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<document id="0AF5041D7D5B3CDFA9129E447D4CB0B5" ID-DOI="10.11646/zootaxa.5383.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10361564" ID-ZooBank="20988E42-C14D-4878-A716-8CEAB5E5EF92" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1702374886811" checkinUser="plazi" docAuthor="Tasaku, Yuto, Ono, Hiroki &amp; Maruyama, Munetoshi" docDate="2023" docId="03FE87D64256FFE082AA389B4EC4F9C2" docLanguage="en" docName="zootaxa.5383.3.1.pdf" docOrigin="Zootaxa 5383 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.3.1/52454" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diaulota submarina Ahn 2023" docType="treatment" docVersion="3" lastPageNumber="268" masterDocId="FFC7FFAE4247FFF1823D3A504E1BFFA2" masterDocTitle="Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="296" masterPageNumber="251" pageNumber="268" updateTime="1702492195905" updateUser="karina" zenodo-license-document="CLOSED">
<mods:mods id="272259A2988F13F5C555D85DA4615BD7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="57FBD5165E7F9CE99C3473AF78602B9E">Review of the intertidal rove beetle tribe Liparocephalini Fenyes (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
</mods:titleInfo>
<mods:name id="64CA7E9838EAAB8C36296F7D749A1CD7" type="personal">
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<mods:roleTerm id="80B044C1F9E1E92E8328268FFBBD2979">Author</mods:roleTerm>
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<mods:namePart id="B7F5CDC6BB4030D0F8151EB055A80A50">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="05D9F0B8F1C197A5E5FA5AF0189AACC3">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan. &amp; Corresponding author</mods:affiliation>
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</mods:role>
<mods:namePart id="3028850C6AA47200C1ECFB97856FFA62">Ono, Hiroki</mods:namePart>
<mods:affiliation id="7FD28F470D5F75307148F52B750608FF">Sounan, Minami-ku, Sagamihara-shi, Kanagawa, 252 - 0312, Japan.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="FBB669E958AD9C356E0F85A026B49E9C">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="73E31D819A1FB083696046B0D27BF2B2">The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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<mods:typeOfResource id="160FBE5AA8806DCC9A929C77471F30C7">text</mods:typeOfResource>
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<mods:title id="810D69A53B9B4A217815F08B457E04EE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8820CFA0BB83B18EE1545C37BA97AA91">
<mods:date id="AA18CA42DEEB9BAFF1CB28F3848BD896">2023</mods:date>
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<mods:number id="E8141314CDD9E061ABFB6D3A2C70699B">2023-12-12</mods:number>
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<mods:number id="70E15B66E04278AF6CA49FA643AE29DB">5383</mods:number>
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<mods:number id="8578EB53F5B19EC641266E55DCEEEC89">3</mods:number>
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<mods:classification id="F5511282DB278992CE33539CEEF56DBF">journal article</mods:classification>
<mods:identifier id="A83138A355C50FDD4C9B4DBB16729905" type="DOI">10.11646/zootaxa.5383.3.1</mods:identifier>
<mods:identifier id="323F359B76FA8FD2FE02BB9845A0C9EE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FE87D64256FFE082AA389B4EC4F9C2" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE082AA389B4EC4F9C2" httpUri="http://treatment.plazi.org/id/03FE87D64256FFE082AA389B4EC4F9C2" lastPageNumber="268" pageId="17" pageNumber="268">
<subSubSection id="C34D654B4256FFE082AA389B4C2EFCA8" pageId="17" pageNumber="268" type="nomenclature">
<paragraph id="8BE836C04256FFE082AA389B4C0BFD44" blockId="17.[151,565,715,815]" box="[151,528,715,742]" pageId="17" pageNumber="268">
<heading id="D0A081AC4256FFE082AA389B4C0BFD44" bold="true" box="[151,528,715,742]" fontSize="11" level="1" pageId="17" pageNumber="268" reason="1">
<taxonomicName id="4C574D434256FFE082AA389B4C0BFD44" authority="Ahn, 2023" authorityName="Ahn" authorityYear="2023" box="[151,528,715,742]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24256FFE082AA389B4C0BFD44" bold="true" box="[151,528,715,742]" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE082AA389B4F91FD44" bold="true" box="[151,394,715,742]" italics="true" pageId="17" pageNumber="268">Diaulota submarina</emphasis>
<bibRefCitation id="EFC64B314256FFE083AC389C4C0BFD44" author="Ahn, K. - J." box="[401,528,716,742]" pageId="17" pageNumber="268" pagination="141 - 147" refId="ref21503" refString="Ahn, K. - J. (2023) Description of Diaulota submarina sp. nov. (Coleoptera: Staphylinidae: Aleocharinae) on Korean coasts. Zootaxa, 5336 (1), 141 - 147. https: // doi. org / 10.11646 / zootaxa. 5336.1.8" type="journal article" year="2023">Ahn, 2023</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE836C04256FFE082AA38A04C2EFCA8" blockId="17.[151,565,715,815]" box="[151,565,752,778]" pageId="17" pageNumber="268">[Chôsen-shiributo-umi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082AA39444FB1FC8D" box="[151,426,788,815]" pageId="17" pageNumber="268" type="description">
<paragraph id="8BE836C04256FFE082AA39444FB1FC8D" blockId="17.[151,565,715,815]" box="[151,426,788,815]" pageId="17" pageNumber="268">
(
<figureCitation id="136C2A454256FFE082A239444EE6FC8C" box="[159,253,788,814]" captionStart="FIGURE 10" captionStartId="15.[151,250,1778,1802]" captionTargetBox="[162,1425,500,1744]" captionTargetId="figure-191@15.[151,1436,489,1755]" captionTargetPageId="15" captionText="FIGURE 10 Habitus and mouthparts of Diaulota submarina. A, habitus, male; B, ditto, female; C, labium, D, mentum. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for C &amp; D." figureDoi="http://doi.org/10.5281/zenodo.10361603" httpUri="https://zenodo.org/record/10361603/files/figure.png" pageId="17" pageNumber="268">Figs. 10</figureCitation>
,
<figureCitation id="136C2A454256FFE0833439444F3CFC8C" box="[265,295,788,814]" captionStart="FIGURE 11" captionStartId="16.[151,250,1917,1941]" captionTargetBox="[160,1427,192,1882]" captionTargetId="figure-22@16.[151,1436,181,1893]" captionTargetPageId="16" captionText="FIGURE 11 Genitalia of Diaulota submarina (A–D, from Tsushima; E–H, from Hiroshima-ken). A &amp; E, median lobe of aedeagus, lateral view; B &amp; F, ditto, ventral view; C &amp; G, internal structure of median lobe, ventral view; D &amp; H, spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361606" httpUri="https://zenodo.org/record/10361606/files/figure.png" pageId="17" pageNumber="268">11</figureCitation>
;
<figureCitation id="136C2A454256FFE0830F39444F55FC8C" box="[306,334,788,814]" captionStart="FIGURE 24" captionStartId="36.[151,250,1780,1804]" captionTargetBox="[151,1436,345,1757]" captionTargetId="figure-99@36.[151,1436,345,1757]" captionTargetPageId="36" captionText="FIGURE 24 Distribution of Diaulota uenoi (red circle (red ring, type locality)), D. submarina (blue square), and D. orientalis sp. n. (yellow triangle)." figureDoi="http://doi.org/10.5281/zenodo.10361669" httpUri="https://zenodo.org/record/10361669/files/figure.png" pageId="17" pageNumber="268">24</figureCitation>
&amp;
<figureCitation id="136C2A454256FFE0834F39444FB9FC8D" box="[370,418,788,815]" captionStart="FIGURE 25" captionStartId="37.[151,250,1830,1854]" captionTargetBox="[151,1436,417,1807]" captionTargetId="figure-130@37.[151,1436,417,1807]" captionTargetPageId="37" captionText="FIGURE 25 Distribution and habitat of Rotundicephala gen. n. spp. of Japan. A, known localities in Japan (red circle, R. pacifica comb. n. (red ring, type locality)); blue square, R. koheii sp. n.; yellow triangle, R. koreana comb. n.); B, habitat of R. koheii sp. n. (Kanagawa-ken, Japan); C, habitat of R. pacifica comb. n. (Tsushima, adults of Diaulota submarina was collected same time)." figureDoi="http://doi.org/10.5281/zenodo.10361673" httpUri="https://zenodo.org/record/10361673/files/figure.png" pageId="17" pageNumber="268">25C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082AA39084CA0FC2D" pageId="17" pageNumber="268" type="reference_group">
<paragraph id="8BE836C04256FFE082AA39084CA0FC2D" blockId="17.[151,1399,856,911]" pageId="17" pageNumber="268">
<treatmentCitationGroup id="AB4711EE4256FFE082AA39084C12FCD3" box="[151,521,856,881]" pageId="17" pageNumber="268">
<treatmentCitation id="0AF610D14256FFE082AA39084C12FCD3" ID-DOI="http://doi.org/10.5281/zenodo.8276634" ID-Zenodo-Dep="8276634" author="Ahn, K. - J." box="[151,521,856,881]" httpUri="http://treatment.plazi.org/id/C7295D6BFFED0C72FF39F8D1FF20F839" page="141" pageId="17" pageNumber="268" year="2023">
<taxonomicName id="4C574D434256FFE082AA39084C12FCD3" authority="Ahn, 2023: 141" authorityName="Ahn" authorityPageNumber="141" authorityYear="2023" box="[151,521,856,881]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="submarina">
<emphasis id="B923EAD24256FFE082AA39084F7FFCD2" box="[151,356,856,880]" italics="true" pageId="17" pageNumber="268">Diaulota submarina</emphasis>
<materialsCitation id="3B3F3C9D4256FFE0835739084C12FCD3" box="[362,521,856,881]" country="South Korea" county="Anmyeon Isl." location="Ahn" municipality="Bangpo Beach" pageId="17" pageNumber="268" specimenCount="1" stateProvince="Chungnam">
<bibRefCitation id="EFC64B314256FFE0835739084C12FCD3" author="Ahn, K. - J." box="[362,521,856,881]" pageId="17" pageNumber="268" pagination="141 - 147" refId="ref21503" refString="Ahn, K. - J. (2023) Description of Diaulota submarina sp. nov. (Coleoptera: Staphylinidae: Aleocharinae) on Korean coasts. Zootaxa, 5336 (1), 141 - 147. https: // doi. org / 10.11646 / zootaxa. 5336.1.8" type="journal article" year="2023">Ahn, 2023: 141</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
<treatmentCitationGroup id="AB4711EE4256FFE0803239084CA0FC2D" pageId="17" pageNumber="268">
(original description;
<typeStatus id="54EC88624256FFE080D639094D0CFCD3" box="[747,791,857,881]" pageId="17" pageNumber="268">type</typeStatus>
locality:
<collectingCountry id="F34076504256FFE0814B39094DA3FCD2" box="[886,952,857,880]" name="South Korea" pageId="17" pageNumber="268">Korea</collectingCountry>
:
<collectingRegion id="4993F8224256FFE081FC39084A2FFCD2" box="[961,1076,856,880]" country="South Korea" name="Chungnam" pageId="17" pageNumber="268">Chungnam</collectingRegion>
:
<collectingCounty id="62894E4C4256FFE0860139094AD3FCD2" box="[1084,1224,856,881]" pageId="17" pageNumber="268">Anmyeon Isl.</collectingCounty>
,
<collectingMunicipality id="6B8CACBA4256FFE086EE39094B77FCD2" box="[1235,1388,856,881]" pageId="17" pageNumber="268">Bangpo Beach</collectingMunicipality>
). “
<taxonomicName id="4C574D434256FFE0829F39274F20FC2D" ID-CoL="35GKM" authorityName="K. Sawada" authorityYear="1955" box="[162,315,887,911]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24256FFE0829F39274F20FC2D" box="[162,315,887,911]" italics="true" pageId="17" pageNumber="268">Diaulota uenoi</emphasis>
</taxonomicName>
”:
<location id="8E88601B4256FFE0836F39274F9BFC2D" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE082AA389B4EC4F9C2:8E88601B4256FFE0836F39274F9BFC2D" box="[338,384,887,911]" country="South Korea" county="Anmyeon Isl." municipality="Bangpo Beach" name="Ahn" pageId="17" pageNumber="268" stateProvince="Chungnam">Ahn</location>
, 1996: 284 (misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082AA39ED4C29FB29" pageId="17" pageNumber="268" type="materials_examined">
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<emphasis id="B923EAD24256FFE082AA39ED4F99FC75" bold="true" box="[151,386,957,983]" pageId="17" pageNumber="268">
<collectingRegion id="4993F8224256FFE082AA39ED4EE4FC75" box="[151,255,957,983]" pageId="17" pageNumber="268">Material</collectingRegion>
examined.
</emphasis>
<materialsCitation id="3B3F3C9D4256FFE083B639ED4FFEFC59" collectingDate="2021-09-19" collectorName="H. Yoshigou" country="Japan" location="Etajima-shi" pageId="17" pageNumber="268" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Honshu">
<emphasis id="B923EAD24256FFE083B639ED4D08FC75" bold="true" box="[395,787,957,984]" pageId="17" pageNumber="268">
<collectingCountry id="F34076504256FFE083B639ED4FC3FC75" box="[395,472,957,983]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
:
<collectingRegion id="4993F8224256FFE083DB39ED4C53FC7A" box="[486,584,957,984]" pageId="17" pageNumber="268">Honshû</collectingRegion>
:
<collectingRegion id="4993F8224256FFE0806839ED4D16FC75" box="[597,781,957,983]" pageId="17" pageNumber="268">Hiroshima-ken</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD494256FFE0812639ED4D70FC75" box="[795,875,957,983]" count="1" pageId="17" pageNumber="268" type="male">1 male</specimenCount>
,
<specimenCount id="9D51FD494256FFE0814539ED4DC5FC75" box="[888,990,957,983]" count="1" pageId="17" pageNumber="268" type="female">1 female</specimenCount>
,
<location id="8E88601B4256FFE081D639ED4A74FC75" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE082AA389B4EC4F9C2:8E88601B4256FFE081D639ED4A74FC75" box="[1003,1135,957,983]" country="Japan" name="Etajima-shi" pageId="17" pageNumber="268" stateProvince="Honshu">Etajima-shi</location>
,
<collectorName id="26A253164256FFE0864639ED4B03FC75" box="[1147,1304,957,983]" pageId="17" pageNumber="268">Noumi-shima</collectorName>
,
<collectorName id="26A253164256FFE0871839EE4B77FC7A" box="[1317,1388,958,984]" pageId="17" pageNumber="268">Iwane</collectorName>
,
<date id="FFE910004256FFE0874439ED4EE5FC59" pageId="17" pageNumber="268" value="2021-09-19">
<collectingDate id="EFADE9E84256FFE0874439ED4EE5FC59" pageId="17" pageNumber="268" value="2021-09-19">19. IX. 2021</collectingDate>
</date>
,
<collectorName id="26A253164256FFE0833739B24F8EFC59" box="[266,405,993,1020]" pageId="17" pageNumber="268">H. Yoshigou</collectorName>
(cYT)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4256FFE083CC39B14B83FC59" box="[497,1432,993,1020]" collectionCode="II" collectorName="Obara-hama" country="Japan" location="Hagi-shi" pageId="17" pageNumber="268" specimenCount="1" specimenCount-male="1" stateProvince="Yamaguchi-ken">
<emphasis id="B923EAD24256FFE083CC39B14CAEFC59" bold="true" box="[497,693,993,1019]" pageId="17" pageNumber="268">
<collectingRegion id="4993F8224256FFE083CC39B14CABFC59" box="[497,688,993,1019]" pageId="17" pageNumber="268">Yamaguchi-ken</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD494256FFE0808139B14D10FC59" box="[700,779,993,1019]" count="1" pageId="17" pageNumber="268" type="male">1 male</specimenCount>
,
<location id="8E88601B4256FFE0812A39B14D61FC59" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE082AA389B4EC4F9C2:8E88601B4256FFE0812A39B14D61FC59" box="[791,890,993,1019]" country="Japan" name="Hagi-shi" pageId="17" pageNumber="268" stateProvince="Yamaguchi-ken">Hagi-shi</location>
,
<collectorName id="26A253164256FFE081BB39B14A08FC59" box="[902,1043,993,1019]" pageId="17" pageNumber="268">Obara-hama</collectorName>
,
<date id="FFE910004256FFE0862239B14ABDFC59" box="[1055,1190,993,1020]" pageId="17" pageNumber="268" value="2016-02-28">
28.
<collectionCode id="ED46AE054256FFE0867439B24A44FC5E" box="[1097,1119,994,1020]" pageId="17" pageNumber="268">II</collectionCode>
. 2016
</date>
, M.
<collectorName id="26A253164256FFE086E539B14B23FC59" box="[1240,1336,993,1019]" pageId="17" pageNumber="268">Matsuda</collectorName>
(
<collectionCode id="ED46AE054256FFE0877539B14B94FC59" box="[1352,1423,993,1019]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="17" pageNumber="268" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4256FFE082AA3E554D36FBBD" box="[151,813,1029,1056]" collectionCode="V" collectorName="Chinto" country="Japan" location="Hagi-shi" pageId="17" pageNumber="268" specimenCount="1" specimenCount-female="1" stateProvince="Yamaguchi-ken">
<specimenCount id="9D51FD494256FFE082AA3E554EE1FBBD" box="[151,250,1029,1055]" count="1" pageId="17" pageNumber="268" type="female">1 female</specimenCount>
,
<location id="8E88601B4256FFE083383E554F73FBBD" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE082AA389B4EC4F9C2:8E88601B4256FFE083383E554F73FBBD" box="[261,360,1029,1055]" country="Japan" name="Hagi-shi" pageId="17" pageNumber="268" stateProvince="Yamaguchi-ken">Hagi-shi</location>
,
<collectorName id="26A253164256FFE0834F3E554FDAFBBD" box="[370,449,1029,1055]" pageId="17" pageNumber="268">Chintô</collectorName>
,
<date id="FFE910004256FFE083F13E554C24FB82" box="[460,575,1029,1056]" pageId="17" pageNumber="268" value="2015-05-04">
4.
<collectionCode id="ED46AE054256FFE083DB3E564FECFB82" box="[486,503,1030,1056]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="17" pageNumber="268" type="Museum">V</collectionCode>
. 2015
</date>
, M.
<collectorName id="26A253164256FFE080523E554CD4FBBD" box="[623,719,1029,1055]" pageId="17" pageNumber="268">Matsuda</collectorName>
(
<collectionCode id="ED46AE054256FFE080E33E554D3EFBBD" box="[734,805,1029,1055]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="17" pageNumber="268" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4256FFE081053E554FCCFBE1" collectionCode="II" country="Japan" location="Shimonoseki-shi" pageId="17" pageNumber="268" specimenCount="1" specimenCount-female="1" stateProvince="Yamaguchi-ken">
<specimenCount id="9D51FD494256FFE081053E554D80FBBD" box="[824,923,1029,1055]" count="1" pageId="17" pageNumber="268" type="female">1 female</specimenCount>
,
<location id="8E88601B4256FFE081983E554A7FFBBD" LSID="urn:lsid:plazi:treatment:03FE87D64256FFE082AA389B4EC4F9C2:8E88601B4256FFE081983E554A7FFBBD" box="[933,1124,1029,1055]" country="Japan" name="Shimonoseki-shi" pageId="17" pageNumber="268" stateProvince="Yamaguchi-ken">Shimonoseki-shi</location>
, Chôfusangen’ya-kaigan,
<date id="FFE910004256FFE087BA3E554EEFFBE6" pageId="17" pageNumber="268" value="2015-02-03">
3.
<collectionCode id="ED46AE054256FFE082AA3E7A4EB6FBE6" box="[151,173,1066,1092]" pageId="17" pageNumber="268">II</collectionCode>
. 2015
</date>
, M. Matsuda (cYT)
</materialsCitation>
.
<materialsCitation id="3B3F3C9D4256FFE083DE3E794EC4FBC5" collectionCode="VI" collectorName="Shikano-shima" country="Japan" location="Kyushu" pageId="17" pageNumber="268" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Kyushu">
<emphasis id="B923EAD24256FFE083DE3E794CE7FBE1" bold="true" box="[483,764,1065,1092]" pageId="17" pageNumber="268">
<collectingRegion id="4993F8224256FFE083DE3E794C5EFBE6" box="[483,581,1065,1092]" pageId="17" pageNumber="268">Kyûshû</collectingRegion>
:
<collectingRegion id="4993F8224256FFE0806F3E794CECFBE1" box="[594,759,1065,1091]" pageId="17" pageNumber="268">Fukuoka-ken</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD494256FFE0813E3E794D46FBE1" box="[771,861,1065,1091]" count="3" pageId="17" pageNumber="268" type="male">3 males</specimenCount>
,
<specimenCount id="9D51FD494256FFE081573E794DD5FBE1" box="[874,974,1065,1091]" count="1" pageId="17" pageNumber="268" type="female">1 female</specimenCount>
,
<collectorName id="26A253164256FFE081E63E794A9DFBE1" box="[987,1158,1065,1091]" pageId="17" pageNumber="268">Shikano-shima</collectorName>
,
<date id="FFE910004256FFE086AF3E794B38FBE1" box="[1170,1315,1065,1092]" pageId="17" pageNumber="268" value="2022-06-17">
17.
<collectionCode id="ED46AE054256FFE086813E7A4AC6FBE6" box="[1212,1245,1066,1092]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="268">VI</collectionCode>
. 2022
</date>
, Y. Tasaku (cYT)
</materialsCitation>
;
<materialsCitation id="3B3F3C9D4256FFE082D63E1D4B5BFBC5" box="[235,1344,1101,1128]" collectionCode="VI" collectorName="Izuhara-cho &amp; Azamo" country="Japan" location="Nagasaki-ken" pageId="17" pageNumber="268" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Nagasaki-ken">
<emphasis id="B923EAD24256FFE082D63E1D4C00FBC5" bold="true" box="[235,539,1101,1127]" pageId="17" pageNumber="268">
<collectingRegion id="4993F8224256FFE082D63E1D4F88FBC5" box="[235,403,1101,1127]" pageId="17" pageNumber="268">Nagasaki-ken</collectingRegion>
:
<collectingRegion id="4993F8224256FFE083A23E1D4C0DFBC5" box="[415,534,1101,1127]" pageId="17" pageNumber="268">Tsushima</collectingRegion>
:
</emphasis>
<specimenCount id="9D51FD494256FFE0801E3E1D4C66FBC5" box="[547,637,1101,1127]" count="2" pageId="17" pageNumber="268" type="male">2 males</specimenCount>
,
<specimenCount id="9D51FD494256FFE080B43E1D4CE2FBC5" box="[649,761,1101,1127]" count="3" pageId="17" pageNumber="268" type="female">3 females</specimenCount>
,
<collectorName id="26A253164256FFE081383E1D4D95FBC5" box="[773,910,1101,1127]" pageId="17" pageNumber="268">Izuhara-chô</collectorName>
,
<collectorName id="26A253164256FFE081A53E1E4DF7FBCA" box="[920,1004,1102,1128]" pageId="17" pageNumber="268">Azamo</collectorName>
,
<date id="FFE910004256FFE081C53E1D4A91FBC5" box="[1016,1162,1101,1128]" pageId="17" pageNumber="268" value="2022-06-15">
15.
<collectionCode id="ED46AE054256FFE0861E3E1E4A5FFBCA" box="[1059,1092,1102,1128]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="268">VI</collectionCode>
. 2022
</date>
, H. Ono (cHO);
</materialsCitation>
<materialsCitation id="3B3F3C9D4256FFE087743E1D4C29FB29" collectionCode="VI" collectorName="Izuhara-cho &amp; Azamo" country="Japan" location="Nagasaki-ken" pageId="17" pageNumber="268" specimenCount="1" specimenCount-male="1" stateProvince="Tsushima">
<specimenCount id="9D51FD494256FFE087743E1D4B83FBC5" box="[1353,1432,1101,1127]" count="1" pageId="17" pageNumber="268" type="male">1 male</specimenCount>
, ditto,
<date id="FFE910004256FFE082E43E214F72FB29" box="[217,361,1137,1164]" pageId="17" pageNumber="268" value="2022-06-19">
19.
<collectionCode id="ED46AE054256FFE0833F3E224F38FB2E" box="[258,291,1138,1164]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="268">VI</collectionCode>
. 2022
</date>
, Y. Tasaku (cYT).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082FA3EC54C75FAC1" pageId="17" pageNumber="268" type="diagnosis">
<paragraph id="8BE836C04256FFE082FA3EC54C75FAC1" blockId="17.[151,1437,957,1632]" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE082FA3EC54F59FB0D" bold="true" box="[199,322,1173,1199]" pageId="17" pageNumber="268">Diagnosis.</emphasis>
It is distinguished from other species of the genus except
<taxonomicName id="4C574D434256FFE081C83EC64A42FB0D" authorityName="K. Sawada" authorityYear="1955" box="[1013,1113,1174,1200]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24256FFE081C83EC64A12FB0D" box="[1013,1033,1174,1199]" italics="true" pageId="17" pageNumber="268">D</emphasis>
.
<emphasis id="B923EAD24256FFE086263EC64A42FB0D" box="[1051,1113,1174,1199]" italics="true" pageId="17" pageNumber="268">uenoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C574D434256FFE086AA3EC64B33FB0D" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[1175,1320,1173,1200]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="orientalis" status="sp. n.">
<emphasis id="B923EAD24256FFE086AA3EC64AB0FB0D" box="[1175,1195,1174,1199]" italics="true" pageId="17" pageNumber="268">D</emphasis>
.
<emphasis id="B923EAD24256FFE086803EC54B33FB0D" box="[1213,1320,1173,1199]" italics="true" pageId="17" pageNumber="268">orientalis</emphasis>
</taxonomicName>
<emphasis id="B923EAD24256FFE0870E3EC54B6DFB0D" bold="true" box="[1331,1398,1173,1199]" pageId="17" pageNumber="268">
<taxonomicNameLabel id="A21057A94256FFE0870E3EC54B6DFB0D" box="[1331,1398,1173,1199]" pageId="17" pageNumber="268" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the following combination of characteristics: ground color reddish brown, abdominal segments VI and VII black (sometimes segment V also black) (
<figureCitation id="136C2A454256FFE080143E8D4CCAFB55" box="[553,721,1245,1271]" captionStart="FIGURE 10" captionStartId="15.[151,250,1778,1802]" captionTargetBox="[162,1425,500,1744]" captionTargetId="figure-191@15.[151,1436,489,1755]" captionTargetPageId="15" captionText="FIGURE 10 Habitus and mouthparts of Diaulota submarina. A, habitus, male; B, ditto, female; C, labium, D, mentum. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for C &amp; D." figureDoi="http://doi.org/10.5281/zenodo.10361603" httpUri="https://zenodo.org/record/10361603/files/figure.png" pageId="17" pageNumber="268">Figs. 10A &amp; B</figureCitation>
); male head larger than that of female (
<figureCitation id="136C2A454256FFE086B03E8D4B31FB55" box="[1165,1322,1245,1271]" captionStart="FIGURE 10" captionStartId="15.[151,250,1778,1802]" captionTargetBox="[162,1425,500,1744]" captionTargetId="figure-191@15.[151,1436,489,1755]" captionTargetPageId="15" captionText="FIGURE 10 Habitus and mouthparts of Diaulota submarina. A, habitus, male; B, ditto, female; C, labium, D, mentum. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for C &amp; D." figureDoi="http://doi.org/10.5281/zenodo.10361603" httpUri="https://zenodo.org/record/10361603/files/figure.png" pageId="17" pageNumber="268">Fig. 10A &amp; B</figureCitation>
); mentum transverse (
<figureCitation id="136C2A454256FFE083263F514F9CFAB9" box="[283,391,1281,1307]" captionStart="FIGURE 10" captionStartId="15.[151,250,1778,1802]" captionTargetBox="[162,1425,500,1744]" captionTargetId="figure-191@15.[151,1436,489,1755]" captionTargetPageId="15" captionText="FIGURE 10 Habitus and mouthparts of Diaulota submarina. A, habitus, male; B, ditto, female; C, labium, D, mentum. Scale bars: 1.0 mm for A &amp; B; 0.1 mm for C &amp; D." figureDoi="http://doi.org/10.5281/zenodo.10361603" httpUri="https://zenodo.org/record/10361603/files/figure.png" pageId="17" pageNumber="268">Fig. 10D</figureCitation>
); infraorbital carina short, shorter than half of head; tarsal formula 4-4-4. Compared to
<taxonomicName id="4C574D434256FFE087BC3F524ECEFA9D" authorityName="K. Sawada" authorityYear="1955" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="B923EAD24256FFE087BC3F524B8EFAB9" box="[1409,1429,1282,1307]" italics="true" pageId="17" pageNumber="268">D</emphasis>
.
<emphasis id="B923EAD24256FFE082AA3F764ECEFA9D" box="[151,213,1318,1343]" italics="true" pageId="17" pageNumber="268">uenoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C574D434256FFE083363F764F83FA9D" authority="Tasaku &amp; Ono &amp; Maruyama, 2023" authorityName="Tasaku &amp; Ono &amp; Maruyama" authorityYear="2023" box="[267,408,1317,1344]" class="Insecta" family="Staphylinidae" genus="Diaulota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="orientalis" status="sp. n.">
<emphasis id="B923EAD24256FFE083363F764F04FA9D" box="[267,287,1318,1343]" italics="true" pageId="17" pageNumber="268">D</emphasis>
.
<emphasis id="B923EAD24256FFE083103F754F83FA9D" box="[301,408,1317,1343]" italics="true" pageId="17" pageNumber="268">orientalis</emphasis>
</taxonomicName>
<emphasis id="B923EAD24256FFE083A23F754FC5FA9D" bold="true" box="[415,478,1317,1343]" pageId="17" pageNumber="268">
<taxonomicNameLabel id="A21057A94256FFE083A23F754FC5FA9D" box="[415,478,1317,1343]" pageId="17" pageNumber="268" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, it can be distinguished by its polygonal apical sclerite, thin suspensonia and shape of spermatheca (
<figureCitation id="136C2A454256FFE0830F3F194C7AFAC1" box="[306,609,1353,1380]" captionStart="FIGURE 11" captionStartId="16.[151,250,1917,1941]" captionTargetBox="[160,1427,192,1882]" captionTargetId="figure-22@16.[151,1436,181,1893]" captionTargetPageId="16" captionText="FIGURE 11 Genitalia of Diaulota submarina (A–D, from Tsushima; E–H, from Hiroshima-ken). A &amp; E, median lobe of aedeagus, lateral view; B &amp; F, ditto, ventral view; C &amp; G, internal structure of median lobe, ventral view; D &amp; H, spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361606" httpUri="https://zenodo.org/record/10361606/files/figure.png" pageId="17" pageNumber="268">Figs. 11A, C, D, E, G &amp; H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082FA3F3D4DB4F9BA" pageId="17" pageNumber="268" type="description">
<paragraph id="8BE836C04256FFE082FA3F3D4F41FA51" blockId="17.[151,1437,957,1632]" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE082FA3F3D4F24FA25" bold="true" box="[199,319,1389,1415]" pageId="17" pageNumber="268">Variation.</emphasis>
In the specimens from Tsushima and Fukuoka-ken, Kyushu: apical half of median lobe widened around apical 1/7–1/6 (
<figureCitation id="136C2A454256FFE083A33FC14C1FFA09" box="[414,516,1425,1451]" captionStart="FIGURE 11" captionStartId="16.[151,250,1917,1941]" captionTargetBox="[160,1427,192,1882]" captionTargetId="figure-22@16.[151,1436,181,1893]" captionTargetPageId="16" captionText="FIGURE 11 Genitalia of Diaulota submarina (A–D, from Tsushima; E–H, from Hiroshima-ken). A &amp; E, median lobe of aedeagus, lateral view; B &amp; F, ditto, ventral view; C &amp; G, internal structure of median lobe, ventral view; D &amp; H, spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361606" httpUri="https://zenodo.org/record/10361606/files/figure.png" pageId="17" pageNumber="268">Fig. 11B</figureCitation>
); suspensonia nearly S-shaped (
<figureCitation id="136C2A454256FFE081533FC14DCFFA09" box="[878,980,1425,1451]" captionStart="FIGURE 11" captionStartId="16.[151,250,1917,1941]" captionTargetBox="[160,1427,192,1882]" captionTargetId="figure-22@16.[151,1436,181,1893]" captionTargetPageId="16" captionText="FIGURE 11 Genitalia of Diaulota submarina (A–D, from Tsushima; E–H, from Hiroshima-ken). A &amp; E, median lobe of aedeagus, lateral view; B &amp; F, ditto, ventral view; C &amp; G, internal structure of median lobe, ventral view; D &amp; H, spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361606" httpUri="https://zenodo.org/record/10361606/files/figure.png" pageId="17" pageNumber="268">Fig. 11C</figureCitation>
). In the specimens from Yamaguchi-ken and Hiroshima-ken, Honshû: apical half of median lobe almost parallel-sided (
<figureCitation id="136C2A454256FFE0863D3FE54A79FA6D" box="[1024,1122,1461,1487]" captionStart="FIGURE 11" captionStartId="16.[151,250,1917,1941]" captionTargetBox="[160,1427,192,1882]" captionTargetId="figure-22@16.[151,1436,181,1893]" captionTargetPageId="16" captionText="FIGURE 11 Genitalia of Diaulota submarina (A–D, from Tsushima; E–H, from Hiroshima-ken). A &amp; E, median lobe of aedeagus, lateral view; B &amp; F, ditto, ventral view; C &amp; G, internal structure of median lobe, ventral view; D &amp; H, spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361606" httpUri="https://zenodo.org/record/10361606/files/figure.png" pageId="17" pageNumber="268">Fig. 11F</figureCitation>
); suspensonia thin rectangle shape (
<figureCitation id="136C2A454256FFE082DB3F894F55FA51" box="[230,334,1497,1523]" captionStart="FIGURE 11" captionStartId="16.[151,250,1917,1941]" captionTargetBox="[160,1427,192,1882]" captionTargetId="figure-22@16.[151,1436,181,1893]" captionTargetPageId="16" captionText="FIGURE 11 Genitalia of Diaulota submarina (A–D, from Tsushima; E–H, from Hiroshima-ken). A &amp; E, median lobe of aedeagus, lateral view; B &amp; F, ditto, ventral view; C &amp; G, internal structure of median lobe, ventral view; D &amp; H, spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10361606" httpUri="https://zenodo.org/record/10361606/files/figure.png" pageId="17" pageNumber="268">Fig. 11G</figureCitation>
).
</paragraph>
<paragraph id="8BE836C04256FFE082FA3FAD4DB4F9BA" blockId="17.[151,1437,957,1632]" box="[199,943,1533,1560]" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE082FA3FAD4F31F9B5" bold="true" box="[199,298,1533,1559]" pageId="17" pageNumber="268">Habitat.</emphasis>
This species was collected from rocky shores (
<figureCitation id="136C2A454256FFE081063FAD4DB9F9BA" box="[827,930,1533,1560]" captionStart="FIGURE 25" captionStartId="37.[151,250,1830,1854]" captionTargetBox="[151,1436,417,1807]" captionTargetId="figure-130@37.[151,1436,417,1807]" captionTargetPageId="37" captionText="FIGURE 25 Distribution and habitat of Rotundicephala gen. n. spp. of Japan. A, known localities in Japan (red circle, R. pacifica comb. n. (red ring, type locality)); blue square, R. koheii sp. n.; yellow triangle, R. koreana comb. n.); B, habitat of R. koheii sp. n. (Kanagawa-ken, Japan); C, habitat of R. pacifica comb. n. (Tsushima, adults of Diaulota submarina was collected same time)." figureDoi="http://doi.org/10.5281/zenodo.10361673" httpUri="https://zenodo.org/record/10361673/files/figure.png" pageId="17" pageNumber="268">Fig. 25C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34D654B4256FFE082FA3C714EC4F9C2" pageId="17" pageNumber="268" type="distribution">
<paragraph id="8BE836C04256FFE082FA3C714EC4F9C2" blockId="17.[151,1437,957,1632]" pageId="17" pageNumber="268">
<emphasis id="B923EAD24256FFE082FA3C714F7AF999" bold="true" box="[199,353,1569,1595]" pageId="17" pageNumber="268">Distribution.</emphasis>
<collectingCountry id="F34076504256FFE0835B3C724FB1F99E" box="[358,426,1570,1596]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
: Honshû, Kyûshû, Tsushima (
<collectingCountry id="F34076504256FFE080C93C714D2DF999" box="[756,822,1569,1595]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
Sea side);
<collectingCountry id="F34076504256FFE081933C714A22F99E" box="[942,1081,1569,1596]" name="South Korea" pageId="17" pageNumber="268">South Korea</collectingCountry>
(
<figureCitation id="136C2A454256FFE086753C714A80F999" box="[1096,1179,1569,1595]" captionStart="FIGURE 24" captionStartId="36.[151,250,1780,1804]" captionTargetBox="[151,1436,345,1757]" captionTargetId="figure-99@36.[151,1436,345,1757]" captionTargetPageId="36" captionText="FIGURE 24 Distribution of Diaulota uenoi (red circle (red ring, type locality)), D. submarina (blue square), and D. orientalis sp. n. (yellow triangle)." figureDoi="http://doi.org/10.5281/zenodo.10361669" httpUri="https://zenodo.org/record/10361669/files/figure.png" pageId="17" pageNumber="268">Fig. 24</figureCitation>
). Newly recorded from
<collectingCountry id="F34076504256FFE082AA3C164EC1F9C2" box="[151,218,1606,1632]" name="Japan" pageId="17" pageNumber="268">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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