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<mods:title id="B5ABC3BD6242EB38E2F891234AD73F32">Flightless litter-dwelling Cotasterosoma (Coleoptera: Curculionidae: Cossoninae) found outside of Japan, with mtDNA phylogeography of a new species from Southwest China</mods:title>
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<mods:namePart id="D1D775573852BD99EE55F6B7F27916AB">Grebennikov, Vasily V.</mods:namePart>
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<mods:namePart id="262CBCA5E6FEF5874830DAA99D64822D">Morimoto, Katsura</mods:namePart>
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<treatment id="03D73818FFE7FF8189D485A6FD8339BC" ID-DOI="http://doi.org/10.5281/zenodo.6063484" ID-GBIF-Taxon="124559796" ID-Zenodo-Dep="6063484" LSID="urn:lsid:plazi:treatment:03D73818FFE7FF8189D485A6FD8339BC" httpUri="http://treatment.plazi.org/id/03D73818FFE7FF8189D485A6FD8339BC" lastPageId="2" lastPageNumber="135" pageId="1" pageNumber="134">
<subSubSection id="C364DA85FFE7FF8289D485A6FA973FBB" pageId="1" pageNumber="134" type="nomenclature">
<paragraph id="8BC1890EFFE7FF8289D485A6FE103FD5" blockId="1.[151,1403,1694,1750]" box="[151,508,1694,1720]" pageId="1" pageNumber="134">
<heading id="D0893E62FFE7FF8289D485A6FE103FD5" bold="true" box="[151,508,1694,1720]" fontSize="11" level="1" pageId="1" pageNumber="134" reason="1">
<taxonomicName id="4C7EF28DFFE7FF8289D485A6FE103FD5" ID-CoL="8HGSX" authority="Konishi, 1962" authorityName="Konishi" authorityYear="1962" box="[151,508,1694,1720]" class="Insecta" family="Curculionidae" genus="Cotasterosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B90A551CFFE7FF8289D485A6FE103FD5" bold="true" box="[151,508,1694,1720]" pageId="1" pageNumber="134">
<emphasis id="B90A551CFFE7FF8289D485A6FEA63FD5" bold="true" box="[151,330,1694,1720]" italics="true" pageId="1" pageNumber="134">Cotasterosoma</emphasis>
<bibRefCitation id="EFEFF4FFFFE7FF82881385A6FE103FD5" author="Konishi" box="[336,508,1694,1720]" pageId="1" pageNumber="134" refString="Konishi, M. (1962) Taxonomic studies on the Cossoninae of Japan (Coleoptera: Curculionidae), Part 1. Insecta Matsumurana, 25, 1 - 17." type="journal article" year="1962">Konishi, 1962</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC1890EFFE7FF8289D485F9FA973FBB" blockId="1.[151,1403,1694,1750]" box="[151,1403,1729,1750]" pageId="1" pageNumber="134">
<bibRefCitation id="EFEFF4FFFFE7FF8289D485F9FEF13FBB" author="Konishi" box="[151,285,1729,1750]" pageId="1" pageNumber="134" refString="Konishi, M. (1962) Taxonomic studies on the Cossoninae of Japan (Coleoptera: Curculionidae), Part 1. Insecta Matsumurana, 25, 1 - 17." type="journal article" year="1962">Konishi 1962</bibRefCitation>
: 3 (species included:
<taxonomicName id="4C7EF28DFFE7FF8288A985F9FDB73FBB" box="[490,603,1729,1750]" class="Insecta" family="Curculionidae" genus="Cotasterosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="omogoensis">
<emphasis id="B90A551CFFE7FF8288A985F9FDB73FBB" box="[490,603,1729,1750]" italics="true" pageId="1" pageNumber="134">omogoensis</emphasis>
</taxonomicName>
). Type species:
<taxonomicName id="4C7EF28DFFE7FF828BB585F9FB613FBB" authority="Konishi, 1962" authorityName="Konishi" authorityYear="1962" box="[758,1165,1729,1750]" class="Insecta" family="Curculionidae" genus="Cotasterosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="omogoensis">
<emphasis id="B90A551CFFE7FF828BB585F9FC113FBB" box="[758,1021,1729,1750]" italics="true" pageId="1" pageNumber="134">Cotasterosoma omogoensis</emphasis>
<bibRefCitation id="EFEFF4FFFFE7FF828D4085F9FB613FBB" author="Konishi" box="[1027,1165,1729,1750]" pageId="1" pageNumber="134" refString="Konishi, M. (1962) Taxonomic studies on the Cossoninae of Japan (Coleoptera: Curculionidae), Part 1. Insecta Matsumurana, 25, 1 - 17." type="journal article" year="1962">Konishi, 1962</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C364DA85FFE7FF8189D48439FD8339BC" lastPageId="2" lastPageNumber="135" pageId="1" pageNumber="134" type="diagnosis">
<paragraph id="8BC1890EFFE7FF8289D48439FA703E85" blockId="1.[151,1436,1792,2024]" pageId="1" pageNumber="134">
<emphasis id="B90A551CFFE7FF8289D48439FEE03E75" bold="true" box="[151,268,1793,1816]" pageId="1" pageNumber="134">Diagnosis.</emphasis>
Species of the genus
<taxonomicName id="4C7EF28DFFE7FF8288BE8438FD723E7A" box="[509,670,1792,1815]" class="Insecta" family="Curculionidae" genus="Cotasterosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="B90A551CFFE7FF8288BE8438FD723E7A" box="[509,670,1792,1815]" italics="true" pageId="1" pageNumber="134">Cotasterosoma</emphasis>
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are recognizable among all weevils by the unique combination of the following nine non-unique adult characters: (1.) small-sized relatively glabrous weevils of characteristic “edaphic
<taxonomicName id="4C7EF28DFFE7FF8289D4847EFEF83E30" box="[151,276,1862,1885]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="134" phylum="Arthropoda" rank="subFamily" subFamily="Cossoninae">Cossoninae</taxonomicName>
” habitus (
<figureCitation id="1345958BFFE7FF8288CE847EFE3E3E30" box="[397,466,1862,1885]" captionStart="FIGURE 1" captionStartId="1.[151,244,1571,1593]" captionTargetBox="[155,1428,506,1541]" captionTargetId="figure@1.[151,1436,506,1550]" captionTargetPageId="1" captionText="FIGURE 1. Cotasterosoma coronus sp. n., holotype, male. A D: habitus; E: genitalia and abdominal tergites 7 and 8, ventral; F H: genitalia (F: dorsal, G: ventral, H: lateral)." httpUri="https://zenodo.org/record/267440/files/figure.png" pageId="1" pageNumber="134">Figs 1</figureCitation>
AD, 2AD); (2.) antennae consisting of scape, funicle with six (
<figureCitation id="1345958BFFE7FF828D92847EFAFA3E30" box="[1233,1302,1862,1885]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="1" pageNumber="134">Fig. 2</figureCitation>
F) or seven antennomeres (the type species, but see Discussion) and seemingly one-segmented club (
<figureCitation id="1345958BFFE7FF828D328451FB583EED" box="[1137,1204,1897,1920]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="1" pageNumber="134">Fig. 2</figureCitation>
F); (3.) all tarsi fivesegmented (
<figureCitation id="1345958BFFE7FF82885484B4FEB03ECE" box="[279,348,1932,1955]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="1" pageNumber="134">Figs 2</figureCitation>
GI); (4.) all tarsomeres subcylindrical in shape (
<figureCitation id="1345958BFFE7FF828A2F84B4FC5D3ECE" box="[876,945,1932,1955]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="1" pageNumber="134">Figs 2</figureCitation>
GI); (5.) fore-legs with unci arising from the middle of apical margin of tibia (
<figureCitation id="1345958BFFE7FF8288B48496FDDB3EA8" box="[503,567,1966,1989]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="1" pageNumber="134">Fig. 2</figureCitation>
G), and not from the dorso-apical corner; (6.) tarsomere 3 not enlarged (
<figureCitation id="1345958BFFE7FF828C748496FA973EA8" box="[1335,1403,1966,1989]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="1" pageNumber="134">Figs 2</figureCitation>
G I); (7.) tarsomere 4 short and inserting at apical surface of tarsomere 3, with a few setae on ventral side (
<figureCitation id="1345958BFFE7FF828DB184E9FADA3E85" box="[1266,1334,2001,2024]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="1" pageNumber="134">Figs 2</figureCitation>
GI); (8.)
</paragraph>
<paragraph id="8BC1890EFFE4FF8189D483A0FD8339BC" blockId="2.[151,1435,152,209]" pageId="2" pageNumber="135">
male genitalia with U- or Y-shaped tegmen broadly abbreviated dorsally (
<figureCitation id="1345958BFFE4FF818AE283A0FC3339C2" box="[929,991,152,175]" captionStart="FIGURE 1" captionStartId="1.[151,244,1571,1593]" captionTargetBox="[155,1428,506,1541]" captionTargetId="figure@1.[151,1436,506,1550]" captionTargetPageId="1" captionText="FIGURE 1. Cotasterosoma coronus sp. n., holotype, male. A D: habitus; E: genitalia and abdominal tergites 7 and 8, ventral; F H: genitalia (F: dorsal, G: ventral, H: lateral)." httpUri="https://zenodo.org/record/267440/files/figure.png" pageId="2" pageNumber="135">Figs 1</figureCitation>
EH); (9.) collum of spermatheca bent at apex and almost parallel to ramus (
<figureCitation id="1345958BFFE4FF818B4F8382FDA039BC" box="[524,588,186,209]" captionStart="FIGURE 2" captionStartId="2.[151,244,1438,1460]" captionTargetBox="[161,1423,253,1410]" captionTargetId="figure@2.[151,1436,251,1417]" captionTargetPageId="2" captionText="FIGURE 2. Cotasterosoma coronus sp. n., paratype, female, specimen # 1110 from Mt. Emei. A D: habitus; E F: head and prothorax (E: ventral, F: lateral); G I: distal part of right legs (G: fore-, H: mid-, I: hind-); J: genitalia, ventral; K: spermatheca." httpUri="https://zenodo.org/record/267441/files/figure.png" pageId="2" pageNumber="135">Fig. 2</figureCitation>
K).
</paragraph>
</subSubSection>
</treatment>
</document>