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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1088.77703" ID-GBIF-Dataset="da371969-c24a-4799-a47c-debfa77ad39a" ID-PMC="PMC8917117" ID-Pensoft-Pub="1313-2970-1088-41" ID-Pensoft-UUID="D9743E2046515A2BB20DE3FE0D5FE9E1" ID-PubMed="35437366" ID-ZooBank="41B17D93ABBB49F3B15AB4F24D18753F" ModsDocID="1313-2970-1088-41" checkinTime="1646454884193" checkinUser="pensoft" docAuthor="Liu 1, Xiaoshuang, Li *, Yixuan &amp; Di, Zhiyong" docDate="2022" docId="60ED176E34165BCE9AFD79035CE69070" docLanguage="en" docName="ZooKeys 1088: 41-52" docOrigin="ZooKeys 1088" docPubDate="2022-03-04" docSource="http://dx.doi.org/10.3897/zookeys.1088.77703" docTitle="Otostigmus (O.) beroni Lewis 2001" docType="treatment" docVersion="5" id="D9743E2046515A2BB20DE3FE0D5FE9E1" lastPageNumber="41" masterDocId="D9743E2046515A2BB20DE3FE0D5FE9E1" masterDocTitle="The revalidation of Otostigmus (O.) lewisi Song et al., 2005 (Scolopendromorpha, Scolopendridae) based on new material from Jiacha County, China" masterLastPageNumber="52" masterPageNumber="41" pageNumber="41" updateTime="1668151526590" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The revalidation of Otostigmus (O.) lewisi Song et al., 2005 (Scolopendromorpha, Scolopendridae) based on new material from Jiacha County, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu 1, Xiaoshuang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Application of Hebei Province, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China &amp; Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li *, Yixuan</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Application of Hebei Province, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China &amp; Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Di, Zhiyong</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Application of Hebei Province, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China &amp; Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
<mods:nameIdentifier type="email">zydi@ustc.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1088</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1088.77703</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1088.77703</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1088-41</mods:identifier>
<mods:identifier type="ZooBank">41B17D93ABBB49F3B15AB4F24D18753F</mods:identifier>
<mods:identifier type="Pensoft-UUID">D9743E2046515A2BB20DE3FE0D5FE9E1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="193874018" LSID="urn:lsid:plazi:treatment:60ED176E34165BCE9AFD79035CE69070" httpUri="http://treatment.plazi.org/id/60ED176E34165BCE9AFD79035CE69070" lastPageNumber="41" pageId="0" pageNumber="41">
<subSubSection pageId="0" pageNumber="41" type="nomenclature">
<paragraph pageId="0" pageNumber="41">
<taxonomicName LSID="60ED176E-3416-5BCE-9AFD-79035CE69070" authority="Lewis, 2001" authorityName="Lewis" authorityYear="2001" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otostigmus (O.) beroni" order="Scolopendromorpha" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="beroni" subGenus="O.">Otostigmus (O.) beroni Lewis, 2001</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">Figs 1</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A habitat of O. (O.) beroni in Jilong Town, Jilong County B habitat of O. (O.) lewisi in Anrao Town, Jiacha County C O. (O.) beroni (Ar. - MHBU-SoJL 21080101) D O. (O.) lewisi (Ar. - MHBU-SoJC 1608 DX 01)." figureDoi="10.3897/zookeys.1088.77703.figure4" httpUri="https://binary.pensoft.net/fig/653522" pageId="0" pageNumber="41">, 4C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="reference_group">
<paragraph pageId="0" pageNumber="41">
<taxonomicName authorityName="Lewis" authorityYear="2001" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otostigmus beroni" order="Scolopendromorpha" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="beroni">Otostigmus beroni</taxonomicName>
Lewis, 2001: 22;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2579.1.1" author="Lewis, JGE" journalOrPublisher="Zootaxa" pageId="0" pageNumber="41" pagination="1 - 29" refId="B4" refString="Lewis, JGE, 2010. A revision of the rugulosus group of Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae). Zootaxa 2579: 1 - 29, DOI: https://doi.org/10.11646/zootaxa.2579.1.1" title="A revision of the rugulosus group of Otostigmus subgenus Otostigmus Porat, 1876 (Chilopoda: Scolopendromorpha: Scolopendridae)." url="https://doi.org/10.11646/zootaxa.2579.1.1" volume="2579" year="2010">Lewis 2010</bibRefCitation>
: 26.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="materials_examined">
<paragraph pageId="0" pageNumber="41">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="41">
<materialsCitation collectingDate="2016-08-05" collectingDateMax="2021-08-01" collectingDateMin="2016-08-05" collectorName="Zhiyong Di. Ar., Zhiyong Di. Housed" country="China" county="Jilong (Gyirong) County" latitude="28.435" location="Jilong Town" longLatPrecision="6" longitude="85.257" municipality="Jilong (Gyirong) County" specimenCount="1" stateProvince="Xizang">
Ar.-MHBU-SoJL1608050101-Ar.-MHBU-SoJL1608050120:
<location LSID="urn:lsid:plazi:treatment:60ED176E34165BCE9AFD79035CE69070:590054A46766BF643576C401A72CE50C" country="China" county="Jilong (Gyirong) County" latitude="28.435" longLatPrecision="6" longitude="85.257" municipality="Jilong (Gyirong) County" name="Jilong Town" stateProvince="Xizang">Jilong Town</location>
,
<collectingCounty>Jilong (Gyirong) County</collectingCounty>
,
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
(
<collectingRegion country="China" name="Xizang">Tibet</collectingRegion>
),
<collectingCountry name="China">China</collectingCountry>
,
<geoCoordinate degrees="28.4370" direction="north" orientation="latitude" precision="5" value="28.437">28.4370°N</geoCoordinate>
,
<geoCoordinate degrees="85.2568" direction="east" orientation="longitude" precision="5" value="85.2568">85.2568°E</geoCoordinate>
,
<collectingDate value="2016-08-05">5/8/2016</collectingDate>
, leg.
<collectorName>Zhiyong Di. Ar.</collectorName>
-MHBU-SoJL21080101:
<location LSID="urn:lsid:plazi:treatment:60ED176E34165BCE9AFD79035CE69070:7D81460BB980B858168221A39B704411" country="China" county="Jilong (Gyirong) County" latitude="28.435" longLatPrecision="6" longitude="85.257" municipality="Jilong (Gyirong) County" name="Jilong Town" stateProvince="Xizang">Jilong Town</location>
,
<collectingMunicipality>Jilong (Gyirong) County</collectingMunicipality>
,
<collectingRegion country="China" name="Xizang">Xizang</collectingRegion>
(
<collectingRegion country="China" name="Xizang">Tibet</collectingRegion>
),
<collectingCountry name="China">China</collectingCountry>
,
<geoCoordinate degrees="28.4350" direction="north" orientation="latitude" precision="5" value="28.435">28.4350°N</geoCoordinate>
,
<geoCoordinate degrees="85.2570" direction="east" orientation="longitude" precision="5" value="85.257">85.2570°E</geoCoordinate>
,
<collectingDate value="2021-08-01">1/8/2021</collectingDate>
, leg.
<collectorName>Zhiyong Di. Housed</collectorName>
in MHBU
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="diagnosis">
<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="41">
Maximum length 57 mm. Antennae 18 articles, the basal 2.25 to 2.40 glabrous dorsally (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1C, E</figureCitation>
). With 4 or 5 teeth (rarely 3) on each tooth plate, the median 3 (or 4) ones more or less fused (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1D, E</figureCitation>
). Sternites with incomplete paramedian sutures occupying at least anterior 66 to 100% in mid and hind body region (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1G</figureCitation>
). Coxopleural process long, typically with 2 apical spines, 1-2 lateral spines and 1-2 dorsal spines (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1I</figureCitation>
). Ultimate prefemur with 4 rows of prominent spines, disposed on swollen bases (10-16 spines in total) (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1H, I</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 1.</emphasis>
<taxonomicName authorityName="Lewis" authorityYear="2001" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otostigmus (O.) beroni" order="Scolopendromorpha" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="beroni" subGenus="O.">Otostigmus (O.) beroni</taxonomicName>
Lewis, 2001
<emphasis bold="true" pageId="0" pageNumber="41">A, B, G</emphasis>
Ar.-MHBU-SoJL1608050101
<emphasis bold="true" pageId="0" pageNumber="41">C, D, F, H, I</emphasis>
Ar.-MHBU-SoJL1608050104
<emphasis bold="true" pageId="0" pageNumber="41">E</emphasis>
Ar.-MHBU-SoJL1608050105
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
cephalic plate, antennae, tergite 1 and legs 1-2, dorsal view
<emphasis bold="true" pageId="0" pageNumber="41">D, E</emphasis>
ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate
<emphasis bold="true" pageId="0" pageNumber="41">F</emphasis>
tergites 16-19
<emphasis bold="true" pageId="0" pageNumber="41">G</emphasis>
sternites 11-14
<emphasis bold="true" pageId="0" pageNumber="41">H</emphasis>
last two segments and ultimate legs, dorsal view
<emphasis bold="true" pageId="0" pageNumber="41">I</emphasis>
last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (
<emphasis bold="true" pageId="0" pageNumber="41">A, B</emphasis>
); 2.0 mm (
<emphasis bold="true" pageId="0" pageNumber="41">C, F-I</emphasis>
); 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="41">D, E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Description</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">
(Ar.-MHBU-SoJL1608050101).
<emphasis italics="true" pageId="0" pageNumber="41">Length</emphasis>
</emphasis>
: 54 mm (measured from anterior margin of cephalic plate to posterior margin of tergite 21).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Pigmentation</emphasis>
(after remaining in alcohol for five years): body color yellow-brown; legs and antennae yellow; cephalic plate, tergites 1-2, tergites 20-21 and penultimate and ultimate legs blue-green (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1A, B</figureCitation>
). Live individual (Ar.-MHBU-SoJL21080101): cephalic plate blue-green, tergites blue-brown, penultimate legs and ultimate legs yellow with blue middle part in each segment, the other legs yellow (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A habitat of O. (O.) beroni in Jilong Town, Jilong County B habitat of O. (O.) lewisi in Anrao Town, Jiacha County C O. (O.) beroni (Ar. - MHBU-SoJL 21080101) D O. (O.) lewisi (Ar. - MHBU-SoJC 1608 DX 01)." figureDoi="10.3897/zookeys.1088.77703.figure4" httpUri="https://binary.pensoft.net/fig/653522" pageId="0" pageNumber="41">4C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Cephalic plate</emphasis>
: wider than long, rounded anteriorly, without sutures or sulci (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Antennae</emphasis>
: 18 articles, basal 2.3 glabrous, the remainder are covered with short hairs (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1C, E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Forcipular segment</emphasis>
: tooth plates with 4 teeth in each plate, the median 3 fused. Process of forcipular trochanteroprefemur with 3 teeth (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Tergites (T)</emphasis>
: with complete paramedian sutures from 3 to 20, marginate from 7 to 21 (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1F</figureCitation>
). Central part of the posterior border of T21 slightly convex (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1H</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Sternites (S)</emphasis>
: smooth, with incomplete paramedian sutures from 5 to 19 (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1G</figureCitation>
); S21 with a slight median longitudinal depression and converging posteriorly (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1I</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Coxopleuron</emphasis>
: with numerous small pores, coxopleural process moderately long and apical border with protuberance (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1I</figureCitation>
). With pore-free longitudinal strips within pore field. Coxopleural process with 2 apical spines, 2 lateral spines, and 1-2 dorsal spines.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Legs (L)</emphasis>
: L1-16 and L18-19 with 2 tarsal spurs, L1-4, left L5 and right L6 with 1 tibial spur and L1 with 1 femoral spur.
</paragraph>
<paragraph pageId="0" pageNumber="41">
Ultimate prefemur with 4 ventro-lateral, 2 ventro-medial, 3 medial, 2 dorso-medial spines and 1 corner spine (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Otostigmus (O.) beroni Lewis, 2001 A, B, G Ar. - MHBU-SoJL 1608050101 C, D, F, H, I Ar. - MHBU-SoJL 1608050104 E Ar. - MHBU-SoJL 1608050105 A dorsal view B ventral view C cephalic plate, antennae, tergite 1 and legs 1 - 2, dorsal view D, E ventral view of head, basal antennal articles, an arrow showing the profile of the tooth plate F tergites 16 - 19 G sternites 11 - 14 H last two segments and ultimate legs, dorsal view I last two segments and ultimate legs, ventral view. Scale bars: 10.0 mm (A, B); 2.0 mm (C, F-I); 1.0 mm (D, E)." figureDoi="10.3897/zookeys.1088.77703.figure1" httpUri="https://binary.pensoft.net/fig/653519" pageId="0" pageNumber="41">1H, I</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="variability">
<paragraph pageId="0" pageNumber="41">Variability.</paragraph>
<paragraph pageId="0" pageNumber="41">
There are multiple differences among individuals as addressed below. The number of teeth of forcipular tooth plates 4+4 (11 specimens), 5+5 (6 specimens), 3+3 (2 specimen) or 3+4 (2 specimens). Coxopleural process with 4-7 spines (2 apical spines, 1-2 lateral spines and 1-3 dorsal spines). One tibial spur on L1-4 (8 specimens), L1-5 (5 specimens), L1-2 (2 specimens), L1-3 (3 specimens), or L1-6 (2 specimens). Two tarsal spurs on 1 to 18 or 19 pairs of legs; 1 tarsal spur on subsequent to penultimate legs. Ultimate legs without tarsal spur. Ultimate legs prefemur with 11-14 spines (4 or 5 ventro-lateral, 1-3 ventro-medial, 3 or 5 medial and 1 or 2 dorso-medial and 1 corner spine, rarely 0 or 3 corner spines) (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Variation in O. (O.) beroni from Jilong County, Xizang (1: Ar. - MHBU-SoJL 1608050101; 2: Ar. - MHBU-SoJL 1608050102; 3: Ar. - MHBU-SoJL 1608050103; 4: Ar. - MHBU-SoJL 1608050104; 5: Ar. - MHBU-SoJL 1608050105) (L / R: Left / Right)." httpUri="http://table.plazi.org/id/E045B19F5D5DA87CFB1BE54EB868C0D3" pageId="0" pageNumber="41" tableUuid="E045B19F5D5DA87CFB1BE54EB868C0D3">1</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="E045B19F5D5DA87CFB1BE54EB868C0D3" httpUri="http://table.plazi.org/id/E045B19F5D5DA87CFB1BE54EB868C0D3" pageId="0" pageNumber="41" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Table 1.</emphasis>
Variation in
<taxonomicName class="Chilopoda" kingdom="Animalia" lsidName="O. (O.) beroni" order="Scolopendromorpha" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="beroni" subGenus="O.">O. (O.) beroni</taxonomicName>
from Jilong County, Xizang (1: Ar.-MHBU-SoJL1608050101; 2: Ar.-MHBU-SoJL1608050102; 3: Ar.-MHBU-SoJL1608050103; 4: Ar.-MHBU-SoJL1608050104; 5: Ar.-MHBU-SoJL1608050105) (L/R: Left/Right).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<table pageId="0" pageNumber="41">
<tr pageId="0" pageNumber="41">
<th colspan="2" isEmpty="true" pageId="0" pageNumber="41" rowspan="2">-</th>
<th colspan="5" pageId="0" pageNumber="41" rowspan="1">Specimens</th>
</tr>
<tr pageId="0" pageNumber="41">
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">1</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">2</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">3</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">4</th>
<th colspan="1" pageId="0" pageNumber="41" rowspan="1">5</th>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Length/mm</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">54</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">43</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">47</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">57</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">39</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Number of antennal articles (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">18/19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">18</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">18</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">18</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">18</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Number of glabrous basal antennal articles</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2.3</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Tooth-plate teeth</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4+4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4+4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5+5</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4+4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5+5</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Paramedian sutures on tergites</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3-20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4-20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4-20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3-20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4-20</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Tergites marginate</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">6-21</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">6-21</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">7-21</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">7-21</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">6-21</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Paramedian sutures on sternites</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">7-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">7-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">7-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">6-19</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="3">Coxopleural process</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Apical spines (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2/3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Dorsal spines (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1/2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2/1</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Lateral spines (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1/2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2/1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="3">Legs</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">With tibial spur</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-5</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-3</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">With 1 tarsal spur</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">17, 20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">20</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">20</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">With 2 tarsal spurs</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-16, 18-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-19</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1-19</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="5">Spines of ultimate prefemur</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Ventro-lateral (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5/4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5/4</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Ventro-medial (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1/2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2/3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Medial (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5/3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3/5</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Dorso-medial (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2/1</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Corner spine (L/R)</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3/1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1/0</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="2" pageId="0" pageNumber="41" rowspan="1">Femoral spur of leg 1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="habitat">
<paragraph pageId="0" pageNumber="41">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="41">
Found under stones in humid mountain bush (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A habitat of O. (O.) beroni in Jilong Town, Jilong County B habitat of O. (O.) lewisi in Anrao Town, Jiacha County C O. (O.) beroni (Ar. - MHBU-SoJL 21080101) D O. (O.) lewisi (Ar. - MHBU-SoJC 1608 DX 01)." figureDoi="10.3897/zookeys.1088.77703.figure4" httpUri="https://binary.pensoft.net/fig/653522" pageId="0" pageNumber="41">4A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="distribution">
<paragraph pageId="0" pageNumber="41">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="41">
China (Xizang) and Nepal (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distributions of Otostigmus in Xizang, China a circle - O. (O.) beroni, Jilong Town, Jilong (Gyirong) County b triangle - O. (O.) lewisi, 2005, Jiacha (Gyaca) County c pentagon - O. (O.) martensi, Nielamu (Nyalam) County d square - O. (O.) xizangensis, Bomi (Bome) County." figureDoi="10.3897/zookeys.1088.77703.figure5" httpUri="https://binary.pensoft.net/fig/653523" pageId="0" pageNumber="41">5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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