC1BB3CF18AF92BFCF5E5C9" blockId="1.[151,1436,1702,2015]" lastBlockId="2.[151,1437,151,501]" lastPageId="2" lastPageNumber="567" pageId="1" pageNumber="566">
<materialsCitation id="3FC12D49FFC1BB3FF18AF92BFD79E2DE" ID-GBIF-Occurrence="1326058834" collectionCode="UFAM" country="Brazil" location="Manaus" pageId="1" pageNumber="566" specimenCount="2" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="BDDDFB06FFC1BB3FF18AF92BFEAEE282" bold="true" box="[151,329,1702,1727]" pageId="1" pageNumber="566">
<typeStatus id="501299B6FFC1BB3FF18AF92BFF35E282" box="[151,210,1702,1727]" pageId="1" pageNumber="566">Type</typeStatus>
material.
</emphasis>
<typeStatus id="501299B6FFC1BB3FF048F92BFE5BE283" box="[341,444,1702,1726]" pageId="1" pageNumber="566" type="holotype">
<emphasis id="BDDDFB06FFC1BB3FF048F92BFE5BE283" box="[341,444,1702,1726]" italics="true" pageId="1" pageNumber="566">Holotype</emphasis>
</typeStatus>
male (INPA-OPI 003417) from the Fazenda da Universidade Federal do
<collectingRegion id="4D6DE9F6FFC1BB3FF43BF92AFA7BE282" box="[1318,1436,1703,1727]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="566">Amazonas</collectingRegion>
(Fazenda
<collectionCode id="E9B8BFD1FFC1BB3FF018F946FEBEE2DE" box="[261,345,1739,1763]" pageId="1" pageNumber="566">UFAM</collectionCode>
),
<location id="8A7671CFFFC1BB3FF076F946FE20E2DF" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC1BB3FF076F946FE20E2DF" box="[363,455,1739,1762]" country="Brazil" name="Manaus" pageId="1" pageNumber="566" stateProvince="Amazonas">Manaus</location>
,
<collectingRegion id="4D6DE9F6FFC1BB3FF0CCF946FDADE2DE" box="[465,586,1739,1763]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="566">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC1BB3FF34BF947FD79E2DE" box="[598,670,1738,1763]" name="Brazil" pageId="1" pageNumber="566">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC1BB3FF3B4F947FB8CE33A" ID-GBIF-Occurrence="1326058833" collectingDate="2011-07-28" collectingDateMax="2011-07-30" collectingDateMin="2011-07-28" collectionCode="UFAM" collectorName="Leg. Larissa Lanca &amp; Winkler" country="Brazil" location="Fazenda" municipality="Manaus" pageId="1" pageNumber="566" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC1BB3FF3B4F947FCB2E2DE" box="[681,853,1738,1763]" pageId="1" pageNumber="566" value="2011-07-28" valueMax="2011-07-30" valueMin="2011-07-28">
<date id="FB1701D4FFC1BB3FF3B4F947FCB2E2DE" box="[681,853,1738,1763]" pageId="1" pageNumber="566" value="2011-07-28" valueMax="2011-07-30" valueMin="2011-07-28">28–30.vii.2011</date>
</collectingDate>
;
<collectorName id="225C42C2FFC1BB3FF27DF946FBDEE2DF" box="[864,1081,1738,1763]" pageId="1" pageNumber="566">Leg. Larissa Lança</collectorName>
;
<collectorName id="225C42C2FFC1BB3FF558F947FB47E2DE" box="[1093,1184,1738,1763]" pageId="1" pageNumber="566">Winkler</collectorName>
technique.
<emphasis id="BDDDFB06FFC1BB3FF43CF946FA7BE2DF" box="[1313,1436,1739,1762]" italics="true" pageId="1" pageNumber="566">
<typeStatus id="501299B6FFC1BB3FF43CF946FA71E2DF" box="[1313,1430,1739,1762]" pageId="1" pageNumber="566" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="99AFEC9DFFC1BB3FF18AF962FF01E33A" box="[151,230,1774,1799]" pageId="1" pageNumber="566" type="male">1 male</specimenCount>
(INPA-OP 1008) from
<location id="8A7671CFFFC1BB3FF31CF963FDB9E33A" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC1BB3FF31CF963FDB9E33A" box="[513,606,1774,1799]" country="Brazil" municipality="Manaus" name="Fazenda" pageId="1" pageNumber="566" stateProvince="Amazonas">Fazenda</location>
<collectionCode id="E9B8BFD1FFC1BB3FF377F962FD5BE33A" box="[618,700,1775,1799]" pageId="1" pageNumber="566">UFAM</collectionCode>
, L2-500,
<collectingMunicipality id="6F72BD6EFFC1BB3FF22DF962FC6BE33B" box="[816,908,1775,1798]" pageId="1" pageNumber="566">Manaus</collectingMunicipality>
,
<collectingRegion id="4D6DE9F6FFC1BB3FF287F962FBF4E33A" box="[922,1043,1775,1799]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="566">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC1BB3FF53EF963FB8CE33A" box="[1059,1131,1774,1799]" name="Brazil" pageId="1" pageNumber="566">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC1BB3FF566F963FA7FE316" ID-GBIF-Occurrence="1326058840" collectingDate="2010-07-28" collectionCode="UFAM" collectorName="Leg. Larissa Lanca &amp; Beating" country="Brazil" location="Fazenda" municipality="Manaus" pageId="1" pageNumber="566" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC1BB3FF566F963FB1AE33A" box="[1147,1277,1774,1799]" pageId="1" pageNumber="566" value="2010-07-28">
<date id="FB1701D4FFC1BB3FF566F963FB1AE33A" box="[1147,1277,1774,1799]" pageId="1" pageNumber="566" value="2010-07-28">28.vii.2010</date>
</collectingDate>
;
<collectorName id="225C42C2FFC1BB3FF410F962FF38E317" pageId="1" pageNumber="566">Leg. Larissa Lança</collectorName>
;
<collectorName id="225C42C2FFC1BB3FF1F1F89FFEA4E316" box="[236,323,1810,1835]" pageId="1" pageNumber="566">Beating</collectorName>
technique.
<specimenCount id="99AFEC9DFFC1BB3FF0D5F89EFDCCE316" box="[456,555,1810,1835]" pageId="1" pageNumber="566" type="female">1 female</specimenCount>
(INPA-OP 1557) from
<location id="8A7671CFFFC1BB3FF226F89FFC7FE316" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC1BB3FF226F89FFC7FE316" box="[827,920,1810,1835]" country="Brazil" municipality="Manaus" name="Fazenda" pageId="1" pageNumber="566" stateProvince="Amazonas">Fazenda</location>
<collectionCode id="E9B8BFD1FFC1BB3FF2BCF89EFC14E316" box="[929,1011,1811,1835]" pageId="1" pageNumber="566">UFAM</collectionCode>
, L3-500,
<collectingMunicipality id="6F72BD6EFFC1BB3FF57CF89EFB5AE317" box="[1121,1213,1811,1834]" pageId="1" pageNumber="566">Manaus</collectingMunicipality>
,
<collectingRegion id="4D6DE9F6FFC1BB3FF5D7F89EFAA4E316" box="[1226,1347,1811,1835]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="566">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC1BB3FF44DF89FFA7FE316" box="[1360,1432,1810,1835]" name="Brazil" pageId="1" pageNumber="566">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC1BB3FF18AF8BBFD77E3AA" ID-GBIF-Occurrence="1326058832" collectingDate="2010-08-23" collectionCode="UFAM" collectorName="Leg. O. &amp; Beating" country="Brazil" location="Fazenda" municipality="Manaus" pageId="1" pageNumber="566" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC1BB3FF18AF8BBFEC6E372" box="[151,289,1846,1871]" pageId="1" pageNumber="566" value="2010-08-23">
<date id="FB1701D4FFC1BB3FF18AF8BBFEC6E372" box="[151,289,1846,1871]" pageId="1" pageNumber="566" value="2010-08-23">23.viii.2010</date>
</collectingDate>
;
<collectorName id="225C42C2FFC1BB3FF032F8BAFE61E372" box="[303,390,1847,1871]" pageId="1" pageNumber="566">Leg. O.</collectorName>
Ferreira da Silva;
<collectorName id="225C42C2FFC1BB3FF343F8BBFD52E372" box="[606,693,1846,1871]" pageId="1" pageNumber="566">Beating</collectorName>
technique.
<specimenCount id="99AFEC9DFFC1BB3FF226F8BAFC74E372" box="[827,915,1846,1871]" pageId="1" pageNumber="566" type="male">2 males</specimenCount>
and
<specimenCount id="99AFEC9DFFC1BB3FF2D3F8BAFBD6E372" box="[974,1073,1846,1871]" pageId="1" pageNumber="566" type="female">1 female</specimenCount>
(INPA-OPI 003491) with same data as holotype (one male with dissected genitalia and appendages).
<specimenCount id="99AFEC9DFFC1BB3FF2DEF8D6FBCFE34E" box="[963,1064,1882,1907]" pageId="1" pageNumber="566" type="female">1 female</specimenCount>
(INPA-OP 1009) from
<location id="8A7671CFFFC1BB3FF422F8D7FA7BE34E" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC1BB3FF422F8D7FA7BE34E" box="[1343,1436,1882,1907]" country="Brazil" municipality="Manaus" name="Fazenda" pageId="1" pageNumber="566" stateProvince="Amazonas">Fazenda</location>
<collectionCode id="E9B8BFD1FFC1BB3FF18AF8F2FF0EE3AA" box="[151,233,1919,1943]" pageId="1" pageNumber="566">UFAM</collectionCode>
, L3–500,
<collectingMunicipality id="6F72BD6EFFC1BB3FF046F8F2FE50E3AB" box="[347,439,1919,1942]" pageId="1" pageNumber="566">Manaus</collectingMunicipality>
,
<collectingRegion id="4D6DE9F6FFC1BB3FF0DEF8F2FDDBE3AA" box="[451,572,1919,1943]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="566">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC1BB3FF355F8F3FD77E3AA" box="[584,656,1918,1943]" name="Brazil" pageId="1" pageNumber="566">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC1BB3FF381F8F3FC38E386" ID-GBIF-Occurrence="1326058842" collectingDate="2010-08-23" collectorName="Leg. Larissa Lanca &amp; Beating &amp; Reserva Ducke, LO" country="Brazil" location="Manaus" pageId="1" pageNumber="566" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC1BB3FF381F8F3FCC0E3AA" box="[668,807,1918,1943]" pageId="1" pageNumber="566" value="2010-08-23">
<date id="FB1701D4FFC1BB3FF381F8F3FCC0E3AA" box="[668,807,1918,1943]" pageId="1" pageNumber="566" value="2010-08-23">23.viii.2010</date>
</collectingDate>
;
<collectorName id="225C42C2FFC1BB3FF22EF8F2FBE8E3AB" box="[819,1039,1918,1943]" pageId="1" pageNumber="566">Leg. Larissa Lança</collectorName>
;
<collectorName id="225C42C2FFC1BB3FF506F8F3FB95E3AA" box="[1051,1138,1918,1943]" pageId="1" pageNumber="566">Beating</collectorName>
technique.
<specimenCount id="99AFEC9DFFC1BB3FF5EBF8F2FAA5E3AA" box="[1270,1346,1918,1943]" pageId="1" pageNumber="566" type="male">1 male</specimenCount>
(INPA- OP 2551) from
<collectorName id="225C42C2FFC1BB3FF04AF82EFDA6E386" box="[343,577,1954,1979]" pageId="1" pageNumber="566">Reserva Ducke, LO</collectorName>
5-2500,
<location id="8A7671CFFFC1BB3FF3BFF82EFD19E387" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC1BB3FF3BFF82EFD19E387" box="[674,766,1955,1978]" country="Brazil" name="Manaus" pageId="1" pageNumber="566" stateProvince="Amazonas">Manaus</location>
,
<collectingRegion id="4D6DE9F6FFC1BB3FF213F82EFC60E386" box="[782,903,1955,1979]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="566">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC1BB3FF28AF82FFC38E386" box="[919,991,1954,1979]" name="Brazil" pageId="1" pageNumber="566">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC1BB3FF2F2F82FFA7FE3E2" ID-GBIF-Occurrence="1326058837" collectingDate="2012-06-06" collectorName="Leg. O. &amp; Reserva Ducke, LO" country="Brazil" location="Manaus" pageId="1" pageNumber="566" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC1BB3FF2F2F82FFB8CE386" box="[1007,1131,1954,1979]" pageId="1" pageNumber="566" value="2012-06-06">
<date id="FB1701D4FFC1BB3FF2F2F82FFB8CE386" box="[1007,1131,1954,1979]" pageId="1" pageNumber="566" value="2012-06-06">06.vi.2012</date>
</collectingDate>
;
<collectorName id="225C42C2FFC1BB3FF566F82EFB32E386" box="[1147,1237,1955,1979]" pageId="1" pageNumber="566">Leg. O.</collectorName>
Ferreira; cryptic manual searching.
<specimenCount id="99AFEC9DFFC1BB3FF063F84AFE34E3E2" box="[382,467,1990,2015]" pageId="1" pageNumber="566" type="male">1 male</specimenCount>
(INPA-OP 3308) from
<collectorName id="225C42C2FFC1BB3FF218F84AFC1DE3E2" box="[773,1018,1990,2015]" pageId="1" pageNumber="566">Reserva Ducke, LO</collectorName>
3-500,
<location id="8A7671CFFFC1BB3FF54CF84AFB4BE3E3" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC1BB3FF54CF84AFB4BE3E3" box="[1105,1196,1991,2014]" country="Brazil" name="Manaus" pageId="1" pageNumber="566" stateProvince="Amazonas">Manaus</location>
,
<collectingRegion id="4D6DE9F6FFC1BB3FF5DCF84AFADDE3E2" box="[1217,1338,1991,2015]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="566">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC1BB3FF44DF84BFA7FE3E2" box="[1360,1432,1990,2015]" name="Brazil" pageId="1" pageNumber="566">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC2BB3CF18AFF1AFD42E4E8" ID-GBIF-Occurrence="1326058835" collectingDate="2013-12-16" collectorName="Leg. P. &amp; A. Tourinho &amp; Reserva Ducke, LO" country="Brazil" location="Manaus" pageId="2" pageNumber="567" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC2BB3CF18AFF1AFEFEE48D" box="[151,281,151,176]" pageId="2" pageNumber="567" value="2013-12-16">
<date id="FB1701D4FFC2BB3CF18AFF1AFEFEE48D" box="[151,281,151,176]" pageId="2" pageNumber="567" value="2013-12-16">16.xii.2013</date>
</collectingDate>
;
<collectorName id="225C42C2FFC2BB3CF039FF15FE95E492" box="[292,370,152,176]" pageId="2" pageNumber="567">Leg. P.</collectorName>
Colmenares &amp;
<collectorName id="225C42C2FFC2BB3CF302FF15FD4CE48D" box="[543,683,151,176]" pageId="2" pageNumber="567">A. Tourinho</collectorName>
; cryptic manual searching.
<specimenCount id="99AFEC9DFFC2BB3CF2C6FF15FBDBE48D" box="[987,1084,151,176]" pageId="2" pageNumber="567" type="female">1 female</specimenCount>
(INPA-OP 4432) from
<collectorName id="225C42C2FFC2BB3CF45FFF15FEF1E4E8" pageId="2" pageNumber="567">Reserva Ducke, LO</collectorName>
7-1500,
<location id="8A7671CFFFC2BB3CF06CFF30FE2AE4E9" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC2BB3CF06CFF30FE2AE4E9" box="[369,461,189,212]" country="Brazil" name="Manaus" pageId="2" pageNumber="567" stateProvince="Amazonas">Manaus</location>
,
<collectingRegion id="4D6DE9F6FFC2BB3CF0C5FF31FDB6E4E9" box="[472,593,188,212]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="567">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC2BB3CF340FF31FD42E4E8" box="[605,677,188,213]" name="Brazil" pageId="2" pageNumber="567">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC2BB3CF3ADFF31FBF6E4C5" ID-GBIF-Occurrence="1326058836" collectingDate="2014-07-27" collectorName="Leg. P." country="Brazil" location="Igapo-Acu" pageId="2" pageNumber="567" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC2BB3CF3ADFF31FCD4E4E8" box="[688,819,188,213]" pageId="2" pageNumber="567" value="2014-07-27">
<date id="FB1701D4FFC2BB3CF3ADFF31FCD4E4E8" box="[688,819,188,213]" pageId="2" pageNumber="567" value="2014-07-27">27.vii.2014</date>
</collectingDate>
;
<collectorName id="225C42C2FFC2BB3CF222FF30FC6AE4E9" box="[831,909,189,213]" pageId="2" pageNumber="567">Leg. P.</collectorName>
Colmenares; cryptic manual searching.
<specimenCount id="99AFEC9DFFC2BB3CF44DFF31FA7BE4E8" box="[1360,1436,188,213]" pageId="2" pageNumber="567" type="male">1 male</specimenCount>
(INPA-OP 3929) from BR-319 road, km 260,
<location id="8A7671CFFFC2BB3CF3AAFF6DFCD5E4C5" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC2BB3CF3AAFF6DFCD5E4C5" box="[695,818,224,248]" country="Brazil" name="Igapo-Acu" pageId="2" pageNumber="567" stateProvince="Amazonas">Igapó-Açu</location>
,
<collectingRegion id="4D6DE9F6FFC2BB3CF25CFF6DFC5DE4C5" box="[833,954,224,248]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="567">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC2BB3CF2D4FF52FBF6E4C5" box="[969,1041,223,248]" name="Brazil" pageId="2" pageNumber="567">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC2BB3CF53DFF52FE81E57D" ID-GBIF-Occurrence="1326058838" collectingDate="2012-12-10" collectorName="Leg. P." country="Brazil" location="Rio Preto da Eva" pageId="2" pageNumber="567" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC2BB3CF53DFF52FB44E4C5" box="[1056,1187,223,248]" pageId="2" pageNumber="567" value="2012-12-10">
<date id="FB1701D4FFC2BB3CF53DFF52FB44E4C5" box="[1056,1187,223,248]" pageId="2" pageNumber="567" value="2012-12-10">10.xii.2012</date>
</collectingDate>
;
<collectorName id="225C42C2FFC2BB3CF5AFFF6DFAE3E4CA" box="[1202,1284,224,248]" pageId="2" pageNumber="567">Leg. P.</collectorName>
Colmenares; cryptic manual searching; in ethanol 100%.
<specimenCount id="99AFEC9DFFC2BB3CF389FE89FD0BE520" box="[660,748,260,285]" pageId="2" pageNumber="567" type="male">3 males</specimenCount>
and
<specimenCount id="99AFEC9DFFC2BB3CF235FE89FC70E520" box="[808,919,260,285]" pageId="2" pageNumber="567" type="female">3 females</specimenCount>
(INPA-OPI 003421) from
<location id="8A7671CFFFC2BB3CF5D2FE89FA70E521" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC2BB3CF5D2FE89FA70E521" box="[1231,1431,260,285]" country="Brazil" name="Rio Preto da Eva" pageId="2" pageNumber="567" stateProvince="Amazonas">Rio Preto da Eva</location>
,
<collectingRegion id="4D6DE9F6FFC2BB3CF18AFEA5FEF7E57D" box="[151,272,296,320]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="567">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC2BB3CF003FEAAFE81E57D" box="[286,358,295,320]" name="Brazil" pageId="2" pageNumber="567">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC2BB3CF069FEAAFD61E558" ID-GBIF-Occurrence="1326058843" collectingDate="2006-01-13" collectorName="Leg. R. &amp; A. Tourinho &amp; Beating" country="Brazil" location="Rio Preto da Eva" pageId="2" pageNumber="567" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC2BB3CF069FEAAFE06E57D" box="[372,481,295,320]" pageId="2" pageNumber="567" value="2006-01-13">
<date id="FB1701D4FFC2BB3CF069FEAAFE06E57D" box="[372,481,295,320]" pageId="2" pageNumber="567" value="2006-01-13">13.i.2006</date>
</collectingDate>
;
<collectorName id="225C42C2FFC2BB3CF0F2FEA5FDA2E502" box="[495,581,296,320]" pageId="2" pageNumber="567">Leg. R.</collectorName>
Saturnino &amp;
<collectorName id="225C42C2FFC2BB3CF3F9FEA5FC94E57D" box="[740,883,295,320]" pageId="2" pageNumber="567">A. Tourinho</collectorName>
;
<collectorName id="225C42C2FFC2BB3CF29CFEAAFC3FE57D" box="[897,984,295,320]" pageId="2" pageNumber="567">Beating</collectorName>
technique.
<specimenCount id="99AFEC9DFFC2BB3CF542FEA5FB4AE57D" box="[1119,1197,295,320]" pageId="2" pageNumber="567" type="male">1 male</specimenCount>
(INPA-OPI 003420) from
<location id="8A7671CFFFC2BB3CF1C5FEC1FE41E559" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC2BB3CF1C5FEC1FE41E559" box="[216,422,332,357]" country="Brazil" name="Rio Preto da Eva" pageId="2" pageNumber="567" stateProvince="Amazonas">Rio Preto da Eva</location>
,
<collectingRegion id="4D6DE9F6FFC2BB3CF0A8FEC1FDC9E559" box="[437,558,332,356]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="567">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC2BB3CF323FEC1FD61E558" box="[574,646,332,357]" name="Brazil" pageId="2" pageNumber="567">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC2BB3CF388FEC1FCDCE5B5" ID-GBIF-Occurrence="1326058839" collectingDate="2006-01-13" collectionCode="PDBFF" collectorName="Leg. R. &amp; A. Tourinho &amp; Beating" country="Brazil" location="Manaus" pageId="2" pageNumber="567" specimenCount="2" specimenCount-female="2" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC2BB3CF388FEC1FCE4E558" box="[661,771,332,357]" pageId="2" pageNumber="567" value="2006-01-13">
<date id="FB1701D4FFC2BB3CF388FEC1FCE4E558" box="[661,771,332,357]" pageId="2" pageNumber="567" value="2006-01-13">13.i.2006</date>
</collectingDate>
;
<collectorName id="225C42C2FFC2BB3CF20FFEC0FC8EE559" box="[786,873,333,357]" pageId="2" pageNumber="567">Leg. R.</collectorName>
Saturnino &amp;
<collectorName id="225C42C2FFC2BB3CF510FEC1FB79E558" box="[1037,1182,332,357]" pageId="2" pageNumber="567">A. Tourinho</collectorName>
;
<collectorName id="225C42C2FFC2BB3CF5B0FEC1FAE3E558" box="[1197,1284,332,357]" pageId="2" pageNumber="567">Beating</collectorName>
technique.
<specimenCount id="99AFEC9DFFC2BB3CF493FEC1FF09E5B5" pageId="2" pageNumber="567" type="female">2 females</specimenCount>
(IB-5799) from
<collectionCode id="E9B8BFD1FFC2BB3CF0B5FEFDFE19E5BA" box="[424,510,368,391]" pageId="2" pageNumber="567">PDBFF</collectionCode>
,
<location id="8A7671CFFFC2BB3CF314FEFDFD82E5BA" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC2BB3CF314FEFDFD82E5BA" box="[521,613,368,391]" country="Brazil" name="Manaus" pageId="2" pageNumber="567" stateProvince="Amazonas">Manaus</location>
,
<collectingRegion id="4D6DE9F6FFC2BB3CF372FEFDFD0FE5B5" box="[623,744,368,392]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="567">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC2BB3CF3EEFEE2FCDCE5B5" box="[755,827,367,392]" name="Brazil" pageId="2" pageNumber="567">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC2BB3CF25BFEFDFD5EE590" ID-GBIF-Occurrence="1326058844" collectingDate="2001" collectingDateMax="2003" collectingDateMin="2001" collectionCode="PDBFF" collectorName="Leg. F." country="Brazil" location="Manaus" pageId="2" pageNumber="567" specimenCount="3" specimenCount-male="3" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC2BB3CF25BFEFDFC20E5B5" box="[838,967,368,392]" pageId="2" pageNumber="567" value="2001" valueMax="2003">
<date id="FB1701D4FFC2BB3CF25BFEFDFC20E5B5" box="[838,967,368,392]" pageId="2" pageNumber="567" value="2001" valueMax="2003">2001–2003</date>
</collectingDate>
;
<collectorName id="225C42C2FFC2BB3CF2CFFEFDFBC7E5BA" box="[978,1056,368,392]" pageId="2" pageNumber="567">Leg. F.</collectorName>
Rego; sample 165A.
<specimenCount id="99AFEC9DFFC2BB3CF40CFEFDFA80E5B5" box="[1297,1383,367,392]" pageId="2" pageNumber="567" type="male">3 males</specimenCount>
(IB- 5818) from
<collectionCode id="E9B8BFD1FFC2BB3CF03DFE18FE90E591" box="[288,375,405,428]" pageId="2" pageNumber="567">PDBFF</collectionCode>
,
<location id="8A7671CFFFC2BB3CF09EFE18FE39E591" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC2BB3CF09EFE18FE39E591" box="[387,478,405,428]" country="Brazil" name="Manaus" pageId="2" pageNumber="567" stateProvince="Amazonas">Manaus</location>
,
<collectingRegion id="4D6DE9F6FFC2BB3CF0F6FE19FD83E591" box="[491,612,404,428]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="567">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC2BB3CF36CFE19FD5EE590" box="[625,697,404,429]" name="Brazil" pageId="2" pageNumber="567">Brazil</collectingCountry>
</materialsCitation>
;
<materialsCitation id="3FC12D49FFC2BB3CF3DBFE19FB54E5ED" ID-GBIF-Occurrence="1326058841" collectingDate="2001" collectingDateMax="2003" collectingDateMin="2001" collectorName="Leg. F. &amp; A. Other &amp; Reserva Ducke, LO" country="Brazil" location="Manaus" pageId="2" pageNumber="567" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Amazonas" typeStatus="paratype">
<collectingDate id="EB53F83CFFC2BB3CF3DBFE19FCAFE591" box="[710,840,404,428]" pageId="2" pageNumber="567" value="2001" valueMax="2003">
<date id="FB1701D4FFC2BB3CF3DBFE19FCAFE591" box="[710,840,404,428]" pageId="2" pageNumber="567" value="2001" valueMax="2003">2001–2003</date>
</collectingDate>
;
<collectorName id="225C42C2FFC2BB3CF249FE18FC43E591" box="[852,932,405,429]" pageId="2" pageNumber="567">Leg. F.</collectorName>
Rego; sample 94
<collectorName id="225C42C2FFC2BB3CF573FE19FB37E590" box="[1134,1232,404,429]" pageId="2" pageNumber="567">A. Other</collectorName>
material:
<specimenCount id="99AFEC9DFFC2BB3CF459FE19FA7BE590" box="[1348,1436,404,429]" pageId="2" pageNumber="567" type="male">3 males</specimenCount>
and
<specimenCount id="99AFEC9DFFC2BB3CF1DAFE35FED3E5ED" box="[199,308,439,464]" pageId="2" pageNumber="567" type="female">2 females</specimenCount>
(INPA-OP 4424) from
<collectorName id="225C42C2FFC2BB3CF35CFE35FCC4E5ED" box="[577,803,439,464]" pageId="2" pageNumber="567">Reserva Ducke, LO</collectorName>
7-1500,
<location id="8A7671CFFFC2BB3CF263FE35FC3DE5F2" LSID="urn:lsid:plazi:treatment:07009602FFC1BB3BF18AF9B1FD6CE520:8A7671CFFFC2BB3CF263FE35FC3DE5F2" box="[894,986,440,463]" country="Brazil" name="Manaus" pageId="2" pageNumber="567" stateProvince="Amazonas">Manaus</location>
,
<collectingRegion id="4D6DE9F6FFC2BB3CF2FBFE35FBB8E5ED" box="[998,1119,440,464]" country="Venezuela" name="Amazonas" pageId="2" pageNumber="567">Amazonas</collectingRegion>
,
<collectingCountry id="F7BE6784FFC2BB3CF576FE3AFB54E5ED" box="[1131,1203,439,464]" name="Brazil" pageId="2" pageNumber="567">Brazil</collectingCountry>
</materialsCitation>
;
<date id="FB1701D4FFC2BB3CF5A2FE3AFAA6E5ED" box="[1215,1345,439,464]" pageId="2" pageNumber="567" value="2014-07-27">27.vii.2014</date>
; Leg. P. Colmenares; cryptic manual searching; in ethanol 100%.
</paragraph>
</subSubSection>
<caption id="DBD6779CFFC2BB3CF18AF8A8FCF6E345" httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="2" pageNumber="567" targetBox="[163,1423,563,1793]" targetPageId="2">
<paragraph id="8F162714FFC2BB3CF18AF8A8FCF6E345" blockId="2.[151,1437,1829,1912]" pageId="2" pageNumber="567">
<emphasis id="BDDDFB06FFC2BB3CF18AF8A8FEF2E307" bold="true" box="[151,277,1829,1851]" pageId="2" pageNumber="567">FIGURE 1.</emphasis>
<taxonomicName id="48A95C97FFC2BB3CF000F8ABFDFDE306" box="[285,538,1829,1851]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="567" phylum="Arthropoda" rank="species" species="isadorae" status="sp. nov.">
<emphasis id="BDDDFB06FFC2BB3CF000F8ABFDFDE306" box="[285,538,1829,1851]" italics="true" pageId="2" pageNumber="567">Maracaynatum isadorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE467DFFC2BB3CF33FF8A8FD92E307" box="[546,629,1829,1850]" pageId="2" pageNumber="567" rank="species">
<emphasis id="BDDDFB06FFC2BB3CF33FF8A8FD92E307" bold="true" box="[546,629,1829,1850]" pageId="2" pageNumber="567">sp. nov.</emphasis>
</taxonomicNameLabel>
Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A–B, E–F: 1 mm; C–D, G: 0.1 mm.
</paragraph>
</caption>
<caption id="DBD6779CFFC3BB3DF18AFAE4FDAAE181" httpUri="https://zenodo.org/record/261023/files/figure.png" pageId="3" pageNumber="568" targetBox="[228,1354,191,1346]" targetPageId="3">
<paragraph id="8F162714FFC3BB3DF18AFAE4FDAAE181" blockId="3.[151,1435,1385,1468]" pageId="3" pageNumber="568">
<emphasis id="BDDDFB06FFC3BB3DF18AFAE4FEF5E143" bold="true" box="[151,274,1385,1407]" pageId="3" pageNumber="568">FIGURE 2.</emphasis>
<taxonomicName id="48A95C97FFC3BB3DF004FAE7FDF4E142" box="[281,531,1385,1407]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="isadorae" status="sp. nov.">
<emphasis id="BDDDFB06FFC3BB3DF004FAE7FDF4E142" box="[281,531,1385,1407]" italics="true" pageId="3" pageNumber="568">Maracaynatum isadorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE467DFFC3BB3DF304FAE4FD8EE143" box="[537,617,1385,1406]" pageId="3" pageNumber="568" rank="species">
<emphasis id="BDDDFB06FFC3BB3DF304FAE4FD8EE143" bold="true" box="[537,617,1385,1406]" pageId="3" pageNumber="568">sp. nov.</emphasis>
</taxonomicNameLabel>
Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C7B3749FFFC3BB3DF1DAFA69FBF8E278" pageId="3" pageNumber="568" type="etymology">
<paragraph id="8F162714FFC3BB3DF1DAFA69FBF8E278" blockId="3.[151,1437,1508,2037]" pageId="3" pageNumber="568">
<emphasis id="BDDDFB06FFC3BB3DF1DAFA69FEA9E1C0" bold="true" box="[199,334,1508,1533]" pageId="3" pageNumber="568">Etymology.</emphasis>
Patronymic in honor to our collaborator Merle Isadora Williams, a graphic designer who has been of great help to our lab, not only by improving the quality of our illustrations with her advice and digital editing, but also by acting as an assistant during field work and English language review.
</paragraph>
</subSubSection>
<subSubSection id="C7B3749FFFC3BB3DF1DAF9C2FD17E2C5" pageId="3" pageNumber="568" type="diagnosis">
<paragraph id="8F162714FFC3BB3DF1DAF9C2FD17E2C5" blockId="3.[151,1437,1508,2037]" pageId="3" pageNumber="568">
<emphasis id="BDDDFB06FFC3BB3DF1DAF9C2FEDDE255" bold="true" box="[199,314,1615,1640]" pageId="3" pageNumber="568">Diagnosis</emphasis>
. The new species is similar to
<taxonomicName id="48A95C97FFC3BB3DF392F9DCFBAEE255" authority="Silhavy, 1979" authorityName="Silhavy" authorityYear="1979" box="[655,1097,1615,1640]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="568" phylum="Arthropoda" rank="species" species="cubanum">
<emphasis id="BDDDFB06FFC3BB3DF392F9DCFC40E255" box="[655,935,1616,1640]" italics="true" pageId="3" pageNumber="568">Maracaynatum cubanum</emphasis>
<bibRefCitation id="EB385AE5FFC3BB3DF2B3F9C2FBAEE255" author="Silhavy" box="[942,1097,1615,1640]" pageId="3" pageNumber="568" refString="Silhavy, V. (1979) New american representatives of the subfamily Samoinae (Opiliones: Phalangodidae). Annotationes zoologicae et botanicae, Slovenske Narodne Muzeum, Bratislava, 130, 1 - 27." type="journal article" year="1979">Šilhavý, 1979</bibRefCitation>
</taxonomicName>
, from which differs by having three strong setiferous tubercles instead of two on the pedipalpal patella. Different from all remaining species by the shape of the stridulatory organ on the mesal face of cheliceral segment I, which is composed by two ellipsoidal tubercles and one small proximal tubercle. Penis easily distinguishable from all remaining species by the capsula interna, which has bifid conductors and a bifid stylus.
</paragraph>
</subSubSection>
<subSubSection id="C7B3749FFFC3BB3AF1DAF889FC6CE1D8" lastPageId="4" lastPageNumber="569" pageId="3" pageNumber="568" type="description">
<paragraph id="8F162714FFC3BB3DF1DAF889FA7AE321" blockId="3.[151,1437,1508,2037]" box="[199,1437,1796,1821]" pageId="3" pageNumber="568">
<emphasis id="BDDDFB06FFC3BB3DF1DAF889FDF7E320" bold="true" box="[199,528,1796,1821]" pageId="3" pageNumber="568">
Description. Male
<typeStatus id="501299B6FFC3BB3DF0BFF889FDECE320" box="[418,523,1796,1821]" pageId="3" pageNumber="568" type="holotype">holotype</typeStatus>
.
</emphasis>
Measurements: DSL: 1.18; PL: 0.31; DSW: 0.88; PW: 0.55; ID: 0.10; MH: 0.74.
</paragraph>
<paragraph id="8F162714FFC3BB3DF1DAF8A5FA71E3C8" blockId="3.[151,1437,1508,2037]" pageId="3" pageNumber="568">
Dorsum (
<figureCitation id="17923B91FFC3BB3DF029F8AAFE65E37D" box="[308,386,1831,1856]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="3" pageNumber="568">Figs. 1</figureCitation>
A–B). Dorsal scutum guitar-shaped in dorsal view, with the narrowest width between the prosoma and opisthosoma. Prosoma: anterior margin with one tubercle on each corner, with two clear cuticle areas on the cheliceral sockets, almost reaching the ocularium. Ocularium round and elevated in lateral view, placed in the middle of the prosoma and densely covered with small tubercles. Opisthosoma: higher than prosoma in lateral view, with four areas barely recognizable, wider at area IV and densely granulated. Sulci barely recognizable. Lateral and posterior margin with a row of minute tubercles. Free tergites with rows of minute setiferous tubercles.
</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFF15FBEEE520" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">Venter: Coxa I with two rows of 3–5 well developed tubercles and one larger apical anterior tubercle; II with two rows of small tubercles and two larger posterior tubercles; III with sparse minute tubercles; IV with sparse minute tubercles, two anterior and two posterior larger tubercles near the stigmatic area. Genital operculum slightly triangular. Free sternites and anal plate with rows of small setiferous tubercles.</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFEAAFD13E590" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
Chelicerae (
<figureCitation id="17923B91FFC4BB3AF053FEAAFE68E57D" box="[334,399,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="4" pageNumber="569">Fig. 1</figureCitation>
C). Not swollen. Segment I with well developed bulla and one stridulatory organ consisting of three elliptical ridges (
<bibRefCitation id="EB385AE5FFC4BB3AF0A7FEC1FDB3E559" author="Silhavy" box="[442,596,332,357]" pageId="4" pageNumber="569" refString="Silhavy, V. (1979) New american representatives of the subfamily Samoinae (Opiliones: Phalangodidae). Annotationes zoologicae et botanicae, Slovenske Narodne Muzeum, Bratislava, 130, 1 - 27." type="journal article" year="1979">Šilhavý 1979</bibRefCitation>
) (
<figureCitation id="17923B91FFC4BB3AF373FEC1FD48E559" box="[622,687,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="4" pageNumber="569">Fig. 1</figureCitation>
D) on the mesal face, consisting of sclerotized projections of the cuticle, elevated and with well defined margins. Segment II with 3–4 frontal and pointed tubercles, some distal setae and five distal teeth. Segment III with 3–4 teeth.
</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFE3AFCDFE65D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
Pedipalps (
<figureCitation id="17923B91FFC4BB3AF05AFE3AFE73E5ED" box="[327,404,439,464]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="4" pageNumber="569">Figs. 1</figureCitation>
E–F). Coxa with one ventral large tubercle. Trochanter with two ventral large tubercles. Femur ventrally with two basal, one medial, and one subapical strong setiferous tubercles, and one mesal subapical strong setiferous tubercle. Patella with one ectal and two mesal strong setiferous tubercles. Tibia with three ectal (IIi) and three mesal (IiI) strong setiferous tubercles. Tarsus with two ectal and two mesal strong setiferous tubercles equal in size and with several setae between them.
</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFDE0FE6BE6CD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
Legs (
<figureCitation id="17923B91FFC4BB3AF011FDE1FEB0E6B9" box="[268,343,620,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="4" pageNumber="569">Figs. 1</figureCitation>
B, 1G). All segments without armature only covered with fine and small setae. Metatarsus III not modified. Tarsal formula: 4(2)/6(3)/5/6; first segment of basitarsus and last segment of distitarsus II longer than remaining segments; first segment of basitarsus III–IV longer than remaining tarsal segment (
<figureCitation id="17923B91FFC4BB3AF5AFFD39FB15E6F1" box="[1202,1266,692,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="4" pageNumber="569">Fig. 1</figureCitation>
B). Scopula on tarsi III–IV (
<figureCitation id="17923B91FFC4BB3AF038FD5AFE83E6CD" box="[293,356,727,752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="4" pageNumber="569">Fig. 1</figureCitation>
G).
</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFD71FC17E009" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
Penis (
<figureCitation id="17923B91FFC4BB3AF008FD71FE83E729" box="[277,356,764,789]" captionStart="FIGURE 2" captionStartId="3.[151,250,1385,1407]" captionTargetBox="[228,1354,191,1346]" captionTargetId="figure@3.[220,1366,179,1364]" captionTargetPageId="3" captionText="FIGURE 2. Maracaynatum isadorae sp. nov. Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/261023/files/figure.png" pageId="4" pageNumber="569">Figs. 2</figureCitation>
A–G). Truncus thinner basally and swollen distally, ventrally with a transversal hump and low dorsal hump, both more recognizable in lateral view and located in the region between pars basalis and pars distalis. Pars distalis slightly compressed dorso-ventrally;
<emphasis id="BDDDFB06FFC4BB3AF22BFCC8FC08E761" box="[822,1007,836,860]" italics="true" pageId="4" pageNumber="569">Lamina apicalis</emphasis>
short, not modified into a calyx; six pairs of ventro-lateral foliar macrosetae, in three groups: three basal, one medial, and two distal; one pair of acuminate ventro-distal macrosetae (
<figureCitation id="17923B91FFC4BB3AF35CFC01FD6EE799" box="[577,649,908,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1385,1407]" captionTargetBox="[228,1354,191,1346]" captionTargetId="figure@3.[220,1366,179,1364]" captionTargetPageId="3" captionText="FIGURE 2. Maracaynatum isadorae sp. nov. Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/261023/files/figure.png" pageId="4" pageNumber="569">Fig. 2</figureCitation>
C). Capsula externa represented by an eversible membranous follis (
<figureCitation id="17923B91FFC4BB3AF182FC22FF0AE7F5" box="[159,237,943,968]" captionStart="FIGURE 2" captionStartId="3.[151,250,1385,1407]" captionTargetBox="[228,1354,191,1346]" captionTargetId="figure@3.[220,1366,179,1364]" captionTargetPageId="3" captionText="FIGURE 2. Maracaynatum isadorae sp. nov. Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/261023/files/figure.png" pageId="4" pageNumber="569">Figs. 2</figureCitation>
A–B). Capsula interna fully eversible, composed of free and flattened stylus, with a bifid tip (
<figureCitation id="17923B91FFC4BB3AF410FC22FAB6E7F5" box="[1293,1361,943,968]" captionStart="FIGURE 2" captionStartId="3.[151,250,1385,1407]" captionTargetBox="[228,1354,191,1346]" captionTargetId="figure@3.[220,1366,179,1364]" captionTargetPageId="3" captionText="FIGURE 2. Maracaynatum isadorae sp. nov. Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/261023/files/figure.png" pageId="4" pageNumber="569">Fig. 2</figureCitation>
B) and two laminar, rigid, and divergent conductors, basally fused and ending in bifid tips that fold on lateral margins of pars distalis when in non-expanded position (
<figureCitation id="17923B91FFC4BB3AF386FC7AFD07E02D" box="[667,736,1015,1040]" captionStart="FIGURE 2" captionStartId="3.[151,250,1385,1407]" captionTargetBox="[228,1354,191,1346]" captionTargetId="figure@3.[220,1366,179,1364]" captionTargetPageId="3" captionText="FIGURE 2. Maracaynatum isadorae sp. nov. Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/261023/files/figure.png" pageId="4" pageNumber="569">Fig. 2</figureCitation>
B). When the penis is expanded the conductors and stylus are projected distally, and the follis is inflated, expanding dorsally (
<figureCitation id="17923B91FFC4BB3AF27DFB91FC49E009" box="[864,942,1052,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1385,1407]" captionTargetBox="[228,1354,191,1346]" captionTargetId="figure@3.[220,1366,179,1364]" captionTargetPageId="3" captionText="FIGURE 2. Maracaynatum isadorae sp. nov. Male paratype (INPA-OP 1008), penis non-expanded: A) dorsal view; B) lateral view; C) ventral view. Male paratype (INPA-OP 3419), penis expanded: D) dorsal view; E) lateral view; F) ventral view; G) truncus, complete view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/261023/files/figure.png" pageId="4" pageNumber="569">Figs. 2</figureCitation>
D–F).
</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFBB2FD42E09D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
Color (
<figureCitation id="17923B91FFC4BB3AF00AFBB2FE87E065" box="[279,352,1087,1112]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[163,1423,563,1793]" captionTargetId="figure@2.[151,1436,553,1807]" captionTargetPageId="2" captionText="FIGURE 1. Maracaynatum isadorae sp. nov. Male holotype (INPA-OPI 003417), habitus: A) dorsal view, B) lateral view. Male paratype (INPA-OP 3419): C) chelicerae, mesal view, D) stridulatory organ, E) Pedipalp, ectal view, F) same, mesal view, G) scopula IV. Scale bars: A – B, E – F: 1 mm; C – D, G: 0.1 mm." httpUri="https://zenodo.org/record/167242/files/figure.png" pageId="4" pageNumber="569">Figs. 1</figureCitation>
A–B). Body yellowish, darker at prosoma and legs, from femora to metatarsi. Tarsi clearer than the rest of the body. Dark reticled areas on prosoma. Lateral portions of scutal areas, free tergites, complete anal plate and last esternite with dark colored areas.
</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFB21FBE4E0D4" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
<emphasis id="BDDDFB06FFC4BB3AF1DAFB21FEF9E0F8" bold="true" box="[199,286,1196,1221]" pageId="4" pageNumber="569">Female</emphasis>
(
<typeStatus id="501299B6FFC4BB3AF031FB21FE69E0F9" box="[300,398,1196,1220]" pageId="4" pageNumber="569" type="paratype">paratype</typeStatus>
INPA-OP 1557): Measurements: DSL: 1.38; PL: 0.42; DSW: 1.36; PW: 0.74; ID: 0.19; MH: 1.21. Very similar to male. Segment I of chelicerae without stridulatory organ.
</paragraph>
<paragraph id="8F162714FFC4BB3AF1DAFB79FC6CE1D8" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
<emphasis id="BDDDFB06FFC4BB3AF1DAFB79FE62E130" bold="true" box="[199,389,1268,1293]" pageId="4" pageNumber="569">Natural history.</emphasis>
This species seems to be a tree climber that spends the day on palms and trees, descending at night. It can be found in the lower parts of tree trunks and occasionally in the litter. The number of specimens found in a few lots obtained by beating technique suggests that multiple individuals of
<taxonomicName id="48A95C97FFC4BB3AF501FAB0FB44E169" box="[1052,1187,1340,1364]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="isadorae" status="sp. nov.">
<emphasis id="BDDDFB06FFC4BB3AF501FAB0FB44E169" box="[1052,1187,1340,1364]" italics="true" pageId="4" pageNumber="569">M. isadorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE467DFFC4BB3AF5B6FAB1FAE3E168" box="[1195,1284,1340,1365]" pageId="4" pageNumber="569" rank="species">
<emphasis id="BDDDFB06FFC4BB3AF5B6FAB1FAE3E168" bold="true" box="[1195,1284,1340,1365]" pageId="4" pageNumber="569">sp. nov.</emphasis>
</taxonomicNameLabel>
can be found in the same place. Another species of the genus,
<taxonomicName id="48A95C97FFC4BB3AF3DBFAECFCA6E145" box="[710,833,1376,1400]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="linaresi">
<emphasis id="BDDDFB06FFC4BB3AF3DBFAECFCA6E145" box="[710,833,1376,1400]" italics="true" pageId="4" pageNumber="569">M. linaresi</emphasis>
</taxonomicName>
, was observed climbing walls in small groups in the mountains of the Sierra de Perijá National Park, state of
<collectingRegion id="4D6DE9F6FFC4BB3AF24FFA09FC77E1A0" box="[850,912,1412,1437]" country="Venezuela" name="Zulia" pageId="4" pageNumber="569">Zulia</collectingRegion>
,
<collectingCountry id="F7BE6784FFC4BB3AF2BFFA09FBF2E1A0" box="[930,1045,1412,1437]" name="Venezuela" pageId="4" pageNumber="569">Venezuela</collectingCountry>
(Colmenares, unpublished data), suggesting some degree of gregarism. Another samoid not identified yet has been seen in groups climbing trees during the night in Reserva Ducke (Colmenares, unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C7B3749FFFC4BB3BF1DAFA62FD6CE520" lastPageId="5" lastPageNumber="570" pageId="4" pageNumber="569" type="distribution">
<paragraph id="8F162714FFC4BB3AF1DAFA62FEC5E211" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="569">
<emphasis id="BDDDFB06FFC4BB3AF1DAFA62FE87E235" bold="true" box="[199,352,1519,1544]" pageId="4" pageNumber="569">Distribution.</emphasis>
<collectingCountry id="F7BE6784FFC4BB3AF07AFA62FE57E235" box="[359,432,1519,1544]" name="Brazil" pageId="4" pageNumber="569">Brazil</collectingCountry>
:
<collectingRegion id="4D6DE9F6FFC4BB3AF0A7FA7DFDD2E235" box="[442,565,1520,1544]" country="Venezuela" name="Amazonas" pageId="4" pageNumber="569">Amazonas</collectingRegion>
: Fazenda UFAM; Reserva Ducke, Manaus; Rio Preto da Eva; Igapó Açu, BR- 319 (
<figureCitation id="17923B91FFC4BB3AF1CDF999FEF3E211" box="[208,276,1556,1581]" captionStart="FIGURE 3" captionStartId="5.[151,250,1246,1268]" captionTargetBox="[163,1427,343,1218]" captionTargetId="figure@5.[151,1436,339,1224]" captionTargetPageId="5" captionText="FIGURE 3. Distribution of Maracaynatum spp." httpUri="https://zenodo.org/record/261024/files/figure.png" pageId="4" pageNumber="569">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8F162714FFC4BB3BF1DAF9BAFD6CE520" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,285]" lastPageId="5" lastPageNumber="570" pageId="4" pageNumber="569">
<emphasis id="BDDDFB06FFC4BB3AF1DAF9BAFE8BE26D" bold="true" box="[199,364,1591,1616]" pageId="4" pageNumber="569">Comparisons.</emphasis>
<taxonomicName id="48A95C97FFC4BB3AF06AF9B4FD69E26D" box="[375,654,1592,1616]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="isadorae" status="sp. nov.">
<emphasis id="BDDDFB06FFC4BB3AF06AF9B4FD69E26D" box="[375,654,1592,1616]" italics="true" pageId="4" pageNumber="569">Maracaynatum isadorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE467DFFC4BB3AF385F9BAFD12E26D" box="[664,757,1591,1616]" pageId="4" pageNumber="569" rank="species">
<emphasis id="BDDDFB06FFC4BB3AF385F9BAFD12E26D" bold="true" box="[664,757,1591,1616]" pageId="4" pageNumber="569">sp. nov.</emphasis>
</taxonomicNameLabel>
seems to be a very unique species within
<taxonomicName id="48A95C97FFC4BB3AF5F7F9B4FA72E26D" box="[1258,1429,1593,1616]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="genus">
<emphasis id="BDDDFB06FFC4BB3AF5F7F9B4FA72E26D" box="[1258,1429,1593,1616]" italics="true" pageId="4" pageNumber="569">Maracaynatum</emphasis>
</taxonomicName>
, regarding the genital morphology. So far, is the only species of the genus with a bifid stylus. Also, the tips of the conductors in this species are bifid. These two features could be autapomorphic for this species. The only other species showing bifid conductors is
<taxonomicName id="48A95C97FFC4BB3AF32FF928FD4AE281" box="[562,685,1700,1724]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="linaresi">
<emphasis id="BDDDFB06FFC4BB3AF32FF928FD4AE281" box="[562,685,1700,1724]" italics="true" pageId="4" pageNumber="569">M. linaresi</emphasis>
</taxonomicName>
, but in that species the bifurcation happens more in the middle of the conductors and not in the tips (see
<bibRefCitation id="EB385AE5FFC4BB3AF359F945FCA7E2DD" author="Soares" box="[580,832,1736,1760]" pageId="4" pageNumber="569" refString="Soares H. E. M. &amp; Avram, S. (1981) Opilionides du Venezuela [I]. Travaux de l´Institut Speologie &quot; Emile Racovitza &quot;, 20, 1 - 21." type="journal article" year="1981">Soares &amp; Avram 1981</bibRefCitation>
). Regarding the external morphology, the new species resembles the Caribbean taxa:
<emphasis id="BDDDFB06FFC4BB3AF311F960FD41E339" box="[524,678,1772,1796]" italics="true" pageId="4" pageNumber="569">
<taxonomicName id="48A95C97FFC4BB3AF311F960FD45E339" box="[524,674,1772,1796]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="cubanum">M. cubanum</taxonomicName>
,
</emphasis>
<taxonomicName id="48A95C97FFC4BB3AF3A8F960FB4BE339" authority="Silhavy, 1979" authorityName="Silhavy" authorityYear="1979" box="[693,1196,1772,1797]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="trinidadense">
<emphasis id="BDDDFB06FFC4BB3AF3A8F960FC1DE339" box="[693,1018,1772,1796]" italics="true" pageId="4" pageNumber="569">Maracaynatum trinidadense</emphasis>
<bibRefCitation id="EB385AE5FFC4BB3AF514F961FB4BE339" author="Silhavy" box="[1033,1196,1772,1797]" pageId="4" pageNumber="569" refString="Silhavy, V. (1979) New american representatives of the subfamily Samoinae (Opiliones: Phalangodidae). Annotationes zoologicae et botanicae, Slovenske Narodne Muzeum, Bratislava, 130, 1 - 27." type="journal article" year="1979">Šilhavý, 1979</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="48A95C97FFC4BB3AF5ECF960FE48E315" authority="Silhavy, 1979" authorityName="Silhavy" authorityYear="1979" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="stridulans">
<emphasis id="BDDDFB06FFC4BB3AF5ECF960FEE0E315" italics="true" pageId="4" pageNumber="569">Maracaynatum stridulans</emphasis>
<bibRefCitation id="EB385AE5FFC4BB3AF012F882FE48E315" author="Silhavy" box="[271,431,1807,1832]" pageId="4" pageNumber="569" refString="Silhavy, V. (1979) New american representatives of the subfamily Samoinae (Opiliones: Phalangodidae). Annotationes zoologicae et botanicae, Slovenske Narodne Muzeum, Bratislava, 130, 1 - 27." type="journal article" year="1979">Šilhavý, 1979</bibRefCitation>
</taxonomicName>
. The males of the species in this group have very similar cheliceral morphology, having stridulatory organs in the shape of ellipsoidal flat tubercles with engrossed margins and a darker central area (
<bibRefCitation id="EB385AE5FFC4BB3AF183F8DAFEDFE34D" author="Silhavy" box="[158,312,1879,1904]" pageId="4" pageNumber="569" refString="Silhavy, V. (1979) New american representatives of the subfamily Samoinae (Opiliones: Phalangodidae). Annotationes zoologicae et botanicae, Slovenske Narodne Muzeum, Bratislava, 130, 1 - 27." type="journal article" year="1979">Šilhavý 1979</bibRefCitation>
).
<taxonomicName id="48A95C97FFC4BB3AF051F8D4FD89E34D" box="[332,622,1880,1904]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="stridulans">
<emphasis id="BDDDFB06FFC4BB3AF051F8D4FD89E34D" box="[332,622,1880,1904]" italics="true" pageId="4" pageNumber="569">Maracaynatum stridulans</emphasis>
</taxonomicName>
has a stridulatory organ composed of seven ridges, whereas the organ of
<taxonomicName id="48A95C97FFC4BB3AF18AF8F0FEAFE3A9" box="[151,328,1916,1940]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="trinidadense">
<emphasis id="BDDDFB06FFC4BB3AF18AF8F0FEAFE3A9" box="[151,328,1916,1940]" italics="true" pageId="4" pageNumber="569">M. trinidadense</emphasis>
</taxonomicName>
has only two, and the organ of
<taxonomicName id="48A95C97FFC4BB3AF3B9F8F0FCD7E3A9" box="[676,816,1916,1940]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="cubanum">
<emphasis id="BDDDFB06FFC4BB3AF3B9F8F0FCD7E3A9" box="[676,816,1916,1940]" italics="true" pageId="4" pageNumber="569">M. cubanum</emphasis>
</taxonomicName>
has two complete ridges and a small proximal tubercle, like
<taxonomicName id="48A95C97FFC4BB3AF1D0F82CFEBEE385" box="[205,345,1952,1976]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="isadorae" status="sp. nov.">
<emphasis id="BDDDFB06FFC4BB3AF1D0F82CFEBEE385" box="[205,345,1952,1976]" italics="true" pageId="4" pageNumber="569">M. isadorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE467DFFC4BB3AF078F82DFE23E384" box="[357,452,1952,1977]" pageId="4" pageNumber="569" rank="species">
<emphasis id="BDDDFB06FFC4BB3AF078F82DFE23E384" bold="true" box="[357,452,1952,1977]" pageId="4" pageNumber="569">sp. nov.</emphasis>
</taxonomicNameLabel>
Another feature is the cheliceral segment II, which is armed with three acuminate tubercles in
<taxonomicName id="48A95C97FFC4BB3AF03EF848FE5DE3E1" box="[291,442,1988,2012]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="stridulans">
<emphasis id="BDDDFB06FFC4BB3AF03EF848FE5DE3E1" box="[291,442,1988,2012]" italics="true" pageId="4" pageNumber="569">M. stridulans</emphasis>
</taxonomicName>
and
<taxonomicName id="48A95C97FFC4BB3AF0E9F848FD9AE3E1" box="[500,637,1988,2012]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="isadorae" status="sp. nov.">
<emphasis id="BDDDFB06FFC4BB3AF0E9F848FD9AE3E1" box="[500,637,1988,2012]" italics="true" pageId="4" pageNumber="569">M. isadorae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6EE467DFFC4BB3AF39BF849FD06E3E0" box="[646,737,1988,2013]" pageId="4" pageNumber="569" rank="species">
<emphasis id="BDDDFB06FFC4BB3AF39BF849FD06E3E0" bold="true" box="[646,737,1988,2013]" pageId="4" pageNumber="569">sp. nov.</emphasis>
</taxonomicNameLabel>
, a few small setiferous tubercles in
<emphasis id="BDDDFB06FFC4BB3AF59DF848FAF2E3E1" box="[1152,1301,1988,2012]" italics="true" pageId="4" pageNumber="569">
<taxonomicName id="48A95C97FFC4BB3AF59DF848FAF6E3E1" box="[1152,1297,1988,2012]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="569" phylum="Arthropoda" rank="species" species="cubanum">M. cubanum</taxonomicName>
,
</emphasis>
and is only covered with setae in
<taxonomicName id="48A95C97FFC5BB3BF08AFF14FDACE48D" box="[407,587,152,176]" class="Arachnida" family="Samoidae" genus="Maracaynatum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="570" phylum="Arthropoda" rank="species" species="trinidadense">
<emphasis id="BDDDFB06FFC5BB3BF08AFF14FDACE48D" box="[407,587,152,176]" italics="true" pageId="5" pageNumber="570">M. trinidadense</emphasis>
</taxonomicName>
. They also share a few characters in the genital morphology, such as the shape of the conductors, which are all laminar and curved distally. Horeover, given the homogeneity in the external morphology of the genus and without more detail about the genital morphology of the remaining species, it is difficult to offer more accurate comparisons.
</paragraph>
</subSubSection>
</treatment>
</document>
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