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<mods:title>Synopsis of the genus Vitex (Lamiaceae) in the Democratic Republic of the Congo</mods:title>
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<mods:namePart>Meerts, Pierre</mods:namePart>
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<emphasis bold="true" box="[842,889,1675,1699]" pageId="7" pageNumber="387">11b.</emphasis>
<taxonomicName ID-CoL="5LS3R" ID-ENA="1899207" authority="(Britten) F. White" authorityName="F. White" baseAuthorityName="Britten" box="[899,1492,1675,1700]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="387" phylum="Tracheophyta" rank="subSpecies" species="madiensis" subSpecies="milanjiensis">
<emphasis bold="true" box="[899,1075,1675,1700]" italics="true" pageId="7" pageNumber="387">Vitex madiensis</emphasis>
subsp.
<emphasis bold="true" box="[1162,1295,1675,1699]" italics="true" pageId="7" pageNumber="387">milanjiensis</emphasis>
(Britten) F.White
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<bibRefCitation author="White F. &amp; Angus A." box="[849,1042,1706,1730]" journalOrPublisher="Oxford University Press, Oxford" pageId="7" pageNumber="387" refId="ref15105" refString="White F. (coll. Angus A.) (1962) Forest flora of Northern Rhodesia. Oxford University Press, Oxford." title="Forest flora of Northern Rhodesia" type="book" year="1962">White 1962: 455</bibRefCitation>
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);
<treatmentCitation author="Verdcourt B." box="[1065,1299,1706,1730]" page="61" pageId="7" pageNumber="387" year="1992">
<bibRefCitation author="Verdcourt B." box="[1065,1299,1706,1730]" editor="Polhill R. M." journalOrPublisher="Rotterdam, A. A. Balkema" pageId="7" pageNumber="387" refId="ref15028" refString="Verdcourt B. (1992) Verbenaceae. In: Polhill R. M. (ed) Flora of Tropical East Africa. Rotterdam, A. A. Balkema." title="Verbenaceae" type="book" volumeTitle="Flora of Tropical East Africa" year="1992">Verdcourt (1992: 61)</bibRefCitation>
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;
<treatmentCitation author="Lebrun J. - P. &amp; Stork A." page="524" pageId="7" pageNumber="387" year="1997">
<bibRefCitation author="Lebrun J. - P. &amp; Stork A." journalOrPublisher="Geneve, Conservatoire et Jardin botaniques de la ville de Geneve" pageId="7" pageNumber="387" refId="ref13855" refString="Lebrun J. - P., Stork A. (1997) Enumeration des plantes a fleurs d'Afrique. IV. Geneve, Conservatoire et Jardin botaniques de la ville de Geneve." title="Enumeration des plantes a fleurs d'Afrique. IV" type="book" year="1997">Lebrun &amp; Stork (1997: 524)</bibRefCitation>
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;
<treatmentCitation author="Sales F." box="[988,1177,1736,1760]" page="197" pageId="7" pageNumber="387" year="2001">
<bibRefCitation author="Sales F." box="[988,1177,1736,1760]" journalOrPublisher="Kew Bulletin" pageId="7" pageNumber="387" pagination="189 - 207" part="56" refId="ref14673" refString="Sales F. (2001) A synopsis of Vitex L. in the Flora Zambesiaca area. Kew Bulletin 56: 189 - 207. https: // doi. org / 10.2307 / 4119435" title="A synopsis of Vitex L. in the Flora Zambesiaca area" type="journal article" year="2001">Sales (2001: 197</bibRefCitation>
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;
<treatmentCitation author="Sales F." box="[1192,1292,1736,1760]" page="84" pageId="7" pageNumber="387" year="2005">
<bibRefCitation author="Sales F." box="[1192,1292,1736,1760]" editor="Pope E. V. &amp; Martins E. S." journalOrPublisher="Kew, Royal Botanic Gardens." pageId="7" pageNumber="387" pagination="73 - 92" part="8" refId="ref14709" refString="Sales F. (2005) Vitex. In: Pope E. V., Martins E. S. (eds) Flora Zam- besiaca 8 (7): 73 - 92. Kew, Royal Botanic Gardens." title="Vitex" type="book chapter" volumeTitle="Flora Zam- besiaca" year="2005">2005: 84</bibRefCitation>
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);
<treatmentCitation author="Coates Palgrave K &amp; Coates Palgrave M." page="983" pageId="7" pageNumber="387" year="2005">
<bibRefCitation author="Coates Palgrave K &amp; Coates Palgrave M." journalOrPublisher="Cape Town, Struik Nature" pageId="7" pageNumber="387" refId="ref12621" refString="Coates Palgrave K, Coates Palgrave M. (2005) Trees of Southern Africa. 3 rd Ed. Cape Town, Struik Nature." title="Trees of Southern Africa" type="book" year="2005">Coates Palgrave &amp; Coates Palgrave (2005: 983)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Meerts P. &amp; Hasson M." page="336" pageId="7" pageNumber="387" year="2016">
<bibRefCitation author="Meerts P. &amp; Hasson M." journalOrPublisher="Agentschap Plantentuin Meise" pageId="7" pageNumber="387" refId="ref14123" refString="Meerts P., Hasson M. (2016) Arbres et arbustes du Haut-Katanga. Meise, Agentschap Plantentuin Meise." title="Arbres et arbustes du Haut-Katanga. Meise" type="book" year="2016">Meerts &amp; Hasson (2016: 336)</bibRefCitation>
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;
<treatmentCitation author="Meerts P." box="[905,1114,1796,1820]" page="231" pageId="7" pageNumber="387" year="2016">
<bibRefCitation author="Meerts P." box="[905,1114,1796,1820]" journalOrPublisher="Phytotaxa" pageId="7" pageNumber="387" pagination="201 - 250" part="258" refId="ref14078" refString="Meerts P. (2016) An annotated checklist to the trees and shrubs of the Upper Katanga (D. R. Congo). Phytotaxa 258: 201 - 250. https: // doi. org / 10.11646 / phytotaxa. 258.3.1" title="An annotated checklist to the trees and shrubs of the Upper Katanga (D. R. Congo)" type="journal article" year="2016">Meerts (2016: 231)</bibRefCitation>
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.
</paragraph>
<paragraph blockId="7.[842,1492,1675,1971]" pageId="7" pageNumber="387">
<treatmentCitationGroup pageId="7" pageNumber="387">
<taxonomicName ID-CoL="5LS3R" authority="(Britten) W. Piep." authorityName="W. Piep." baseAuthorityName="Britten" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="387" phylum="Tracheophyta" rank="variety" species="madiensis" variety="milanjiensis">
<emphasis box="[1144,1310,1796,1820]" italics="true" pageId="7" pageNumber="387">Vitex madiensis</emphasis>
var.
<emphasis box="[1362,1491,1796,1820]" italics="true" pageId="7" pageNumber="387">milanjiensis</emphasis>
(Britten) W.Piep.
</taxonomicName>
<treatmentCitation author="Pieper W." box="[1037,1234,1826,1851]" page="63" pageId="7" pageNumber="387" year="1928">
(
<bibRefCitation author="Pieper W." bookContentInfo="XIII tables" box="[1044,1226,1826,1851]" journalOrPublisher="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzenge- ographie" pageId="7" pageNumber="387" pagination="1 - 91" part="142" refId="ref14553" refString="Pieper W. (1928) Vorarbeiten zu einer Revision der afrikanischen Vitex - Arten mit Berucksichtigung der ubrigen. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzenge- ographie 62, Beibl. 141 (&quot; 142 &quot;): 1 - 91 (+ XIII tables)." title="Vorarbeiten zu einer Revision der afrikanischen Vitex - Arten mit Berucksichtigung der ubrigen" type="book chapter" year="1928">Pieper 1928: 63</bibRefCitation>
)
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;
<treatmentCitation author="De Wildeman E." page="12" pageId="7" pageNumber="387" year="1929">
<bibRefCitation author="De Wildeman E." journalOrPublisher="Paris, Lechevalier" pageId="7" pageNumber="387" part="5" refId="ref13190" refString="De Wildeman E. (1929 a) Plantae Bequaertianae 5 (1). Paris, Lechevalier." title="Plantae Bequaertianae" type="book" year="1929">De Wildeman (1929a: 12</bibRefCitation>
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;
<treatmentCitation author="De Wildeman E." box="[885,1012,1856,1881]" page="105" pageId="7" pageNumber="387" year="1929">
<bibRefCitation author="De Wildeman E." box="[885,1012,1856,1881]" journalOrPublisher="Bruxelles, Comite special du Katanga" pageId="7" pageNumber="387" refId="ref13209" refString="De Wildeman E. (1929 b) Contribution a la flore du Katanga. Sup- plement 2. Bruxelles, Comite special du Katanga." title="Contribution a la flore du Katanga. Sup- plement 2" type="book" year="1929">1929b: 105</bibRefCitation>
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);
<treatmentCitation author="Delevoy G." box="[1034,1265,1857,1881]" page="483" pageId="7" pageNumber="387" year="1929">
<bibRefCitation author="Delevoy G." box="[1034,1265,1857,1881]" journalOrPublisher="Bruxelles, Comite Special du Katanga" pageId="7" pageNumber="387" refId="ref12647" refString="Delevoy G. (1929) La question forestiere au Katanga. II. Les essences forestieres au Katanga. Bruxelles, Comite Special du Katanga." title="La question forestiere au Katanga. II. Les essences forestieres au Katanga" type="book" year="1929">Delevoy (1929: 483)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph blockId="7.[842,1492,1675,1971]" pageId="7" pageNumber="387">
<treatmentCitationGroup pageId="7" pageNumber="387">
<taxonomicName ID-CoL="5BL92" authority="Britten" authorityName="Britten" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="387" phylum="Tracheophyta" rank="species" species="milanjiensis">
<emphasis box="[1300,1492,1857,1881]" italics="true" pageId="7" pageNumber="387">Vitex milanjiensis</emphasis>
Britten
</taxonomicName>
<treatmentCitation author="Britten J." box="[931,1141,1887,1911]" page="36" pageId="7" pageNumber="387" year="1894">
(
<bibRefCitation author="Britten J." box="[938,1133,1887,1911]" journalOrPublisher="Transactions of the Linnean Society of London" pageId="7" pageNumber="387" pagination="16 - 37" part="4" refId="ref12526" refString="Britten J. (1894) Monopetalae. In: The Plants of Milanji, Nyasaland. Transactions of the Linnean Society of London, Series 2, Botany 4: 16 - 37." title="Monopetalae" type="journal article" volumeTitle="The Plants of Milanji, Nyasaland" year="1894">Britten 1894: 36</bibRefCitation>
)
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;
<treatmentCitation author="Baker J. G." box="[1161,1375,1887,1911]" page="330" pageId="7" pageNumber="387" year="1900">
<bibRefCitation author="Baker J. G." box="[1161,1375,1887,1911]" editor="Thiselton-Dyer W. T." journalOrPublisher="London, Lovell Reeve &amp; Co." pageId="7" pageNumber="387" pagination="315 - 332" part="5" refId="ref12379" refString="Baker J. G. (1900) Vitex. In: Thiselton-Dyer W. T. (ed.) Flora of tropical Africa 5 (2): 315 - 332. London, Lovell Reeve &amp; Co." title="Vitex" type="journal article" volumeTitle="Flora of tropical Africa" year="1900">Baker (1900: 330)</bibRefCitation>
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.
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<emphasis bold="true" box="[1393,1406,1887,1911]" pageId="7" pageNumber="387"></emphasis>
<materialsCitation accessionNumber="BM001124551" collectingDate="1891-10-31" collectorName="Whyte" country="Malawi" elevation="1800" municipality="Mulanje" pageId="7" pageNumber="387" typeStatus="syntype">
<typeStatus box="[1421,1489,1887,1911]" pageId="7" pageNumber="387">Types</typeStatus>
: Malawi, Milanji [Mulanje], 6000 ft [
<quantity box="[1258,1351,1917,1941]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="387" unit="m" value="1800.0">1800 m</quantity>
],
<date box="[1374,1487,1917,1941]" pageId="7" pageNumber="387" value="1891-10">Oct. 1891</date>
,
<emphasis box="[842,909,1947,1971]" italics="true" pageId="7" pageNumber="387">Whyte</emphasis>
138 (syn-: BM barcode BM001124551);
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<materialsCitation accessionNumber="BM000566972, K000249031" collectingDate="1891-09-30" collectorName="White" country="Malawi" lastPageId="8" lastPageNumber="388" municipality="Zomba" pageId="7" pageNumber="387" typeStatus="syntype">
Malawi, Zomba,
<date box="[201,334,189,213]" pageId="8" pageNumber="388" value="1891-09">Sept. 1891</date>
,
<emphasis box="[361,428,189,213]" italics="true" pageId="8" pageNumber="388">Whyte</emphasis>
s.n. (syn-:
<pageTitle box="[596,637,190,213]" pageId="8" pageNumber="388">BM</pageTitle>
barcode BM000566972; Kbarcode K000249031
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).
</paragraph>
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<paragraph blockId="8.[95,746,263,1971]" box="[96,503,263,287]" pageId="8" pageNumber="388">
<treatmentCitationGroup box="[96,503,263,287]" pageId="8" pageNumber="388">
<taxonomicName ID-CoL="5BL56" authority="Baker" authorityName="Baker" box="[96,293,263,287]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="grisea">
<emphasis box="[96,223,264,287]" italics="true" pageId="8" pageNumber="388">Vitex grisea</emphasis>
Baker
</taxonomicName>
<treatmentCitation author="Baker J. G." box="[300,499,263,287]" page="325" pageId="8" pageNumber="388" year="1900">
(
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)
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.
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</paragraph>
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<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<materialsCitation accessionNumber="K001067050" collectorName="Welwitsch" country="Angola" location="Welwitsch" pageId="8" pageNumber="388" specimenCount="1" stateProvince="Distr. Huilla" typeStatus="lectotype">
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:
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,
<collectingRegion pageId="8" pageNumber="388">Distr. Huilla</collectingRegion>
, in silvestribus alte dumetosis de Catumba,
<emphasis box="[639,746,293,317]" italics="true" pageId="8" pageNumber="388">
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5759 (lecto-:
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001067050,
<emphasis bold="true" box="[551,738,322,346]" pageId="8" pageNumber="388">designated here</emphasis>
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;
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isolecto-:
<collectionCode box="[202,243,352,375]" pageId="8" pageNumber="388">BM</collectionCode>
barcode
<collectionCode box="[341,383,351,375]" pageId="8" pageNumber="388">BM</collectionCode>
000839718),
<emphasis bold="true" box="[525,647,351,375]" pageId="8" pageNumber="388">synon. nov</emphasis>
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<emphasis bold="true" box="[647,652,351,375]" pageId="8" pageNumber="388">.</emphasis>
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<subSubSection pageId="8" pageNumber="388" type="reference_group">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<treatmentCitationGroup pageId="8" pageNumber="388">
<taxonomicName ID-CoL="5BL5Y" authority="Hiern" authorityName="Hiern" box="[96,342,388,412]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="huillensis">
<emphasis box="[96,266,388,412]" italics="true" pageId="8" pageNumber="388">Vitex huillensis</emphasis>
Hiern
</taxonomicName>
(
<treatmentCitation author="Hiern W. P." box="[364,562,388,412]" page="837" pageId="8" pageNumber="388" year="1900">
<bibRefCitation author="Hiern W. P." box="[364,562,388,412]" journalOrPublisher="London, Longmans" pageId="8" pageNumber="388" refId="ref13678" refString="Hiern W. P. (1900) Catalogue of African plants collected by Dr Friedrich Welwitsch in 1853 - 1861. Dicotyledons. IV. London, Longmans." title="Catalogue of African plants collected by Dr Friedrich Welwitsch in 1853 - 1861. Dicotyledons. IV" type="book" year="1900">Hiern 1900: 837</bibRefCitation>
</treatmentCitation>
);
<treatmentCitation author="De Wildeman E." page="191" pageId="8" pageNumber="388" year="1914">
<bibRefCitation author="De Wildeman E." journalOrPublisher="Bulletin du Jardin botanique de l'Etat a Bruxelles" pageId="8" pageNumber="388" pagination="1 - 241" part="4" refId="ref13065" refString="De Wildeman E. (1914 b) Additions a la Flore du Congo. Bulletin du Jardin botanique de l'Etat a Bruxelles 4: 1 - 241. https: // doi. org / 10.2307 / 3666479" title="Additions a la Flore du Congo" type="journal article" year="1914">De Wildeman (1914b: 191)</bibRefCitation>
</treatmentCitation>
nom. illeg.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<treatmentCitationGroup pageId="8" pageNumber="388">
<taxonomicName ID-CoL="5BL5R" authority="De Wild." authorityName="De Wild." box="[96,342,455,479]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="hockii">
<emphasis box="[96,226,455,479]" italics="true" pageId="8" pageNumber="388">Vitex hockii</emphasis>
De Wild.
</taxonomicName>
(
<treatmentCitation author="De Wildeman E." box="[361,656,455,479]" page="113" pageId="8" pageNumber="388" year="1914">
<bibRefCitation author="De Wildeman E." box="[361,656,455,479]" journalOrPublisher="Bulletin du Jardin botanique de l'Etat a Bruxelles" pageId="8" pageNumber="388" pagination="1 - 241" part="4" refId="ref13065" refString="De Wildeman E. (1914 b) Additions a la Flore du Congo. Bulletin du Jardin botanique de l'Etat a Bruxelles 4: 1 - 241. https: // doi. org / 10.2307 / 3666479" title="Additions a la Flore du Congo" type="journal article" year="1914">De Wildeman 1914b: 113</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="De Wildeman E." page="403" pageId="8" pageNumber="388" year="1914">
<bibRefCitation author="De Wildeman E." journalOrPublisher="Annales de la Societe scientifique de Bruxelles" pageId="8" pageNumber="388" pagination="353 - 465" part="38" refId="ref13104" refString="De Wildeman E. (1914 c) Notes sur la flore du Katanga. Annales de la Societe scientifique de Bruxelles 38: 353 - 465." title="Notes sur la flore du Katanga" type="journal article" year="1914">1914c: 403</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="De Wildeman E." box="[152,265,484,508]" page="164" pageId="8" pageNumber="388" year="1921">
<bibRefCitation author="De Wildeman E." box="[152,265,484,508]" journalOrPublisher="Bruxelles, Comite special du Katanga" pageId="8" pageNumber="388" refId="ref13132" refString="De Wildeman E. (1921) Contribution a l'etude de la flore du Ka- tanga. Bruxelles, Comite special du Katanga. https: // doi. org / 10.5962 / bhl. title. 11020" title="Contribution a l'etude de la flore du Ka- tanga" type="book" year="1921">1921: 164</bibRefCitation>
</treatmentCitation>
);
<treatmentCitation author="Pieper W." box="[286,479,484,508]" page="61" pageId="8" pageNumber="388" year="1928">
<bibRefCitation author="Pieper W." bookContentInfo="XIII tables" box="[286,479,484,508]" journalOrPublisher="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzenge- ographie" pageId="8" pageNumber="388" pagination="1 - 91" part="142" refId="ref14553" refString="Pieper W. (1928) Vorarbeiten zu einer Revision der afrikanischen Vitex - Arten mit Berucksichtigung der ubrigen. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzenge- ographie 62, Beibl. 141 (&quot; 142 &quot;): 1 - 91 (+ XIII tables)." title="Vorarbeiten zu einer Revision der afrikanischen Vitex - Arten mit Berucksichtigung der ubrigen" type="book chapter" year="1928">Pieper (1928: 61)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Delevoy G." box="[491,720,484,508]" page="483" pageId="8" pageNumber="388" year="1929">
<bibRefCitation author="Delevoy G." box="[491,720,484,508]" journalOrPublisher="Bruxelles, Comite Special du Katanga" pageId="8" pageNumber="388" refId="ref12647" refString="Delevoy G. (1929) La question forestiere au Katanga. II. Les essences forestieres au Katanga. Bruxelles, Comite Special du Katanga." title="La question forestiere au Katanga. II. Les essences forestieres au Katanga" type="book" year="1929">Delevoy (1929: 483)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="materials_examined">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<materialsCitation collectingDate="1911-10-31" collectorName="Hock" country="Democratic Republic of the Congo" municipality="Manika" pageId="8" pageNumber="388" specimenCount="1" stateProvince="Haut-Katanga" typeStatus="holotype">
<typeStatus box="[98,155,514,538]" pageId="8" pageNumber="388">Type</typeStatus>
:
<collectingCountry box="[166,298,513,537]" pageId="8" pageNumber="388" value="Democratic Republic of the Congo">D.R. Congo</collectingCountry>
,
<collectingMunicipality box="[310,462,514,538]" pageId="8" pageNumber="388">Haut-Katanga</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03D1532200561E46FC8C56CEFA49FEB3:8EA7B4EF00591E46FE1C5244FDFAFD9F" box="[474,558,513,537]" country="D. R. Congo" municipality="Haut-Katanga" name="Manika" pageId="8" pageNumber="388">Manika</location>
,
<date box="[571,679,513,537]" pageId="8" pageNumber="388" value="1911-10">
<collectingDate box="[571,679,513,537]" pageId="8" pageNumber="388" value="1911-10">Oct. 1911</collectingDate>
</date>
,
<emphasis box="[690,746,513,537]" italics="true" pageId="8" pageNumber="388">
<location LSID="urn:lsid:plazi:treatment:03D1532200561E46FC8C56CEFA49FEB3:8EA7B4EF00591E46FD745244FD3EFD9F" box="[690,746,513,537]" country="D. R. Congo" municipality="Haut-Katanga" name="Hock" pageId="8" pageNumber="388">Hock</location>
</emphasis>
s.n. (holo-:
<collectionCode box="[219,255,543,566]" pageId="8" pageNumber="388">BR</collectionCode>
barcode
<collectionCode box="[352,387,543,567]" pageId="8" pageNumber="388">BR</collectionCode>
0000008906557)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="reference_group">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<treatmentCitationGroup pageId="8" pageNumber="388">
<taxonomicName ID-CoL="5BLBW" authority="De Wild." authorityName="De Wild." box="[96,358,580,604]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="ringoetii">
<emphasis box="[96,250,580,603]" italics="true" pageId="8" pageNumber="388">Vitex ringoetii</emphasis>
De Wild.
</taxonomicName>
(
<treatmentCitation author="De Wildeman E." box="[374,659,580,604]" page="143" pageId="8" pageNumber="388" year="1914">
<bibRefCitation author="De Wildeman E." box="[374,659,580,604]" journalOrPublisher="Repertorium specierum novarum regni vegetabilis" pageId="8" pageNumber="388" pagination="137 - 147" part="13" refId="ref13023" refString="De Wildeman E. (1914 a) Decades novarum specierum florae Katan- gensis. XIX - XXI. Repertorium specierum novarum regni vegetabilis 13: 137 - 147. https: // doi. org / 10.1002 / fedr. 19140130904" title="Decades novarum specierum florae Katan- gensis. XIX - XXI" type="journal article" year="1914">De Wildeman 1914a: 143</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="De Wildeman E." page="407" pageId="8" pageNumber="388" year="1914">
<bibRefCitation author="De Wildeman E." journalOrPublisher="Annales de la Societe scientifique de Bruxelles" pageId="8" pageNumber="388" pagination="353 - 465" part="38" refId="ref13104" refString="De Wildeman E. (1914 c) Notes sur la flore du Katanga. Annales de la Societe scientifique de Bruxelles 38: 353 - 465." title="Notes sur la flore du Katanga" type="journal article" year="1914">1914c: 407</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="De Wildeman E." box="[157,276,609,633]" page="165" pageId="8" pageNumber="388" year="1921">
<bibRefCitation author="De Wildeman E." box="[157,276,609,633]" journalOrPublisher="Bruxelles, Comite special du Katanga" pageId="8" pageNumber="388" refId="ref13132" refString="De Wildeman E. (1921) Contribution a l'etude de la flore du Ka- tanga. Bruxelles, Comite special du Katanga. https: // doi. org / 10.5962 / bhl. title. 11020" title="Contribution a l'etude de la flore du Ka- tanga" type="book" year="1921">1921: 165</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="materials_examined">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<materialsCitation accessionNumber="BR0000005571505" collectingDate="1912-10-31" collectionCode="BR" collectorName="Ringoet" country="Democratic Republic of the Congo" municipality="Nieuwdorp" pageId="8" pageNumber="388" specimenCount="1" typeStatus="holotype">
<typeStatus box="[329,386,610,634]" pageId="8" pageNumber="388">Type</typeStatus>
:
<collectingCountry box="[403,542,609,633]" pageId="8" pageNumber="388" value="Democratic Republic of the Congo">D.R. Congo</collectingCountry>
,
<collectingMunicipality box="[559,682,609,633]" pageId="8" pageNumber="388">Nieuwdorp</collectingMunicipality>
,
<date pageId="8" pageNumber="388" value="1912-10">
<collectingDate pageId="8" pageNumber="388" value="1912-10">Oct. 1912</collectingDate>
</date>
,
<emphasis box="[163,246,639,662]" italics="true" pageId="8" pageNumber="388">
<location LSID="urn:lsid:plazi:treatment:03D1532200561E46FC8C56CEFA49FEB3:8EA7B4EF00591E46FF65523AFF22FD10" box="[163,246,639,662]" country="D. R. Congo" municipality="Nieuwdorp" name="Ringoet" pageId="8" pageNumber="388">Ringoet</location>
</emphasis>
1 (holo-:
<collectionCode box="[352,388,639,662]" pageId="8" pageNumber="388">BR</collectionCode>
barcode
<collectionCode box="[485,520,638,662]" pageId="8" pageNumber="388">BR</collectionCode>
0000005571505)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="reference_group">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<treatmentCitationGroup pageId="8" pageNumber="388">
<treatmentCitationGroup pageId="8" pageNumber="388">
<taxonomicName ID-CoL="5BL43" authority="Mildbr. ex W. Piep." authorityName="W. Piep." box="[96,492,675,699]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="epidictyodes">
<emphasis box="[96,287,675,699]" italics="true" pageId="8" pageNumber="388">Vitex epidictyodes</emphasis>
Mildbr. ex W.Piep.
</taxonomicName>
(
<treatmentCitation author="Pieper W." box="[504,674,675,700]" page="61" pageId="8" pageNumber="388" year="1928">
<bibRefCitation author="Pieper W." bookContentInfo="XIII tables" box="[504,674,675,700]" journalOrPublisher="Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzenge- ographie" pageId="8" pageNumber="388" pagination="1 - 91" part="142" refId="ref14553" refString="Pieper W. (1928) Vorarbeiten zu einer Revision der afrikanischen Vitex - Arten mit Berucksichtigung der ubrigen. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzenge- ographie 62, Beibl. 141 (&quot; 142 &quot;): 1 - 91 (+ XIII tables)." title="Vorarbeiten zu einer Revision der afrikanischen Vitex - Arten mit Berucksichtigung der ubrigen" type="book chapter" year="1928">Pieper 1928: 61</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Pieper W." page="164" pageId="8" pageNumber="388" year="1929">
<bibRefCitation author="Pieper W." journalOrPublisher="Feddes Repertorium Specierum no- varum regni vegetabilis" pageId="8" pageNumber="388" pagination="161 - 166" part="26" refId="ref14602" refString="Pieper W. (1929) Neue und noch nicht beschriebene Arten der Gattung Vitex aus Afrika. Feddes Repertorium Specierum no- varum regni vegetabilis 26: 161 - 166. https: // doi. org / 10.1002 / fedr. 19290260704" title="Neue und noch nicht beschriebene Arten der Gattung Vitex aus Afrika" type="journal article" year="1929">1929: 164</bibRefCitation>
)
</treatmentCitation>
</treatmentCitationGroup>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<treatmentCitationGroup pageId="8" pageNumber="388">
<treatmentCitationGroup pageId="8" pageNumber="388">
<taxonomicName ID-CoL="5S433" authority="(W. Piep.) Verdc." authorityName="Verdc." baseAuthorityName="W. Piep." class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="variety" species="madiensis" subSpecies="milanjiensis" variety="epidictyodes">
<emphasis box="[187,358,705,729]" italics="true" pageId="8" pageNumber="388">Vitex madiensis</emphasis>
subsp.
<emphasis box="[448,577,705,729]" italics="true" pageId="8" pageNumber="388">milanjiensis</emphasis>
var.
<emphasis italics="true" pageId="8" pageNumber="388">epidictyodes</emphasis>
(W.Piep.) Verdc.
</taxonomicName>
<treatmentCitation author="Verdcourt B." box="[328,559,734,758]" page="61" pageId="8" pageNumber="388" year="1992">
(
<bibRefCitation author="Verdcourt B." box="[335,549,734,758]" editor="Polhill R. M." journalOrPublisher="Rotterdam, A. A. Balkema" pageId="8" pageNumber="388" refId="ref15028" refString="Verdcourt B. (1992) Verbenaceae. In: Polhill R. M. (ed) Flora of Tropical East Africa. Rotterdam, A. A. Balkema." title="Verbenaceae" type="book" volumeTitle="Flora of Tropical East Africa" year="1992">Verdcourt 1992: 61</bibRefCitation>
)
</treatmentCitation>
;
<treatmentCitation author="Lebrun J. - P. &amp; Stork A." page="525" pageId="8" pageNumber="388" year="1997">
<bibRefCitation author="Lebrun J. - P. &amp; Stork A." journalOrPublisher="Geneve, Conservatoire et Jardin botaniques de la ville de Geneve" pageId="8" pageNumber="388" refId="ref13855" refString="Lebrun J. - P., Stork A. (1997) Enumeration des plantes a fleurs d'Afrique. IV. Geneve, Conservatoire et Jardin botaniques de la ville de Geneve." title="Enumeration des plantes a fleurs d'Afrique. IV" type="book" year="1997">Lebrun &amp; Stork (1997: 525)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="materials_examined">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<materialsCitation accessionNumber="B100279988" collectorName="Stolz" country="Malawi" location="am nordende des Nyassa" municipality="Langenburg" pageId="8" pageNumber="388" specimenCount="1" typeStatus="lectotype">
Type:
<collectingCountry box="[325,409,763,787]" name="Malawi" pageId="8" pageNumber="388" value="Malawi">Malawi</collectingCountry>
,
<collectingMunicipality box="[420,551,763,787]" pageId="8" pageNumber="388">Langenburg</collectingMunicipality>
, am nordende des
<location LSID="urn:lsid:plazi:treatment:03D1532200561E46FC8C56CEFA49FEB3:8EA7B4EF00591E46FFA7535CFF65FCB7" box="[97,177,793,817]" country="Malawi" municipality="Langenburg" name="Nyassa" pageId="8" pageNumber="388">Nyassa</location>
,
<emphasis box="[190,242,793,817]" italics="true" pageId="8" pageNumber="388">
<location LSID="urn:lsid:plazi:treatment:03D1532200561E46FC8C56CEFA49FEB3:8EA7B4EF00591E46FF78535CFF26FCB7" box="[190,242,793,817]" country="Malawi" municipality="Langenburg" name="Stolz" pageId="8" pageNumber="388">Stolz</location>
</emphasis>
556 (First-step
<typeStatus box="[421,519,793,817]" pageId="8" pageNumber="388">lectotype</typeStatus>
designated by Pieper (1929); second-step lectotype: Bbarcode B 100279988,
<emphasis bold="true" box="[96,283,851,876]" pageId="8" pageNumber="388">designated here</emphasis>
</materialsCitation>
;
<materialsCitation accessionNumber="BM001125198, G00366697, P00713461, U0040942, Z000067396" collectorName="Stolz" country="Malawi" location="am nordende des Nyassa" municipality="Langenburg" pageId="8" pageNumber="388" specimenCount="1" typeStatus="isolectotype">isolecto-: BM barcode BM001125198, Gbarcode G00366697, Pbarcode P00713461, Ubarcode U0040942, Zbarcode Z-000067396)</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="description">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
Differs from the
<typeStatus box="[318,364,948,972]" pageId="8" pageNumber="388">type</typeStatus>
subspecies by the following traits. A shrub. Twigs stout, often wine red. Leaves 5 foliolate; lamina abruptly contracted at tip and cuspidate, very coriaceous, entire; upper surface very scabrid, reticulum prominulent.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="distribution">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<emphasis bold="true" box="[96,235,1072,1096]" pageId="8" pageNumber="388">Distribution</emphasis>
Zambezian region, from Angola eastwards to Tanzania and Mozambique.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="388" type="discussion">
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
<emphasis bold="true" box="[96,160,1139,1163]" pageId="8" pageNumber="388">Notes</emphasis>
Dwarf, geofrutescent forms of
<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="subSpecies" species="madiensis" subSpecies="milanjiensis">
<emphasis box="[533,669,1139,1163]" italics="true" pageId="8" pageNumber="388">V. madiensis</emphasis>
subsp.
<emphasis box="[96,225,1168,1192]" italics="true" pageId="8" pageNumber="388">milanjiensis</emphasis>
</taxonomicName>
occur throughout the distribution range of
<taxonomicName box="[96,305,1198,1222]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="subSpecies" species="madiensis" subSpecies="milanjiensis">
subsp.
<emphasis box="[176,305,1198,1222]" italics="true" pageId="8" pageNumber="388">milanjiensis</emphasis>
</taxonomicName>
. They have been repeatedly described as
<taxonomicName authority="De Wild." authorityName="De Wild." box="[127,330,1227,1251]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="hockii">
<emphasis box="[127,220,1227,1251]" italics="true" pageId="8" pageNumber="388">V. hockii</emphasis>
De Wild.
</taxonomicName>
,
<taxonomicName authority="De Wild." authorityName="De Wild." box="[343,572,1227,1251]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="ringoetii">
<emphasis box="[343,464,1228,1251]" italics="true" pageId="8" pageNumber="388">V. ringoetii</emphasis>
De Wild.
</taxonomicName>
,
<taxonomicName authority="W. Piep." authorityName="W. Piep." class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="epidictyodes">
<emphasis box="[585,746,1227,1251]" italics="true" pageId="8" pageNumber="388">V. epidictyodes</emphasis>
W.Piep.
</taxonomicName>
,
<taxonomicName authority="Exell. Such" authorityName="Exell. Such" box="[195,469,1256,1280]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="caespitosa">
<emphasis box="[195,336,1257,1280]" italics="true" pageId="8" pageNumber="388">V. caespitosa</emphasis>
Exell. Such
</taxonomicName>
forms pose two different taxonomic problems. First, it is unclear if they are genetically fixed or if, alternatively, they represent pyrophytic forms induced by burning;
<bibRefCitation author="Verdcourt B." box="[328,515,1344,1369]" editor="Polhill R. M." journalOrPublisher="Rotterdam, A. A. Balkema" pageId="8" pageNumber="388" refId="ref15028" refString="Verdcourt B. (1992) Verbenaceae. In: Polhill R. M. (ed) Flora of Tropical East Africa. Rotterdam, A. A. Balkema." title="Verbenaceae" type="book" volumeTitle="Flora of Tropical East Africa" year="1992">Verdcourt (1992)</bibRefCitation>
favoured the former hypothesis. Secondly, assuming it is genetically fixed, it is not clear if that morph is monophyletic or, alternatively, if it has evolved several times independently in response to selection by fire or other environmental constraints. Ihave not been able to find differences between dwarf and tall morphs except size.
</paragraph>
<paragraph blockId="8.[95,746,263,1971]" pageId="8" pageNumber="388">
Lectotypification of
<taxonomicName authority="Mildbr. ex W. Piep." authorityName="W. Piep." class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="epidictyodes">
<emphasis box="[385,594,1558,1582]" italics="true" pageId="8" pageNumber="388">Vitex epidictyodes</emphasis>
Mildbr. ex W.Piep.
</taxonomicName>
Three
<typeStatus box="[265,358,1588,1612]" pageId="8" pageNumber="388">syntypes</typeStatus>
are cited in the protologue (
<emphasis box="[693,745,1587,1611]" italics="true" pageId="8" pageNumber="388">Stolz</emphasis>
556;
<emphasis box="[152,204,1616,1640]" italics="true" pageId="8" pageNumber="388">Stolz</emphasis>
1478;
<emphasis box="[283,353,1616,1640]" italics="true" pageId="8" pageNumber="388">Trotha</emphasis>
95).
<bibRefCitation author="Pieper W." box="[413,562,1616,1641]" journalOrPublisher="Feddes Repertorium Specierum no- varum regni vegetabilis" pageId="8" pageNumber="388" pagination="161 - 166" part="26" refId="ref14602" refString="Pieper W. (1929) Neue und noch nicht beschriebene Arten der Gattung Vitex aus Afrika. Feddes Repertorium Specierum no- varum regni vegetabilis 26: 161 - 166. https: // doi. org / 10.1002 / fedr. 19290260704" title="Neue und noch nicht beschriebene Arten der Gattung Vitex aus Afrika" type="journal article" year="1929">Pieper (1929)</bibRefCitation>
designated
<emphasis box="[693,745,1616,1640]" italics="true" pageId="8" pageNumber="388">Stolz</emphasis>
556 as the
<typeStatus box="[216,314,1646,1670]" pageId="8" pageNumber="388">lectotype</typeStatus>
(see
<bibRefCitation author="Verdcourt B." box="[373,544,1646,1670]" editor="Polhill R. M." journalOrPublisher="Rotterdam, A. A. Balkema" pageId="8" pageNumber="388" refId="ref15028" refString="Verdcourt B. (1992) Verbenaceae. In: Polhill R. M. (ed) Flora of Tropical East Africa. Rotterdam, A. A. Balkema." title="Verbenaceae" type="book" volumeTitle="Flora of Tropical East Africa" year="1992">Verdcourt 1992</bibRefCitation>
). Many specimens of that gathering have been distributed and
<bibRefCitation author="Pieper W." box="[559,705,1675,1699]" journalOrPublisher="Feddes Repertorium Specierum no- varum regni vegetabilis" pageId="8" pageNumber="388" pagination="161 - 166" part="26" refId="ref14602" refString="Pieper W. (1929) Neue und noch nicht beschriebene Arten der Gattung Vitex aus Afrika. Feddes Repertorium Specierum no- varum regni vegetabilis 26: 161 - 166. https: // doi. org / 10.1002 / fedr. 19290260704" title="Neue und noch nicht beschriebene Arten der Gattung Vitex aus Afrika" type="journal article" year="1929">Pieper (1929)</bibRefCitation>
did not indicate a specific herbarium; therefore second-step lectotypification is necessary (art 9.17 of ICN;
<bibRefCitation author="McNeill J. &amp; Turland N. J. &amp; Barrie F. R. &amp; Buck W. R. &amp; Greuter W. &amp; Wiersema J. H." journalOrPublisher="Konigstein, Koeltz Scientific Books" pageId="8" pageNumber="388" refId="ref14022" refString="McNeill J., Turland N. J., Barrie F. R., Buck W. R., Greuter W., Wiersema J. H. (2012) International Code of Nomenclature for Algae, Fungi, and Plants. Regnum Vegetabile 154. Konigstein, Koeltz Scientific Books." title="International Code of Nomenclature for Algae, Fungi, and Plants" type="book" volumeTitle="Regnum Vegetabile 154" year="2012">McNeill et al. 2012</bibRefCitation>
). Since Pieper worked in Berlin, Itherefore select the specimen kept in Berlin as the
<typeStatus box="[425,526,1793,1817]" pageId="8" pageNumber="388">lectotype</typeStatus>
.
</paragraph>
<paragraph blockId="8.[95,746,263,1971]" lastBlockId="8.[786,1437,189,309]" pageId="8" pageNumber="388">
Lectotypification of
<taxonomicName authority="Baker." authorityName="Baker." box="[354,559,1830,1854]" class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="species" species="grisea">
<emphasis box="[354,482,1830,1853]" italics="true" pageId="8" pageNumber="388">Vitex grisea</emphasis>
Baker.
</taxonomicName>
The specimen of
<emphasis box="[97,204,1859,1883]" italics="true" pageId="8" pageNumber="388">Welwitsch</emphasis>
5759 deposited in Kis chosen as the
<typeStatus box="[609,707,1859,1883]" pageId="8" pageNumber="388">lectotype</typeStatus>
for two reasons. First it shows better the acute leaflet tips mentioned in the protologue; secondly, Baker worked at Kew Botanic Gardens after Welwitschs collections were split and, therefore, almost certainly used the duplicates deposited in K. This morphotype is a somewhat extreme variant in the phenetic variation space of the polymorphic
<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Vitex" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="388" phylum="Tracheophyta" rank="subSpecies" species="madiensis" subSpecies="milanjiensis">
<emphasis box="[1218,1384,253,277]" italics="true" pageId="8" pageNumber="388">Vitex madiensis</emphasis>
subsp.
<emphasis box="[824,953,285,309]" italics="true" pageId="8" pageNumber="388">milanjiensis</emphasis>
</taxonomicName>
not deserving formal taxonomic recognition.
</paragraph>
</subSubSection>
</treatment>
</document>