530 lines
78 KiB
XML
530 lines
78 KiB
XML
<document id="6DDA6319D9F081016F9A454C4B21E9C1" ID-DOI="10.11646/zootaxa.2901.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5284936" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629727970278" checkinUser="felipe" docAuthor="Yanes, Yurena, Santana, Jesús, Artiles, Miguel, Deniz, Francisco, Martín, Javier, Alonso, María R. & Ibáñez, Miguel" docDate="2011" docId="8146737DFFE0FFD0FF838E0CA3B1E3A5" docLanguage="en" docName="zt02901p051.pdf" docOrigin="Zootaxa 2901 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2901.1.3" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Napaeus josei Santana, Alonso & Ibanez 2011, n. sp." docType="treatment" docVersion="5" lastPageNumber="43" masterDocId="7D7F0B05FFE4FFD8FF148B76A25FE52B" masterDocTitle="Five new Napaeus species (Gastropoda: Pulmonata: Enidae) from Gran Canaria and El Hierro (Canary Islands)" masterLastPageNumber="51" masterPageNumber="35" pageNumber="39" updateTime="1699102865397" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="0518AD55E864D0CCFCEB25FA39DC3BB2">
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<mods:title id="4F9F499E664DCF288CC8173FA23F26BE">Five new Napaeus species (Gastropoda: Pulmonata: Enidae) from Gran Canaria and El Hierro (Canary Islands)</mods:title>
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<mods:name id="7C98A4A12552551C3F761E16943608A9" type="personal">
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<mods:namePart id="4DD6D0D3432FAD125E2FAD8BD940FE55">Yanes, Yurena</mods:namePart>
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<mods:affiliation id="434C11E542D34C7246F5EDA67A9B3140">Instituto Andaluz de Ciencias de la Tierra (CSIC-Universidad de Granada), Camino del Jueves s / n, 18100, Armilla, Granada, Spain. E-mail: yurenayanes @ ugr. es</mods:affiliation>
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<mods:roleTerm id="8038C083AF124DB210BBBBE5670D60D7">Author</mods:roleTerm>
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<mods:namePart id="7F4EBF540853650C6DA22D8B06286D86">Santana, Jesús</mods:namePart>
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<mods:affiliation id="1E8870732C683DB5EF8F0566C2B918E4">Las Palmas de Gran Canaria, Gran Canaria, Canary Islands, Spain. E-mail: CASIOPEA _ 69 @ terra. es; franciscodeniz @ yahoo. es Arinaga, Gran Canaria, Canary Islands, Spain. E-mail: miguel _ artiles @ yahoo. es</mods:affiliation>
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<mods:nameIdentifier id="D2CDB7F505367EB9C745ECE1AD7FE23C" type="email">_69@terra.es</mods:nameIdentifier>
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<mods:roleTerm id="CCEA1A8777C6985706F42EB8D27401A6">Author</mods:roleTerm>
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<mods:namePart id="936251D889C37A00875B0F8E8F392D6B">Artiles, Miguel</mods:namePart>
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<mods:affiliation id="17E9E05C0CC32696739B42CAE593DE0E">Las Palmas de Gran Canaria, Gran Canaria, Canary Islands, Spain. E-mail: CASIOPEA _ 69 @ terra. es; franciscodeniz @ yahoo. es Arinaga, Gran Canaria, Canary Islands, Spain. E-mail: miguel _ artiles @ yahoo. es</mods:affiliation>
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<mods:nameIdentifier id="C78D9E972CDC17B58A55A6BD0C2CB9FD" type="email">_69@terra.es</mods:nameIdentifier>
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<mods:roleTerm id="CCF11B9F295653B7CEFA9B347A4A5560">Author</mods:roleTerm>
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<mods:namePart id="4C87333B74F77F9DFA1453749092C8A1">Deniz, Francisco</mods:namePart>
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<mods:namePart id="435F7D2D8A6AC99CA4C7FB18A072BB3C">Martín, Javier</mods:namePart>
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<mods:affiliation id="B5384E123D28DBEC884D81B5042C3592">El Coromoto, La Laguna, Tenerife, Canary Islands, Spain. E-mail: javimarba @ hotmail. com</mods:affiliation>
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<mods:namePart id="66DD1F9100F6BCFC4645AA8519B6C44D">Alonso, María R.</mods:namePart>
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<mods:affiliation id="82256193EEA2C87036DDF0112B1B788C">Departamento de Biologia Animal, Universidad de La Laguna, E- 38206 La Laguna, Tenerife, Islas Canarias, Spain.</mods:affiliation>
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<mods:namePart id="86F438A36A8379E670A0E15E789B4304">Ibáñez, Miguel</mods:namePart>
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<mods:affiliation id="1C48E6CCFA4716E9F0E337E0CD6C49A6">Departamento de Biologia Animal, Universidad de La Laguna, E- 38206 La Laguna, Tenerife, Islas Canarias, Spain.</mods:affiliation>
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<mods:nameIdentifier id="3A4C7DCB24511BE13D979DBAA1AFD1BD" type="email">mibanez@ull.es</mods:nameIdentifier>
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<mods:title id="2ED9504DB07910895C55C712BFA0D86D">Zootaxa</mods:title>
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<mods:part id="6483627778F71B7002838AF0BDFD51D8">
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<mods:date id="6E7CB950860884444250AE3501CAF970">2011</mods:date>
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<mods:number id="0F8E53165ED627BC5AB79A87B3948431">2011-06-01</mods:number>
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<mods:detail id="C743716C6C5629D43F7BE64ED0CB46D4" type="volume">
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<mods:number id="8E748709261B6C0006107EA3D98A3BE2">2901</mods:number>
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<mods:number id="2F51D006E3A8C8B1F425A26217C3F926">1</mods:number>
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<mods:start id="951073EFECA478B043292B588531B19E">35</mods:start>
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<mods:url id="623EA4798DCCB972C626D2621F35250C">https://biotaxa.org/Zootaxa/article/view/zootaxa.2901.1.3</mods:url>
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<mods:classification id="F745E27658B32F23528F9C9DF0C24724">journal article</mods:classification>
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<mods:identifier id="4AA2FABFBDF3FDCC1CA30607E8078008" type="DOI">10.11646/zootaxa.2901.1.3</mods:identifier>
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<mods:identifier id="277A95F2CBB7AAD8C99DD8BF271F84B4" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="51F603529F2FCA9F354B4DB3D6D29C2F" type="Zenodo-Dep">5284936</mods:identifier>
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<treatment id="8146737DFFE0FFD0FF838E0CA3B1E3A5" ID-DOI="http://doi.org/10.5281/zenodo.5292362" ID-Zenodo-Dep="5292362" LSID="urn:lsid:plazi:treatment:8146737DFFE0FFD0FF838E0CA3B1E3A5" httpUri="http://treatment.plazi.org/id/8146737DFFE0FFD0FF838E0CA3B1E3A5" lastPageId="8" lastPageNumber="43" pageId="4" pageNumber="39">
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<subSubSection id="41F591E0FFE0FFDCFF838E0CA095E0B8" box="[151,714,1401,1428]" pageId="4" pageNumber="39" type="nomenclature">
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<paragraph id="0950C26BFFE0FFDCFF838E0CA095E0B8" blockId="4.[151,714,1401,1461]" box="[151,714,1401,1428]" pageId="4" pageNumber="39">
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<heading id="52187507FFE0FFDCFF838E0CA095E0B8" bold="true" box="[151,714,1401,1428]" fontSize="11" level="1" pageId="4" pageNumber="39" reason="1">
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<emphasis id="3B9B1E79FFE0FFDCFF838E0CA095E0B8" bold="true" box="[151,714,1401,1428]" pageId="4" pageNumber="39">
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<taxonomicName id="CEEFB9E8FFE0FFDCFF838E0CA020E0B8" ID-CoL="45NQ6" authority="Santana, Alonso & Ibanez" authorityName="Santana, Alonso & Ibanez" authorityYear="2011" box="[151,639,1401,1428]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="39" phylum="Mollusca" rank="species" species="josei" status="sp. nov.">
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<emphasis id="3B9B1E79FFE0FFDCFF838E0CA365E0BF" bold="true" box="[151,314,1402,1428]" italics="true" pageId="4" pageNumber="39">Napaeus josei</emphasis>
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Santana, Alonso & Ibáñez
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</taxonomicName>
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,
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<taxonomicNameLabel id="20A8A302FFE0FFDCFD9F8E0FA095E0B8" box="[651,714,1401,1427]" pageId="4" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="41F591E0FFE0FFDCFF838EEBA025E09E" box="[151,634,1437,1461]" pageId="4" pageNumber="39" type="description">
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<paragraph id="0950C26BFFE0FFDCFF838EEBA025E09E" blockId="4.[151,714,1401,1461]" box="[151,634,1437,1461]" pageId="4" pageNumber="39">
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<figureCitation id="91D4DEEEFFE0FFDCFF838EEBA2BCE09E" box="[151,227,1437,1461]" captionStart="FIGURE 1" captionStartId="3.[151,248,1648,1671]" captionTargetBox="[156,1430,207,1614]" captionTargetId="figure-40@3.[151,1436,195,1624]" captionTargetPageId="3" captionText="FIGURE 1. Geographical distribution of the new species." figureDoi="http://doi.org/10.5281/zenodo.5284942" httpUri="https://zenodo.org/record/5284942/files/figure.png" pageId="4" pageNumber="39">Figs. 1</figureCitation>
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,
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||
<figureCitation id="91D4DEEEFFE0FFDCFFE58EEBA342E09E" box="[241,285,1437,1461]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="4" pageNumber="39">3 A</figureCitation>
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,
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<figureCitation id="91D4DEEEFFE0FFDCFE3C8EEBA32AE09E" box="[296,373,1437,1461]" captionStart="FIGURE 4" captionStartId="6.[151,248,1647,1670]" captionTargetBox="[151,1436,177,1626]" captionTargetId="figure-181@6.[151,1436,175,1626]" captionTargetPageId="6" captionText="FIGURE 4. Napaeus josei n. sp. A. Paratype from the type locality (AIT). B. Three paratypes from the type locality, attached to a stone and disguised with a soil covering (JSGC). C. Disguised paratype with a very thick soil covering (JSGC)." figureDoi="http://doi.org/10.5281/zenodo.5284954" httpUri="https://zenodo.org/record/5284954/files/figure.png" pageId="4" pageNumber="39">4 A–C</figureCitation>
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,
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<figureCitation id="91D4DEEEFFE0FFDCFE948EEBA393E09F" box="[384,460,1437,1461]" captionStart="FIGURE 5" captionStartId="7.[151,248,1552,1575]" captionTargetBox="[151,1436,709,1528]" captionTargetId="figure-451@7.[151,1436,709,1528]" captionTargetPageId="7" captionText="FIGURE 5. Details of shell ornamentation of holotypes (A, C: penultimate whorl; B, D: body whorl). A–B. Napaeus josei n. sp. C–D. N. venegueraensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5284956" httpUri="https://zenodo.org/record/5284956/files/figure.png" pageId="4" pageNumber="39">5 A, B</figureCitation>
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,
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<figureCitation id="91D4DEEEFFE0FFDCFECC8EEBA07BE09F" box="[472,548,1437,1461]" captionStart="FIGURE 6" captionStartId="8.[151,248,1204,1227]" captionTargetBox="[151,1436,287,1187]" captionTargetId="figure-398@8.[151,1436,287,1187]" captionTargetPageId="8" captionText="FIGURE 6. Napaeus josei n. sp., paratype from Barranco del Lechugal, Gran Canaria (AIT). A. Genital system. B. internal anatomy of penis, showing the penial papilla; A1–A5, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; d, diverticulum; e, epiphallus; ec, epiphallic caecum; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; p, penis; pp, penial papilla; pr, penis retractor; pw, penis wall; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5284960" httpUri="https://zenodo.org/record/5284960/files/figure.png" pageId="4" pageNumber="39">6 A, B</figureCitation>
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,
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<figureCitation id="91D4DEEEFFE0FFDCFD248EEBA025E09E" box="[560,634,1437,1461]" captionStart="FIGURE 7" captionStartId="9.[151,248,1358,1381]" captionTargetBox="[171,1416,198,1320]" captionTargetId="figure-301@9.[151,1436,193,1337]" captionTargetPageId="9" captionText="FIGURE 7. Scatter plots of some shell measurements for the new species and the most similar species. AS, aperture surface area (plane view); BS, body whorl surface area (plane view); FS, first whorls surface area (plane view); PS, penultimate whorl surface area (plane view); SB, shell breadth; SH, shell height; SS, shell surface area (plane view). The dimensions are in mm (lengths) or mm2 (surface areas)." figureDoi="http://doi.org/10.5281/zenodo.5284962" httpUri="https://zenodo.org/record/5284962/files/figure.png" pageId="4" pageNumber="39">7 A–C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="41F591E0FFE0FFDCFF838E93A34CE3A6" pageId="4" pageNumber="39" type="materials_examined">
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<paragraph id="0950C26BFFE0FFDCFF838E93A6D4E0D6" blockId="4.[151,1437,1508,2003]" box="[151,1163,1508,1533]" pageId="4" pageNumber="39">
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<emphasis id="3B9B1E79FFE0FFDCFF838E93A36AE0D6" bold="true" box="[151,309,1508,1533]" inLineHeading="true" pageId="4" pageNumber="39" reason="1">Type locality.</emphasis>
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Barranco del Lechugal (Gran Canaria;
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<collectionCode id="6FFE5AAEFFE0FFDCFDE48E93A16DE0D7" box="[752,818,1509,1532]" pageId="4" pageNumber="39">UTM</collectionCode>
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: 28RDR
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<quantity id="CE176F8EFFE0FFDCFC818E93A640E0D7" box="[917,1055,1509,1533]" metricMagnitude="3" metricUnit="m" metricValue="3.0751" pageId="4" pageNumber="39" unit="m" value="3075.1">3075, 100 m</quantity>
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altitude).
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</paragraph>
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<paragraph id="0950C26BFFE0FFDCFFD18D7FA6D9E30B" blockId="4.[151,1437,1508,2003]" box="[197,1158,1544,1569]" pageId="4" pageNumber="39">
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<materialsCitation id="B987C836FFE0FFDCFFD18D7FA6D9E30B" box="[197,1158,1544,1569]" collectingDate="2009-01-26" collectionCode="TFMC" collectorName="Leg. J. Santana" pageId="4" pageNumber="39" specimenCode="MT 0415" specimenCount="1" typeStatus="holotype">
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<emphasis id="3B9B1E79FFE0FFDCFFD18D7FA068E30B" box="[197,567,1544,1569]" inLineHeading="true" pageId="4" pageNumber="39" reason="1">
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<emphasis id="3B9B1E79FFE0FFDCFFD18D7FA370E30A" bold="true" box="[197,303,1545,1569]" pageId="4" pageNumber="39">
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<typeStatus id="D6547CC9FFE0FFDCFFD18D7FA370E30A" box="[197,303,1545,1569]" pageId="4" pageNumber="39" type="holotype">Holotype</typeStatus>
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</emphasis>
|
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(
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||
<figureCitation id="91D4DEEEFFE0FFDCFE298D7EA3C5E30B" box="[317,410,1544,1568]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="4" pageNumber="39">Fig. 3 A</figureCitation>
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; empty shell).
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</emphasis>
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<collectionCode id="6FFE5AAEFFE0FFDCFD298D7EA0D6E30B" box="[573,649,1544,1568]" pageId="4" pageNumber="39">TFMC</collectionCode>
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(
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<specimenCode id="59496A10FFE0FFDCFD8C8D7EA157E30B" box="[664,776,1544,1568]" collectionCode="TFMC" pageId="4" pageNumber="39">MT 0415</specimenCode>
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).
|
||
<collectorName id="A41AA7BDFFE0FFDCFC0E8D7FA19DE30B" box="[794,962,1544,1569]" pageId="4" pageNumber="39">Leg. J. Santana</collectorName>
|
||
,
|
||
<date id="7D51E4ABFFE0FFDCFCDF8D7EA6DDE30B" box="[971,1154,1544,1569]" pageId="4" pageNumber="39" value="2009-01-26">
|
||
<collectingDate id="6D151D43FFE0FFDCFCDF8D7EA6DDE30B" box="[971,1154,1544,1569]" pageId="4" pageNumber="39" value="2009-01-26">26 January 2009</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFE0FFDCFFD18D5BA34CE3A6" blockId="4.[151,1437,1508,2003]" pageId="4" pageNumber="39">
|
||
<materialsCitation id="B987C836FFE0FFDCFFD18D5BA350E3A6" collectionCode="AIT, JSGC, FDGC, MAGC" location="Gran Canaria" pageId="4" pageNumber="39" specimenCount="12" typeStatus="paratype">
|
||
<emphasis id="3B9B1E79FFE0FFDCFFD18D5BA31BE36F" box="[197,324,1581,1605]" inLineHeading="true" pageId="4" pageNumber="39" reason="1">
|
||
<emphasis id="3B9B1E79FFE0FFDCFFD18D5BA362E36E" bold="true" box="[197,317,1581,1605]" pageId="4" pageNumber="39">
|
||
<typeStatus id="D6547CC9FFE0FFDCFFD18D5BA362E36E" box="[197,317,1581,1605]" pageId="4" pageNumber="39" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="1FE909E2FFE0FFDCFE588D5BA388E36E" box="[332,471,1581,1605]" pageId="4" pageNumber="39" type="generic">6 specimens</specimenCount>
|
||
and 102 shells, collected between 1998 and 2010 from the southwest of
|
||
<location id="0C3094B0FFE0FFDCFA168D5BA7C7E36E" LSID="urn:lsid:plazi:treatment:8146737DFFE0FFD0FF838E0CA3B1E3A5:0C3094B0FFE0FFDCFA168D5BA7C7E36E" box="[1282,1432,1581,1605]" name="Gran Canaria" pageId="4" pageNumber="39">Gran Canaria</location>
|
||
, deposited in
|
||
<collectionCode id="6FFE5AAEFFE0FFDCFE308D26A30DE343" box="[292,338,1616,1640]" pageId="4" pageNumber="39">AIT</collectionCode>
|
||
(
|
||
<specimenCount id="1FE909E2FFE0FFDCFE768D26A3B3E343" box="[354,492,1616,1640]" pageId="4" pageNumber="39" type="generic">6 specimens</specimenCount>
|
||
, 5 shells) and the following collections:
|
||
<collectionCode id="6FFE5AAEFFE0FFDCFCBF8D26A1B4E343" box="[939,1003,1616,1640]" pageId="4" pageNumber="39">JSGC</collectionCode>
|
||
(68 shells);
|
||
<collectionCode id="6FFE5AAEFFE0FFDCFB6F8D26A699E343" box="[1147,1222,1616,1640]" pageId="4" pageNumber="39">FDGC</collectionCode>
|
||
(9 shells);
|
||
<collectionCode id="6FFE5AAEFFE0FFDCFA5C8D26A7C3E343" box="[1352,1436,1616,1640]" pageId="4" pageNumber="39">MAGC</collectionCode>
|
||
(20 shells)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFE0FFDCFFD18DEFA370E3FE" pageId="4" pageNumber="39" type="etymology">
|
||
<paragraph id="0950C26BFFE0FFDCFFD18DEFA370E3FE" blockId="4.[151,1437,1508,2003]" pageId="4" pageNumber="39">
|
||
<emphasis id="3B9B1E79FFE0FFDCFFD18DEFA314E39A" bold="true" box="[197,331,1689,1713]" inLineHeading="true" pageId="4" pageNumber="39" reason="1">Etymology.</emphasis>
|
||
The specific name is dedicated to our late friend Jose María Hernández, marine malacologist from Gran Canaria.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFE0FFDEFFD18D95A0DDE264" lastPageId="6" lastPageNumber="41" pageId="4" pageNumber="39" type="distribution">
|
||
<paragraph id="0950C26BFFE0FFDEFFD18D95A0DDE264" blockId="4.[151,1437,1508,2003]" lastBlockId="6.[151,1436,1739,2021]" lastPageId="6" lastPageNumber="41" pageId="4" pageNumber="39">
|
||
<emphasis id="3B9B1E79FFE0FFDCFFD18D95A017E3D0" box="[197,584,1763,1788]" inLineHeading="true" pageId="4" pageNumber="39" reason="1">
|
||
<emphasis id="3B9B1E79FFE0FFDCFFD18D95A3B8E3D7" bold="true" box="[197,487,1763,1788]" pageId="4" pageNumber="39">Distribution and habitat</emphasis>
|
||
(
|
||
<figureCitation id="91D4DEEEFFE0FFDCFEE28D95A068E3D0" box="[502,567,1763,1787]" captionStart="FIGURE 1" captionStartId="3.[151,248,1648,1671]" captionTargetBox="[156,1430,207,1614]" captionTargetId="figure-40@3.[151,1436,195,1624]" captionTargetPageId="3" captionText="FIGURE 1. Geographical distribution of the new species." figureDoi="http://doi.org/10.5281/zenodo.5284942" httpUri="https://zenodo.org/record/5284942/files/figure.png" pageId="4" pageNumber="39">Fig. 1</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Endemic to Gran Canaria, occupying an area of about
|
||
<quantity id="CE176F8EFFE0FFDCFBB98D95A75EE3D0" box="[1197,1281,1763,1787]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="39" unit="km" value="100.0">100 km</quantity>
|
||
<superScript id="FE9A6F23FFE0FFDCFA158D96A756E3C5" attach="left" box="[1281,1289,1760,1774]" fontSize="6" pageId="4" pageNumber="39">2</superScript>
|
||
of the southwest of the island, with lowland vegetation, within an altitudinal range between 40 and
|
||
<specimenCount id="1FE909E2FFE0FFDCFB7B8C71A6E1E234" box="[1135,1214,1799,1823]" pageId="4" pageNumber="39" type="generic">
|
||
<quantity id="CE176F8EFFE0FFDCFB7B8C71A6E5E234" box="[1135,1210,1799,1823]" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="39" unit="m" value="460.0">460 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
<taxonomicName id="CEEFB9E8FFE0FFDCFBD18C7FA73EE234" authorityName="Santana, Alonso & Ibanez" authorityYear="2011" box="[1221,1377,1800,1824]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="39" phylum="Mollusca" rank="species" species="josei">
|
||
<emphasis id="3B9B1E79FFE0FFDCFBD18C7FA73EE234" box="[1221,1377,1800,1824]" italics="true" pageId="4" pageNumber="39">Napaeus josei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFE0FFDCFA648C71A29DE268" captionStart="FIGURE 4" captionStartId="6.[151,248,1647,1670]" captionTargetBox="[151,1436,177,1626]" captionTargetId="figure-181@6.[151,1436,175,1626]" captionTargetPageId="6" captionText="FIGURE 4. Napaeus josei n. sp. A. Paratype from the type locality (AIT). B. Three paratypes from the type locality, attached to a stone and disguised with a soil covering (JSGC). C. Disguised paratype with a very thick soil covering (JSGC)." figureDoi="http://doi.org/10.5281/zenodo.5284954" httpUri="https://zenodo.org/record/5284954/files/figure.png" pageId="4" pageNumber="39">Fig. 4 A</figureCitation>
|
||
) is mainly a ground dweller, living associated with large
|
||
<taxonomicName id="CEEFB9E8FFE0FFDCFC428C5AA675E268" box="[854,1066,1835,1860]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="3B9B1E79FFE0FFDCFC428C5AA191E26F" box="[854,974,1836,1860]" italics="true" pageId="4" pageNumber="39">Euphorbia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, sheltering during the day under stones or between
|
||
<taxonomicName id="CEEFB9E8FFE0FFDCFE708C26A383E243" box="[356,476,1872,1896]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="3B9B1E79FFE0FFDCFE708C26A383E243" box="[356,476,1872,1896]" italics="true" pageId="4" pageNumber="39">Euphorbia</emphasis>
|
||
</taxonomicName>
|
||
roots. The majority of the specimens collected had the shell disguised by a covering of soil, which was sometimes very thick (
|
||
<figureCitation id="91D4DEEEFFE0FFDCFD468C05A080E2A0" box="[594,735,1907,1931]" captionStart="FIGURE 4" captionStartId="6.[151,248,1647,1670]" captionTargetBox="[151,1436,177,1626]" captionTargetId="figure-181@6.[151,1436,175,1626]" captionTargetPageId="6" captionText="FIGURE 4. Napaeus josei n. sp. A. Paratype from the type locality (AIT). B. Three paratypes from the type locality, attached to a stone and disguised with a soil covering (JSGC). C. Disguised paratype with a very thick soil covering (JSGC)." figureDoi="http://doi.org/10.5281/zenodo.5284954" httpUri="https://zenodo.org/record/5284954/files/figure.png" pageId="4" pageNumber="39">Figs. 4 B, C</figureCitation>
|
||
), altering the appearance of the shell considerably as in other disguised
|
||
<taxonomicName id="CEEFB9E8FFE0FFDCFE138CEFA39FE284" box="[263,448,1943,1968]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="39" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="3B9B1E79FFE0FFDCFE138CEFA336E29B" box="[263,361,1945,1968]" italics="true" pageId="4" pageNumber="39">Napaeus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="6D7EBF9AFFE0FFDCFEDA8CE1A0CDE284" author="Yanes" box="[462,658,1943,1968]" etAl="et al." firstAuthor="Yanes" pageId="4" pageNumber="39" pagination="143 - 148" refId="ref9431" refString="Yanes, Y., Martin, J., Delgado, J. D., Alonso, M. R. & Ibanez, M. (2010) Active disguise in land snails: Napaeus badiosus (Gastropoda, Pulmonata, Enidae) from the Canary Islands. Journal of Conchology, 40, 143 - 148." type="journal article" year="2010">
|
||
Yanes
|
||
<emphasis id="3B9B1E79FFE0FFDCFD0E8CEFA017E29B" box="[538,584,1944,1968]" italics="true" pageId="4" pageNumber="39">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
). This renders most living animals very inconspicuous in their normal resting places, presumably reducing the risk of visual detection by predators. The snails are part of the diet of the Southern Grey Shrike,
|
||
<taxonomicName id="CEEFB9E8FFE2FFDEFE8E8DBAA0A5E3C8" authority="Temminck" authorityName="Temminck" authorityYear="1820" box="[410,762,1739,1764]" class="Aves" family="Laniidae" genus="Lanius" kingdom="Animalia" order="Passeriformes" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="meridionalis">
|
||
<emphasis id="3B9B1E79FFE2FFDEFE8E8DBAA024E3CF" box="[410,635,1740,1764]" italics="true" pageId="6" pageNumber="41">Lanius meridionalis</emphasis>
|
||
Temminck
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="CEEFB9E8FFE2FFDEFC1D8DBDA11AE3C8" authorityName="Linnaeus" authorityYear="1758" box="[777,837,1739,1763]" class="Aves" kingdom="Animalia" pageId="6" pageNumber="41" phylum="Chordata" rank="class">Aves</taxonomicName>
|
||
,
|
||
<taxonomicName id="CEEFB9E8FFE2FFDEFC458DBDA1AFE3C8" box="[849,1008,1739,1763]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="6" pageNumber="41" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="6D7EBF9AFFE2FFDEFB128DBDA68AE3C8" author="Padilla" box="[1030,1237,1739,1764]" etAl="et al." firstAuthor="Padilla" pageId="6" pageNumber="41" pagination="557 - 568" refId="ref9204" refString="Padilla, D. P., Gonzalez-Castro, A., Nieves, C. & Nogales, M. (2009) Trophic ecology of the southern grey shrike (Lanius meridionalis) in insular environments: the influence of altitude and seasonality. Journal of Ornithology, 150, 557 - 568." type="journal article" year="2009">
|
||
Padilla
|
||
<emphasis id="3B9B1E79FFE2FFDEFB498DBAA6D2E3CF" box="[1117,1165,1740,1764]" italics="true" pageId="6" pageNumber="41">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
) and the omnivorous lizard,
|
||
<taxonomicName id="CEEFB9E8FFE2FFDEFE0E8D99A0FBE22C" authority="(Peters & Doria)" baseAuthorityName="Peters & Doria" box="[282,676,1775,1799]" class="Reptilia" family="Lacertidae" genus="Gallotia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="41" phylum="Chordata" rank="species" species="atlantica">
|
||
<emphasis id="3B9B1E79FFE2FFDEFE0E8D99A3BDE22C" box="[282,482,1775,1799]" italics="true" pageId="6" pageNumber="41">Gallotia atlantica</emphasis>
|
||
(Peters & Doria)
|
||
</taxonomicName>
|
||
, endemic to the eastern islands (
|
||
<bibRefCitation id="6D7EBF9AFFE2FFDEFB1D8D99A74AE22C" author="Valido & Nogales" box="[1033,1301,1775,1799]" firstAuthor="Valido" pageId="6" pageNumber="41" pagination="331 - 344" refId="ref9328" refString="Valido, A. & Nogales, M. (2003) Digestive ecology of two omnivorous Canarian lizard species (Gallotia, Lacertidae). Amphibia-Reptilia, 24, 331 - 344." type="journal article" year="2003">Valido & Nogales 2003</bibRefCitation>
|
||
). Other species of
|
||
<taxonomicName id="CEEFB9E8FFE2FFDEFFFD8C62A319E207" box="[233,326,1812,1836]" class="Reptilia" family="Lacertidae" genus="Gallotia" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="41" phylum="Chordata" rank="genus">
|
||
<emphasis id="3B9B1E79FFE2FFDEFFFD8C62A319E207" box="[233,326,1812,1836]" italics="true" pageId="6" pageNumber="41">Gallotia</emphasis>
|
||
</taxonomicName>
|
||
, which as a genus is present in all the islands of the Archipelago, probably also consume snails as part of their regular diet (
|
||
<bibRefCitation id="6D7EBF9AFFE2FFDEFEA58C41A029E264" author="Yanes" box="[433,630,1847,1871]" etAl="et al." firstAuthor="Yanes" pageId="6" pageNumber="41" pagination="143 - 148" refId="ref9431" refString="Yanes, Y., Martin, J., Delgado, J. D., Alonso, M. R. & Ibanez, M. (2010) Active disguise in land snails: Napaeus badiosus (Gastropoda, Pulmonata, Enidae) from the Canary Islands. Journal of Conchology, 40, 143 - 148." type="journal article" year="2010">
|
||
Yanes
|
||
<emphasis id="3B9B1E79FFE2FFDEFEE98C4EA073E264" box="[509,556,1847,1871]" italics="true" pageId="6" pageNumber="41">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="5D9092E3FFE1FFDDFF838C4FA742E2C2" ID-DOI="http://doi.org/10.5281/zenodo.5284948" ID-Zenodo-Dep="5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="5" pageNumber="40" startId="5.[151,250,1849,1872]" targetBox="[176,1412,193,1835]" targetPageId="5">
|
||
<paragraph id="0950C26BFFE1FFDDFF838C4FA742E2C2" blockId="5.[151,1437,1849,2025]" pageId="5" pageNumber="40">
|
||
<emphasis id="3B9B1E79FFE1FFDDFF838C4FA34DE27B" bold="true" box="[151,274,1849,1872]" pageId="5" pageNumber="40">FIGURE 3.</emphasis>
|
||
Shells of: A.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFE8B8C4DA071E27B" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[415,558,1851,1872]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="josei" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE1FFDDFE8B8C4DA071E27B" box="[415,558,1851,1872]" italics="true" pageId="5" pageNumber="40">Napaeus josei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFE1FFDDFD208C4CA034E264" bold="true" box="[564,619,1850,1871]" pageId="5" pageNumber="40">
|
||
<taxonomicNameLabel id="20A8A302FFE1FFDDFD208C4CA034E264" box="[564,619,1850,1871]" pageId="5" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. B.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFDEE8C4DA133E27B" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[762,876,1850,1872]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="nanodes">
|
||
<emphasis id="3B9B1E79FFE1FFDDFDEE8C4DA133E27B" box="[762,876,1850,1872]" italics="true" pageId="5" pageNumber="40">N. nanodes</emphasis>
|
||
</taxonomicName>
|
||
, from Barranco del Agua, Tenerife (AIT). C.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFA288C4DA2E8E244" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="3B9B1E79FFE1FFDDFA288C4DA2E8E244" italics="true" pageId="5" pageNumber="40">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
, from Barranco de Valle Seco, Tenerife. D.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFD948C2CA164E244" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[640,827,1882,1903]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="venegueraensis" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE1FFDDFD948C2CA164E244" box="[640,827,1882,1903]" italics="true" pageId="5" pageNumber="40">N. venegueraensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFE1FFDDFC578C2FA124E245" bold="true" box="[835,891,1881,1902]" pageId="5" pageNumber="40">
|
||
<taxonomicNameLabel id="20A8A302FFE1FFDDFC578C2FA124E245" box="[835,891,1881,1902]" pageId="5" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. E.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFB198C2CA6F3E244" baseAuthorityName="Wollaston" baseAuthorityYear="1878" box="[1037,1196,1881,1903]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="chrysaloides">
|
||
<emphasis id="3B9B1E79FFE1FFDDFB198C2CA6F3E244" box="[1037,1196,1881,1903]" italics="true" pageId="5" pageNumber="40">N. chrysaloides</emphasis>
|
||
</taxonomicName>
|
||
, from Tamadaba, Gran Canaria (AIT). F.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFE5F8C0EA3F1E2A6" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[331,430,1911,1933]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="validoi" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE1FFDDFE5F8C0EA3F1E2A6" box="[331,430,1911,1933]" italics="true" pageId="5" pageNumber="40">N. validoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFE1FFDDFEA08C01A3B4E2A7" bold="true" box="[436,491,1911,1932]" pageId="5" pageNumber="40">
|
||
<taxonomicNameLabel id="20A8A302FFE1FFDDFEA08C01A3B4E2A7" box="[436,491,1911,1932]" pageId="5" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. G.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFD638C0EA0B2E2A6" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" box="[631,749,1911,1933]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="obesatus">
|
||
<emphasis id="3B9B1E79FFE1FFDDFD638C0EA0B2E2A6" box="[631,749,1911,1933]" italics="true" pageId="5" pageNumber="40">N. obesatus</emphasis>
|
||
</taxonomicName>
|
||
, from Barranco del Pagador, Gran Canaria (AIT). H.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFA1E8C0EA2EDE287" authorityName="Ibanez & Alonso" authorityYear="2009" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="bajamarensis">
|
||
<emphasis id="3B9B1E79FFE1FFDDFA1E8C0EA2EDE287" italics="true" pageId="5" pageNumber="40">N. bajamarensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype (from
|
||
<bibRefCitation id="6D7EBF9AFFE1FFDDFE768CE1A048E287" author="Yanes" box="[354,535,1942,1964]" etAl="et al." firstAuthor="Yanes" pageId="5" pageNumber="40" pagination="2179 - 2207" refId="ref9492" refString="Yanes, Y., Martin, J., Moro, L., Alonso, M. R. & Ibanez, M. (2009) On the relationships of the genus Napaeus (Gastropoda: Pulmonata: Enidae), with the description of four new species from the Canary Islands. Journal of Natural History, 43, 2179 - 2207." type="journal article" year="2009">
|
||
Yanes
|
||
<emphasis id="3B9B1E79FFE1FFDDFEB28CE1A38DE287" box="[422,466,1942,1964]" italics="true" pageId="5" pageNumber="40">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
). J.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFD568CE1A0C8E287" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[578,663,1942,1964]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="grohi" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE1FFDDFD568CE1A0C8E287" box="[578,663,1942,1964]" italics="true" pageId="5" pageNumber="40">N. grohi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFE1FFDDFD8A8CE0A088E280" bold="true" box="[670,727,1942,1963]" pageId="5" pageNumber="40">
|
||
<taxonomicNameLabel id="20A8A302FFE1FFDDFD8A8CE0A088E280" box="[670,727,1942,1963]" pageId="5" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. K.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFC788CE1A1ADE287" baseAuthorityName="Wollaston" baseAuthorityYear="1878" box="[876,1010,1943,1964]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="osoriensis">
|
||
<emphasis id="3B9B1E79FFE1FFDDFC788CE1A1ADE287" box="[876,1010,1943,1964]" italics="true" pageId="5" pageNumber="40">N. osoriensis</emphasis>
|
||
</taxonomicName>
|
||
, from Osorio, Gran Canaria (AIT). L.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFA908CE1A2AFE2E1" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="myosotis">
|
||
<emphasis id="3B9B1E79FFE1FFDDFA908CE1A2AFE2E1" italics="true" pageId="5" pageNumber="40">N. myosotis</emphasis>
|
||
</taxonomicName>
|
||
, from Montaña de Guía, Gran Canaria (AIT). M.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFDE28CC3A114E2E0" authorityName="Henriquez" authorityYear="1995" box="[758,843,1973,1995]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="exilis">
|
||
<emphasis id="3B9B1E79FFE1FFDDFDE28CC3A114E2E0" box="[758,843,1973,1995]" italics="true" pageId="5" pageNumber="40">N. exilis</emphasis>
|
||
</taxonomicName>
|
||
, from Altos de Gáldar, Gran Canaria (AIT). N.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFA298CC3A2BCE2C2" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="arinagaensis" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE1FFDDFA298CC3A2BCE2C2" italics="true" pageId="5" pageNumber="40">N. arinagaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFE1FFDDFFFD8CA2A37FE2C2" bold="true" box="[233,288,2004,2025]" pageId="5" pageNumber="40">
|
||
<taxonomicNameLabel id="20A8A302FFE1FFDDFFFD8CA2A37FE2C2" box="[233,288,2004,2025]" pageId="5" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype. O.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFEA58CA2A00EE2C2" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" box="[433,593,2004,2025]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="moquinianus">
|
||
<emphasis id="3B9B1E79FFE1FFDDFEA58CA2A00EE2C2" box="[433,593,2004,2025]" italics="true" pageId="5" pageNumber="40">N. moquinianus</emphasis>
|
||
</taxonomicName>
|
||
, from Brezal del Palmital, Gran Canaria (AIT). P.
|
||
<taxonomicName id="CEEFB9E8FFE1FFDDFB458CA2A6EEE2C2" authorityName="Groh & Ibanez" authorityYear="1992" box="[1105,1201,2003,2025]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="40" phylum="Mollusca" rank="species" species="isletae">
|
||
<emphasis id="3B9B1E79FFE1FFDDFB458CA2A6EEE2C2" box="[1105,1201,2003,2025]" italics="true" pageId="5" pageNumber="40">N. isletae</emphasis>
|
||
</taxonomicName>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5D9092E3FFE2FFDEFF838D19A752E38F" ID-DOI="http://doi.org/10.5281/zenodo.5284954" ID-Zenodo-Dep="5284954" httpUri="https://zenodo.org/record/5284954/files/figure.png" pageId="6" pageNumber="41" startId="6.[151,248,1647,1670]" targetBox="[151,1436,177,1626]" targetPageId="6">
|
||
<paragraph id="0950C26BFFE2FFDEFF838D19A752E38F" blockId="6.[151,1437,1647,1700]" pageId="6" pageNumber="41">
|
||
<emphasis id="3B9B1E79FFE2FFDEFF838D19A350E3AD" bold="true" box="[151,271,1647,1670]" pageId="6" pageNumber="41">FIGURE 4.</emphasis>
|
||
<taxonomicName id="CEEFB9E8FFE2FFDEFE008D06A3FDE3AE" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[276,418,1648,1669]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="41" phylum="Mollusca" rank="species" species="josei" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE2FFDEFE008D06A3FDE3AE" box="[276,418,1648,1669]" italics="true" pageId="6" pageNumber="41">Napaeus josei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFE2FFDEFEBC8D19A381E3AF" bold="true" box="[424,478,1647,1668]" pageId="6" pageNumber="41">
|
||
<taxonomicNameLabel id="20A8A302FFE2FFDEFEBC8D19A381E3AF" box="[424,478,1647,1668]" pageId="6" pageNumber="41" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Paratype from the type locality (AIT). B. Three paratypes from the type locality, attached to a stone and disguised with a soil covering (JSGC). C. Disguised paratype with a very thick soil covering (JSGC).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="41F591E0FFE2FFDEFFD18C2DA0F1E2CE" pageId="6" pageNumber="41" type="diagnosis">
|
||
<paragraph id="0950C26BFFE2FFDEFFD18C2DA0F1E2CE" blockId="6.[151,1436,1739,2021]" pageId="6" pageNumber="41">
|
||
<emphasis id="3B9B1E79FFE2FFDEFFD18C2DA364E25F" bold="true" box="[197,315,1883,1908]" inLineHeading="true" pageId="6" pageNumber="41" reason="1">Diagnosis.</emphasis>
|
||
Shell very obese, conic-ovate, brown-coloured, with first whorls slightly darker than rest. Teleoconch granulated. Penis short and thick, with large penial papilla. System of penial appendix parts A
|
||
<subScript id="956BC02EFFE2FFDEFBFC8CF8A6AFE2B7" attach="left" box="[1256,1264,1934,1948]" fontSize="6" pageId="6" pageNumber="41">1</subScript>
|
||
–A
|
||
<subScript id="956BC02EFFE2FFDEFA068CF8A745E2B7" attach="left" box="[1298,1306,1934,1948]" fontSize="6" pageId="6" pageNumber="41">3</subScript>
|
||
very short. Appendicular retractor muscle inserts laterally in proximal tip of part A
|
||
<subScript id="956BC02EFFE2FFDEFCA18CC3A1E2E2E8" attach="left" box="[949,957,1973,1987]" fontSize="6" pageId="6" pageNumber="41">3</subScript>
|
||
. Epiphallus with epiphallic caecum. Bursa copulatrix duct inflated, with long diverticulum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFE3FFD0FFD18BE1A3B1E3A5" lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42" type="description">
|
||
<paragraph id="0950C26BFFE3FFDFFFD18BE1A74EE4D8" blockId="7.[151,1436,151,644]" pageId="7" pageNumber="42">
|
||
<emphasis id="3B9B1E79FFE3FFDFFFD18BE1A308E59B" bold="true" box="[197,343,151,176]" inLineHeading="true" pageId="7" pageNumber="42" reason="1">Description.</emphasis>
|
||
Body pale brown-greyish, covered with brown spots, denser in cephalic area (
|
||
<figureCitation id="91D4DEEEFFE3FFDFFBF28BE1A713E584" box="[1254,1356,151,175]" captionStart="FIGURE 4" captionStartId="6.[151,248,1647,1670]" captionTargetBox="[151,1436,177,1626]" captionTargetId="figure-181@6.[151,1436,175,1626]" captionTargetPageId="6" captionText="FIGURE 4. Napaeus josei n. sp. A. Paratype from the type locality (AIT). B. Three paratypes from the type locality, attached to a stone and disguised with a soil covering (JSGC). C. Disguised paratype with a very thick soil covering (JSGC)." figureDoi="http://doi.org/10.5281/zenodo.5284954" httpUri="https://zenodo.org/record/5284954/files/figure.png" pageId="7" pageNumber="42">Fig. 4 A</figureCitation>
|
||
). Shell (
|
||
<figureCitation id="91D4DEEEFFE3FFDFFF8B8BCAA35EE5FF" box="[159,257,188,212]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="7" pageNumber="42">Fig. 3 A</figureCitation>
|
||
) dextral, robust, very obese (SB/SH index), conic-ovate, with sharp apex, irregularly increasing whorls (penultimate whorl and body whorl giving oval appearance), 5¾–6 convex whorls and deeply marked suture. Body whorl large (SB/SH index) occupying more than ⅔ of shell surface area (BS/SS index). Protoconch smooth, with 1¾–2 whorls. Aperture long, very wide (AH/SH and AB/SB indices), rounded at palatal side, more curved at junction of columellar and palatal edges. Angle (
|
||
<figureCitation id="91D4DEEEFFE3FFDFFD9F8A3AA093E44F" box="[651,716,332,356]" captionStart="FIGURE 2" captionStartId="4.[151,248,908,931]" captionTargetBox="[158,1428,208,867]" captionTargetId="figure-435@4.[151,1436,193,887]" captionTargetPageId="4" captionText="FIGURE 2. Drawings of the shell of the holotype of Napaeus arinagaensis n. sp., showing the placement of the measurements obtained (in º, 2 mm or mm). α, upper palatal angle (i.e. the angle between the columella and the upper palatal margin of the aperture); AB, aperture breadth; AH, aperture height; AP, aperture perimeter; AS, aperture surface area (plane view); BH, body whorl height (at columella level); BP, body whorl perimeter; BS, body whorl surface area (plane view); FB, first whorls breadth; FH, first whorls height; FP, first whorls perimeter; FS, first whorls surface area (plane view); PB, penultimate whorl breadth; PH, penultimate whorl height; PP, penultimate whorl perimeter; PS, penultimate whorl surface area (plane view); SB, shell breadth; SH, shell height; SP, shell perimeter; SS, shell surface area (plane view); T, distance between columella and periphery of penultimate whorl above aperture; U, highest distance between columella and palatal edge of aperture." figureDoi="http://doi.org/10.5281/zenodo.5284944" httpUri="https://zenodo.org/record/5284944/files/figure.png" pageId="7" pageNumber="42">Fig. 2</figureCitation>
|
||
) between columella and upper palatal margin about 122º. Peristome discontinuous, expanded as a wide, whitish lip, more developed in lower part of the palatal and columellar edges, reflected on columellar edge, where it partly covers umbilical slit. Older specimens with a callosity between peristome edges and a nodule at junction of parietal and palatal margins. Aperture extending laterally beyond penultimate whorl by about 32% (
|
||
<emphasis id="3B9B1E79FFE3FFDFFD028AABA06FE4DF" box="[534,560,477,500]" italics="true" pageId="7" pageNumber="42">ca</emphasis>
|
||
.
|
||
<quantity id="CE176F8EFFE3FFDFFD2B8AAAA0DBE4DF" box="[575,644,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="42" unit="mm" value="1.0">1 mm</quantity>
|
||
). Colour brown, with first whorls slightly darker than rest.
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFE3FFDFFFD18A89A765E7AF" blockId="7.[151,1436,151,644]" pageId="7" pageNumber="42">
|
||
Ornamentation of first teleoconch whorl characterised by numerous, thin, radial, oblique ribs with small, irregular protuberances. Protuberances from second teleoconch whorl to body whorl more developed, and those of penultimate whorl (
|
||
<figureCitation id="91D4DEEEFFE3FFDFFFF28931A31DE774" box="[230,322,583,607]" captionStart="FIGURE 5" captionStartId="7.[151,248,1552,1575]" captionTargetBox="[151,1436,709,1528]" captionTargetId="figure-451@7.[151,1436,709,1528]" captionTargetPageId="7" captionText="FIGURE 5. Details of shell ornamentation of holotypes (A, C: penultimate whorl; B, D: body whorl). A–B. Napaeus josei n. sp. C–D. N. venegueraensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5284956" httpUri="https://zenodo.org/record/5284956/files/figure.png" pageId="7" pageNumber="42">Fig. 5 A</figureCitation>
|
||
) broader and usually equidistantly placed, giving appearance of granular rows perpendicular to radial shell ribs. Body whorl mainly with radial ribs provided with irregular granulation on the upper side (
|
||
<figureCitation id="91D4DEEEFFE3FFDFFBC6891AA772E7AF" box="[1234,1325,620,644]" captionStart="FIGURE 5" captionStartId="7.[151,248,1552,1575]" captionTargetBox="[151,1436,709,1528]" captionTargetId="figure-451@7.[151,1436,709,1528]" captionTargetPageId="7" captionText="FIGURE 5. Details of shell ornamentation of holotypes (A, C: penultimate whorl; B, D: body whorl). A–B. Napaeus josei n. sp. C–D. N. venegueraensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5284956" httpUri="https://zenodo.org/record/5284956/files/figure.png" pageId="7" pageNumber="42">Fig. 5 B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="5D9092E3FFE3FFDFFF838D66A394E36F" ID-DOI="http://doi.org/10.5281/zenodo.5284956" ID-Zenodo-Dep="5284956" httpUri="https://zenodo.org/record/5284956/files/figure.png" pageId="7" pageNumber="42" startId="7.[151,248,1552,1575]" targetBox="[151,1436,709,1528]" targetPageId="7">
|
||
<paragraph id="0950C26BFFE3FFDFFF838D66A394E36F" blockId="7.[151,1437,1552,1605]" pageId="7" pageNumber="42">
|
||
<emphasis id="3B9B1E79FFE3FFDFFF838D66A350E30D" bold="true" box="[151,271,1552,1575]" pageId="7" pageNumber="42">FIGURE 5.</emphasis>
|
||
Details of shell ornamentation of holotypes (A, C: penultimate whorl; B, D: body whorl). A–B.
|
||
<taxonomicName id="CEEFB9E8FFE3FFDFFBC48D67A700E30D" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[1232,1375,1553,1574]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="42" phylum="Mollusca" rank="species" species="josei" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE3FFDFFBC48D67A700E30D" box="[1232,1375,1553,1574]" italics="true" pageId="7" pageNumber="42">Napaeus josei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFE3FFDFFA718D67A7C2E30D" bold="true" box="[1381,1437,1553,1574]" pageId="7" pageNumber="42">
|
||
<taxonomicNameLabel id="20A8A302FFE3FFDFFA718D67A7C2E30D" box="[1381,1437,1553,1574]" pageId="7" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
C–D.
|
||
<taxonomicName id="CEEFB9E8FFE3FFDFFFC78D46A3D2E36E" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[211,397,1584,1605]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="42" phylum="Mollusca" rank="species" species="venegueraensis" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE3FFDFFFC78D46A3D2E36E" box="[211,397,1584,1605]" italics="true" pageId="7" pageNumber="42">N. venegueraensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="20A8A302FFE3FFDFFE878D59A394E36F" box="[403,459,1583,1604]" pageId="7" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0950C26BFFE3FFDFFFD18D0DA335E2F4" blockId="7.[151,1437,1659,2015]" pageId="7" pageNumber="42">
|
||
Genital system (
|
||
<figureCitation id="91D4DEEEFFE3FFDFFE688D0DA056E3B9" box="[380,521,1659,1683]" captionStart="FIGURE 6" captionStartId="8.[151,248,1204,1227]" captionTargetBox="[151,1436,287,1187]" captionTargetId="figure-398@8.[151,1436,287,1187]" captionTargetPageId="8" captionText="FIGURE 6. Napaeus josei n. sp., paratype from Barranco del Lechugal, Gran Canaria (AIT). A. Genital system. B. internal anatomy of penis, showing the penial papilla; A1–A5, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; d, diverticulum; e, epiphallus; ec, epiphallic caecum; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; p, penis; pp, penial papilla; pr, penis retractor; pw, penis wall; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5284960" httpUri="https://zenodo.org/record/5284960/files/figure.png" pageId="7" pageNumber="42">Figs. 6 A, B</figureCitation>
|
||
;
|
||
<specimenCount id="1FE909E2FFE3FFDFFD028D0DA0E2E3B8" box="[534,701,1659,1683]" pageId="7" pageNumber="42" type="generic">two specimens</specimenCount>
|
||
dissected). Atrium very short. Penis short and thick, without evident division into separate portions and about a third of the length of the epiphallus. Penial papilla large, welldeveloped, occupying almost all of penial cavity. Penial retractor muscle inserts apically into proximal penis tip, next to epiphallus-penis connection. Epiphallus opening distally into penis, tubular, slender, with epiphallic caecum inserting in its proximal third. Flagellum very short, globular. Vas deferens opens sub-terminally on proximal end of epiphallus. Penial appendix long, arising obliquely in distal portion of penis near atrium, longer than penis and epiphallus together. System of penial appendix parts A
|
||
<subScript id="956BC02EFFE3FFDFFC1E8C14A14DE25B" attach="left" box="[778,786,1890,1904]" fontSize="6" pageId="7" pageNumber="42">1</subScript>
|
||
–A
|
||
<subScript id="956BC02EFFE3FFDFFC208C14A163E25B" attach="left" box="[820,828,1890,1904]" fontSize="6" pageId="7" pageNumber="42">3</subScript>
|
||
very short, boundary between them scarcely visible. Parts A
|
||
<subScript id="956BC02EFFE3FFDFFFFE8CFFA2ADE2BC" attach="left" box="[234,242,1929,1943]" fontSize="6" pageId="7" pageNumber="42">3</subScript>
|
||
and A
|
||
<subScript id="956BC02EFFE3FFDFFE2F8CFFA31CE2BC" attach="left" box="[315,323,1929,1943]" fontSize="6" pageId="7" pageNumber="42">4</subScript>
|
||
clearly separated from each other, A
|
||
<subScript id="956BC02EFFE3FFDFFDCA8CFFA0B9E2BC" attach="left" box="[734,742,1929,1943]" fontSize="6" pageId="7" pageNumber="42">4</subScript>
|
||
long and thin, passing gradually into expanded A
|
||
<subScript id="956BC02EFFE3FFDFFA048CFFA747E2BC" attach="left" box="[1296,1304,1929,1943]" fontSize="6" pageId="7" pageNumber="42">5</subScript>
|
||
. Appendicular retractor muscle inserts laterally in proximal tip of short part A
|
||
<subScript id="956BC02EFFE3FFDFFC848CC6A1C7E295" attach="left" box="[912,920,1968,1982]" fontSize="6" pageId="7" pageNumber="42">3</subScript>
|
||
, joined to penial retractor just before insertion on lower lung wall.
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFECFFD0FFD18BE1A3CDE5DC" blockId="8.[151,1437,151,247]" pageId="8" pageNumber="43">Vagina tubiform, similar in length to free oviduct and proximal third of epiphallus, respectively. Bursa copulatrix duct inflated, with diverticulum longer than duct and branching off in proximal quarter of duct, surmounting the vesicle of the bursa copulatrix.</paragraph>
|
||
<caption id="5D9092E3FFECFFD0FF838FC2A1B9E002" ID-DOI="http://doi.org/10.5281/zenodo.5284960" ID-Zenodo-Dep="5284960" httpUri="https://zenodo.org/record/5284960/files/figure.png" pageId="8" pageNumber="43" startId="8.[151,248,1204,1227]" targetBox="[151,1436,287,1187]" targetPageId="8">
|
||
<paragraph id="0950C26BFFECFFD0FF838FC2A1B9E002" blockId="8.[151,1436,1204,1321]" pageId="8" pageNumber="43">
|
||
<emphasis id="3B9B1E79FFECFFD0FF838FC2A34EE1E0" bold="true" box="[151,273,1204,1227]" pageId="8" pageNumber="43">FIGURE 6.</emphasis>
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FE0D8FC0A3F4E1E0" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[281,427,1206,1227]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="josei" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFECFFD0FE0D8FC0A3F4E1E0" box="[281,427,1206,1227]" italics="true" pageId="8" pageNumber="43">Napaeus josei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFECFFD0FEA78FC3A3B3E1E1" bold="true" box="[435,492,1205,1226]" pageId="8" pageNumber="43">
|
||
<taxonomicNameLabel id="20A8A302FFECFFD0FEA78FC3A3B3E1E1" box="[435,492,1205,1226]" pageId="8" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype from Barranco del Lechugal, Gran Canaria (AIT). A. Genital system. B. internal anatomy of penis, showing the penial papilla; A
|
||
<subScript id="956BC02EFFECFFD0FD9E8F97A0CEE1C6" attach="left" box="[650,657,1249,1261]" fontSize="5" pageId="8" pageNumber="43">1</subScript>
|
||
–A
|
||
<subScript id="956BC02EFFECFFD0FDBB8F97A0E9E1C6" attach="left" box="[687,694,1249,1261]" fontSize="5" pageId="8" pageNumber="43">5</subScript>
|
||
, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; d, diverticulum; e, epiphallus; ec, epiphallic caecum; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; p, penis; pp, penial papilla; pr, penis retractor; pw, penis wall; v, vagina; vd, vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0950C26BFFECFFD0FFD18E23A13AE36D" blockId="8.[151,1436,1365,1678]" pageId="8" pageNumber="43">
|
||
<emphasis id="3B9B1E79FFECFFD0FFD18E23A041E045" bold="true" box="[197,542,1365,1390]" inLineHeading="true" pageId="8" pageNumber="43" reason="1">Comparison with other taxa.</emphasis>
|
||
Shell morphology: The shell of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FC908E21A187E045" authority="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez, 2011" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[900,984,1367,1390]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="josei" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFECFFD0FC908E21A187E045" box="[900,984,1367,1390]" italics="true" pageId="8" pageNumber="43">N. josei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="3B9B1E79FFECFFD0FCCB8E20A643E045" bold="true" box="[991,1052,1366,1390]" pageId="8" pageNumber="43">
|
||
<taxonomicNameLabel id="20A8A302FFECFFD0FCCB8E20A643E045" box="[991,1052,1366,1390]" pageId="8" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="91D4DEEEFFECFFD0FB388E20A6D3E045" box="[1068,1164,1366,1390]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Fig. 3 A</figureCitation>
|
||
) is comparable in shape and size mainly to that of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FED58E0DA16BE0B9" authority="(Shuttleworth, 1852)" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[449,820,1402,1426]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="nanodes">
|
||
<emphasis id="3B9B1E79FFECFFD0FED58E0DA01FE0B9" box="[449,576,1402,1426]" italics="true" pageId="8" pageNumber="43">N. nanodes</emphasis>
|
||
(
|
||
<bibRefCitation id="6D7EBF9AFFECFFD0FD458E0CA174E0B9" author="Shuttleworth" box="[593,811,1402,1426]" firstAuthor="Shuttleworth" pageId="8" pageNumber="43" pagination="289 - 304" refId="ref9300" refString="Shuttleworth, R. J. (1852) Diagnosen neuer Mollusken. Mittheilungen der naturforschenden Gesellschaft in Bern, 260 / 261, 289 - 304." type="journal article" year="1852">Shuttleworth, 1852</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFECFFD0FC508E0CA1FCE0B9" box="[836,931,1402,1426]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Fig. 3 B</figureCitation>
|
||
) and
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FCF18E0DA7C3E0B9" authority="(Webb & Berthelot, 1833)" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" box="[997,1436,1402,1426]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="3B9B1E79FFECFFD0FCF18E0DA635E0B9" box="[997,1130,1402,1426]" italics="true" pageId="8" pageNumber="43">N. badiosus</emphasis>
|
||
(Webb & Berthelot, 1833)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFECFFD0FF8B8EE8A2A3E09D" box="[159,252,1438,1462]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Fig. 3 C</figureCitation>
|
||
), both from Tenerife. The shell of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FD6E8EE9A092E09D" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[634,717,1439,1462]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="josei">
|
||
<emphasis id="3B9B1E79FFECFFD0FD6E8EE9A092E09D" box="[634,717,1439,1462]" italics="true" pageId="8" pageNumber="43">N. josei</emphasis>
|
||
</taxonomicName>
|
||
is clearly more obese than that of both species (
|
||
<figureCitation id="91D4DEEEFFECFFD0FBDA8EE8A706E09D" box="[1230,1369,1438,1462]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Figs. 3 A–C</figureCitation>
|
||
,
|
||
<figureCitation id="91D4DEEEFFECFFD0FA778EE9A7D0E09D" box="[1379,1423,1438,1462]" captionStart="FIGURE 7" captionStartId="9.[151,248,1358,1381]" captionTargetBox="[171,1416,198,1320]" captionTargetId="figure-301@9.[151,1436,193,1337]" captionTargetPageId="9" captionText="FIGURE 7. Scatter plots of some shell measurements for the new species and the most similar species. AS, aperture surface area (plane view); BS, body whorl surface area (plane view); FS, first whorls surface area (plane view); PS, penultimate whorl surface area (plane view); SB, shell breadth; SH, shell height; SS, shell surface area (plane view). The dimensions are in mm (lengths) or mm2 (surface areas)." figureDoi="http://doi.org/10.5281/zenodo.5284962" httpUri="https://zenodo.org/record/5284962/files/figure.png" pageId="8" pageNumber="43">7 A</figureCitation>
|
||
). The shell aperture of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FE928EB5A386E0F1" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[390,473,1475,1498]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="josei">
|
||
<emphasis id="3B9B1E79FFECFFD0FE928EB5A386E0F1" box="[390,473,1475,1498]" italics="true" pageId="8" pageNumber="43">N. josei</emphasis>
|
||
</taxonomicName>
|
||
is similar to that of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FDA58EB5A16BE0F1" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" box="[689,820,1474,1498]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="3B9B1E79FFECFFD0FDA58EB5A16BE0F1" box="[689,820,1474,1498]" italics="true" pageId="8" pageNumber="43">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
and bigger than that of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FB548EB5A6E2E0F1" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[1088,1213,1474,1498]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="nanodes">
|
||
<emphasis id="3B9B1E79FFECFFD0FB548EB5A6E2E0F1" box="[1088,1213,1474,1498]" italics="true" pageId="8" pageNumber="43">N. nanodes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFECFFD0FBD98EB4A707E0F1" box="[1229,1368,1474,1498]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Figs. 3 A, B</figureCitation>
|
||
,
|
||
<figureCitation id="91D4DEEEFFECFFD0FA708EB5A7D3E0F1" box="[1380,1420,1474,1498]" captionStart="FIGURE 7" captionStartId="9.[151,248,1358,1381]" captionTargetBox="[171,1416,198,1320]" captionTargetId="figure-301@9.[151,1436,193,1337]" captionTargetPageId="9" captionText="FIGURE 7. Scatter plots of some shell measurements for the new species and the most similar species. AS, aperture surface area (plane view); BS, body whorl surface area (plane view); FS, first whorls surface area (plane view); PS, penultimate whorl surface area (plane view); SB, shell breadth; SH, shell height; SS, shell surface area (plane view). The dimensions are in mm (lengths) or mm2 (surface areas)." figureDoi="http://doi.org/10.5281/zenodo.5284962" httpUri="https://zenodo.org/record/5284962/files/figure.png" pageId="8" pageNumber="43">7 C</figureCitation>
|
||
). The shell ornamentation is clearly different in the three species, the granulation of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FB388E91A6DEE0D5" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[1068,1153,1511,1534]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="josei">
|
||
<emphasis id="3B9B1E79FFECFFD0FB388E91A6DEE0D5" box="[1068,1153,1511,1534]" italics="true" pageId="8" pageNumber="43">N. josei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFECFFD0FB858E90A6A0E0D5" box="[1169,1279,1510,1534]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Figs. 3 A</figureCitation>
|
||
,
|
||
<figureCitation id="91D4DEEEFFECFFD0FA188E90A706E0D5" box="[1292,1369,1510,1534]" captionStart="FIGURE 5" captionStartId="7.[151,248,1552,1575]" captionTargetBox="[151,1436,709,1528]" captionTargetId="figure-451@7.[151,1436,709,1528]" captionTargetPageId="7" captionText="FIGURE 5. Details of shell ornamentation of holotypes (A, C: penultimate whorl; B, D: body whorl). A–B. Napaeus josei n. sp. C–D. N. venegueraensis n. sp." figureDoi="http://doi.org/10.5281/zenodo.5284956" httpUri="https://zenodo.org/record/5284956/files/figure.png" pageId="8" pageNumber="43">5 A, B</figureCitation>
|
||
) contrasting with the sinuous, undulating ribs surrounding large depressions in
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FCD68D7DA660E309" baseAuthorityName="Shuttleworth" baseAuthorityYear="1852" box="[962,1087,1546,1570]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="nanodes">
|
||
<emphasis id="3B9B1E79FFECFFD0FCD68D7DA660E309" box="[962,1087,1546,1570]" italics="true" pageId="8" pageNumber="43">N. nanodes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFECFFD0FB5B8D7CA6F4E309" box="[1103,1195,1546,1570]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Fig. 3 B</figureCitation>
|
||
) and the more or less uniformly ribbed, not granulated shell of
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FD718D59A0B6E36D" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" box="[613,745,1582,1606]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="3B9B1E79FFECFFD0FD718D59A0B6E36D" box="[613,745,1582,1606]" italics="true" pageId="8" pageNumber="43">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFECFFD0FDED8D58A107E36D" box="[761,856,1582,1606]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="8" pageNumber="43">Fig. 3 C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFECFFD0FFD18D24A3B1E3A5" blockId="8.[151,1436,1365,1678]" pageId="8" pageNumber="43">
|
||
Genital anatomy:
|
||
<taxonomicName id="CEEFB9E8FFECFFD0FE938D25A384E341" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[391,475,1619,1642]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="josei">
|
||
<emphasis id="3B9B1E79FFECFFD0FE938D25A384E341" box="[391,475,1619,1642]" italics="true" pageId="8" pageNumber="43">N. josei</emphasis>
|
||
</taxonomicName>
|
||
has an enormous penial papilla (
|
||
<figureCitation id="91D4DEEEFFECFFD0FC5A8D24A1F2E341" box="[846,941,1618,1642]" captionStart="FIGURE 6" captionStartId="8.[151,248,1204,1227]" captionTargetBox="[151,1436,287,1187]" captionTargetId="figure-398@8.[151,1436,287,1187]" captionTargetPageId="8" captionText="FIGURE 6. Napaeus josei n. sp., paratype from Barranco del Lechugal, Gran Canaria (AIT). A. Genital system. B. internal anatomy of penis, showing the penial papilla; A1–A5, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix; d, diverticulum; e, epiphallus; ec, epiphallic caecum; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; p, penis; pp, penial papilla; pr, penis retractor; pw, penis wall; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5284960" httpUri="https://zenodo.org/record/5284960/files/figure.png" pageId="8" pageNumber="43">Fig. 6 B</figureCitation>
|
||
), the largest known in the genus, occupying almost the entire penial cavity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |