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<document id="28C8BD838DCBEC27271B4AC89B4E8361" ID-DOI="10.5281/zenodo.5377199" ID-ISSN="1638-9395" ID-Zenodo-Dep="5377199" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630282681036" checkinUser="marcus" docAuthor="Popov, Vasil V." docDate="2004" docId="03B287E9FFAEFFB0FF0B661BB853FE48" docLanguage="en" docName="Geodiversitas.2004.26.3.403-491.pdf" docOrigin="Geodiversitas 26 (3)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Allocricetus bursae Schaub 1930" docType="treatment" docVersion="4" lastPageNumber="460" masterDocId="FF8BFF91FF97FF8AFFC5650BB968FFA1" masterDocTitle="Pliocene small mammals (Mammalia, Lipotyphla, Chiroptera, Lagomorpha, Rodentia) from Muselievo (North Bulgaria)" masterLastPageNumber="491" masterPageNumber="403" pageNumber="460" updateTime="1699212936602" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="6B61F4E9F091FC31670A212284DEFF5A">Pliocene small mammals (Mammalia, Lipotyphla, Chiroptera, Lagomorpha, Rodentia) from Muselievo (North Bulgaria)</mods:title>
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<mods:namePart id="6AA7C795E1F94D0F72D0E9B49BA0EE65">Popov, Vasil V.</mods:namePart>
<mods:affiliation id="6E627A7DC0977DD044469B65B329FE34">Institute of Zoology, Bulgarian Academy of Sciences, Boul. Tsar Osvoboditel 1, 1000 Sofia (Republic of Bulgaria) popov @ zoology. bas. bg</mods:affiliation>
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<subSubSection id="C3016574FFAEFFB3FF0B661BB8A2FCEB" pageId="57" pageNumber="460" type="nomenclature">
<paragraph id="8BA436FFFFAEFFB3FF0B661BB8A2FCEB" blockId="57.[206,564,784,842]" pageId="57" pageNumber="460">
<heading id="D0EC8193FFAEFFB3FF0B661BB8A2FCEB" centered="true" fontSize="11" level="2" pageId="57" pageNumber="460" reason="2">
<taxonomicName id="4C1B4D7CFFAEFFB3FF0B661BBB5CFC8B" authority="Schaub, 1930" authorityName="Schaub" authorityYear="1930" box="[206,564,784,810]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFAEFFB3FF0B661BB8FFFC8B" bold="true" box="[206,407,784,810]" italics="true" pageId="57" pageNumber="460">Allocricetus bursae</emphasis>
<bibRefCitation id="EF8A4B0EFFAEFFB3FE5A661BBB5CFC8B" author="SCHAUB S." box="[415,564,784,810]" pageId="57" pageNumber="460" pagination="1 - 49" refId="ref46460" refString="SCHAUB S. 1930. - Quartare und jungtertiare Hamster. Abhandlung der Schweizerischen Palaonologischen Gesellschaft 49 (6): 1 - 49, pls 1, 2." type="book chapter" year="1930">Schaub, 1930</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13202A7AFFAEFFB3FE85663BB8E5FCEB" box="[320,397,816,842]" captionStart="FIG" captionStartId="44.[119,130,1625,1642]" captionTargetBox="[199,1133,306,1619]" captionTargetId="figure-22@44.[199,1133,306,1631]" captionTargetPageId="44" captionText="FIG. 24. — A, Pliopetaurista dehneli (Sulimski, 1964), left p4, Ms141, occlusal view; B, Sciurus sp., left M1-2, Ms142, occlusal view; C,?Spermophilus sp., right p4, Ms143, occlusal view; D-G, Allocricetus bursae Schaub, 1930; D, right m1, Ms309, occlusal view; E, right m1, Ms310, occlusal view; F, left M1, Ms313, occlusal view; G, left M1, Ms311, occlusal view; H, Tamias orlovi (Sulimski, 1964), left M3, Ms144, occlusal view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5377239" httpUri="https://zenodo.org/record/5377239/files/figure.png" pageId="57" pageNumber="460">Fig. 24</figureCitation>
D-G)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016574FFAEFFB3FFB36679B8B4FC2B" box="[118,476,882,906]" pageId="57" pageNumber="460" type="reference_group">
<paragraph id="8BA436FFFFAEFFB3FFB36679B8B4FC2B" blockId="57.[118,651,882,1038]" box="[118,476,882,906]" pageId="57" pageNumber="460">
<treatmentCitationGroup id="AB0B11D1FFAEFFB3FFB36679B8B4FC2B" box="[118,476,882,906]" pageId="57" pageNumber="460">
<treatmentCitation id="0ABA10EEFFAEFFB3FFB36679B8B0FC2B" author="SCHAUB S." box="[118,472,882,906]" page="33" pageId="57" pageNumber="460" year="1930">
<taxonomicName id="4C1B4D7CFFAEFFB3FFB36679B8B0FC2B" authority="Schaub, 1930: 33" authorityName="Schaub" authorityPageNumber="33" authorityYear="1930" box="[118,472,882,906]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFAEFFB3FFB36679B84BFC2B" box="[118,291,882,906]" italics="true" pageId="57" pageNumber="460">Allocricetus bursae</emphasis>
<bibRefCitation id="EF8A4B0EFFAEFFB3FEEC6679B8B0FC2B" author="SCHAUB S." box="[297,472,882,906]" pageId="57" pageNumber="460" pagination="1 - 49" refId="ref46460" refString="SCHAUB S. 1930. - Quartare und jungtertiare Hamster. Abhandlung der Schweizerischen Palaonologischen Gesellschaft 49 (6): 1 - 49, pls 1, 2." type="book chapter" year="1930">Schaub, 1930: 33</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3016574FFAEFFB3FFB36691B869FC6D" pageId="57" pageNumber="460" type="materials_examined">
<paragraph id="8BA436FFFFAEFFB3FFB36691B869FC6D" blockId="57.[118,651,882,1038]" pageId="57" pageNumber="460">
<collectionCode id="ED0AAE3AFFAEFFB3FFB36691B821FC11" box="[118,329,922,945]" pageId="57" pageNumber="460">MATERIAL EXAMINED</collectionCode>
. — 2 M1 (Ms311, 313),
<quantity id="4CE39B1AFFAEFFB3FD936691BBEBFC13" box="[598,643,922,946]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="57" pageNumber="460" unit="m" value="2.0">2 m</quantity>
1 (Ms309-310).
</paragraph>
</subSubSection>
<subSubSection id="C3016574FFAEFFB3FFB366D0BB03F90B" pageId="57" pageNumber="460" type="description">
<paragraph id="8BA436FFFFAEFFB3FFB366D0B890FBAF" blockId="57.[118,651,882,1038]" pageId="57" pageNumber="460">MEASUREMENTS (L × W). — M1 = 1.82 × 1. 30; 1.95 × 1.29; m1 = 2.00 × 1.22; 1.97 × 1.27.</paragraph>
<paragraph id="8BA436FFFFAEFFB3FFB36144B86FFBC6" blockId="57.[118,652,1103,1706]" box="[118,263,1103,1129]" pageId="57" pageNumber="460">DESCRIPTION</paragraph>
<paragraph id="8BA436FFFFAEFFB3FFB3617BB9D2FA6B" blockId="57.[118,652,1103,1706]" pageId="57" pageNumber="460">m1: the anteroconid consists of two poorly individualized cusps, which are either in close connection (unworn specimen) or widely associated but with separated dentine fields (slightly worn specimen). Although there is a short spur on the anterolophid, directed postero-labially, the anterosinusid remains open. The anterolophulid is single. The main tubercles alternate. The mesolophid is lacking. The posterolophid is well developed but it does not close the posterosinusid.</paragraph>
<paragraph id="8BA436FFFFAEFFB3FFB360DBBB03F90B" blockId="57.[118,652,1103,1706]" pageId="57" pageNumber="460">M1: the anterocone is bicuspide but its parts are closely situated, so, in some cases, their anterior parts connect and form an anterior pit. The main cusps are opposite and the inner parts of the respective labial valleys soon become reduced by the conversion of their inner portions into enamel pits. The posterior cingulum is well developed.</paragraph>
</subSubSection>
<subSubSection id="C3016574FFAEFFB0FD6F65C4B853FE48" lastPageId="58" lastPageNumber="461" pageId="57" pageNumber="460" type="discussion">
<paragraph id="8BA436FFFFAEFFB3FD6F65C4BA66FF46" blockId="57.[682,1215,207,1706]" box="[682,782,207,233]" pageId="57" pageNumber="460">REMARKS</paragraph>
<paragraph id="8BA436FFFFAEFFB3FD6F65FBBA54FC4B" blockId="57.[682,1215,207,1706]" pageId="57" pageNumber="460">
The early evolutionary stages of small hamsters of modern
<typeStatus id="54A0885DFFAEFFB3FCD56404BA2EFE88" box="[784,838,271,297]" pageId="57" pageNumber="460">type</typeStatus>
, such as
<taxonomicName id="4C1B4D7CFFAEFFB3FC71641BBD5FFE8B" authorityName="Schaub" authorityYear="1930" box="[948,1079,272,298]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFAEFFB3FC71641BBD5FFE8B" box="[948,1079,272,298]" italics="true" pageId="57" pageNumber="460">Allocricetus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1B4D7CFFAEFFB3FB8F641BBACBFEEB" authority="Milne-Edwards, 1867" authorityName="Milne-Edwards" authorityYear="1867" class="Mammalia" family="Cricetidae" genus="Cricetulus" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFAEFFB3FB8F641BBDD6FE8B" box="[1098,1214,272,298]" italics="true" pageId="57" pageNumber="460">Cricetulus</emphasis>
Milne-Edwards, 1867
</taxonomicName>
,
<emphasis id="B96FEAEDFFAEFFB3FC77643BBD71FEEB" box="[946,1049,304,330]" italics="true" pageId="57" pageNumber="460">Tscherkia</emphasis>
Ognev, 1914, and
<taxonomicName id="4C1B4D7CFFAEFFB3FD1F6444BA87FECB" authority="Zdansky, 1928" authorityName="Zdansky" authorityYear="1928" box="[730,1007,335,362]" class="Mammalia" family="Cricetidae" genus="Cricetinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFAEFFB3FD1F6444BA2CFEC8" box="[730,836,335,361]" italics="true" pageId="57" pageNumber="460">Cricetinus</emphasis>
Zdansky, 1928
</taxonomicName>
are poorly known. Four forms,
<taxonomicName id="4C1B4D7CFFAEFFB3FCFB647BBA87FE28" box="[830,1007,367,394]" class="Mammalia" family="Cricetidae" genus="Cricetulus" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="undefined-i">
<emphasis id="B96FEAEDFFAEFFB3FCFB647BBAC7FE2B" box="[830,943,368,394]" italics="true" pageId="57" pageNumber="460">Cricetulus</emphasis>
sp. I
</taxonomicName>
,
<taxonomicName id="4C1B4D7CFFAEFFB3FC3B647BBDF1FE28" box="[1022,1177,367,394]" class="Mammalia" family="Cricetidae" genus="Cricetulus" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B96FEAEDFFAEFFB3FC3B647BBD07FE2B" box="[1022,1135,368,394]" italics="true" pageId="57" pageNumber="460">Cricetulus</emphasis>
sp.
</taxonomicName>
II,
<taxonomicName id="4C1B4D7CFFAEFFB3FD6F6484BD75FE0B" authority="Kretzoi, 1959" authorityName="Kretzoi" authorityYear="1959" box="[682,1053,399,426]" class="Mammalia" family="Cricetidae" genus="Cricetinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="europaeus">
<emphasis id="B96FEAEDFFAEFFB3FD6F6484BA17FE09" box="[682,895,399,425]" italics="true" pageId="57" pageNumber="460">Cricetinus europaeus</emphasis>
<bibRefCitation id="EF8A4B0EFFAEFFB3FC42649BBD75FE0B" author="KRETZOI M." box="[903,1053,400,426]" pageId="57" pageNumber="460" pagination="237 - 246" refId="ref44793" refString="KRETZOI M. 1959. - Insectivoren, Nagetiere und Lagomorphen der jungstpliozanen Fauna von Csarnota im Villayer Gebirge (Sudungarn). Vertebrata Hungarica 1 (2): 237 - 246." type="journal article" year="1959">Kretzoi, 1959</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1B4D7CFFAEFFB3FB906484BAACFE6B" authority="Hir, 1994" authorityName="Hir" authorityYear="1994" class="Mammalia" family="Cricetidae" genus="Cricetinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="beremendensis">
<emphasis id="B96FEAEDFFAEFFB3FB906484BA2BFE6B" italics="true" pageId="57" pageNumber="460">Cricetinus beremendensis</emphasis>
<bibRefCitation id="EF8A4B0EFFAEFFB3FC8864BBBAACFE6B" author="HIR J." box="[845,964,432,458]" pageId="57" pageNumber="460" pagination="71 - 89" refId="ref44200" refString="HIR J. 1994. - Cricetinus beremendensis n. sp. (Rodentia, Mammalia) from the Pliocene fauna of Beremend 15 (S. Hungary). Fragmenta Mineralogica et Palaeontologica 17: 71 - 89." type="journal article" year="1994">Hir, 1994</bibRefCitation>
</taxonomicName>
, similar in size to the material from Muselievo, are known from some Vallesian (Sumeg, MN10) and Pliocene (Osztramos 1, Csarnóta, Beremend 15, MN14- 16) localities of
<collectingCountry id="F30C766FFFAEFFB3FCA86724BABDFDE8" box="[877,981,559,585]" name="Hungary" pageId="57" pageNumber="460">Hungary</collectingCountry>
(
<bibRefCitation id="EF8A4B0EFFAEFFB3FC22673BBDEFFDEB" author="KORDOS L." box="[999,1159,560,586]" pageId="57" pageNumber="460" pagination="65 - 88" refId="ref44407" refString="KORDOS L. 1987. - Karstocricetus skofleki gen. n., sp. n. and the evolution of the Late Neogene Cricetidae in the Carpatian Basin. Fragmenta Mineralogica et Palaeontologica 13: 65 - 88." type="journal article" year="1987">Kordos 1987</bibRefCitation>
;
<bibRefCitation id="EF8A4B0EFFAEFFB3FB50673BBB80FDCB" author="HIR J." pageId="57" pageNumber="460" pagination="71 - 89" refId="ref44200" refString="HIR J. 1994. - Cricetinus beremendensis n. sp. (Rodentia, Mammalia) from the Pliocene fauna of Beremend 15 (S. Hungary). Fragmenta Mineralogica et Palaeontologica 17: 71 - 89." type="journal article" year="1994">Hir 1994</bibRefCitation>
). These forms differ from the population under study in having an undivided anteroconid on m1. This feature may be considered as a primitive one. More over
<taxonomicName id="4C1B4D7CFFAEFFB3FC5867A4BD17FD69" authorityName="Kretzoi" authorityYear="1959" box="[925,1151,687,713]" class="Mammalia" family="Cricetidae" genus="Cricetinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="europaeus">
<emphasis id="B96FEAEDFFAEFFB3FC5867A4BD17FD69" box="[925,1151,687,713]" italics="true" pageId="57" pageNumber="460">Cricetinus europaeus</emphasis>
</taxonomicName>
from Csarnóta-2 shows a double anterolophulid on this tooth. On the other hand, the occlusal pattern of M1 of
<taxonomicName id="4C1B4D7CFFAEFFB3FC886604BABBFC89" authorityName="Kretzoi" authorityYear="1959" box="[845,979,783,809]" class="Mammalia" family="Cricetidae" genus="Cricetinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="europaeus">
<emphasis id="B96FEAEDFFAEFFB3FC886604BABBFC89" box="[845,979,783,809]" italics="true" pageId="57" pageNumber="460">C. europaeus</emphasis>
</taxonomicName>
is very similar to the material from Muselievo and to the Pleistocene populations of
<taxonomicName id="4C1B4D7CFFAEFFB3FC93665BBD74FCCB" authorityName="Schaub" authorityYear="1930" box="[854,1052,848,874]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFAEFFB3FC93665BBD74FCCB" box="[854,1052,848,874]" italics="true" pageId="57" pageNumber="460">Allocricetus bursae</emphasis>
</taxonomicName>
. The Far East Pleistocene species
<taxonomicName id="4C1B4D7CFFAEFFB3FC4F6664BDD6FC2B" authority="Zdansky" authorityName="Zdansky" box="[906,1214,879,906]" class="Mammalia" family="Cricetidae" genus="Cricetinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="varians">
<emphasis id="B96FEAEDFFAEFFB3FC4F6664BD3CFC29" box="[906,1108,879,905]" italics="true" pageId="57" pageNumber="460">Cricetinus varians</emphasis>
Zdansky
</taxonomicName>
(
<bibRefCitation id="EF8A4B0EFFAEFFB3FD75669BBA50FC0B" author="FEJFAR O." box="[688,824,912,938]" pageId="57" pageNumber="460" pagination="227 - 297" refId="ref43555" refString="FEJFAR O. 1970. - Die plio-pleistozanen Wirbeltierfaunen von Hajnacka und Ivanovce (Slowakei, CSSR). VI. Cricetidae, Rodentia, Mammalia. Mitteilungen der Bayerische Staatssammlung fur Palaontologie und historische Geologie Munich 10: 227 - 297." type="journal article" year="1970">Fejfar 1970</bibRefCitation>
;
<bibRefCitation id="EF8A4B0EFFAEFFB3FC806684BA95FC0B" author="VORONTSOV N. N." box="[837,1021,911,938]" pageId="57" pageNumber="460" refId="ref47715" refString="VORONTSOV N. N. 1982. - Nizschie khomijakoobraznie (Cricetidae) Mirovoj fauni. I. Morphologia I ekologia. [= Lower cricetids (Cricetidae) of the world fauna. Morphology and ecology], in Fauna of the USSR, Mammals 3 (6). Nauka, Moscow, 450 p. (in Russian)." type="book" year="1982">Vorontsov 1982</bibRefCitation>
) differs from the Hungarian species by having mesolophid on the lower molars.
</paragraph>
<paragraph id="8BA436FFFFAEFFB3FD6F66FBBA0DFAAB" blockId="57.[682,1215,207,1706]" pageId="57" pageNumber="460">
The material from the late Pliocene locality Maritsa (the isle of Rhodes,
<collectingCountry id="F30C766FFFAEFFB3FC1E611BBD42FB8B" box="[987,1066,1040,1066]" name="Greece" pageId="57" pageNumber="460">Greece</collectingCountry>
), described as?
<taxonomicName id="4C1B4D7CFFAEFFB3FD76613BBA28FBE8" box="[691,832,1071,1098]" class="Mammalia" family="Cricetidae" genus="Cricetulus" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B96FEAEDFFAEFFB3FD76613BBA74FBEB" box="[691,796,1072,1098]" italics="true" pageId="57" pageNumber="460">Cricetulus</emphasis>
sp.
</taxonomicName>
(
<bibRefCitation id="EF8A4B0EFFAEFFB3FC8B613BBD5CFBEB" author="DE BRUIJN H. &amp; DAWSON M. R. &amp; MEIN P." box="[846,1076,1071,1098]" pageId="57" pageNumber="460" pagination="535 - 584" refId="ref43163" refString="DE BRUIJN H., DAWSON M. R. &amp; MEIN P. 1970. - Upper Pliocene Rodentia, Lagomorpha and Insectivora (Mammalia) from the isle of Rhodes (Greece). Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B 73 (5): 535 - 584." type="journal article" year="1970">
De Bruijn
<emphasis id="B96FEAEDFFAEFFB3FC076124BA9DFBEB" box="[962,1013,1071,1098]" italics="true" pageId="57" pageNumber="460">et al.</emphasis>
1970
</bibRefCitation>
) is similar in size with the specimens from Muselievo but it differs in the presence of a short mesolophid on m1. According to
<bibRefCitation id="EF8A4B0EFFAEFFB3FCB9619BBA96FB0B" author="SEN S." box="[892,1022,1168,1194]" pageId="57" pageNumber="460" pagination="90 - 170" refId="ref46490" refString="SEN S. 1977. - La faune de Rongeurs pliocenes de Calta (Ankara, Turkie). Bulletin du Museum national d'Histoire naturelle ser. 3, Sciences de la Terre 61, 465: 90 - 170." type="journal article" year="1977">Sen (1977)</bibRefCitation>
, these specimens should be referred to
<taxonomicName id="4C1B4D7CFFAEFFB3FC5861A4BD63FB4B" authority="De Bruijn, Dawson &amp; Mein, 1970" authorityName="De Bruijn, Dawson &amp; Mein" authorityYear="1970" class="Mammalia" family="Cricetidae" genus="Mesocricetus" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="primitivus">
<emphasis id="B96FEAEDFFAEFFB3FC5861A4BDFCFB69" box="[925,1172,1199,1224]" italics="true" pageId="57" pageNumber="460">Mesocricetus primitivus</emphasis>
<bibRefCitation id="EF8A4B0EFFAEFFB3FB5861A4BD63FB4B" author="DE BRUIJN H. &amp; DAWSON M. R. &amp; MEIN P." pageId="57" pageNumber="460" pagination="535 - 584" refId="ref43163" refString="DE BRUIJN H., DAWSON M. R. &amp; MEIN P. 1970. - Upper Pliocene Rodentia, Lagomorpha and Insectivora (Mammalia) from the isle of Rhodes (Greece). Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B 73 (5): 535 - 584." type="journal article" year="1970">De Bruijn, Dawson &amp; Mein, 1970</bibRefCitation>
</taxonomicName>
, described from the same locality.
</paragraph>
<paragraph id="8BA436FFFFAEFFB3FD6F601BBAEEF98B" blockId="57.[682,1215,207,1706]" pageId="57" pageNumber="460">
The teeth from the Slovakian locality Ivanovce (MN15), described as
<taxonomicName id="4C1B4D7CFFAEFFB3FC62603BBDE6FAEB" authorityName="Schaub" authorityYear="1930" box="[935,1166,1328,1354]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFAEFFB3FC62603BBD76FAEB" box="[935,1054,1328,1354]" italics="true" pageId="57" pageNumber="460">Allocricetus</emphasis>
cf.
<emphasis id="B96FEAEDFFAEFFB3FB8E603BBDE6FAEB" box="[1099,1166,1328,1354]" italics="true" pageId="57" pageNumber="460">bursae</emphasis>
</taxonomicName>
, are smaller than the specimens from Muselievo. In this respect, they differ also from the comparative material of
<taxonomicName id="4C1B4D7CFFAEFFB3FC8A6084BAA0FA0B" authorityName="Schaub" authorityYear="1930" box="[847,968,1423,1450]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFAEFFB3FC8A6084BAA0FA0B" box="[847,968,1423,1450]" italics="true" pageId="57" pageNumber="460">A. bursae</emphasis>
</taxonomicName>
from Zirany and Hundsheim (
<bibRefCitation id="EF8A4B0EFFAEFFB3FC8560BBBAA2FA6B" author="FEJFAR O." box="[832,970,1456,1482]" pageId="57" pageNumber="460" pagination="227 - 297" refId="ref43555" refString="FEJFAR O. 1970. - Die plio-pleistozanen Wirbeltierfaunen von Hajnacka und Ivanovce (Slowakei, CSSR). VI. Cricetidae, Rodentia, Mammalia. Mitteilungen der Bayerische Staatssammlung fur Palaontologie und historische Geologie Munich 10: 227 - 297." type="journal article" year="1970">Fejfar 1970</bibRefCitation>
). Moreover nearly all M1s from Ivanovce show undivided anterocone, while this tubercle is clearly divided in the material from Muselievo.
</paragraph>
<paragraph id="8BA436FFFFAEFFB0FD6F633BB853FE48" blockId="57.[682,1215,207,1706]" lastBlockId="58.[119,652,207,490]" lastPageId="58" lastPageNumber="461" pageId="57" pageNumber="460">
The available teeth are similar in size and overall occlusal pattern to the Pleistocene populations of
<taxonomicName id="4C1B4D7CFFAEFFB3FD6F637BBA19F92B" authorityName="Schaub" authorityYear="1930" box="[682,881,1648,1674]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="57" pageNumber="460" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFAEFFB3FD6F637BBA19F92B" box="[682,881,1648,1674]" italics="true" pageId="57" pageNumber="460">Allocricetus bursae</emphasis>
</taxonomicName>
from
<collectingCountry id="F30C766FFFAEFFB3FC7E637BBD74F92B" box="[955,1052,1648,1674]" name="Bulgaria" pageId="57" pageNumber="460">Bulgaria</collectingCountry>
. Although the anteroconid of the m1s from Muselievo is clearly divided, both parts are closely set, giving in this way a more primitive appearance of the population under consideration. However, this feature occurs also in some Pleistocene samples of
<taxonomicName id="4C1B4D7CFFADFFB0FFB2645BB855FECB" authorityName="Schaub" authorityYear="1930" box="[119,317,336,362]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="461" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFADFFB0FFB2645BB855FECB" box="[119,317,336,362]" italics="true" pageId="58" pageNumber="461">Allocricetus bursae</emphasis>
</taxonomicName>
from
<collectingCountry id="F30C766FFFADFFB0FE43645BB88AFECB" box="[390,482,336,362]" name="Bulgaria" pageId="58" pageNumber="461">Bulgaria</collectingCountry>
(Varbeshnitsa, Morovitsa) (
<bibRefCitation id="EF8A4B0EFFADFFB0FF3B647BB8E1FE2B" author="POPOV V. V." box="[254,393,368,394]" pageId="58" pageNumber="461" pagination="193 - 234" refId="ref45554" refString="POPOV V. V. 1988. - Middle Pleistocene small mammals (Mammalia: Insectivora, Lagomorpha, Rodentia) from Varbeshnitsa (Bulgaria). Acta zoologica cracoviensia 31 (5): 193 - 234." type="journal article" year="1988">Popov 1988</bibRefCitation>
,
<bibRefCitation id="EF8A4B0EFFADFFB0FE53647BB8BAFE2B" author="POPOV V. V." box="[406,466,368,394]" pageId="58" pageNumber="461" pagination="561 - 588" refId="ref45593" refString="POPOV V. V. 1989. - Middle Pleistocene small mammals (Insectivora, Lagomorpha, Rodentia) from Morovitsa Cave (North Bulgaria). Acta zoologica cracoviensia 32 (13): 561 - 588." type="journal article" year="1989">1989</bibRefCitation>
). These comparisons suggest that the teeth from Muselievo most probably belong to a primitive form of
<taxonomicName id="4C1B4D7CFFADFFB0FFB264DBB85DFE4B" authorityName="Schaub" authorityYear="1930" box="[119,309,464,490]" class="Mammalia" family="Cricetidae" genus="Allocricetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="58" pageNumber="461" phylum="Chordata" rank="species" species="bursae">
<emphasis id="B96FEAEDFFADFFB0FFB264DBB85DFE4B" box="[119,309,464,490]" italics="true" pageId="58" pageNumber="461">Allocricetus bursae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>