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<document id="8C35C7CA11E8D4DAF8BE8E9815BF2335" ID-CLB-Dataset="265957" ID-DOI="10.11646/zootaxa.5343.1.3" ID-GBIF-Dataset="2fdc0a71-7108-489f-a878-b23d3437bddf" ID-ISSN="1175-5326" ID-Zenodo-Dep="8324250" ID-ZooBank="20BF9AB4-F86D-42EA-AEAA-7C03EBA44EB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694069790509" checkinUser="plazi" docAuthor="Yamasaki, Takeshi, Hashimoto, Yoshiaki, Endo, Tomoji, Hyodo, Fujio, Itioka, Takao, Mohamed, Maryati &amp; Meleng, Paulus" docDate="2023" docId="03FA87E7FFB1FFD3FF7D3F87FD568BA2" docLanguage="en" docName="zootaxa.5343.1.3.pdf" docOrigin="Zootaxa 5343 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Utivarachna rubra Deeleman-Reinhold 2001" docType="treatment" docVersion="5" lastPageNumber="63" masterDocId="FFC3FF9FFFB7FFDBFFEA3E15FFEC8816" masterDocTitle="Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species" masterLastPageNumber="73" masterPageNumber="55" pageNumber="61" updateTime="1694104750127" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E7C0B8C9BEDB5F0F2707F342E5294B39">Taxonomic study of Bornean species of Utivarachna Kishida, 1940 (Araneae: Trachelidae), with the description of a new species</mods:title>
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<mods:namePart id="55D41688F9329E534E18663B5A94589D">Yamasaki, Takeshi</mods:namePart>
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<mods:affiliation id="229C10491089DE72D03EC54558C08226">Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; k 0468874 @ gmail. com; https: // orcid. org / 0000 - 0002 - 2419 - 188 X</mods:affiliation>
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<mods:namePart id="E0CE389EA47EB3303335DC5CDBBE16A8">Hashimoto, Yoshiaki</mods:namePart>
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<mods:affiliation id="F0E44C95A93A997833877786BAA78F92">Institute of Natural and Environmental Sciences, University of Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; Museum of Nature and Human Activities, Hyogo, Yayoigaoka 6, Sanda-shi, Hyogo 669 - 1546, Japan &amp; aenictus @ mac. com; https: // orcid. org / 0000 - 0002 - 6696 - 5011</mods:affiliation>
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<mods:namePart id="B4ED2F92C838C06050407D3BCDF63157">Endo, Tomoji</mods:namePart>
<mods:nameIdentifier id="D9B6DFBF85E905541F53539E7F203940" type="ORCID">0009-0007-8279-9897</mods:nameIdentifier>
<mods:affiliation id="A0D637A20D247CED1A7442FB7CF24B13">School of Human Sciences, Kobe College, Okadayama 4 - 1, Nishinomiya, Hyogo, 662 - 8505, Japan &amp; endo @ mail. kobe-c. ac. jp; https: // orcid. org / 0009 - 0007 - 8279 - 9897</mods:affiliation>
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<mods:namePart id="F55EB237C2881D579CAB2FB4D234BAEB">Hyodo, Fujio</mods:namePart>
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<mods:affiliation id="97B5FCB3017903A69DD36568F002E9C8">Research Core for Interdisciplinary Sciences, Okayama University, 3 - 1 - 1, Tsushimanaka, Okayama, 700 - 8530, Japan &amp; fhyodo @ cc. okayama-u. ac. jp; https: // orcid. org / 0000 - 0001 - 8344 - 9353</mods:affiliation>
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<mods:namePart id="FCEC33D2D74095EA06032EBB132E2A7E">Itioka, Takao</mods:namePart>
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<mods:affiliation id="A301C65009AE64CDDB306E28AB89DD1B">Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-nihonmatsu, Kyoto 606 - 8501, Japan &amp; ichioka. takao. 5 m @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 8295 - 5657</mods:affiliation>
<mods:nameIdentifier id="7D773B95743A97B53662859D30CB525E" type="email">ichioka.takao.5m@kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="60356806C493994A4710CFD0E6FA8612">Mohamed, Maryati</mods:namePart>
<mods:affiliation id="16808E4B44B6CF2C4658D5EF75C7D860">Tun Hussein Onn Malaysia, Hab Pendidikan Tinggi Pagoh, KM 1, Jalan Panchor, 84600 Panchor, Johor, Malaysia &amp; maryatihayati 52 @ gmail. com</mods:affiliation>
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<mods:namePart id="71EE742FECF5A0EF90DE6D5AA90F379B">Meleng, Paulus</mods:namePart>
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<mods:affiliation id="0242B4A96889594D5260A5116EC1E793">Research Development and Innovation Division, Forest Department Sarawak, Kuching, KM 10, Jalan Datuk Amar Stephen Kalong Ningkan, 93250 Kuching, Sarawak, Malaysia &amp; paulusm @ sarawak. gov. my; https: // orcid. org / 0000 - 0003 - 3777 - 0597</mods:affiliation>
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<mods:date id="E20A10005380546311BE2AACD688578A">2023</mods:date>
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<treatment id="03FA87E7FFB1FFD3FF7D3F87FD568BA2" ID-DOI="http://doi.org/10.5281/zenodo.8324639" ID-Zenodo-Dep="8324639" LSID="urn:lsid:plazi:treatment:03FA87E7FFB1FFD3FF7D3F87FD568BA2" httpUri="http://treatment.plazi.org/id/03FA87E7FFB1FFD3FF7D3F87FD568BA2" lastPageId="8" lastPageNumber="63" pageId="6" pageNumber="61">
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<heading id="D0A4819DFFB1FFDDFF7D3F87FD2F89BA" bold="true" box="[151,707,402,429]" fontSize="11" level="1" pageId="6" pageNumber="61" reason="1">
<taxonomicName id="4C534D72FFB1FFDDFF7D3F87FD2F89BA" ID-CoL="7F633" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[151,707,402,429]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B927EAE3FFB1FFDDFF7D3F87FD2F89BA" bold="true" box="[151,707,402,429]" pageId="6" pageNumber="61">
<emphasis id="B927EAE3FFB1FFDDFF7D3F87FE9689BB" bold="true" box="[151,378,402,429]" italics="true" pageId="6" pageNumber="61">Utivarachna rubra</emphasis>
<bibRefCitation id="EFC24B00FFB1FFDDFE683F87FD2F89BA" author="Deeleman-Reinhold, C. L." box="[386,707,402,428]" pageId="6" pageNumber="61" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
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<figureCitation id="13682A74FFB1FFDDFF7D3FA2FEEA89C7" box="[151,262,439,465]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStart-2="FIGURE 10" captionStart-3="FIGURE 11" captionStartId-0="6.[151,250,1471,1495]" captionStartId-1="7.[151,250,922,946]" captionStartId-2="7.[151,250,1935,1959]" captionStartId-3="8.[151,250,491,515]" captionTargetBox-0="[169,1435,699,1447]" captionTargetBox-1="[151,1433,419,899]" captionTargetBox-2="[151,1436,1295,1912]" captionTargetBox-3="[151,1435,182,468]" captionTargetId-0="figure-115@6.[151,1435,699,1447]" captionTargetId-1="figure-114@7.[151,1435,417,899]" captionTargetId-2="figure-228@7.[151,1436,1295,1912]" captionTargetId-3="figure-17@8.[151,1435,181,468]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 8. Utivarachna rubra Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15184). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: A and B, 1 mm; C and D, 0.50 mm." captionText-1="FIGURE 9. Utivarachna rubra, male, holotype (RNMH.ARA.15184). A left palp, prolateral view; B same, ventral view; C same, retrolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." captionText-2="FIGURE 10. Utivarachna rubra Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15184). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D left leg I, prolateral view. Scale bars: A and B, 1 mm; C and D, 0.50 mm." captionText-3="FIGURE 11. Utivarachna rubra, female, paratype (RNMH.ARA.15184). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.8324274" figureDoi-1="http://doi.org/10.5281/zenodo.8324276" figureDoi-2="http://doi.org/10.5281/zenodo.8324278" figureDoi-3="http://doi.org/10.5281/zenodo.8324280" httpUri-0="https://zenodo.org/record/8324274/files/figure.png" httpUri-1="https://zenodo.org/record/8324276/files/figure.png" httpUri-2="https://zenodo.org/record/8324278/files/figure.png" httpUri-3="https://zenodo.org/record/8324280/files/figure.png" pageId="6" pageNumber="61">Figs 8–11</figureCitation>
</paragraph>
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<paragraph id="8BEC36F1FFB1FFDDFF7D3FEEFCD98A05" blockId="6.[151,821,507,531]" box="[151,821,507,531]" pageId="6" pageNumber="61">
<treatmentCitationGroup id="AB4311DFFFB1FFDDFF7D3FEEFCD98A05" box="[151,821,507,531]" pageId="6" pageNumber="61">
<treatmentCitation id="0AF210E0FFB1FFDDFF7D3FEEFD728A05" author="Deeleman-Reinhold, C. L." box="[151,670,507,531]" page="391" pageId="6" pageNumber="61" year="2001">
<taxonomicName id="4C534D72FFB1FFDDFF7D3FEEFD728A05" ID-CoL="7F633" authority="Deeleman-Reinhold, 2001: 391" authorityName="Deeleman-Reinhold" authorityPageNumber="391" authorityYear="2001" box="[151,670,507,531]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B927EAE3FFB1FFDDFF7D3FEEFEB48A05" box="[151,344,507,531]" italics="true" pageId="6" pageNumber="61">Utivarachna rubra</emphasis>
<bibRefCitation id="EFC24B00FFB1FFDDFEB43FEEFD728A05" author="Deeleman-Reinhold, C. L." box="[350,670,507,531]" pageId="6" pageNumber="61" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001: 391</bibRefCitation>
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, figs 620–622.
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</paragraph>
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<subSubSection id="C349657AFFB1FFDDFF7D3C54FE3A8EE1" pageId="6" pageNumber="61" type="materials_examined">
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<emphasis id="B927EAE3FFB1FFDDFF7D3C54FE538A4D" bold="true" box="[151,447,577,603]" pageId="6" pageNumber="61">Type material examined</emphasis>
.
<materialsCitation id="3B3B3CACFFB1FFDDFE3B3C54FDA98AB5" collectingDate="1985-09-02" collectingDateMax="1985-09-16" collectingDateMin="1985-09-02" collectionCode="RMNH" latitude="-2.0333333" location="Kalimantan" longLatPrecision="1307" longitude="113.666664" municipality="Deeleman-Reinhold" pageId="6" pageNumber="61" specimenCode="RMNH 15184" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B927EAE3FFB1FFDDFE3B3C54FDB18A49" bold="true" box="[465,605,577,607]" pageId="6" pageNumber="61">
<typeStatus id="54E88853FFB1FFDDFE3B3C54FDD28A4D" box="[465,574,577,603]" pageId="6" pageNumber="61">Holotype</typeStatus>
♁
</emphasis>
(
<specimenCode id="DBF59E8AFFB1FFDDFD853C54FCF48A4A" box="[623,792,577,604]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="61" type="Museum">RMNH 15184</specimenCode>
) and
<emphasis id="B927EAE3FFB1FFDDFCB73C54FC048A49" bold="true" box="[861,1000,577,607]" pageId="6" pageNumber="61">
paratype
<specimenCount id="9D55FD78FFB1FFDDFC393C56FC048A49" box="[979,1000,579,607]" pageId="6" pageNumber="61" type="female">♀</specimenCount>
</emphasis>
(
<specimenCode id="DBF59E8AFFB1FFDDFC103C54FB4A8A4A" box="[1018,1190,577,604]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="61" type="Museum">RMNH 15184</specimenCode>
).
<collectorName id="26A65327FFB1FFDDFB563C54FB108A4D" box="[1212,1276,577,603]" pageId="6" pageNumber="61">Label</collectorName>
data for both shown in
<figureCitation id="13682A74FFB1FFDDFEE13C70FE888A69" box="[267,356,613,639]" captionStart="FIGURE 8" captionStartId="6.[151,250,1471,1495]" captionTargetBox="[169,1435,699,1447]" captionTargetId="figure-115@6.[151,1435,699,1447]" captionTargetPageId="6" captionText="FIGURE 8. Utivarachna rubra Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15184). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: A and B, 1 mm; C and D, 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.8324274" httpUri="https://zenodo.org/record/8324274/files/figure.png" pageId="6" pageNumber="61">Fig. 8E</figureCitation>
.
<collectorName id="26A65327FFB1FFDDFE983C73FE4B8A96" box="[370,423,614,640]" pageId="6" pageNumber="61">Data</collectorName>
according to
<bibRefCitation id="EFC24B00FFB1FFDDFDA03C70FC688A69" author="Deeleman-Reinhold, C. L." box="[586,900,613,639]" pageId="6" pageNumber="61" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">
<collectingMunicipality id="6B88AC8BFFB1FFDDFDA03C70FCC08A69" box="[586,812,613,639]" pageId="6" pageNumber="61">Deeleman-Reinhold</collectingMunicipality>
(2001)
</bibRefCitation>
:
<geoCoordinate id="EE675036FFB1FFDDFC783C70FC028A69" box="[914,1006,613,639]" degrees="02" direction="south" minutes="02" orientation="latitude" pageId="6" pageNumber="61" precision="925" value="-2.0333333">02º02’S</geoCoordinate>
,
<geoCoordinate id="EE675036FFB1FFDDFC173C70FB848A69" box="[1021,1128,613,639]" degrees="113" direction="east" minutes="40" orientation="longitude" pageId="6" pageNumber="61" precision="925" value="113.666664">113º40’E</geoCoordinate>
,
<collectorName id="26A65327FFB1FFDDFB9D3C70FAE98A69" box="[1143,1285,613,639]" pageId="6" pageNumber="61">Kaharian, C</collectorName>
.
<location id="8E8C602AFFB1FFDDFAF93C70FA758A69" LSID="urn:lsid:plazi:treatment:03FA87E7FFB1FFD3FF7D3F87FD568BA2:8E8C602AFFB1FFDDFAF93C70FA758A69" box="[1299,1433,613,639]" latitude="-2.0333333" longLatPrecision="1307" longitude="113.666664" municipality="Deeleman-Reinhold" name="Kalimantan" pageId="6" pageNumber="61">Kalimantan</location>
,
<date id="FFED1031FFB1FFDDFF7D3C9CFED98AB2" box="[151,309,649,676]" pageId="6" pageNumber="61" value="1985-09-02" valueMax="1985-09-16" valueMin="1985-09-02">
<collectingDate id="EFA9E9D9FFB1FFDDFF7D3C9CFED98AB2" box="[151,309,649,676]" pageId="6" pageNumber="61" value="1985-09-02" valueMax="1985-09-16" valueMin="1985-09-02">2–16.IX.1985</collectingDate>
</date>
, leg.
<collectorName id="26A65327FFB1FFDDFE9B3C9CFDA98AB5" box="[369,581,649,675]" pageId="6" pageNumber="61">S. Djojosudharmo.</collectorName>
</materialsCitation>
</paragraph>
<caption id="DF2C6679FFB1FFDDFF7D3BAAFE628E08" ID-DOI="http://doi.org/10.5281/zenodo.8324274" ID-Zenodo-Dep="8324274" httpUri="https://zenodo.org/record/8324274/files/figure.png" pageId="6" pageNumber="61" startId="6.[151,250,1471,1495]" targetBox="[169,1435,699,1447]" targetPageId="6" targetType="figure">
<paragraph id="8BEC36F1FFB1FFDDFF7D3BAAFE628E08" blockId="6.[151,1437,1471,1567]" pageId="6" pageNumber="61">
<emphasis id="B927EAE3FFB1FFDDFF7D3BAAFEF98DC1" bold="true" box="[151,277,1471,1495]" pageId="6" pageNumber="61">FIGURE 8.</emphasis>
<taxonomicName id="4C534D72FFB1FFDDFEF73BAAFD148DC1" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[285,760,1471,1495]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B927EAE3FFB1FFDDFEF73BAAFE0C8DC1" box="[285,480,1471,1495]" italics="true" pageId="6" pageNumber="61">Utivarachna rubra</emphasis>
<bibRefCitation id="EFC24B00FFB1FFDDFE023BAAFD148DC1" author="Deeleman-Reinhold, C. L." box="[488,760,1471,1495]" pageId="6" pageNumber="61" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, male, holotype (RNMH.ARA.15184).
<emphasis id="B927EAE3FFB1FFDDFB783BAAFB488DC1" bold="true" box="[1170,1188,1471,1495]" pageId="6" pageNumber="61">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFB1FFDDFA613BAAFA708DC1" bold="true" box="[1419,1436,1471,1495]" pageId="6" pageNumber="61">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFB1FFDDFE8C3BF6FE948DED" bold="true" box="[358,376,1507,1531]" pageId="6" pageNumber="61">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFB1FFDDFCDE3BF6FCAA8DED" bold="true" box="[820,838,1507,1531]" pageId="6" pageNumber="61">D</emphasis>
leg I, left, prolateral view;
<emphasis id="B927EAE3FFB1FFDDFB8B3BF6FB9E8DED" bold="true" box="[1121,1138,1507,1531]" pageId="6" pageNumber="61">E</emphasis>
label. Scale bars: A and B, 1 mm; C and D, 0.50 mm.
</paragraph>
</caption>
<paragraph id="8BEC36F1FFB1FFDDFF2D3864FE3A8EE1" blockId="6.[151,1437,1649,2036]" pageId="6" pageNumber="61">
<emphasis id="B927EAE3FFB1FFDDFF2D3864FE6E8E9D" bold="true" box="[199,386,1649,1675]" pageId="6" pageNumber="61">Other material.</emphasis>
<materialsCitation id="3B3B3CACFFB1FFDDFE603864FE3F8EE1" collectingDate="2009-02-27" collectingDateMax="2018-08-29" collectingDateMin="2009-02-27" collectionCode="T, MNHAH" country="Malaysia" county="Endo" location="Sungai Liku" municipality="Lambir Hills National Park" pageId="6" pageNumber="61" specimenCode="LMB-TY20180829" specimenCount="2" specimenCount-female="2">
2 ♁ (
<specimenCode id="DBF59E8AFFB1FFDDFE2E3864FD488E9D" box="[452,676,1649,1675]" collectionCode="LMB-TY" pageId="6" pageNumber="61">LMB-TY20180829</specimenCode>
-TR04,
<specimenCode id="DBF59E8AFFB1FFDDFD133864FC378E9A" box="[761,987,1649,1676]" collectionCode="LMB-TY" pageId="6" pageNumber="61">LMB-TY20180829</specimenCode>
-TR05),
<specimenCount id="9D55FD78FFB1FFDDFBDF3864FB8C8E9A" box="[1077,1120,1649,1676]" pageId="6" pageNumber="61" type="female">2 ♀</specimenCount>
(
<specimenCode id="DBF59E8AFFB1FFDDFB853864FABC8E9D" box="[1135,1360,1649,1675]" collectionCode="LMB-TY" pageId="6" pageNumber="61">LMB-TY20180829</specimenCode>
-TR02,
<specimenCode id="DBF59E8AFFB1FFDDFF7D3880FE958EB9" box="[151,377,1685,1711]" collectionCode="LMB-TY" pageId="6" pageNumber="61">LMB-TY20180829</specimenCode>
-TR03), Sungai Liku [Liku River], Lambir Hills National Park, Sarawak,
<date id="FFED1031FFB1FFDDFB5F3880FAA58EB9" box="[1205,1353,1685,1711]" pageId="6" pageNumber="61" value="2018-08-29">29.VIII.2018</date>
, leg. T. Yamasaki; 1 ♁ (LCo20090227-E3, MNHAH), Walk-way (beyond second tower),
<date id="FFED1031FFB1FFDDFBCF38ACFB708EC5" box="[1061,1180,1721,1747]" pageId="6" pageNumber="61" value="2009-02-27">27.II.2009</date>
, Lambir Hills National Park, Sarawak, leg. T. Endo
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFB1FFDDFF2D3914FC968F0A" box="[199,890,1793,1820]" pageId="6" pageNumber="61" type="diagnosis">
<paragraph id="8BEC36F1FFB1FFDDFF2D3914FC968F0A" blockId="6.[151,1437,1649,2036]" box="[199,890,1793,1820]" pageId="6" pageNumber="61">
<emphasis id="B927EAE3FFB1FFDDFF2D3914FEAE8F0D" bold="true" box="[199,322,1793,1819]" pageId="6" pageNumber="61">Diagnosis.</emphasis>
See Diagnosis of
<taxonomicName id="4C534D72FFB1FFDDFDE73917FD9B8F0D" authority="Yamasaki &amp; Hashimoto &amp; Endo &amp; Hyodo &amp; Itioka &amp; Mohamed &amp; Meleng, 2023" authorityName="Yamasaki &amp; Hashimoto &amp; Endo &amp; Hyodo &amp; Itioka &amp; Mohamed &amp; Meleng" authorityYear="2023" box="[525,631,1793,1819]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="itiokai" status="sp. nov.">
<emphasis id="B927EAE3FFB1FFDDFDE73917FD9B8F0D" box="[525,631,1793,1819]" italics="true" pageId="6" pageNumber="61">U. itiokai</emphasis>
</taxonomicName>
<emphasis id="B927EAE3FFB1FFDDFD943914FD3B8F0D" bold="true" box="[638,727,1793,1819]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="A2145798FFB1FFDDFD943914FD3B8F0D" box="[638,727,1793,1819]" pageId="6" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C534D72FFB1FFDDFCE43917FC9F8F0D" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[782,883,1793,1819]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="dusun">
<emphasis id="B927EAE3FFB1FFDDFCE43917FC9F8F0D" box="[782,883,1793,1819]" italics="true" pageId="6" pageNumber="61">U. dusun</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFB1FFD3FF2D3930FB3D8B86" lastPageId="8" lastPageNumber="63" pageId="6" pageNumber="61" type="description">
<paragraph id="8BEC36F1FFB1FFDCFF2D3930FCF988C3" blockId="6.[151,1437,1649,2036]" lastBlockId="7.[151,1437,150,393]" lastPageId="7" lastPageNumber="62" pageId="6" pageNumber="61">
<emphasis id="B927EAE3FFB1FFDDFF2D3930FE9A8F29" bold="true" box="[199,374,1829,1855]" pageId="6" pageNumber="61">Measurements</emphasis>
(
<typeStatus id="54E88853FFB1FFDDFE663930FE1C8F29" box="[396,496,1829,1855]" pageId="6" pageNumber="61">holotype</typeStatus>
♁/
<typeStatus id="54E88853FFB1FFDDFDFF3932FD908F54" box="[533,636,1831,1858]" pageId="6" pageNumber="61">paratype</typeStatus>
<specimenCount id="9D55FD78FFB1FFDDFD603930FD728F29" box="[650,670,1829,1855]" pageId="6" pageNumber="61" type="female">♀</specimenCount>
). Carapace length 1.52/1.35; cephalic width 0.78/0.68; thoracic width 1.10/0.96. Diameter of eyes: AME 0.11/0.08; ALE 0.12/0.12; PME 0.10/0.11; PLE 0.11/0.11. Interdistances between eyes: AME–AME 0.03/0.02; AME–ALE 0.02/0.01; ALE–ALE 0.28/0.23; PME–PME 0.10/0.08; PME– PLE 0.10/0.07; PLE–PLE 0.47/0.43; ALE–PLE 0.08/0.06; AME–PME 0.05/0.03. MOA: length 0.28/0.26; anterior width 0.25/0.22; posterior width 0.30/0.28. Clypeus height 0.27/0.21. Abdomen length 1.45/1.47; width 1.01/1.13. Legs: I 3.83 (1.25, 0.50, 0.93, 0.63, 0.52)/3.22 (1.01, 0.37, 0.75, 0.61, 0.48); II 3.25 (0.96, 0.37, 0.72, 0.73, 0.47)/2.95 (0.88, 0.31, 0.66, 0.65, 0.45); III 2.56 (0.75, 0.30, 0.52, 0.62, 0.37)/2.29 (0.66, 0.28, 0.47, 0.55, 0.33); IV 3.23 (0.90, 0.33, 0.72, 0.85, 0.43)/3.02 (0.85, 0.28, 0.68, 0.80, 0.41).
</paragraph>
<paragraph id="8BEC36F1FFB0FFDCFF2D3ECAFBC9899F" blockId="7.[151,1437,150,393]" pageId="7" pageNumber="62">
<emphasis id="B927EAE3FFB0FFDCFF2D3ECAFEE888EF" bold="true" box="[199,260,223,249]" pageId="7" pageNumber="62">Male</emphasis>
(
<figureCitation id="13682A74FFB0FFDCFEFF3ECAFE7988EF" box="[277,405,223,249]" captionStart="FIGURE 8" captionStartId="6.[151,250,1471,1495]" captionTargetBox="[169,1435,699,1447]" captionTargetId="figure-115@6.[151,1435,699,1447]" captionTargetPageId="6" captionText="FIGURE 8. Utivarachna rubra Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15184). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: A and B, 1 mm; C and D, 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.8324274" httpUri="https://zenodo.org/record/8324274/files/figure.png" pageId="7" pageNumber="62">Fig. 8A–D</figureCitation>
). Carapace oval and superficially granulated with truncated posterior end. Thoracic lateral margins hardly undulating. AER slightly recurved. PER longer than AER, almost straight. Fovea shallow, indistinct. Clypeus margin with small projection that truncated or weakly bifurcated. Chelicerae superficially granulated, with three teeth on each prolateral and retrolateral margin. Pedicel short. Abdomen oval; entire dorsum weakly sclerotized. Legs superficially granulated; leg I enlarged, bearing cusps ventrally.
</paragraph>
<caption id="DF2C6679FFB0FFDCFF7D3D8FFAC58BC0" ID-DOI="http://doi.org/10.5281/zenodo.8324276" ID-Zenodo-Dep="8324276" httpUri="https://zenodo.org/record/8324276/files/figure.png" pageId="7" pageNumber="62" startId="7.[151,250,922,946]" targetBox="[151,1433,419,899]" targetPageId="7" targetType="figure">
<paragraph id="8BEC36F1FFB0FFDCFF7D3D8FFAC58BC0" blockId="7.[151,1436,922,982]" pageId="7" pageNumber="62">
<emphasis id="B927EAE3FFB0FFDCFF7D3D8FFEF88BA4" bold="true" box="[151,276,922,946]" pageId="7" pageNumber="62">FIGURE 9.</emphasis>
<taxonomicName id="4C534D72FFB0FFDCFEF73D8FFE338BA4" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[285,479,922,946]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B927EAE3FFB0FFDCFEF73D8FFE338BA4" box="[285,479,922,946]" italics="true" pageId="7" pageNumber="62">Utivarachna rubra</emphasis>
</taxonomicName>
, male, holotype (RNMH.ARA.15184).
<emphasis id="B927EAE3FFB0FFDCFC933D8FFC678BA4" bold="true" box="[889,907,922,946]" pageId="7" pageNumber="62">A</emphasis>
left palp, prolateral view;
<emphasis id="B927EAE3FFB0FFDCFB4A3D8FFB5D8BA4" bold="true" box="[1184,1201,922,946]" pageId="7" pageNumber="62">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFB0FFDCFA603D8FFA708BA4" bold="true" box="[1418,1436,922,946]" pageId="7" pageNumber="62">C</emphasis>
same, retrolateral view;
<emphasis id="B927EAE3FFB0FFDCFE663DABFE728BC0" bold="true" box="[396,414,958,982]" pageId="7" pageNumber="62">D</emphasis>
same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum.
</paragraph>
</caption>
<paragraph id="8BEC36F1FFB0FFDCFF2D3A3DFE418CB8" blockId="7.[151,1437,1064,1271]" pageId="7" pageNumber="62">
Palp (
<figureCitation id="13682A74FFB0FFDCFEE53A3DFE7E8C55" box="[271,402,1064,1091]" captionStart="FIGURE 9" captionStartId="7.[151,250,922,946]" captionTargetBox="[151,1433,419,899]" captionTargetId="figure-114@7.[151,1435,417,899]" captionTargetPageId="7" captionText="FIGURE 9. Utivarachna rubra, male, holotype (RNMH.ARA.15184). A left palp, prolateral view; B same, ventral view; C same, retrolateral view; D same, dorsal view. Scale bars: 0.50 mm. E—embolus, Sd—sperm duct, St—subtegulum." figureDoi="http://doi.org/10.5281/zenodo.8324276" httpUri="https://zenodo.org/record/8324276/files/figure.png" pageId="7" pageNumber="62">Fig. 9A–D</figureCitation>
). Cymbium oval. Bulb oval. Sperm duct U-shaped, visible through posterior tegulum. Embolus coiled at horizontal plane; embolus coil plane inclining dorsally, covered by cymbium; basal embolus part accompanied by sclerotized plate located in middle and anterior parts of tegulum. RTA long; anterior half of RTA strongly curved dorsally.
</paragraph>
<paragraph id="8BEC36F1FFB0FFDCFF2D3AADFB1E8CE0" blockId="7.[151,1437,1064,1271]" pageId="7" pageNumber="62">
Coloration and setation (
<figureCitation id="13682A74FFB0FFDCFE083AADFD8D8CC5" box="[482,609,1208,1235]" captionStart="FIGURE 8" captionStartId="6.[151,250,1471,1495]" captionTargetBox="[169,1435,699,1447]" captionTargetId="figure-115@6.[151,1435,699,1447]" captionTargetPageId="6" captionText="FIGURE 8. Utivarachna rubra Deeleman-Reinhold, 2001, male, holotype (RNMH.ARA.15184). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D leg I, left, prolateral view; E label. Scale bars: A and B, 1 mm; C and D, 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.8324274" httpUri="https://zenodo.org/record/8324274/files/figure.png" pageId="7" pageNumber="62">Fig. 8A–D</figureCitation>
). Carapace brown. Chelicerae brown. Abdomen covered with fine setae; dorsal sclerotized surface pale amber-colored; lateral and ventral surfaces cream, tinged with black.
</paragraph>
<caption id="DF2C6679FFB0FFDCFF7D399AFEAE8FF9" ID-DOI="http://doi.org/10.5281/zenodo.8324278" ID-Zenodo-Dep="8324278" httpUri="https://zenodo.org/record/8324278/files/figure.png" pageId="7" pageNumber="62" startId="7.[151,250,1935,1959]" targetBox="[151,1436,1295,1912]" targetPageId="7" targetType="figure">
<paragraph id="8BEC36F1FFB0FFDCFF7D399AFEAE8FF9" blockId="7.[151,1436,1935,2031]" pageId="7" pageNumber="62">
<emphasis id="B927EAE3FFB0FFDCFF7D399AFECD8FB1" bold="true" box="[151,289,1935,1959]" pageId="7" pageNumber="62">FIGURE 10.</emphasis>
<taxonomicName id="4C534D72FFB0FFDCFEC3399AFCEE8FB1" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[297,770,1935,1959]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B927EAE3FFB0FFDCFEC3399AFE078FB1" box="[297,491,1935,1959]" italics="true" pageId="7" pageNumber="62">Utivarachna rubra</emphasis>
<bibRefCitation id="EFC24B00FFB0FFDCFE19399AFCEE8FB1" author="Deeleman-Reinhold, C. L." box="[499,770,1935,1959]" pageId="7" pageNumber="62" refId="ref8882" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill, Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
, female, paratype (RNMH.ARA.15184).
<emphasis id="B927EAE3FFB0FFDCFB47399AFB538FB1" bold="true" box="[1197,1215,1935,1959]" pageId="7" pageNumber="62">A</emphasis>
habitus, dorsal view;
<emphasis id="B927EAE3FFB0FFDCFF7D39A6FF448FDD" bold="true" box="[151,168,1971,1995]" pageId="7" pageNumber="62">B</emphasis>
same, ventral view;
<emphasis id="B927EAE3FFB0FFDCFE9739A6FE638FDD" bold="true" box="[381,399,1971,1995]" pageId="7" pageNumber="62">C</emphasis>
cephalic part and chelicerae, frontal view;
<emphasis id="B927EAE3FFB0FFDCFCA339A6FCB78FDD" bold="true" box="[841,859,1971,1995]" pageId="7" pageNumber="62">D</emphasis>
left leg I, prolateral view. Scale bars: A and B, 1 mm; C and D, 0.50 mm.
</paragraph>
</caption>
<caption id="DF2C6679FFBFFFD3FF7D3FFEFB748A3E" ID-DOI="http://doi.org/10.5281/zenodo.8324280" ID-Zenodo-Dep="8324280" httpUri="https://zenodo.org/record/8324280/files/figure.png" pageId="8" pageNumber="63" startId="8.[151,250,491,515]" targetBox="[151,1435,182,468]" targetPageId="8" targetType="figure">
<paragraph id="8BEC36F1FFBFFFD3FF7D3FFEFB748A3E" blockId="8.[151,1436,491,552]" pageId="8" pageNumber="63">
<emphasis id="B927EAE3FFBFFFD3FF7D3FFEFEF38A12" bold="true" box="[151,287,491,516]" pageId="8" pageNumber="63">FIGURE 11.</emphasis>
<taxonomicName id="4C534D72FFBFFFD3FECD3FF9FE048A12" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[295,488,492,516]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B927EAE3FFBFFFD3FECD3FF9FE048A12" box="[295,488,492,516]" italics="true" pageId="8" pageNumber="63">Utivarachna rubra</emphasis>
</taxonomicName>
, female, paratype (RNMH.ARA.15184).
<emphasis id="B927EAE3FFBFFFD3FC793FFEFC498A15" bold="true" box="[915,933,491,515]" pageId="8" pageNumber="63">A</emphasis>
epigyne, ventral view;
<emphasis id="B927EAE3FFBFFFD3FB723FF9FB458A12" bold="true" box="[1176,1193,492,516]" pageId="8" pageNumber="63">B</emphasis>
internal organs, ventral view;
<emphasis id="B927EAE3FFBFFFD3FF3C3C1AFF048A31" bold="true" box="[214,232,527,551]" pageId="8" pageNumber="63">C</emphasis>
same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca.
</paragraph>
</caption>
<paragraph id="8BEC36F1FFBFFFD3FF2D3C6FFDAC8B16" blockId="8.[151,1436,634,948]" pageId="8" pageNumber="63">
<emphasis id="B927EAE3FFBFFFD3FF2D3C6FFEF28A82" bold="true" box="[199,286,634,660]" pageId="8" pageNumber="63">Female</emphasis>
(
<figureCitation id="13682A74FFBFFFD3FEC53C6FFE578A82" box="[303,443,634,660]" captionStart="FIGURE 10" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[151,1436,1295,1912]" captionTargetId="figure-228@7.[151,1436,1295,1912]" captionTargetPageId="7" captionText="FIGURE 10. Utivarachna rubra Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15184). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D left leg I, prolateral view. Scale bars: A and B, 1 mm; C and D, 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.8324278" httpUri="https://zenodo.org/record/8324278/files/figure.png" pageId="8" pageNumber="63">Fig. 10A–D</figureCitation>
). Somatic characteristics almost identical to diagnosis characteristics of
<taxonomicName id="4C534D72FFBFFFD3FB073C6EFABF8A82" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[1261,1363,634,660]" class="Arachnida" family="Trachelidae" genus="Utivarachna" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="dusun">
<emphasis id="B927EAE3FFBFFFD3FB073C6EFABF8A82" box="[1261,1363,634,660]" italics="true" pageId="8" pageNumber="63">U. dusun</emphasis>
</taxonomicName>
group (described above). Chelicerae with three prolateral and two retrolateral teeth. Abdomen oval, without distinct dorsal scutum. Leg I robust but weaker than that of male; ventral side covered with sparse cusps; ventral depression at proximal end of metatarsus indistinct.
</paragraph>
<paragraph id="8BEC36F1FFBFFFD3FF2D3D1FFA808B7A" blockId="8.[151,1436,634,948]" pageId="8" pageNumber="63">
Copulatory organs (
<figureCitation id="13682A74FFBFFFD3FE4A3D1FFDCA8B32" box="[416,550,778,804]" captionStart="FIGURE 11" captionStartId="8.[151,250,491,515]" captionTargetBox="[151,1435,182,468]" captionTargetId="figure-17@8.[151,1435,181,468]" captionTargetPageId="8" captionText="FIGURE 11. Utivarachna rubra, female, paratype (RNMH.ARA.15184). A epigyne, ventral view; B internal organs, ventral view; C same, dorsal view. Scale bars: 0.50 mm. Br—bursa, Fd—fertilization duct, S—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.8324280" httpUri="https://zenodo.org/record/8324280/files/figure.png" pageId="8" pageNumber="63">Fig. 11A–C</figureCitation>
). Copulatory atrium wider than long, with sclerotized lateral margins.Copulatory ducts wide anteriorly, gradually tapering posteriorly, forming complicated loops. Bursae slender pear-shaped, with spherical terminals. Spermathecae tube-like shaped, with long narrow fertilization ducts extending posteriorly.
</paragraph>
<paragraph id="8BEC36F1FFBFFFD3FF2D3D63FB3D8B86" blockId="8.[151,1436,634,948]" box="[199,1233,886,912]" pageId="8" pageNumber="63">
Coloration and setation (
<figureCitation id="13682A74FFBFFFD3FE363D63FD8B8B86" box="[476,615,886,912]" captionStart="FIGURE 10" captionStartId="7.[151,250,1935,1959]" captionTargetBox="[151,1436,1295,1912]" captionTargetId="figure-228@7.[151,1436,1295,1912]" captionTargetPageId="7" captionText="FIGURE 10. Utivarachna rubra Deeleman-Reinhold, 2001, female, paratype (RNMH.ARA.15184). A habitus, dorsal view; B same, ventral view; C cephalic part and chelicerae, frontal view; D left leg I, prolateral view. Scale bars: A and B, 1 mm; C and D, 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.8324278" httpUri="https://zenodo.org/record/8324278/files/figure.png" pageId="8" pageNumber="63">Fig. 10A–D</figureCitation>
). Almost same as in male, except for grayish abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C349657AFFBFFFD3FF2D3D8FFD568BA2" box="[199,698,922,948]" pageId="8" pageNumber="63" type="distribution">
<paragraph id="8BEC36F1FFBFFFD3FF2D3D8FFD568BA2" blockId="8.[151,1436,634,948]" box="[199,698,922,948]" pageId="8" pageNumber="63">
<emphasis id="B927EAE3FFBFFFD3FF2D3D8FFE8D8BA2" bold="true" box="[199,353,922,948]" pageId="8" pageNumber="63">Distribution.</emphasis>
Central Kalimantan,
<collectingCountry id="F3447661FFBFFFD3FDB83D8FFD5A8BA2" box="[594,694,922,948]" name="Malaysia" pageId="8" pageNumber="63">Sarawak</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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