254 lines
32 KiB
XML
254 lines
32 KiB
XML
<document id="7FEA7AB2C9B1A3D5646ACE3F6258772A" ID-CLB-Dataset="8140" ID-DOI="10.2478/aemnp-2019-0021" ID-GBIF-Dataset="726682f3-9eac-4961-ac4d-d8b8679e8211" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488950" ID-ZooBank="ABBA2B4F-8B60-41E2-B80B-45861F974B23" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219567173" checkinUser="felipe" docAuthor="Ribera, Ignacio, Hernando, Carles & Cieslak, Alexandra" docDate="2019" docId="03DD87A0FFD4FFD6FF4EFE6FDDDFFC8B" docLanguage="en" docName="ActaEntMusNatPra.59.1.253-272.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Ochthebius (Ochthebius) alhajarensis Ribera & Hernando & Cieslak 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="260" masterDocId="FFE4FFD8FFD3FFD1FF86FF93D922FFFE" masterDocTitle="Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae" masterLastPageNumber="272" masterPageNumber="253" pageNumber="260" updateTime="1698926098198" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods id="817551EA43114F5FF018CAFC555F39E7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B438E73B7DE147019AE259F38460E21F">
|
||
<mods:title id="251113BC16A7090201A7931B105F0066">Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="DC93F3E4F269073456DC11A960718CA7" type="personal">
|
||
<mods:role id="4B180CF1FC8AE290BED73561CC495F99">
|
||
<mods:roleTerm id="2AEE92CAC9F57E19EDB776BD844416DF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8A78FD81F14A44DCCB5EAAEF6A696ACA">Ribera, Ignacio</mods:namePart>
|
||
<mods:affiliation id="5F4B3B984941E716451DD72FC52228DE">Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37, E- 08003 Barcelona, Spain;</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="5A8274A3879682D6FFFFFCFC519C7A95" type="personal">
|
||
<mods:role id="B5EE7DF8EC0CA4B651E51B1F4110DC1C">
|
||
<mods:roleTerm id="58D0D975BC77CD6CC2CA79A9196A2AE6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="023F54A857D631565AF13938470F8059">Hernando, Carles</mods:namePart>
|
||
<mods:affiliation id="1E6C24C28FDC37422DD0083B6AB22102">P. O. box 118, E- 08911 Badalona, Catalonia, Spain; e-mail: leptotyphlus @ gmail. com</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="8B66AA38C981D417E4F6D2FD1377A500" type="personal">
|
||
<mods:role id="D918DCF01E7E411E89483DB7A2E7A295">
|
||
<mods:roleTerm id="7F91C0625ABAB30F6C43B9F18D5A901C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1DDD250D3A19A821DE2F12E306E846C0">Cieslak, Alexandra</mods:namePart>
|
||
<mods:affiliation id="12FC6B13BC381BC351705EB272088BC4">Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37, E- 08003 Barcelona, Spain;</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A9C0F005BCF36EC0A6535CAC92654A2C">text</mods:typeOfResource>
|
||
<mods:relatedItem id="9F9E500B41D41AE69E0E6AEB9B9CB315" type="host">
|
||
<mods:titleInfo id="05B3A8FD15BE5986DED767F0B0DC4F21">
|
||
<mods:title id="5A979C2A5D72BC56863E07324FA727A8">Acta Entomologica Musei Nationalis Pragae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="42B6E1C9C8C95893022D59A86D2992A5">
|
||
<mods:date id="7449DF5EE5BA8C63CC468DC5C2E4D890">2019</mods:date>
|
||
<mods:detail id="432DD88FE38796E4AE7D347AE4A8DF96" type="pubDate">
|
||
<mods:number id="B8290033C5AD33566B17AB2C3F0B43DE">2019-06-24</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="466CE58E9F6C285DFD88A4B20AEDC8D4" type="volume">
|
||
<mods:number id="0146780EF207443F2CC32DC697ADDBA4">59</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AB742201B1178F9AD2FBF3940B13F362" type="issue">
|
||
<mods:number id="187B4902C17B980411EE052CDB30F781">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="1FE5D92D00DA947DA634C31FD39A7999" unit="page">
|
||
<mods:start id="D1645E4E04BACDB6AF2C32866CD67BAB">253</mods:start>
|
||
<mods:end id="B0279492C98F24898C7E732D99D8B04F">272</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="7AF448837B5CC92CE557AABBF4DA9E84">journal article</mods:classification>
|
||
<mods:identifier id="6DBFD4405AC135FA88C350F66205D712" type="CLB-Dataset">8140</mods:identifier>
|
||
<mods:identifier id="F8D58649828D48DC99DC6B8AD3A375AE" type="DOI">10.2478/aemnp-2019-0021</mods:identifier>
|
||
<mods:identifier id="A0E3C44CFB2DE6A565ACF1F9D8D57C5B" type="GBIF-Dataset">726682f3-9eac-4961-ac4d-d8b8679e8211</mods:identifier>
|
||
<mods:identifier id="1A65CAC51A9AEBFC53F34484050933A6" type="ISSN">1804-6487</mods:identifier>
|
||
<mods:identifier id="BA3A806CA8934F989F395B4B0BA79ECC" type="Zenodo-Dep">4488950</mods:identifier>
|
||
<mods:identifier id="047BF2ADA04D41EAE3D414EBA0B03EDF" type="ZooBank">ABBA2B4F-8B60-41E2-B80B-45861F974B23</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03DD87A0FFD4FFD6FF4EFE6FDDDFFC8B" ID-DOI="http://doi.org/10.5281/zenodo.4549014" ID-GBIF-Taxon="177423647" ID-Zenodo-Dep="4549014" LSID="urn:lsid:plazi:treatment:03DD87A0FFD4FFD6FF4EFE6FDDDFFC8B" httpUri="http://treatment.plazi.org/id/03DD87A0FFD4FFD6FF4EFE6FDDDFFC8B" lastPageNumber="260" pageId="7" pageNumber="260">
|
||
<subSubSection id="C36E653DFFD4FFD6FF4EFE6FDBE5FDED" box="[200,711,508,531]" pageId="7" pageNumber="260" type="nomenclature">
|
||
<paragraph id="8BCB36B6FFD4FFD6FF4EFE6FDBE5FDED" blockId="7.[200,711,508,531]" box="[200,711,508,531]" pageId="7" pageNumber="260">
|
||
<heading id="D08381DAFFD4FFD6FF4EFE6FDBE5FDED" box="[200,711,508,531]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="260" reason="3">
|
||
<emphasis id="B900EAA4FFD4FFD6FF4EFE6FDBE5FDED" bold="true" box="[200,711,508,531]" pageId="7" pageNumber="260">
|
||
<taxonomicName id="4C744D35FFD4FFD6FF4EFE6FDB48FDED" ID-CoL="8XTVG" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[200,618,508,531]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="alhajarensis" status="sp. nov." subGenus="Ochthebius">
|
||
<emphasis id="B900EAA4FFD4FFD6FF4EFE6FD864FDED" bold="true" box="[200,326,508,531]" italics="true" pageId="7" pageNumber="260">Ochthebius</emphasis>
|
||
(
|
||
<emphasis id="B900EAA4FFD4FFD6FED0FE6FD8F6FDED" bold="true" box="[342,468,508,531]" italics="true" pageId="7" pageNumber="260">Ochthebius</emphasis>
|
||
)
|
||
<emphasis id="B900EAA4FFD4FFD6FE65FE6FDB48FDED" bold="true" box="[483,618,508,531]" italics="true" pageId="7" pageNumber="260">alhajarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357DFFFD4FFD6FDF7FE6FDBE5FDED" box="[625,711,508,531]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD4FFD6FE17FD89D8DDFDD3" box="[401,511,538,557]" pageId="7" pageNumber="260" type="description">
|
||
<paragraph id="8BCB36B6FFD4FFD6FE17FD89D8DDFDD3" blockId="7.[401,511,538,557]" box="[401,511,538,557]" pageId="7" pageNumber="260">
|
||
(
|
||
<figureCitation id="134F2A33FFD4FFD6FE11FD89D8FBFDD3" box="[407,473,538,557]" captionStart="Figs 14–17" captionStartId="8.[151,188,2019,2038]" captionTargetBox="[179,1369,223,1980]" captionTargetId="figure-13@8.[151,1436,189,1999]" captionTargetPageId="8" captionText="Figs 14–17. Habitus of new species: 14 – Hydraena naja sp. nov.; 15 – Ochthebius alhajarensis sp. nov.; 16 – Ochthebius bernard sp. nov.; 17 – Agraphydrus elongatus sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488958" httpUri="https://zenodo.org/record/4488958/files/figure.png" pageId="7" pageNumber="260">Figs 15</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A33FFD4FFD6FE64FD89D8DAFDD3" box="[482,504,538,557]" captionStart="Figs 18–21" captionStartId="9.[151,188,2012,2031]" captionTargetBox="[257,1306,228,1943]" captionTargetId="figure-12@9.[224,1364,189,1988]" captionTargetPageId="9" captionText="Figs 18–21. Aedeagus of new species: 18 – Hydraena naja sp. nov.; 19 – Ochthebius alhajarensis sp. nov.; 20 – Ochthebius bernard sp. nov.; 21 – Agraphydrus elongatus sp. nov. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4488962" httpUri="https://zenodo.org/record/4488962/files/figure.png" pageId="7" pageNumber="260">19</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD4FFD6FF11FDD7D859FC9F" pageId="7" pageNumber="260" type="materials_examined">
|
||
<paragraph id="8BCB36B6FFD4FFD6FF11FDD7D838FD8F" blockId="7.[151,761,580,865]" pageId="7" pageNumber="260">
|
||
<emphasis id="B900EAA4FFD4FFD6FF11FDD7D832FDA9" bold="true" box="[151,272,580,599]" pageId="7" pageNumber="260">
|
||
<typeStatus id="54CF8814FFD4FFD6FF11FDD7D9E6FDA9" box="[151,196,580,599]" pageId="7" pageNumber="260">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Source of wadi Bani Awf in Jebel Al-Hajar,
|
||
<collectingCountry id="F3637626FFD4FFD6FD16FDD7DBE6FDA9" box="[656,708,580,599]" name="Oman" pageId="7" pageNumber="260">Oman</collectingCountry>
|
||
(Loc. 4;
|
||
<figureCitation id="134F2A33FFD4FFD6FF2BFDCDD9C0FD8F" box="[173,226,606,625]" captionStart="Fig" captionStartId="1.[1088,1120,2029,2048]" captionTargetBox="[152,1071,1387,2079]" captionTargetId="figure-691@1.[152,1074,1387,2079]" captionTargetPageId="1" captionText="Fig. 1. Sampled localities (see Table 1 for details)." figureDoi="http://doi.org/10.5281/zenodo.4488952" httpUri="https://zenodo.org/record/4488952/files/figure.png" pageId="7" pageNumber="260">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A33FFD4FFD6FF6AFDCDD82DFD8F" box="[236,271,606,625]" captionStart="Figs 2–7" captionStartId="2.[151,188,2009,2028]" captionTargetBox="[151,1436,546,1992]" captionTargetId="figure-13@2.[151,1436,546,1992]" captionTargetPageId="2" captionText="Figs 2–7. Some of the sampled localities in north Oman. 2 – Loc. No. 1: Ghul spring with pools; 3 – No. 2: residual pools in wadi in Bahla city; 4 – Loc. No. 3: residual pools in wadi; 5 – Loc. No. 4: source of wadi Bani Awf; 6 – Loc. No. 4: source of wadi Bani Awf; 7 – Loc. No. 5: wadi Bani Awf. See Table 1 for details on the localities, and Fig. 1 for their geographical location." figureDoi="http://doi.org/10.5281/zenodo.4488954" httpUri="https://zenodo.org/record/4488954/files/figure.png" pageId="7" pageNumber="260">5, 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FF11FDEAD859FC9F" blockId="7.[151,761,580,865]" pageId="7" pageNumber="260">
|
||
<emphasis id="B900EAA4FFD4FFD6FF11FDEAD838FD72" bold="true" box="[151,282,633,652]" pageId="7" pageNumber="260">Type material.</emphasis>
|
||
<materialsCitation id="3B1C3CEBFFD4FFD6FE98FDEAD830FCEF" ID-GBIF-Occurrence="3038225302" collectingDate="2010-04-06" collectionCode="NHMW" collectorName="J. Al-Akhdar & Hernando" country="Oman" elevation="1300" latitude="23.176722" location="wadi Bani Awf" longLatPrecision="1" longitude="57.409473" pageId="7" pageNumber="260" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="54CF8814FFD4FFD6FE98FDEAD853FD74" box="[286,369,633,652]" pageId="7" pageNumber="260" type="holotype">HOLOTYPE</typeStatus>
|
||
: ♁ (
|
||
<collectionCode id="ED65AE73FFD4FFD6FE13FDEAD8FDFD72" box="[405,479,633,652]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="260" type="Museum">NHMW</collectionCode>
|
||
), “4
|
||
<collectingCountry id="F3637626FFD4FFD6FD82FDEADB1AFD72" box="[516,568,633,652]" name="Oman" pageId="7" pageNumber="260">Oman</collectingCountry>
|
||
<date id="FFCA1076FFD4FFD6FDBAFDEADBA5FD72" box="[572,647,633,652]" pageId="7" pageNumber="260" value="2010-04-06">
|
||
<collectingDate id="EF8EE99EFFD4FFD6FDBAFDEADBA5FD72" box="[572,647,633,652]" pageId="7" pageNumber="260" value="2010-04-06">6.4.2010</collectingDate>
|
||
</date>
|
||
<collectorName id="26815360FFD4FFD6FD0DFDEADBD5FD72" box="[651,759,633,652]" pageId="7" pageNumber="260">J.Al-Akhdar</collectorName>
|
||
// source of
|
||
<location id="8EAB606DFFD4FFD6FF7BFD07D85DFD59" LSID="urn:lsid:plazi:treatment:03DD87A0FFD4FFD6FF4EFE6FDDDFFC8B:8EAB606DFFD4FFD6FF7BFD07D85DFD59" box="[253,383,660,679]" country="Oman" latitude="23.176722" longLatPrecision="1" longitude="57.409473" name="wadi Bani Awf" pageId="7" pageNumber="260">wadi Bani Awf</location>
|
||
, on rock //
|
||
<geoCoordinate id="EE405071FFD4FFD6FE59FD07DB6BFD59" box="[479,585,660,679]" degrees="23" direction="north" minutes="10" orientation="latitude" pageId="7" pageNumber="260" precision="1" seconds="36.2" value="23.176722">N23 10 36.2</geoCoordinate>
|
||
<geoCoordinate id="EE405071FFD4FFD6FDC9FD07DB95FD59" box="[591,695,660,679]" degrees="57" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="260" precision="1" seconds="34.1" value="57.409473">E57 24 34.1</geoCoordinate>
|
||
<quantity id="4C8C9B53FFD4FFD6FD3AFD07DBD5FD59" box="[700,759,660,679]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="260" unit="m" value="1300.0">
|
||
<elevation id="0059D185FFD4FFD6FD3AFD07DBD5FD59" box="[700,759,660,679]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="260" unit="m" value="1300.0">1300m</elevation>
|
||
</quantity>
|
||
// Ribera, Cieslak &
|
||
<collectorName id="26815360FFD4FFD6FECEFD3DD8BEFD3F" box="[328,412,686,705]" pageId="7" pageNumber="260">Hernando</collectorName>
|
||
leg.”, used for DNA extraction, voucher number IBE-RA104 (sequences published in
|
||
<bibRefCitation id="EFE54B47FFD4FFD6FDA7FD5ADBCDFD22" author="VILLASTRIGO" box="[545,751,713,732]" etAl="et al." firstAuthor="VILLASTRIGO" pageId="7" pageNumber="260" pagination="273 - 288" refId="ref18145" refString="VILLASTRIGO A., JACH M. A., CARDOSO A., VALLADARES L. F. & RIBERA I. 2019: A molecular phylogeny of the tribe Ochthebiini (Coleoptera, Hydraenidae, Ochthebiinae). Systematic Entomology 44: 273 - 288." type="journal article" year="2019">VILLASTRIGO et al. 2019</bibRefCitation>
|
||
), aedeagus dissected and mounted in DMHF on a transparent card, with
|
||
<typeStatus id="54CF8814FFD4FFD6FF11FD6DD9C0FCEF" box="[151,226,766,785]" pageId="7" pageNumber="260" type="holotype">holotype</typeStatus>
|
||
label
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="54CF8814FFD4FFD6FE9CFD6DD853FCEE" box="[282,369,766,785]" pageId="7" pageNumber="260" type="paratype">PARATYPES</typeStatus>
|
||
(
|
||
<specimenCount id="9D72FD3FFFD4FFD6FEFAFD6DD899FCEF" box="[380,443,766,785]" pageId="7" pageNumber="260" type="generic">4 spec.</specimenCount>
|
||
): (
|
||
<collectionCode id="ED65AE73FFD4FFD6FE48FD6DDB29FCEF" box="[462,523,766,785]" pageId="7" pageNumber="260">CCHB</collectionCode>
|
||
,
|
||
<collectionCode id="ED65AE73FFD4FFD6FD95FD6DDB6AFCEF" box="[531,584,766,785]" pageId="7" pageNumber="260">IBEB</collectionCode>
|
||
):
|
||
<specimenCount id="9D72FD3FFFD4FFD6FDD3FD6DDBA0FCEC" box="[597,642,763,786]" pageId="7" pageNumber="260" type="female">3 ♀♀</specimenCount>
|
||
, same data as
|
||
<typeStatus id="54CF8814FFD4FFD6FF11FC8AD9C6FCD2" box="[151,228,793,812]" pageId="7" pageNumber="260" type="holotype">holotype</typeStatus>
|
||
, with
|
||
<typeStatus id="54CF8814FFD4FFD6FE9FFC8AD840FCD2" box="[281,354,793,812]" pageId="7" pageNumber="260" type="paratype">paratype</typeStatus>
|
||
labels; 1 ♁, “6
|
||
<collectingCountry id="F3637626FFD4FFD6FE6AFC8ADB02FCD2" box="[492,544,793,812]" name="Oman" pageId="7" pageNumber="260">Oman</collectingCountry>
|
||
<date id="FFCA1076FFD4FFD6FDA0FC8ADB53FCD2" box="[550,625,793,812]" pageId="7" pageNumber="260" value="2010-04-06">6.4.2010</date>
|
||
J. Al-Akhdar // wadi Bani Awf residual pools //
|
||
<geoCoordinate id="EE405071FFD4FFD6FE27FCA7DB2AFCB9" box="[417,520,820,839]" degrees="23" direction="north" minutes="13" orientation="latitude" pageId="7" pageNumber="260" precision="1" seconds="42.9" value="23.228584">N23 13 42.9</geoCoordinate>
|
||
<geoCoordinate id="EE405071FFD4FFD6FD8AFCA7DB53FCB9" box="[524,625,820,839]" degrees="57" direction="east" minutes="25" orientation="longitude" pageId="7" pageNumber="260" precision="1" seconds="25.8" value="57.423836">E57 25 25.8</geoCoordinate>
|
||
<quantity id="4C8C9B53FFD4FFD6FDF3FCA7DB84FCB9" box="[629,678,820,839]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="7" pageNumber="260" unit="m" value="660.0">660m</quantity>
|
||
// Ribera, Cieslak & Hernando leg.”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD4FFD6FF11FCE9D871F86F" pageId="7" pageNumber="260" type="description">
|
||
<paragraph id="8BCB36B6FFD4FFD6FF11FCE9D9C1FC0F" blockId="7.[151,761,890,2065]" pageId="7" pageNumber="260">
|
||
<emphasis id="B900EAA4FFD4FFD6FF11FCE9D802FC6F" bold="true" box="[151,288,890,913]" pageId="7" pageNumber="260">Description.</emphasis>
|
||
Habitus as in
|
||
<figureCitation id="134F2A33FFD4FFD6FE29FCE9D8D8FC6F" box="[431,506,890,913]" captionStart="Figs 14–17" captionStartId="8.[151,188,2019,2038]" captionTargetBox="[179,1369,223,1980]" captionTargetId="figure-13@8.[151,1436,189,1999]" captionTargetPageId="8" captionText="Figs 14–17. Habitus of new species: 14 – Hydraena naja sp. nov.; 15 – Ochthebius alhajarensis sp. nov.; 16 – Ochthebius bernard sp. nov.; 17 – Agraphydrus elongatus sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488958" httpUri="https://zenodo.org/record/4488958/files/figure.png" pageId="7" pageNumber="260">Fig. 15</figureCitation>
|
||
. Body length:
|
||
<quantity id="4C8C9B53FFD4FFD6FD08FCE9D9E6FC4F" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.25" metricValueMin="2.15" pageId="7" pageNumber="260" unit="mm" value="2.2" valueMax="2.25" valueMin="2.15">2.15–2.25 mm</quantity>
|
||
; width:
|
||
<quantity id="4C8C9B53FFD4FFD6FEA7FC09D8E6FC4F" box="[289,452,922,945]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="7" pageNumber="260" unit="mm" value="0.925" valueMax="0.95" valueMin="0.9">0.90–0.95 mm</quantity>
|
||
. Dorsal surface black (dark brown in immature specimens); palpi, antennae and legs brown.
|
||
</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FF33FC69DB58FB4F" blockId="7.[151,761,890,2065]" pageId="7" pageNumber="260">Upper surface of head with very long, adpressed fine whitish setae. Labrum only slightly incised, surface between punctures smooth. Frontoclypeal suture distinct, strongly arched. Surface of head largely smooth, with microreticulated areas around eyes and on neck area. With two large foveae on vertex; ocelli not visible.</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FF33FB29DBDBFA2F" blockId="7.[151,761,890,2065]" pageId="7" pageNumber="260">
|
||
Pronotum trapezoidal, almost as wide as long; surface covered with same
|
||
<typeStatus id="54CF8814FFD4FFD6FEDAFB49D8ABFB0F" box="[348,393,1242,1265]" pageId="7" pageNumber="260">type</typeStatus>
|
||
of setae as on head but more sparse; anterior margin straight in middle; anterior angles roughly arcuated; lateral margin irregularly defined. Hyaline membrane narrow at anterior and posterior margins, very wide at posterior corners. Surface of disk smooth, with sparse punctures and some microreticulation on margins; with shallow longitudinal and lateral furrows; sides of pronotum strongly microreticulated, with granulose appearance.
|
||
</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FF33FA49D8A5F9AF" blockId="7.[151,761,890,2065]" pageId="7" pageNumber="260">Elytra oval; with very irregular rows of shallow punctures, with long and thin whitish seta on anterior part of each puncture. Lateral rim smooth, not explanate. Membranous wings well developed.</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FF33F9C9D81DF96F" blockId="7.[151,761,890,2065]" pageId="7" pageNumber="260">Legs relatively long, with rows of strong setae, without natatorial setae.</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FF33F909DBCAF90F" blockId="7.[151,761,890,2065]" pageId="7" pageNumber="260">Ventral surface black, surface shagreened, covered with long, thin, whitish pubescence. Margins of metaventrite and abdominal ventrites with shorter, more dense setae.</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FF33F969D871F86F" blockId="7.[151,761,890,2065]" pageId="7" pageNumber="260">
|
||
Aedeagus (
|
||
<figureCitation id="134F2A33FFD4FFD6FEA9F969D85CF8EF" box="[303,382,1786,1809]" captionStart="Figs 18–21" captionStartId="9.[151,188,2012,2031]" captionTargetBox="[257,1306,228,1943]" captionTargetId="figure-12@9.[224,1364,189,1988]" captionTargetPageId="9" captionText="Figs 18–21. Aedeagus of new species: 18 – Hydraena naja sp. nov.; 19 – Ochthebius alhajarensis sp. nov.; 20 – Ochthebius bernard sp. nov.; 21 – Agraphydrus elongatus sp. nov. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4488962" httpUri="https://zenodo.org/record/4488962/files/figure.png" pageId="7" pageNumber="260">Fig. 19</figureCitation>
|
||
) with main piece strongly bend in almost straight angle; medial part narrow, apex dilated and sinuated. Distal lobe regularly expanded, with lower part hyaline. Parameres inserted near base of main piece, not reaching its apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD4FFD6FF11F809DC6BFD8B" pageId="7" pageNumber="260" type="diagnosis">
|
||
<paragraph id="8BCB36B6FFD4FFD6FF11F809DC6BFD8B" blockId="7.[151,761,890,2065]" lastBlockId="7.[828,1437,190,885]" pageId="7" pageNumber="260">
|
||
<emphasis id="B900EAA4FFD4FFD6FF11F809D8B0F84F" bold="true" box="[151,402,1946,1969]" pageId="7" pageNumber="260">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C744D35FFD4FFD6FE1FF809DBBFF84F" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[409,669,1946,1969]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="alhajarensis" status="sp. nov.">
|
||
<emphasis id="B900EAA4FFD4FFD6FE1FF809DBBFF84F" box="[409,669,1946,1969]" italics="true" pageId="7" pageNumber="260">Ochthebius alhajarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357DFFFD4FFD6FD23F809DBDAF84F" box="[677,760,1946,1969]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the
|
||
<taxonomicName id="4C744D35FFD4FFD6FEB4F829D898F82F" authorityName="Stephens" authorityYear="1829" box="[306,442,1978,2001]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="punctatus">
|
||
<emphasis id="B900EAA4FFD4FFD6FEB4F829D898F82F" box="[306,442,1978,2001]" italics="true" pageId="7" pageNumber="260">O. punctatus</emphasis>
|
||
</taxonomicName>
|
||
group. Among the species for which we have molecular data, it is most closely related to
|
||
<taxonomicName id="4C744D35FFD4FFD6FF33F869D8F2F7EF" authority="Jach, 1989" authorityName="Jach" authorityYear="1989" box="[181,464,2042,2065]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="klapperichi">
|
||
<emphasis id="B900EAA4FFD4FFD6FF33F869D870F7EF" box="[181,338,2042,2065]" italics="true" pageId="7" pageNumber="260">O. klapperichi</emphasis>
|
||
<bibRefCitation id="EFE54B47FFD4FFD6FEDDF869D8F2F7EF" author="Jach" box="[347,464,2042,2065]" firstAuthor="Jach" pageId="7" pageNumber="260" pagination="95 - 126" refId="ref16674" refString="JACH M. A. 1989: Revision of the Palearctic species of the genus Ochthebius Leach 1. The so-called subgenus " Bothochius " (Hydraenidae, Coleoptera). Koleopterologische Rundschau 59: 95 - 126." type="journal article" year="1989">Jäch, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
from central Asia (studied specimen voucher No. MNCN-AI1269;
|
||
<bibRefCitation id="EFE54B47FFD4FFD6FB65FF2DDA51FF0B" author="VILLASTRIGO" etAl="et al." firstAuthor="VILLASTRIGO" pageId="7" pageNumber="260" pagination="273 - 288" refId="ref18145" refString="VILLASTRIGO A., JACH M. A., CARDOSO A., VALLADARES L. F. & RIBERA I. 2019: A molecular phylogeny of the tribe Ochthebiini (Coleoptera, Hydraenidae, Ochthebiinae). Systematic Entomology 44: 273 - 288." type="journal article" year="2019">VILLASTRIGO et al. 2019</bibRefCitation>
|
||
) at a genetic distance of 7% in the COI-3 gene, and both to another unnamed species from
|
||
<collectingCountry id="F3637626FFD4FFD6FB5FFF6DDC60FEEB" box="[1241,1346,254,277]" name="Tajikistan" pageId="7" pageNumber="260">Tajikistan</collectingCountry>
|
||
(unpublished data). They are in turn related to
|
||
<taxonomicName id="4C744D35FFD4FFD6FB4AFE8DDAC6FEAB" authority="Wollaston, 1854" authorityName="Wollaston" authorityYear="1854" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="quadrifoveolatus">
|
||
<emphasis id="B900EAA4FFD4FFD6FB4AFE8DDCBEFECB" box="[1228,1436,286,309]" italics="true" pageId="7" pageNumber="260">O. quadrifoveolatus</emphasis>
|
||
Wollaston, 1854
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D35FFD4FFD6FB95FEADDC38FEAB" authority="Kuwert, 1877" authorityName="Kuwert" authorityYear="1877" box="[1043,1306,318,341]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="ragusae">
|
||
<emphasis id="B900EAA4FFD4FFD6FB95FEADDDA7FEAB" box="[1043,1157,318,341]" italics="true" pageId="7" pageNumber="260">O. ragusae</emphasis>
|
||
Kuwert, 1877
|
||
</taxonomicName>
|
||
, both widely distributed in the Mediterranean region (
|
||
<bibRefCitation id="EFE54B47FFD4FFD6FB79FECDDA57FE6B" author="JACH & SKALE" firstAuthor="JACH" pageId="7" pageNumber="260" pagination="130 - 162" refId="ref16997" refString="JACH M. A. & SKALE A. 2015: Family Hydraenidae. Pp. 130 - 162. In: LOBL I. & LOBL D. (eds): Catalogue of Palaearctic Coleoptera. Volume 2 / 1, Revised and updated edition. Hydrophiloidea-Staphylinoidea. Brill, Leiden, 900 pp." type="book chapter" year="2015">JÄCH & SKALE 2015</bibRefCitation>
|
||
). From its known closest relative,
|
||
<taxonomicName id="4C744D35FFD4FFD6FB52FEEDDC49FE6B" authorityName="Jach" authorityYear="1989" box="[1236,1387,382,405]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="klapperichi">
|
||
<emphasis id="B900EAA4FFD4FFD6FB52FEEDDC49FE6B" box="[1236,1387,382,405]" italics="true" pageId="7" pageNumber="260">O. klapperichi</emphasis>
|
||
</taxonomicName>
|
||
, and from
|
||
<taxonomicName id="4C744D35FFD4FFD6FCF2FE0DDD64FE4B" authorityName="Wollaston" authorityYear="1854" box="[884,1094,414,437]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="quadrifoveolatus">
|
||
<emphasis id="B900EAA4FFD4FFD6FCF2FE0DDAA5FE4B" box="[884,903,414,437]" italics="true" pageId="7" pageNumber="260">O</emphasis>
|
||
.
|
||
<emphasis id="B900EAA4FFD4FFD6FC15FE0DDD64FE4B" box="[915,1094,414,437]" italics="true" pageId="7" pageNumber="260">quadrifoveolatus</emphasis>
|
||
</taxonomicName>
|
||
the new species differs clearly in the shape of the pronotum, with a much narrower base and stronger fossae. The external morphology is most similar to
|
||
<taxonomicName id="4C744D35FFD4FFD6FCD0FE6DDAE9FDEB" authorityName="Kuwert" authorityYear="1877" box="[854,971,510,533]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="ragusae">
|
||
<emphasis id="B900EAA4FFD4FFD6FCD0FE6DDAE9FDEB" box="[854,971,510,533]" italics="true" pageId="7" pageNumber="260">O. ragusae</emphasis>
|
||
</taxonomicName>
|
||
, from which based on the available material apparently only differs in the stronger punctuation of the elytra. The shape of the aedeagus is, however, clearly different to that of all these species (see
|
||
<bibRefCitation id="EFE54B47FFD4FFD6FB4AFDCDDC1FFD8B" author="JACH" box="[1228,1341,606,629]" firstAuthor="JACH" pageId="7" pageNumber="260" pagination="95 - 126" refId="ref16674" refString="JACH M. A. 1989: Revision of the Palearctic species of the genus Ochthebius Leach 1. The so-called subgenus " Bothochius " (Hydraenidae, Coleoptera). Koleopterologische Rundschau 59: 95 - 126." type="journal article" year="1989">JÄCH 1989</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD4FFD6FCBAFDEDDDA3FCCB" pageId="7" pageNumber="260" type="etymology">
|
||
<paragraph id="8BCB36B6FFD4FFD6FCBAFDEDDCBFFD4B" blockId="7.[828,1437,190,885]" pageId="7" pageNumber="260">
|
||
<emphasis id="B900EAA4FFD4FFD6FCBAFDEDDA9EFD6B" bold="true" box="[828,956,638,661]" pageId="7" pageNumber="260">Etymology.</emphasis>
|
||
Named after the mountain range of Al Hajar, in north
|
||
<collectingCountry id="F3637626FFD4FFD6FC14FD0DDAF4FD4B" box="[914,982,670,693]" name="Oman" pageId="7" pageNumber="260">Oman</collectingCountry>
|
||
, were the new species was found; adjective.
|
||
</paragraph>
|
||
<paragraph id="8BCB36B6FFD4FFD6FCBAFD2DDDA3FCCB" blockId="7.[828,1437,190,885]" pageId="7" pageNumber="260">
|
||
<emphasis id="B900EAA4FFD4FFD6FCBAFD2DDD18FD2B" bold="true" box="[828,1082,702,725]" pageId="7" pageNumber="260">Notes on the habitat.</emphasis>
|
||
The species was found in two localities, the source of the wadi Bani Awf (
|
||
<figureCitation id="134F2A33FFD4FFD6FA88FD4DDC4DFD0B" box="[1294,1391,734,757]" captionStart="Figs 2–7" captionStartId="2.[151,188,2009,2028]" captionTargetBox="[151,1436,546,1992]" captionTargetId="figure-13@2.[151,1436,546,1992]" captionTargetPageId="2" captionText="Figs 2–7. Some of the sampled localities in north Oman. 2 – Loc. No. 1: Ghul spring with pools; 3 – No. 2: residual pools in wadi in Bahla city; 4 – Loc. No. 3: residual pools in wadi; 5 – Loc. No. 4: source of wadi Bani Awf; 6 – Loc. No. 4: source of wadi Bani Awf; 7 – Loc. No. 5: wadi Bani Awf. See Table 1 for details on the localities, and Fig. 1 for their geographical location." figureDoi="http://doi.org/10.5281/zenodo.4488954" httpUri="https://zenodo.org/record/4488954/files/figure.png" pageId="7" pageNumber="260">Figs 5, 6</figureCitation>
|
||
; see habitat notes for
|
||
<taxonomicName id="4C744D35FFD4FFD6FC60FD6DDDA0FCEB" authority="Ribera & Hernando & Cieslak, 2019" authorityName="Ribera & Hernando & Cieslak" authorityYear="2019" box="[998,1154,766,789]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="naja" status="sp. nov.">
|
||
<emphasis id="B900EAA4FFD4FFD6FC60FD6DDDA0FCEB" box="[998,1154,766,789]" italics="true" pageId="7" pageNumber="260">Hydraena naja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23357DFFFD4FFD6FB00FD6DDDF5FCEB" box="[1158,1239,766,789]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) and residual pools downstream of the same wadi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E653DFFD4FFD6FCBAFCADDDDFFC8B" pageId="7" pageNumber="260" type="distribution">
|
||
<paragraph id="8BCB36B6FFD4FFD6FCBAFCADDDDFFC8B" blockId="7.[828,1437,190,885]" pageId="7" pageNumber="260">
|
||
<emphasis id="B900EAA4FFD4FFD6FCBAFCADDAECFCAB" bold="true" box="[828,974,830,853]" pageId="7" pageNumber="260">Distribution.</emphasis>
|
||
Only known from two localities in the same stream in the Al-Hajar mountains (
|
||
<figureCitation id="134F2A33FFD4FFD6FB28FCCDDDCFFC8B" box="[1198,1261,862,885]" captionStart="Fig" captionStartId="1.[1088,1120,2029,2048]" captionTargetBox="[152,1071,1387,2079]" captionTargetId="figure-691@1.[152,1074,1387,2079]" captionTargetPageId="1" captionText="Fig. 1. Sampled localities (see Table 1 for details)." figureDoi="http://doi.org/10.5281/zenodo.4488952" httpUri="https://zenodo.org/record/4488952/files/figure.png" pageId="7" pageNumber="260">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |