treatments-xml/data/F0/65/D1/F065D133FA5B513492E5129C7FD5B9F4.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.140.48865" ID-GBIF-Dataset="9a7c2d83-da94-49d7-b9e1-098103510b5a" ID-PMC="PMC7021731" ID-Pensoft-Pub="1314-2003-140-1" ID-Pensoft-UUID="7F4140B884A95B12B17EA292C84361E2" ID-PubMed="32095101" ModsDocID="1314-2003-140-1" checkinTime="1581156392183" checkinUser="pensoft" docAuthor="Canelon, Daniela S., Nino, Santos M., Dorr, Laurence J. &amp; Caraballo-Ortiz, Marcos A." docDate="2020" docId="F065D133FA5B513492E5129C7FD5B9F4" docLanguage="en" docName="PhytoKeys 140: 1-10" docOrigin="PhytoKeys 140" docSource="http://dx.doi.org/10.3897/phytokeys.140.48865" docTitle="Dendrophthora coronata Canelon, S. M. Nino, Dorr &amp; Caraballo 2020, sp. nov." docType="treatment" docVersion="4" id="7F4140B884A95B12B17EA292C84361E2" lastPageNumber="1" masterDocId="7F4140B884A95B12B17EA292C84361E2" masterDocTitle="Two new species of Dendrophthora (Viscaceae) from the Venezuelan Andes" masterLastPageNumber="10" masterPageNumber="1" pageNumber="1" updateTime="1668139648958" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Two new species of Dendrophthora (Viscaceae) from the Venezuelan Andes</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Canelon, Daniela S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nino, Santos M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dorr, Laurence J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caraballo-Ortiz, Marcos A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>140</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>10</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.140.48865</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.140.48865</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-140-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">7F4140B884A95B12B17EA292C84361E2</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="162058968" LSID="urn:lsid:plazi:treatment:F065D133FA5B513492E5129C7FD5B9F4" httpUri="http://treatment.plazi.org/id/F065D133FA5B513492E5129C7FD5B9F4" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
2. 
<taxonomicName LSID="F065D133-FA5B-5134-92E5-129C7FD5B9F4" authority="Canelón, S. M. Niño, Dorr &amp; Caraballo" authorityName="Canelon, S. M. Nino, Dorr &amp; Caraballo" authorityYear="2020" class="Magnoliopsida" family="Santalaceae" genus="Dendrophthora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dendrophthora coronata" order="Santalales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="coronata" status="sp. nov.">
Dendrophthora coronata 
<normalizedToken originalValue="Canelón">Canelon</normalizedToken>
, S.M. 
<normalizedToken originalValue="Niño">Nino</normalizedToken>
, Dorr &amp; Caraballo
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dendrophthora coronata. A Leaf and terminal inflorescences B Cataphylls at the base of a node C Coroniform trichomes D Complete pistillate inflorescence E-G Segments of a pistillate inflorescence H, I Flowers with vestigial anthers J-L Mature fruits. (Source: Dorr et al. 8988, US)." figureDoi="10.3897/phytokeys.140.48865.figure4" httpUri="https://binary.pensoft.net/fig/378019" pageId="0" pageNumber="1">Figure 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Type.</paragraph>
<paragraph pageId="0" pageNumber="1">
Venezuela. Trujillo: Municipio 
<normalizedToken originalValue="Boconó">Bocono</normalizedToken>
, Parque Nacional Guaramacal, trail from antennas on the summit of 
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
de Guaramacal, NE to Fila Los Recostaderos [sic, Recortaderos] (UTM: 19 369344E, 1023140N) [
<geoCoordinate degrees="9.253891" direction="north" orientation="latitude" precision="1" value="9.253891">9.253891N</geoCoordinate>
, - 
<geoCoordinate degrees="70.189471" direction="west" orientation="longitude" precision="1" value="-70.18947">70.189471W</geoCoordinate>
], 
<normalizedToken originalValue="páramo">paramo</normalizedToken>
and 
<normalizedToken originalValue="subpáramo">subparamo</normalizedToken>
vegetation, 2677-3100 m, 14 June 2001, 
<emphasis italics="true" pageId="0" pageNumber="1">L.J. Dorr 8988</emphasis>
with S.M. 
<normalizedToken originalValue="Niño">Nino</normalizedToken>
&amp; R. Caracas (holotype: US-00662772!; isotype: PORT).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Canelon, S. M. Nino, Dorr &amp; Caraballo" authorityYear="2020" class="Magnoliopsida" family="Viscaceae" genus="Dendrophthora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dendrophthora coronata" order="Santalales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">Dendrophthora coronata</emphasis>
</taxonomicName>
is distinguished from congeners by its stems and young branches with longitudinal parallel striations; coroniform trichomes (i.e., papillae crowned by 2-6 minute, simple hairs) covering the entire plant; small, ca. 0.5 
<normalizedToken originalValue="×">x</normalizedToken>
1 mm cataphylls present at all nodes and sometimes found 1-2 cm above nodes on older branches; petioles 0.5-1 mm long, leaf blades 1-1.5 
<normalizedToken originalValue="×">x</normalizedToken>
1-1.2 cm, apex rounded and margin slightly crenulate and papillose; uniseriate inflorescences with 5-9 flowers per series; and fruits globose-compressed, 2-3 
<normalizedToken originalValue="×">x</normalizedToken>
2 mm when mature, white.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Aerial parasitic shrubs</emphasis>
, monoecious; yellow-orange when fresh, drying blackish or dark green. 
<emphasis italics="true" pageId="0" pageNumber="1">Stems</emphasis>
woody, with multiple branches, 30-45 cm long, terete, surface coarse with parallel striations, with a dense layer of coroniform trichomes covering the entire plant; mature nodes separated by 2-3.5 cm long intervals, dichotomous; cataphylls 0.5-1 mm long at nodes, found 1-2 cm above nodes as branches became older. 
<emphasis italics="true" pageId="0" pageNumber="1">Leaves</emphasis>
opposite, coriaceous; petioles flattened, 0.5-2 mm long; blades orbicular to elliptic, 1-1.5 
<normalizedToken originalValue="×">x</normalizedToken>
1-1.2 cm, base slightly cuneate, apex rounded, margin indistinctly crenulate and papillose, surface rough on both sides, veins obscure in dry leaves (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dendrophthora coronata. A Leaf and terminal inflorescences B Cataphylls at the base of a node C Coroniform trichomes D Complete pistillate inflorescence E-G Segments of a pistillate inflorescence H, I Flowers with vestigial anthers J-L Mature fruits. (Source: Dorr et al. 8988, US)." figureDoi="10.3897/phytokeys.140.48865.figure4" httpUri="https://binary.pensoft.net/fig/378019" pageId="0" pageNumber="1">4</figureCitation>
). 
<emphasis italics="true" pageId="0" pageNumber="1">Inflorescences</emphasis>
completely pistillate within a branch, occasionally with staminate flowers at the base of the inflorescence (completely staminate inflorescences not seen); 1 per leaf axil, 1-3 fertile segments, each 4-10 mm long, uniseriate, greenish when dry; peduncles simple, 6-9 mm long, cup subtending inflorescence 1-1.5 
<normalizedToken originalValue="×">x</normalizedToken>
3-3.5 mm, almost always forked with a papillate apical edge. 
<emphasis italics="true" pageId="0" pageNumber="1">Flowers</emphasis>
3-9 per segment, 1-1.4 
<normalizedToken originalValue="×">x</normalizedToken>
0.7-1 mm. 
<emphasis italics="true" pageId="0" pageNumber="1">Fruits</emphasis>
globose-compressed, 2-3 
<normalizedToken originalValue="×">x</normalizedToken>
2 mm, ripening white- or purplish-translucent, surface granulose, tip protruding and crowned with persistent petals.
</paragraph>
<caption doi="10.3897/phytokeys.140.48865.figure4" httpUri="https://binary.pensoft.net/fig/378019" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName authorityName="Canelon, S. M. Nino, Dorr &amp; Caraballo" authorityYear="2020" class="Magnoliopsida" family="Viscaceae" genus="Dendrophthora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dendrophthora coronata" order="Santalales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">Dendrophthora coronata</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Leaf and terminal inflorescences 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Cataphylls at the base of a node 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
Coroniform trichomes 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
Complete pistillate inflorescence 
<emphasis bold="true" pageId="0" pageNumber="1">E-G</emphasis>
Segments of a pistillate inflorescence 
<emphasis bold="true" pageId="0" pageNumber="1">H, I</emphasis>
Flowers with vestigial anthers 
<emphasis bold="true" pageId="0" pageNumber="1">J-L</emphasis>
Mature fruits. (Source: 
<emphasis italics="true" pageId="0" pageNumber="1">Dorr et al. 8988</emphasis>
, US).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species has been found in the transition between cloud forest and 
<normalizedToken originalValue="subpáramo">subparamo</normalizedToken>
in Guaramcal. This vegetation is influenced by multiple factors including high rainfall (3200+ mm/year), elevation above sea level (2400-3100 m), as well as relative humidity (100% for most of the year) (
<bibRefCitation author="Cuello, N" journalOrPublisher="Stain Technology" pageId="0" pageNumber="1" refId="B2" refString="Cuello, N, Cleef, A, 2011. Bosques de los Andes de Venezuela: Caso el Ramal de Guaramacal. BioLlania Ed. Espec.10: 74-105." title="Bosques de los Andes de Venezuela: Caso el Ramal de Guaramacal. BioLlania Ed. Espec. 10: 74 - 105." year="2011">Cuello and Cleef 2011</bibRefCitation>
). 
<taxonomicName authorityName="Canelon, S. M. Nino, Dorr &amp; Caraballo" authorityYear="2020" class="Magnoliopsida" family="Viscaceae" genus="Dendrophthora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dendrophthora coronata" order="Santalales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">Dendrophthora coronata</emphasis>
</taxonomicName>
is found in these Andean/high Andean forests characterized by woody vegetation of low stature with numerous individual shrubs and small trees, and a thin understory with a carpet of thick leaf litter. The canopy can reach between 6-14 m tall, with some emerging trees that reach 16 m, among them: 
<taxonomicName authorityName="Cuello &amp; Aymard" authorityYear="2008" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex guaramacalensis" order="Aquifoliales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="guaramacalensis">
<emphasis italics="true" pageId="0" pageNumber="1">Ilex guaramacalensis</emphasis>
</taxonomicName>
Cuello &amp; Aymard (
<taxonomicName family="Aquifoliaceae" lsidName="" pageId="0" pageNumber="1" rank="family">Aquifoliaceae</taxonomicName>
), 
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Miconia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Miconia jahnii" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="jahnii">
<emphasis italics="true" pageId="0" pageNumber="1">Miconia jahnii</emphasis>
</taxonomicName>
Pittier (
<taxonomicName family="Melastomataceae" lsidName="" pageId="0" pageNumber="1" rank="family">Melastomataceae</taxonomicName>
), 
<taxonomicName class="Magnoliopsida" family="Primulaceae" genus="Myrsine" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Myrsine dependens" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="dependens">
<emphasis italics="true" pageId="0" pageNumber="1">Myrsine dependens</emphasis>
</taxonomicName>
(Ruiz &amp; Pav.) Spreng. (
<taxonomicName family="Primulaceae" lsidName="" pageId="0" pageNumber="1" rank="family">Primulaceae</taxonomicName>
), and 
<taxonomicName authorityName="Steyerm" authorityYear="1953" class="Magnoliopsida" family="Symplocaceae" genus="Symplocos" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Symplocos tamana" order="Ericales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tamana">
<emphasis italics="true" pageId="0" pageNumber="1">Symplocos tamana</emphasis>
</taxonomicName>
Steyerm. (
<taxonomicName family="Symplocaceae" lsidName="" pageId="0" pageNumber="1" rank="family">Symplocaceae</taxonomicName>
) (
<bibRefCitation author="Cuello, N" journalOrPublisher="Stain Technology" pageId="0" pageNumber="1" refId="B2" refString="Cuello, N, Cleef, A, 2011. Bosques de los Andes de Venezuela: Caso el Ramal de Guaramacal. BioLlania Ed. Espec.10: 74-105." title="Bosques de los Andes de Venezuela: Caso el Ramal de Guaramacal. BioLlania Ed. Espec. 10: 74 - 105." year="2011">Cuello and Cleef 2011</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="phenology">
<paragraph pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="1">
As with most other species of tropical mistletoe, 
<taxonomicName authorityName="Canelon, S. M. Nino, Dorr &amp; Caraballo" authorityYear="2020" class="Magnoliopsida" family="Viscaceae" genus="Dendrophthora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dendrophthora coronata" order="Santalales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">Dendrophthora coronata</emphasis>
</taxonomicName>
can be observed bearing flowers and fruits throughout the year. Its white fleshy fruits seem to be an important food source for forest birds.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species name is derived from the coroniform trichomes (i.e., papillae crowned by 2-6 minute, simple hairs) that cover the entire plant and that resemble small crowns.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
At first sight, 
<taxonomicName authorityName="Canelon, S. M. Nino, Dorr &amp; Caraballo" authorityYear="2020" class="Magnoliopsida" family="Viscaceae" genus="Dendrophthora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dendrophthora coronata" order="Santalales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">Dendrophthora coronata</emphasis>
</taxonomicName>
resembles 
<taxonomicName lsidName="D. apiculata" pageId="0" pageNumber="1" rank="species" species="apiculata">
<emphasis italics="true" pageId="0" pageNumber="1">D. apiculata</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="D. lindeniana" pageId="0" pageNumber="1" rank="species" species="lindeniana">
<emphasis italics="true" pageId="0" pageNumber="1">D. lindeniana</emphasis>
</taxonomicName>
. However, 
<taxonomicName lsidName="D. coronata" pageId="0" pageNumber="1" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">D. coronata</emphasis>
</taxonomicName>
can be readily recognized by its marked parallel striations along the stems, small, 0.5-1 mm long basal cataphylls, which sometimes are found 1-2 cm above nodes in old branches, and its dense layer of predominantly coroniform trichomes that cover the entire plant. In contrast, 
<taxonomicName lsidName="D. apiculata" pageId="0" pageNumber="1" rank="species" species="apiculata">
<emphasis italics="true" pageId="0" pageNumber="1">D. apiculata</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="D. lindeniana" pageId="0" pageNumber="1" rank="species" species="lindeniana">
<emphasis italics="true" pageId="0" pageNumber="1">D. lindeniana</emphasis>
</taxonomicName>
have stems sparsely covered by simple trichomes and lack basal cataphylls (or rarely have a few very small, 0.5 mm long cataphylls and then always at the nodes) (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters distinguishing Dendrophthora apiculata, D. coronata, and D. lindeniana." httpUri="http://table.plazi.org/id/24BBC26265B94F1D6E11FF1F14ACD093" pageId="0" pageNumber="1" tableUuid="24BBC26265B94F1D6E11FF1F14ACD093">1</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="24BBC26265B94F1D6E11FF1F14ACD093" httpUri="http://table.plazi.org/id/24BBC26265B94F1D6E11FF1F14ACD093" pageId="0" pageNumber="1" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 1.</emphasis>
Morphological characters distinguishing 
<taxonomicName authorityName="Canelon, S. M. Nino, Dorr &amp; Caraballo" authorityYear="2020" class="Magnoliopsida" family="Viscaceae" genus="Dendrophthora" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dendrophthora apiculata" order="Santalales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="apiculata">
<emphasis italics="true" pageId="0" pageNumber="1">Dendrophthora apiculata</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="D. coronata" pageId="0" pageNumber="1" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">D. coronata</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="D. lindeniana" pageId="0" pageNumber="1" rank="species" species="lindeniana">
<emphasis italics="true" pageId="0" pageNumber="1">D. lindeniana</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="D. apiculata" pageId="0" pageNumber="1" rank="species" species="apiculata">
<emphasis italics="true" pageId="0" pageNumber="1">D. apiculata</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="D. coronata" pageId="0" pageNumber="1" rank="species" species="coronata">
<emphasis italics="true" pageId="0" pageNumber="1">D. coronata</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="D. lindeniana" pageId="0" pageNumber="1" rank="species" species="lindeniana">
<emphasis italics="true" pageId="0" pageNumber="1">D. lindeniana</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Plant height</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">20-30+ cm.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30-45 cm.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Up to 100 cm.</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Indumentum</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Entire plant sparsely papillate.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Entire plant abundantly covered with coroniform trichomes.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Entire plant abundantly papillate.</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Stem</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">With longitudinal striations (not pronounced).</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">With pronounced striations (furrows).</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Without striations.</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Cataphylls</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Cataphylls surrounding nodes 0.2-0.5 mm long; only located at the base of a node.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Cataphylls in basal branches 0.5-1 mm long; found 1-2 cm above a node.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Cataphylls not present.</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Leaf apex</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Apiculate; apiculum 0.2-0.5 mm long.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Not apiculate.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Not apiculate.</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Petiole</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Petiole 2-3 mm long.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Petiole 0.5-2 mm long.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Petiole up to 1 mm long.</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Leaf margin</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Entire.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Slightly crenulate.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Entire.</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">Staminate inflorescence</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Triseriate.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Uniseriate.</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Uniseriate.</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Venezuela. Trujillo</emphasis>
: Municipio 
<normalizedToken originalValue="Boconó">Bocono</normalizedToken>
, Sector El Campamento, UTM: 19 368148 E, 1022056 N [
<geoCoordinate degrees="9.244052" direction="north" orientation="latitude" precision="1" value="9.244052">9.244052N</geoCoordinate>
, - 
<geoCoordinate degrees="70.200324" direction="west" orientation="longitude" precision="1" value="-70.200325">70.200324W</geoCoordinate>
], 2600, 13 Apr. 2019, 
<emphasis italics="true" pageId="0" pageNumber="1">
S. 
<normalizedToken originalValue="Niño">Nino</normalizedToken>
&amp; D. 
<normalizedToken originalValue="Canelón">Canelon</normalizedToken>
6111
</emphasis>
(US); Parque Nacional Guaramacal, sector Vertiente Sur, carretera al 
<normalizedToken originalValue="caserío">caserio</normalizedToken>
Guaramacal, 2000-2750 m, Dec. 1996, 
<emphasis italics="true" pageId="0" pageNumber="1">B. Stergios &amp; A. Licata 16813</emphasis>
(US-00656274).
</paragraph>
</subSubSection>
</treatment>
</document>