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<document id="5165416E1923F275511B51FCC447A186" ID-CLB-Dataset="51481" ID-DOI="10.5169/seals-403072" ID-GBIF-Dataset="f141fb3f-9860-4c7e-b2d2-5424d915a835" ID-Zenodo-Dep="270241" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.metadata_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1459365803748" checkinUser="plazi" docAuthor="Jean-Paul Haenni" docDate="2013" docId="0A78FC5B22495813C4930282FC01FD2A" docLanguage="en" docName="seg-001_2013_86_3-4_a_007_d_text_and_pictures_only.pdf" docOrigin="Mitteilungen Der Schweizerischen Entomologischen Gesellschaft 86" docStyle="DocumentStyle{}" docTitle="Colobostema griseinerve Jean-Paul Haenni 2013, comb. n." docType="treatment" docVersion="8" lastPageNumber="217" masterDocId="F641842322465801C56B0417FFF3FFF5" masterDocTitle="A revision of the West Palaearctic species of Colobostema Enderlein, 1926 (Diptera, Scatopsidae). Part I. European subregion" masterLastPageNumber="242" masterPageNumber="199" pageNumber="214" updateTime="1698199847837" updateUser="admin">
<mods:mods id="8357C87812BAF495469D790D902D3E7D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DBB9D9E90156988EF489C8806311FF11">
<mods:title id="7BC56DC12A000B050E0BD5E3046F7A2C">A revision of the West Palaearctic species of Colobostema Enderlein, 1926 (Diptera, Scatopsidae). Part I. European subregion</mods:title>
</mods:titleInfo>
<mods:name id="98346FC11CA77DF9BAA1E5FF477757EF" type="personal">
<mods:role id="0CF04964CD3DA07D1F5B3C65AA271AF8">
<mods:roleTerm id="27D92BBA87F7B7D2065AD3B80F9E4C34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A61995EBF6DA25D57FA7F26AC498714B">Jean-Paul Haenni</mods:namePart>
</mods:name>
<mods:typeOfResource id="F902D50A49F802E0C3F013D14BFA73EA">text</mods:typeOfResource>
<mods:relatedItem id="E87E42649710537076E1E9E168D9E65C" type="host">
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<mods:title id="977A37D2063B0A0976ABE2BB85A9049D">Mitteilungen Der Schweizerischen Entomologischen Gesellschaft</mods:title>
</mods:titleInfo>
<mods:part id="FF8CE6BAA0C31AD3B98F185937F39FA6">
<mods:date id="00691BE0E5163C60A23B1A56C475B34F">2013</mods:date>
<mods:detail id="5B652048A13CB91C46D35E5EBE6434D5" type="pubDate">
<mods:number id="777069D9CFF2E2D0645FF171F890B630">2013-12-31</mods:number>
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<mods:detail id="B67361C19F7242C6C7A6B0110263336B" type="volume">
<mods:number id="C60EFAE889E139C6C08B31A111A4C258">86</mods:number>
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<mods:start id="A884569784D0CC174C01DB4AC653B931">199</mods:start>
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<mods:classification id="B7166491EB9C0A45FC0AA51D6C0FDA9C">journal article</mods:classification>
<mods:identifier id="7AC29E08D9696B81B6EB7DF304105B35" type="CLB-Dataset">51481</mods:identifier>
<mods:identifier id="581858AE93146FA13641961AB122FF2F" type="DOI">10.5169/seals-403072</mods:identifier>
<mods:identifier id="17F0509264D026263766B71793B3829C" type="GBIF-Dataset">f141fb3f-9860-4c7e-b2d2-5424d915a835</mods:identifier>
<mods:identifier id="97D4F23F76F6764ECFFA63142DEC2BA3" type="Zenodo-Dep">270241</mods:identifier>
</mods:mods>
<treatment id="0A78FC5B22495813C4930282FC01FD2A" ID-DOI="http://doi.org/10.5281/zenodo.6157675" ID-GBIF-Taxon="119329165" ID-Zenodo-Dep="6157675" LSID="urn:lsid:plazi:treatment:0A78FC5B22495813C4930282FC01FD2A" httpUri="http://treatment.plazi.org/id/0A78FC5B22495813C4930282FC01FD2A" lastPageId="18" lastPageNumber="217" pageId="15" pageNumber="214">
<subSubSection id="CACB1EC62249580EC4930282FB83F947" box="[504,1136,1685,1714]" pageId="15" pageNumber="214" type="nomenclature">
<paragraph id="826E4D4D2249580EC4930282FB83F947" blockId="15.[504,1136,1685,1714]" box="[504,1136,1685,1714]" pageId="15" pageNumber="214">
<heading id="D926FA212249580EC4930282FB83F947" bold="true" box="[504,1136,1685,1714]" centered="true" fontSize="12" level="2" pageId="15" pageNumber="214" reason="2">
<emphasis id="B0A5915F2249580EC4930282FB83F947" bold="true" box="[504,1136,1685,1714]" pageId="15" pageNumber="214">
<taxonomicName id="45D136CE2249580EC4930282FC07F947" authority="(Duda, 1928)" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Duda" baseAuthorityYear="1928" box="[504,1012,1685,1714]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="griseinerve" status="comb. nov.">
<emphasis id="B0A5915F2249580EC4930282FCCEF947" bold="true" box="[504,829,1685,1714]" italics="true" pageId="15" pageNumber="214">Colobostema griseinerve</emphasis>
(
<bibRefCitation id="E64030BC2249580EC6260282FC1FF947" author="Duda, 0." bookContentInfo="3 Taf" box="[845,1004,1685,1714]" editor="Lindner, E." journalOrPublisher="Schweizerbart, Stuttgart" pageId="15" pageNumber="214" pagination="1 - 62" refId="ref26368" refString="Duda, 0. 1928. Scatopsidae. In: Lindner, E. (ed.) Die Fliegen der palaarktischen Region 2 (1), 5: 1 - 62, 3 Taf. - Schweizerbart, Stuttgart." title="Scatopsidae" type="book chapter" volumeTitle="Die Fliegen der palaarktischen Region 2 (1), 5" year="1928">Duda, 1928</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="AB962C242249580EC6970282FB83F947" box="[1020,1136,1685,1714]" pageId="15" pageNumber="214">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CACB1EC62249580EC79F02D0FC87F92A" box="[756,884,1735,1759]" pageId="15" pageNumber="214" type="description">
<paragraph id="826E4D4D2249580EC79F02D0FC87F92A" blockId="15.[756,884,1735,1759]" box="[756,884,1735,1759]" pageId="15" pageNumber="214">
(
<figureCitation id="1AEA51C82249580EC79002D0FCCEF92A" box="[763,829,1735,1759]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="15" pageNumber="214">Figs 5</figureCitation>
A-E)
</paragraph>
</subSubSection>
<subSubSection id="CACB1EC62249580EC4680316FB3CF8ED" box="[259,1231,1792,1817]" pageId="15" pageNumber="214" type="reference_group">
<paragraph id="826E4D4D2249580EC4680316FB3CF8ED" blockId="15.[259,1231,1792,1817]" box="[259,1231,1792,1817]" pageId="15" pageNumber="214">
<taxonomicName id="45D136CE2249580EC4680316FCEBF8ED" ID-CoL="4V493" authority="Duda, 1928" authorityName="Duda" authorityYear="1928" box="[259,792,1792,1816]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="214" phylum="Arthropoda" rank="variety" species="tristis" variety="griseinervis">
<emphasis id="B0A5915F2249580EC4680316FE56F8ED" box="[259,421,1793,1816]" italics="true" pageId="15" pageNumber="214">Scatopse tristis</emphasis>
Zett. var.
<emphasis id="B0A5915F2249580EC7780316FD62F8ED" box="[531,657,1793,1816]" italics="true" pageId="15" pageNumber="214">griseinervis</emphasis>
<bibRefCitation id="E64030BC2249580EC7F30317FCEBF8ED" author="Duda, 0." bookContentInfo="3 Taf" box="[664,792,1792,1816]" editor="Lindner, E." journalOrPublisher="Schweizerbart, Stuttgart" pageId="15" pageNumber="214" pagination="1 - 62" refId="ref26368" refString="Duda, 0. 1928. Scatopsidae. In: Lindner, E. (ed.) Die Fliegen der palaarktischen Region 2 (1), 5: 1 - 62, 3 Taf. - Schweizerbart, Stuttgart." title="Scatopsidae" type="book chapter" volumeTitle="Die Fliegen der palaarktischen Region 2 (1), 5" year="1928">Duda, 1928</bibRefCitation>
</taxonomicName>
: Scatopsidae, Fliegen pal. Reg. 2(1)5:39.
</paragraph>
</subSubSection>
<subSubSection id="CACB1EC622495811C43A0357FCC0FD7A" lastPageId="16" lastPageNumber="215" pageId="15" pageNumber="214" type="materials_examined">
<paragraph id="826E4D4D2249580EC43A0357FBA9F8A9" blockId="15.[259,1381,1855,1996]" box="[337,1114,1855,1884]" pageId="15" pageNumber="214">
<emphasis id="B0A5915F2249580EC43A0357FDF3F8A9" box="[337,512,1855,1884]" italics="true" pageId="15" pageNumber="214">
<typeStatus id="5D6AF3EF2249580EC43A0357FE7CF8A9" box="[337,399,1856,1884]" pageId="15" pageNumber="214">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="FAC60DDD2249580EC7630328FD98F8A9" box="[520,619,1855,1884]" name="Italy" pageId="15" pageNumber="214">ITALY</collectingCountry>
, prov. Novara: Macugnaga, Mte Rosa.
</paragraph>
<paragraph id="826E4D4D22495811C43B0373FE22FECA" blockId="15.[259,1381,1855,1996]" lastBlockId="16.[191,1310,216,1998]" lastPageId="16" lastPageNumber="215" pageId="15" pageNumber="214">
<emphasis id="B0A5915F2249580EC43B0373FDA3F874" box="[336,592,1892,1921]" italics="true" pageId="15" pageNumber="214">Material examined.</emphasis>
<emphasis id="B0A5915F2249580EC7370372FD2BF877" bold="true" box="[604,728,1893,1922]" pageId="15" pageNumber="214">
<typeStatus id="5D6AF3EF2249580EC7370372FD2BF877" box="[604,728,1893,1922]" pageId="15" pageNumber="214" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="94D786C42249580EC78E0372FD0CF874" box="[741,767,1893,1921]" pageId="15" pageNumber="214">♂</specimenCount>
, labelled: «Mte Rosa 4.8.00» [in Oldenberg’s handwriting] / «Macugnaga Oldenberg» / «Coli. Oldenberg» / «Typus» /
<emphasis id="B0A5915F2249580EC188039CFACFF852" box="[1251,1340,1931,1959]" italics="true" pageId="15" pageNumber="214">
«
<taxonomicName id="45D136CE2249580EC199039CFACFF852" authorityName="Zetterstedt" authorityYear="1850" box="[1266,1340,1931,1959]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="214" phylum="Arthropoda" rank="species" species="tristis">tristis</taxonomicName>
</emphasis>
Z.
<taxonomicName id="45D136CE2249580EC46F03A7FE25F839" authorityName="Duda" authorityYear="1928" box="[260,470,1968,1996]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="214" phylum="Arthropoda" rank="variety" species="tristis" variety="griseinervis">
var.
<emphasis id="B0A5915F2249580EC45403A7FE25F839" box="[319,470,1968,1996]" italics="true" pageId="15" pageNumber="214">griseinervis</emphasis>
</taxonomicName>
oder n. sp.
<specimenCount id="94D786C42249580EC71C03A7FD7CF839" box="[631,655,1968,1996]" pageId="15" pageNumber="214">♂</specimenCount>
d. Duda» [in Duda’s handwriting] / «coll.
<collectionCode id="E4C0D5882249580EC1A203A7FAF1F839" box="[1225,1282,1968,1996]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="15" pageNumber="214" type="Other type of research institution/biorepository">DEI</collectionCode>
. Eberswalde»; I have labelled it
<emphasis id="B0A5915F22565811C74004CFFC71FF00" box="[555,898,216,245]" italics="true" pageId="16" pageNumber="215">
«
<taxonomicName id="45D136CE22565811C75204CFFC71FF00" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Duda" baseAuthorityYear="1928" box="[569,898,216,245]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="griseinerve" status="comb. nov.">Colobostema griseinerve</taxonomicName>
</emphasis>
<taxonomicNameLabel id="AB962C2422565811C6FB04CFFC04FF00" box="[912,1015,216,245]" pageId="16" pageNumber="215">comb.n.</taxonomicNameLabel>
JP Haenni rev. 1990
<typeStatus id="5D6AF3EF22565811C5D404EAFEC5FEEF" box="[191,310,253,282]" pageId="16" pageNumber="215" type="holotype">Holotype</typeStatus>
<specimenCount id="94D786C422565811C42F04E9FEAFFEEF" box="[324,348,254,282]" pageId="16" pageNumber="215">♂</specimenCount>
»; type specimen pinned, in good state, preserved in the collections of the
<collectionCode id="E4C0D58822565811C5850534FED5FECA" box="[238,294,291,319]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="16" pageNumber="215" type="Other type of research institution/biorepository">DEI</collectionCode>
, Eberswalde.
</paragraph>
<paragraph id="826E4D4D22565811C4670550FB26FE5A" blockId="16.[191,1310,216,1998]" pageId="16" pageNumber="215">
<emphasis id="B0A5915F22565811C4670550FE2EFE91" box="[268,477,327,356]" italics="true" pageId="16" pageNumber="215">Other material.</emphasis>
<collectingCountry id="FAC60DDD22565811C4840550FD8DFE91" box="[495,638,327,356]" name="Austria" pageId="16" pageNumber="215">AUSTRIA</collectingCountry>
.
<collectingRegion id="401583AF22565811C7FF055FFCF3FE91" box="[660,768,328,356]" country="Austria" name="Karnten" pageId="16" pageNumber="215">Kärnten</collectingRegion>
: Heiligenbild, 4.VTI1.1927,
<specimenCount id="94D786C422565811C1E40550FB38FE91" box="[1167,1227,327,356]" count="1" pageId="16" pageNumber="215" type="male">1 ♂</specimenCount>
, coll. Oldenberg,
<collectionCode id="E4C0D58822565811C4130579FE42FE7F" box="[376,433,366,394]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="16" pageNumber="215" type="Other type of research institution/biorepository">DEI</collectionCode>
. —
<collectingCountry id="FAC60DDD22565811C74E057AFD54FE7F" box="[549,679,365,394]" name="France" pageId="16" pageNumber="215">FRANCE</collectingCountry>
, Alpes-de-Haute-Provence: Thorame-Haute (Orgeas),
<quantity id="4529E0A822565811C42A0585FE56FE5A" box="[321,421,402,431]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="215" unit="m" value="2000.0">2000 m</quantity>
,
<specimenCount id="94D786C422565811C4D90585FE17FE5A" box="[434,484,402,431]" count="1" pageId="16" pageNumber="215" type="male">1 ♂</specimenCount>
,
<date id="F66F6B8D22565811C4990585FD77FE5A" box="[498,644,402,431]" pageId="16" pageNumber="215" value="1999-05-21">21.V. 1999</date>
, L. Micas,
<collectionCode id="E4C0D58822565811C6790585FC8EFE5A" box="[786,893,402,431]" pageId="16" pageNumber="215">INRAM</collectionCode>
(
<bibRefCitation id="E64030BC22565811C6E50585FB34FE5A" author="Haenni. J. - P. &amp; Withers, P" box="[910,1223,402,431]" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="16" pageNumber="215" pagination="17 - 27" part="112" refId="ref27411" refString="Haenni. J. - P. &amp; Withers, P. 2007. Liste commentee des Scatopsidae de France continentale (Diptera). - Bulletin de la Societe entomologique de France 112 (1): 17 - 27." title="Liste commentee des Scatopsidae de France continentale (Diptera)" type="journal article" year="2007">Haenni &amp; Withers 2007</bibRefCitation>
).
</paragraph>
<paragraph id="826E4D4D22565811C5A805AFFB1FFE21" blockId="16.[191,1310,216,1998]" box="[195,1260,439,468]" pageId="16" pageNumber="215">
-
<collectingCountry id="FAC60DDD22565811C58305A0FE3EFE21" box="[232,461,439,468]" name="Serbia" pageId="16" pageNumber="215">MONTENEGRO</collectingCountry>
: Durmitor,
<quantity id="4529E0A822565811C70B05A0FD37FE21" box="[608,708,439,468]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="215" unit="m" value="1500.0">1500 m</quantity>
,
<date id="F66F6B8D22565811C7BA05A0FC39FE21" box="[721,970,439,468]" pageId="16" pageNumber="215" value="1988-07-30" valueMax="1988-08-05" valueMin="1988-07-30">30.VII-5.VIII.1988</date>
,
<specimenCount id="94D786C422565811C6BC05A0FBFAFE21" box="[983,1033,439,468]" count="1" pageId="16" pageNumber="215" type="male">1 ♂</specimenCount>
, G. Bächli,
<collectionCode id="E4C0D58822565811C1C905A0FB1BFE21" box="[1186,1256,439,468]" pageId="16" pageNumber="215">CGB</collectionCode>
.
</paragraph>
<paragraph id="826E4D4D22565811C5A805C9FCC0FD7A" blockId="16.[191,1310,216,1998]" pageId="16" pageNumber="215">
-
<collectingCountry id="FAC60DDD22565811C59D05CBFE2DFE0C" box="[246,478,476,505]" name="Switzerland" pageId="16" pageNumber="215">SWITZERLAND</collectingCountry>
. LU: Menzberg,
<quantity id="4529E0A822565811C7BA05CBFCCDFE0F" box="[721,830,476,506]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="215" unit="m" value="1000.0">1000 m</quantity>
,
<date id="F66F6B8D22565811C63F05CBFBFAFE0C" box="[852,1033,476,505]" pageId="16" pageNumber="215" value="1983-08-03" valueMax="1983-08-06" valueMin="1983-08-03">3-6.VIII.1983</date>
,
<specimenCount id="94D786C422565811C17405CBFB81FE0F" box="[1055,1138,476,506]" count="2" pageId="16" pageNumber="215" type="male">2 ♂♂</specimenCount>
, G. Bächli,
<collectionCode id="E4C0D58822565811C5AA0615FEF4FDEA" box="[193,263,514,543]" pageId="16" pageNumber="215">CGB</collectionCode>
; TT: Bolle di Magadino, 17-20.VT.1995,
<specimenCount id="94D786C422565811C6530615FC9DFDEA" box="[824,878,514,543]" count="1" pageId="16" pageNumber="215" type="male">1 ♂</specimenCount>
, B. Merz &amp; G. Bächli,
<collectionCode id="E4C0D58822565811C1DB0614FAE5FDEA" box="[1200,1302,515,543]" country="France" httpUri="http://grbio.org/cool/1hrs-xwta" name="Musee d'Histoire Naturalle" pageId="16" pageNumber="215">MHNN</collectionCode>
; Locarno env., (Ronco s/Ascona, Locarno, Gordola, Orselina, Minusio, Cugnasco), dates ranging from 25.111 to
<date id="F66F6B8D22565811C700065BFCE6FD9C" box="[619,789,588,617]" pageId="16" pageNumber="215" value="1997-08-12">12.VIII.1997</date>
,
<specimenCount id="94D786C422565811C647065BFC62FD9C" box="[812,913,588,617]" count="32" pageId="16" pageNumber="215" type="male">32 ♂♂</specimenCount>
, M. Moretti,
<collectionCode id="E4C0D58822565811C13C065BFB5CFD9C" box="[1111,1199,588,617]" pageId="16" pageNumber="215">MSNL</collectionCode>
/
<collectionCode id="E4C0D58822565811C1D3065BFAE9FD9C" box="[1208,1306,588,617]" country="France" httpUri="http://grbio.org/cool/1hrs-xwta" name="Musee d'Histoire Naturalle" pageId="16" pageNumber="215">MHNN</collectionCode>
(detail of captures in Haenni &amp; Moretti in prep.).
</paragraph>
</subSubSection>
<subSubSection id="CACB1EC622565811C460068FFCEBFC9B" pageId="16" pageNumber="215" type="diagnosis">
<paragraph id="826E4D4D22565811C460068FFCEBFC9B" blockId="16.[191,1310,216,1998]" pageId="16" pageNumber="215">
<emphasis id="B0A5915F22565811C460068FFE66FD41" box="[267,405,664,692]" italics="true" pageId="16" pageNumber="215">Diagnosis.</emphasis>
Males of
<taxonomicName id="45D136CE22565811C776068FFD26FD41" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Duda" baseAuthorityYear="1928" box="[541,725,664,692]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="griseinerve">
<emphasis id="B0A5915F22565811C776068FFD26FD41" box="[541,725,664,692]" italics="true" pageId="16" pageNumber="215">C. griseinerve</emphasis>
</taxonomicName>
can be separated from those of other European species of the
<taxonomicName id="45D136CE22565811C4B206AAFD97FD2C" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[473,612,701,729]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="nigripenne">
<emphasis id="B0A5915F22565811C4B206AAFD97FD2C" box="[473,612,701,729]" italics="true" pageId="16" pageNumber="215">nigripenne</emphasis>
</taxonomicName>
-group (that have tergite 7 strongly produced posteriorly medially) by the shape of the epandrium, which has posterior projections slightly but clearly asymmetrical, horn-like, pointed and rather heavily sclerotized (
<figureCitation id="1AEA51C822565811C5A0073CFEEFFCBD" box="[203,284,811,840]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="16" pageNumber="215">Fig. 5</figureCitation>
D), and by the elongate gonocoxites, appearing either pointed or rounded at apex, according to the angle of view (
<figureCitation id="1AEA51C822565811C7CF0746FD03FC9B" box="[676,752,849,878]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="16" pageNumber="215">Fig. 5</figureCitation>
C).
</paragraph>
</subSubSection>
<subSubSection id="CACB1EC622565811C4600760FE03F98A" pageId="16" pageNumber="215" type="description">
<paragraph id="826E4D4D22565811C4600760FE67FBDD" blockId="16.[191,1310,216,1998]" pageId="16" pageNumber="215">
<emphasis id="B0A5915F22565811C4600760FE59FC66" box="[267,426,887,915]" italics="true" pageId="16" pageNumber="215">Description.</emphasis>
1.9 mm long; brown black in general colour (
<typeStatus id="5D6AF3EF22565811C1240761FB7BFC66" box="[1103,1160,886,915]" pageId="16" pageNumber="215">type</typeStatus>
somewhat faded), dull except lower pleurae shining as usual; antennae entirely dark; wings greyish brown with a lighter zone above angle of M, that extends up to apex of R4+5; veins dirty brownish grey; legs concolourous with body except basal third of tibiae and tarsi lighter.
</paragraph>
<paragraph id="826E4D4D22565811C4600026FD82FB11" blockId="16.[191,1310,216,1998]" pageId="16" pageNumber="215">Head. Eyes approximated on frons above antennae, separated by less than half width of anterior ocellus; antennae slightly widening towards apex, flagellomeres comparatively closer to each other than is usual for the genus; first flagellomere longer than wide, second as long as wide, following wider than long, last flagellomere almost twice preceding one.</paragraph>
<paragraph id="826E4D4D22565811C46000FCFDADFAA1" blockId="16.[191,1310,216,1998]" pageId="16" pageNumber="215">
Thorax. Notum hardly longer than wide; wing (
<figureCitation id="1AEA51C822565811C6E300FCFC24FAFD" box="[904,983,1259,1288]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="16" pageNumber="215">Fig. 5</figureCitation>
E) densely covered with microtrichia; anteriorly directed stem spurious vein on M, conspicuous, long; M|+2 clearly shorter than medial fork.
</paragraph>
<paragraph id="826E4D4D22565811C467014CFC7AF9AC" blockId="16.[191,1310,216,1998]" pageId="16" pageNumber="215">
Abdomen. Basal sclerotized ring of segment 7 incomplete; tergite 7 (
<figureCitation id="1AEA51C822565811C1C2014CFB0BFA8D" box="[1193,1272,1371,1400]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="16" pageNumber="215">Fig. 5</figureCitation>
A) with narrow, apically pointed posterior projection and a pair of comparatively wide lateral expansions, inner fold well developed; sternite 7 (
<figureCitation id="1AEA51C822565811C6D801B1FBF2FA36" box="[947,1025,1446,1475]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="16" pageNumber="215">Fig. 5</figureCitation>
B) with a shallow Vshaped posterior emargination; genital capsule (
<figureCitation id="1AEA51C822565811C62201DCFC68FA1D" box="[841,923,1483,1512]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="16" pageNumber="215">Fig. 5</figureCitation>
C) with gonocoxites slightly asymmetrical, apically pointed or rounded, according to angle of view; parameres elongate; epandrium (
<figureCitation id="1AEA51C822565811C4880202FDC1F9C7" box="[483,562,1557,1586]" captionStart="Fig. 5" captionStartId="15.[259,298,1516,1540]" captionTargetBox="[509,1147,1235,1463]" captionTargetId="figure@15.[509,1148,1235,1463]" captionTargetPageId="15" captionText="Fig. 5, Colobostema griseinerve (Duda): A. Tergite 7, ♂. - B. Sternite 7, ♂. - C. Génital capsule, ♂, ventral. - D. Epandrium, distal part, ♂, dorsal. - E. Wing venation, ♂." httpUri="https://zenodo.org/record/270246/files/figure.png" pageId="16" pageNumber="215">Fig. 5</figureCitation>
D) with heavily sclerotized, pointed, asymmetrical posterior projections, aedeagus of medium length, sinuous.
</paragraph>
<paragraph id="826E4D4D22565811C4600275FE03F98A" blockId="16.[191,1310,216,1998]" box="[267,496,1634,1663]" pageId="16" pageNumber="215">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="CACB1EC622565813C4660290FB94FE40" lastPageId="18" lastPageNumber="217" pageId="16" pageNumber="215" type="discussion">
<paragraph id="826E4D4D22565813C4660290FB94FE40" blockId="16.[191,1310,216,1998]" lastBlockId="18.[189,1307,222,735]" lastPageId="18" lastPageNumber="217" pageId="16" pageNumber="215">
<emphasis id="B0A5915F22565811C4660290FDEAF956" box="[269,537,1670,1699]" italics="true" pageId="16" pageNumber="215">Taxonomic remarks.</emphasis>
<bibRefCitation id="E64030BC22565811C74C0291FD3DF956" author="Duda, 0." bookContentInfo="3 Taf" box="[551,718,1670,1699]" editor="Lindner, E." journalOrPublisher="Schweizerbart, Stuttgart" pageId="16" pageNumber="215" pagination="1 - 62" refId="ref26368" refString="Duda, 0. 1928. Scatopsidae. In: Lindner, E. (ed.) Die Fliegen der palaarktischen Region 2 (1), 5: 1 - 62, 3 Taf. - Schweizerbart, Stuttgart." title="Scatopsidae" type="book chapter" volumeTitle="Die Fliegen der palaarktischen Region 2 (1), 5" year="1928">Duda (1928)</bibRefCitation>
erected this «variety» for some supposedly aberrant specimens of C.
<taxonomicName id="45D136CE22565811C77202BAFD5EF93D" authority="Zett." authorityName="Zetterstedt" authorityYear="1850" box="[537,685,1708,1737]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B0A5915F22565811C77202BAFD91F93C" box="[537,610,1709,1737]" italics="true" pageId="16" pageNumber="215">tristis</emphasis>
Zett.
</taxonomicName>
, characterized by more or less grey wings and veins. As can be seen from his identification label of the
<typeStatus id="5D6AF3EF22565811C6AA02C6FC08F918" box="[961,1019,1745,1773]" pageId="16" pageNumber="215">type</typeStatus>
, he hesitated on what status to give to this form that he considered as «eine Uebergangsform von
<taxonomicName id="45D136CE22565811C1BA02E0FEE7F8CC" authorityName="Meigen" authorityYear="1830" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="nigripennis">
<emphasis id="B0A5915F22565811C1BA02E0FEE7F8CC" italics="true" pageId="16" pageNumber="215">nigripennis</emphasis>
</taxonomicName>
zu
<emphasis id="B0A5915F22565811C420030AFE57F8CC" box="[331,420,1821,1849]" italics="true" pageId="16" pageNumber="215">
<taxonomicName id="45D136CE22565811C420030AFE66F8CC" authority="Duda 1928" authorityName="Zetterstedt" authorityYear="1850" box="[331,405,1821,1849]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="tristis">tristis</taxonomicName>
»
</emphasis>
(
<bibRefCitation id="E64030BC22565811C4D2030CFDA1F8CD" author="Duda, 0." bookContentInfo="3 Taf" box="[441,594,1819,1848]" editor="Lindner, E." journalOrPublisher="Schweizerbart, Stuttgart" pageId="16" pageNumber="215" pagination="1 - 62" refId="ref26368" refString="Duda, 0. 1928. Scatopsidae. In: Lindner, E. (ed.) Die Fliegen der palaarktischen Region 2 (1), 5: 1 - 62, 3 Taf. - Schweizerbart, Stuttgart." title="Scatopsidae" type="book chapter" volumeTitle="Die Fliegen der palaarktischen Region 2 (1), 5" year="1928">Duda 1928</bibRefCitation>
). This taxon has never been referred to since its description. It corresponds, however, to a very distinctive species belonging to the
<emphasis id="B0A5915F22565811C5A80370FEBCF876" box="[195,335,1895,1923]" italics="true" pageId="16" pageNumber="215">Mgripenne</emphasis>
-group of species. The wing coloration features used by
<bibRefCitation id="E64030BC22565811C1290371FB1BF876" author="Duda, 0." bookContentInfo="3 Taf" box="[1090,1256,1894,1923]" editor="Lindner, E." journalOrPublisher="Schweizerbart, Stuttgart" pageId="16" pageNumber="215" pagination="1 - 62" refId="ref26368" refString="Duda, 0. 1928. Scatopsidae. In: Lindner, E. (ed.) Die Fliegen der palaarktischen Region 2 (1), 5: 1 - 62, 3 Taf. - Schweizerbart, Stuttgart." title="Scatopsidae" type="book chapter" volumeTitle="Die Fliegen der palaarktischen Region 2 (1), 5" year="1928">Duda (1928)</bibRefCitation>
are evidently insufficient to allow identification, as is demonstrated by the fact that the only other specimen of C.
<taxonomicName id="45D136CE22565811C77C03A5FD54F83B" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Duda" baseAuthorityYear="1928" box="[535,679,1970,1998]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="215" phylum="Arthropoda" rank="species" species="griseinerve">
<emphasis id="B0A5915F22565811C77C03A5FD54F83B" box="[535,679,1970,1998]" italics="true" pageId="16" pageNumber="215">griseinerve</emphasis>
</taxonomicName>
(from Heiligenblut,
<collectingCountry id="FAC60DDD22565811C6C403A7FBFCF838" box="[943,1039,1968,1997]" name="Austria" pageId="16" pageNumber="215">Austria</collectingCountry>
) seen by Duda had been labelled by him as
<taxonomicName id="45D136CE22545813C72F04C8FD25FF0E" authority="Zett." authorityName="Zett." box="[580,726,223,251]" class="Insecta" family="Scatopsidae" genus="Scatopse" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="217" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B0A5915F22545813C72F04C8FD7EFF0E" box="[580,653,223,251]" italics="true" pageId="18" pageNumber="217">tristis</emphasis>
Zett.
</taxonomicName>
Duda attributed also to the variety
<taxonomicName id="45D136CE22545813C1D204C8FEF7FED5" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Duda" baseAuthorityYear="1928" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="217" phylum="Arthropoda" rank="species" species="griseinerve">
<emphasis id="B0A5915F22545813C1D204C8FEF7FED5" italics="true" pageId="18" pageNumber="217">griseinerve</emphasis>
</taxonomicName>
<specimenCount id="94D786C422545813C47A0514FE10FED5" box="[273,483,259,288]" count="3" pageId="18" pageNumber="217" type="generic">three specimens</specimenCount>
from
<collectingCountry id="FAC60DDD22545813C7500514FD6EFED5" box="[571,669,259,288]" name="Finland" pageId="18" pageNumber="217">Finland</collectingCountry>
and
<collectingRegion id="401583AF22545813C78E0514FCADFED5" box="[741,862,259,288]" country="Finland" name="Lappland" pageId="18" pageNumber="217">Lappland</collectingRegion>
based on rather trivial wing and legs coloration characters (which are variable and clearly of no taxonomic value in this genus); two of these specimens have been recognized since then as the
<typeStatus id="5D6AF3EF22545813C1DA0558FB06FE9E" box="[1201,1269,335,363]" pageId="18" pageNumber="217">types</typeStatus>
of a distinct species (C.
<taxonomicName id="45D136CE22545813C48A0565FD25FE7A" authority="Krivosheina" authorityName="Krivosheina" authorityYear="2001" box="[481,726,370,399]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="217" phylum="Arthropoda" rank="species" species="dudai">
<emphasis id="B0A5915F22545813C48A0565FDD9FE7A" box="[481,554,370,399]" italics="true" pageId="18" pageNumber="217">dudai</emphasis>
Krivosheina
</taxonomicName>
, see above under this species) and the latter specimen as belonging to C.
<taxonomicName id="45D136CE22545813C732058FFCC5FE40" authority="(Hal.)" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Haliday" baseAuthorityYear="1833" box="[601,822,408,437]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="217" phylum="Arthropoda" rank="species" species="infumatum">
<emphasis id="B0A5915F22545813C732058FFD11FE40" box="[601,738,408,437]" italics="true" pageId="18" pageNumber="217">infumatum</emphasis>
(Hal.)
</taxonomicName>
by
<bibRefCitation id="E64030BC22545813C60E058FFB90FE40" author="Krivosheina, N. P." box="[869,1123,408,437]" journalOrPublisher="Entomologicheskoe Obozrenie 79" pageId="18" pageNumber="217" pagination="487 - 498" part="2" refId="ref27591" refString="Krivosheina, N. P. 2000. [On the systematics of the species of the genera Colobostema Enderlein, Holoplagia Enderlein and Cookella Freeman (Diptera, Scatopsidae) from Russia and neighbouring countries.]. - Entomologicheskoe Obozrenie 79 (2): 487 - 498. [in Russian]" title="On the systematics of the species of the genera Colobostema Enderlein, Holoplagia Enderlein and Cookella Freeman (Diptera, Scatopsidae) from Russia and neighbouring countries." type="journal article" year="2000">Krivosheina (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="D6AE1DC522575810C47B02F3FD67F8C3" httpUri="https://zenodo.org/record/270247/files/figure.png" pageId="17" pageNumber="216" targetBox="[274,1393,222,1708]" targetPageId="17">
<paragraph id="826E4D4D22575810C47B02F3FD67F8C3" blockId="17.[272,1394,1764,1846]" pageId="17" pageNumber="216">
Fig. 6.
<taxonomicName id="45D136CE22575810C40902F3FD2CF909" authority="(Haliday)" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Haliday" baseAuthorityYear="1833" box="[354,735,1764,1788]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="216" phylum="Arthropoda" rank="species" species="infumatum">
<emphasis id="B0A5915F22575810C40902F3FD98F909" box="[354,619,1764,1788]" italics="true" pageId="17" pageNumber="216">Colobostema infumatum</emphasis>
(Haliday)
</taxonomicName>
: A. Tergite 7,
<specimenCount id="94D786C422575810C6EE02F2FC6AF909" box="[901,921,1765,1788]" pageId="17" pageNumber="216">♂</specimenCount>
. - B. Sternite 7,
<emphasis id="B0A5915F22575810C10802F3FB63F909" box="[1123,1168,1764,1788]" italics="true" pageId="17" pageNumber="216">
<specimenCount id="94D786C422575810C10802F3FB84F909" box="[1123,1143,1764,1788]" pageId="17" pageNumber="216">♂</specimenCount>
. -
</emphasis>
C. Génital capsule,
<specimenCount id="94D786C422575810C4780315FED4F8EC" box="[275,295,1794,1817]" pageId="17" pageNumber="216">♂</specimenCount>
, ventral. - D. Epandrium, distal part,
<specimenCount id="94D786C422575810C78B0315FD07F8EC" box="[736,756,1794,1817]" pageId="17" pageNumber="216">♂</specimenCount>
, dorsal. - E. Wing venation,
<specimenCount id="94D786C422575810C1290315FBAAF8EC" box="[1090,1113,1794,1817]" pageId="17" pageNumber="216">♂</specimenCount>
. - F. Tergite 7,
<specimenCount id="94D786C422575810C0780315FAD4F8EC" box="[1299,1319,1794,1817]" pageId="17" pageNumber="216">♀</specimenCount>
. - G. Sternite 8,
<specimenCount id="94D786C422575810C4EF0308FE6BF8C3" box="[388,408,1823,1846]" pageId="17" pageNumber="216">♀</specimenCount>
(detail), with valvifers.
</paragraph>
</caption>
<subSubSection id="CACB1EC622545813C46205AAFEE3FD9A" pageId="18" pageNumber="217" type="distribution">
<paragraph id="826E4D4D22545813C46205AAFEE3FD9A" blockId="18.[189,1307,222,735]" pageId="18" pageNumber="217">
<emphasis id="B0A5915F22545813C46205AAFD83FE2F" box="[265,624,445,474]" italics="true" pageId="18" pageNumber="217">
Distribution.
<taxonomicName id="45D136CE22545813C4D305A9FD83FE2F" authorityName="Jean-Paul Haenni" authorityYear="2013" baseAuthorityName="Duda" baseAuthorityYear="1928" box="[440,624,446,474]" class="Insecta" family="Scatopsidae" genus="Colobostema" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="217" phylum="Arthropoda" rank="species" species="griseinerve">C. griseinerve</taxonomicName>
</emphasis>
has been recorded from
<collectingCountry id="FAC60DDD22545813C6D005A9FBE1FE2F" box="[955,1042,446,474]" name="France" pageId="18" pageNumber="217">France</collectingCountry>
(
<bibRefCitation id="E64030BC22545813C14D05AAFF0CFE0A" author="Haenni. J. - P. &amp; Withers, P" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="18" pageNumber="217" pagination="17 - 27" part="112" refId="ref27411" refString="Haenni. J. - P. &amp; Withers, P. 2007. Liste commentee des Scatopsidae de France continentale (Diptera). - Bulletin de la Societe entomologique de France 112 (1): 17 - 27." title="Liste commentee des Scatopsidae de France continentale (Diptera)" type="journal article" year="2007">Haenni &amp; Withers 2007</bibRefCitation>
) and
<collectingCountry id="FAC60DDD22545813C42505F5FE14FE0A" box="[334,487,482,511]" name="Switzerland" pageId="18" pageNumber="217">Switzerland</collectingCountry>
(
<bibRefCitation id="E64030BC22545813C49005F5FD58FE0A" author="Haenni, J. - P." box="[507,683,482,511]" editor="Merz, B." journalOrPublisher="Fauna Helvetica" pageId="18" pageNumber="217" pagination="141 - 144" part="1" refId="ref26757" refString="Haenni, J. - P. 1998. 24. Scatopsidae. In: Merz, B., Bachli, G., Haenni, J. - P. &amp; Gonseth, Y. (eds.), Diptera - Checklist. Fauna Helvetica 1: 141 - 144. - CSCF / SEG, Neuchatel." title="24. Scatopsidae" type="journal article" volumeTitle="Diptera - Checklist" year="1998">Haenni 1998</bibRefCitation>
). It is apparently restricted to the mountainous ranges of Central and South-Eastern Europe, with about 15 localities from the Alps of
<collectingCountry id="FAC60DDD22545813C58E063AFEB4FDBF" box="[229,327,557,586]" name="Austria" pageId="18" pageNumber="217">Austria</collectingCountry>
,
<collectingCountry id="FAC60DDD22545813C4330639FE41FDBF" box="[344,434,558,586]" name="France" pageId="18" pageNumber="217">France</collectingCountry>
,
<collectingCountry id="FAC60DDD22545813C4AF063AFDF1FDBF" box="[452,514,557,586]" name="Italy" pageId="18" pageNumber="217">Italy</collectingCountry>
, and
<collectingCountry id="FAC60DDD22545813C725063AFD19FDBF" box="[590,746,557,586]" name="Switzerland" pageId="18" pageNumber="217">Switzerland</collectingCountry>
, and from the Durmitor range in
<collectingCountry id="FAC60DDD22545813C1D90639FEFFFD9A" name="Serbia" pageId="18" pageNumber="217">Montenegro</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CACB1EC622545813C4630660FC01FD2A" pageId="18" pageNumber="217" type="biology_ecology">
<paragraph id="826E4D4D22545813C4630660FC01FD2A" blockId="18.[189,1307,222,735]" pageId="18" pageNumber="217">
<emphasis id="B0A5915F22545813C4630660FE8AFD61" box="[264,377,631,660]" italics="true" pageId="18" pageNumber="217">Ecology.</emphasis>
Most occurences are from mountainous level, but there are also low altitude catches, especially from
<collectingCountry id="FAC60DDD22545813C7EB068BFCEAFD4C" box="[640,793,668,697]" name="Switzerland" pageId="18" pageNumber="217">Switzerland</collectingCountry>
south of the Alps (
<collectingRegion id="401583AF22545813C15A068BFB7FFD4C" box="[1073,1164,668,697]" country="Switzerland" name="Ticino" pageId="18" pageNumber="217">Ticino</collectingRegion>
). Dates of capture when known range from March to beginning of August.
</paragraph>
</subSubSection>
</treatment>
</document>
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