treatments-xml/data/03/D6/12/03D6126CFF83FFC1FF00FF59FE3894C9.xml

<document id="813C80285E452DB0045921467FDC62E8" ID-DOI="10.5281/zenodo.175139" ID-GBIF-Dataset="df43e153-62dc-4880-8ef8-da196a30f873" ID-ISSN="1175­5326" ID-Zenodo-Dep="175139" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459801013595" checkinUser="plazi" docAuthor="Niogret, Jerome, Nicot, Antoine, Stordeur, Eric De &amp; Bertrand, Michel" docDate="2007" docId="03D6126CFF83FFC1FF00FF59FE3894C9" docLanguage="en" docName="zt01386p029.pdf" docOrigin="Zootaxa 1386" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Macrocheles seraphim Niogret &amp; Nicot, sp. nov." docType="treatment" docVersion="10" lastPageNumber="24" masterDocId="FFEF6A14FF80FFC4FF97FFCEFFD3965B" masterDocTitle="Combination of morphological characters and ITS­sequence to characterize a new species of Macrocheles (Acari: Macrochelidae)" masterLastPageNumber="29" masterPageNumber="19" pageNumber="22" updateTime="1698219919517" updateUser="plazi">
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<mods:title id="930C25F6AFD6E8F930CFF589DE7AA1E6">Combination of morphological characters and ITS­sequence to characterize a new species of Macrocheles (Acari: Macrochelidae)</mods:title>
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<mods:namePart id="079B176036E947183C17A953297F6F18">Niogret, Jerome</mods:namePart>
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<mods:namePart id="C04C92ED8659432458D5AFE3EF76B0F3">Nicot, Antoine</mods:namePart>
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<mods:namePart id="F3E367F330A5F0216B861A31B4867338">Stordeur, Eric De</mods:namePart>
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<mods:namePart id="C439375E965AA95AC614E09A9458F535">Bertrand, Michel</mods:namePart>
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<treatment id="03D6126CFF83FFC1FF00FF59FE3894C9" ID-DOI="http://doi.org/10.5281/zenodo.5630203" ID-GBIF-Taxon="119351719" ID-Zenodo-Dep="5630203" LSID="urn:lsid:plazi:treatment:03D6126CFF83FFC1FF00FF59FE3894C9" httpUri="http://treatment.plazi.org/id/03D6126CFF83FFC1FF00FF59FE3894C9" lastPageId="5" lastPageNumber="24" pageId="3" pageNumber="22">
<subSubSection id="C365F0F1FF83FFC7FF00FF59FD3796EA" box="[151,740,151,177]" pageId="3" pageNumber="22" type="nomenclature">
<paragraph id="8BC0A37AFF83FFC7FF00FF59FD3796EA" blockId="3.[151,740,151,177]" box="[151,740,151,177]" pageId="3" pageNumber="22">
<heading id="D0881416FF83FFC7FF00FF59FD3796EA" bold="true" box="[151,740,151,177]" fontSize="11" level="1" pageId="3" pageNumber="22" reason="1">
<emphasis id="B90B7F68FF83FFC7FF00FF59FD3796EA" bold="true" box="[151,740,151,177]" pageId="3" pageNumber="22">
<taxonomicName id="4C7FD8F9FF83FFC7FF00FF59FDA996EA" authority="Niogret &amp; Nicot" authorityName="Niogret &amp; Nicot" box="[151,634,151,177]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="seraphim" status="sp. nov.">
<emphasis id="B90B7F68FF83FFC7FF00FF59FE7996EA" bold="true" box="[151,426,151,177]" italics="true" pageId="3" pageNumber="22">Macrocheles seraphim</emphasis>
Niogret &amp; Nicot
</taxonomicName>
, 
<taxonomicNameLabel id="A238C213FF83FFC7FD10FF59FD3796EA" box="[647,740,151,177]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365F0F1FF83FFC7FF00FF2CFBD2979F" pageId="3" pageNumber="22" type="materials_examined">
<paragraph id="8BC0A37AFF83FFC7FF00FF2CFBD2979F" blockId="3.[151,1437,226,2012]" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF00FF2CFE4296A7" bold="true" box="[151,401,226,252]" pageId="3" pageNumber="22">Material examined.</emphasis>
<typeStatus id="54C41DD8FF83FFC7FE09FF2CFDD896A7" box="[414,523,226,252]" pageId="3" pageNumber="22" type="holotype">Holotype</typeStatus>
female, Trucy l’Orgueilleux, 
<collectingCountry id="F368E3EAFF83FFC7FCE8FF2CFC0196A7" box="[895,978,226,252]" name="France" pageId="3" pageNumber="22">France</collectingCountry>
, 
<geoCoordinate id="EE4BC5BDFF83FFC7FC73FF2CFB9496A7" box="[996,1095,226,252]" direction="north" orientation="latitude" pageId="3" pageNumber="22" precision="925" value="47.75">47°45'N</geoCoordinate>
, 
<geoCoordinate id="EE4BC5BDFF83FFC7FBCEFF2CFB7996A7" box="[1113,1194,226,252]" direction="east" orientation="longitude" pageId="3" pageNumber="22" precision="925" value="3.6833334">3°41'E</geoCoordinate>
, clay soil, pastured meadow, continental climate, elevation 
<quantity id="4C870E9FFF83FFC7FDFFFEC4FD66977F" box="[616,693,266,292]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="22" unit="m" value="160.0">160 m</quantity>
, 
<date id="FFC185BAFF83FFC7FD56FEC4FC96977F" box="[705,837,266,292]" pageId="3" pageNumber="22" value="2006-04">April 2006</date>
, on 
<taxonomicName id="4C7FD8F9FF83FFC7FCE1FEC4FB819778" box="[886,1106,266,291]" class="Insecta" family="Geotrupidae" genus="Geotrupes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="mutator">
<emphasis id="B90B7F68FF83FFC7FCE1FEC4FB819778" box="[886,1106,266,291]" italics="true" pageId="3" pageNumber="22">Geotrupes mutator</emphasis>
</taxonomicName>
. 
<typeStatus id="54C41DD8FF83FFC7FBF6FEC4FB05977F" box="[1121,1238,266,292]" pageId="3" pageNumber="22" type="paratype">Paratypes</typeStatus>
, 
<specimenCount id="9D7968F3FF83FFC7FB75FEC4FA80977F" box="[1250,1363,266,292]" pageId="3" pageNumber="22" type="female">6 females</specimenCount>
and 
<specimenCount id="9D7968F3FF83FFC7FA1BFEC4FF009717" pageId="3" pageNumber="22" type="male">1 male</specimenCount>
, same data as 
<typeStatus id="54C41DD8FF83FFC7FE13FEFCFE3D9717" box="[388,494,306,332]" pageId="3" pageNumber="22" type="holotype">holotype</typeStatus>
; deposited in collection of the Muséum National d’Histoire Naturelle de Paris (
<collectingCountry id="F368E3EAFF83FFC7FF08FE94FF21972F" box="[159,242,346,372]" name="France" pageId="3" pageNumber="22">France</collectingCountry>
) (
<collectionCode id="ED6E3BBFFF83FFC7FE9DFE94FEB5972F" LSID="urn:lsid:biocol.org:col:34988" box="[266,358,346,372]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="22">MNHN</collectionCode>
Ac1105, Ac1106, Ac 
<quantity id="4C870E9FFF83FFC7FDF0FE94FD6C972F" box="[615,703,346,372]" metricMagnitude="1" metricUnit="m" metricValue="2.8117799999999997" pageId="3" pageNumber="22" unit="in" value="1107.0">1107 in</quantity>
respect); 5 
<typeStatus id="54C41DD8FF83FFC7FCDDFE94FC7C972F" box="[842,943,346,372]" pageId="3" pageNumber="22" type="paratype">paratype</typeStatus>
females in the Institut royal des Sciences naturelles de 
<collectingCountry id="F368E3EAFF83FFC7FEAFFE4CFE7297C7" box="[312,417,386,412]" name="Belgium" pageId="3" pageNumber="22">Belgique</collectingCountry>
(Bruxelles, 
<collectingCountry id="F368E3EAFF83FFC7FDA1FE4CFD4E97C7" box="[566,669,386,412]" name="Belgium" pageId="3" pageNumber="22">Belgium</collectingCountry>
) (No 30.653). Other 
<typeStatus id="54C41DD8FF83FFC7FC0AFE4CFBD197C7" box="[925,1026,386,412]" pageId="3" pageNumber="22" type="paratype">paratype</typeStatus>
specimens (15 individuals) are in the collections of the Laboratoire de Zoogéographie, Montpellier (
<collectingCountry id="F368E3EAFF83FFC7FC09FE64FC27979F" box="[926,1012,426,452]" name="France" pageId="3" pageNumber="22">France</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C365F0F1FF83FFC1FF51FE1CFF1F9441" lastPageId="5" lastPageNumber="24" pageId="3" pageNumber="22" type="description">
<paragraph id="8BC0A37AFF83FFC7FF51FE1CFCD297B7" blockId="3.[151,1437,226,2012]" box="[198,769,466,492]" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF51FE1CFEFA97B7" bold="true" box="[198,297,466,492]" pageId="3" pageNumber="22">Female.</emphasis>
Yellowish­brown species, medium size.
</paragraph>
<paragraph id="8BC0A37AFF83FFC7FF51FE34FE729577" blockId="3.[151,1437,226,2012]" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF51FE34FEBA9448" box="[198,361,506,531]" italics="true" pageId="3" pageNumber="22">Dorsal shield</emphasis>
(figure 1): similar in shape to 
<taxonomicName id="4C7FD8F9FF83FFC7FD7DFE34FCB39448" box="[746,864,506,531]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="B90B7F68FF83FFC7FD7DFE34FCB39448" box="[746,864,506,531]" italics="true" pageId="3" pageNumber="22">M. glaber</emphasis>
</taxonomicName>
, longer than wide with distinct ornamentation, length 805.6 ± 6.5 µm, width 439.7 ± 6.9 µm between setae 
<emphasis id="B90B7F68FF83FFC7FCCFFDECFCA19460" box="[856,882,546,571]" italics="true" pageId="3" pageNumber="22">r3</emphasis>
(n = 10 individuals), clearly demarcated by the procurved line, reticular pattern underlined by punctures around the margins of rather large but irregular pentagonal or hexagonal cells, with irregular separation lines. Shield with 28 pairs of setae, with 
<emphasis id="B90B7F68FF83FFC7FB7DFDBCFAD294D0" box="[1258,1281,626,651]" italics="true" pageId="3" pageNumber="22">j1</emphasis>
directed forward, pilose in the distal half (figure 3a), 
<emphasis id="B90B7F68FF83FFC7FDE8FD54FD4A94E8" box="[639,665,666,691]" italics="true" pageId="3" pageNumber="22">z1</emphasis>
smooth, reduced in length, 
<emphasis id="B90B7F68FF83FFC7FC77FD54FC2894E8" box="[992,1019,666,691]" italics="true" pageId="3" pageNumber="22">z6</emphasis>
longer than 
<emphasis id="B90B7F68FF83FFC7FB19FD54FB7694E8" box="[1166,1189,666,691]" italics="true" pageId="3" pageNumber="22">j6</emphasis>
. Except for the short and bi­serrate setae 
<emphasis id="B90B7F68FF83FFC7FE15FD0CFE4E9480" box="[386,413,706,731]" italics="true" pageId="3" pageNumber="22">J5</emphasis>
, the median setae are simple (
<emphasis id="B90B7F68FF83FFC7FC92FD0CFC139480" box="[773,960,706,731]" italics="true" pageId="3" pageNumber="22">j5, j6, z5, z6, J2</emphasis>
), others are distally irregular (
<emphasis id="B90B7F68FF83FFC7FABEFD0CFAB89480" box="[1321,1387,706,731]" italics="true" pageId="3" pageNumber="22">i.e. j2</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FAECFD0CFA479480" box="[1403,1428,706,731]" italics="true" pageId="3" pageNumber="22">s5</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FF00FD24FF669558" box="[151,181,746,771]" italics="true" pageId="3" pageNumber="22">Z1</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FF53FD24FF3E9558" box="[196,237,746,771]" italics="true" pageId="3" pageNumber="22">Z2)</emphasis>
(figure 3b) or pilose (
<emphasis id="B90B7F68FF83FFC7FE78FD24FDE09558" box="[495,563,746,771]" italics="true" pageId="3" pageNumber="22">i.e. s2</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FDD5FD24FD8F9558" box="[578,604,746,771]" italics="true" pageId="3" pageNumber="22">r2</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FDFCFD24FD549558" box="[619,647,746,771]" italics="true" pageId="3" pageNumber="22">S5</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FD01FD24FD659558" box="[662,694,746,771]" italics="true" pageId="3" pageNumber="22">Z4</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FD53FD24FD309558" box="[708,739,746,771]" italics="true" pageId="3" pageNumber="22">Z5</emphasis>
,) (figure 3a) and distally pilose (
<emphasis id="B90B7F68FF83FFC7FBF3FD24FED89570" italics="true" pageId="3" pageNumber="22">i.e. j3, j4, z2, z4, s4, s6, r4, S1, S2, S4</emphasis>
) (figure 3c).
</paragraph>
<paragraph id="8BC0A37AFF83FFC7FF51FCF4FD459307" blockId="3.[151,1437,226,2012]" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF51FCF4FEAA9508" box="[198,377,826,851]" italics="true" pageId="3" pageNumber="22">Ventral shields</emphasis>
(figure 2): Tritosternum and peritremes normal for genus. Sternal shield rather large (length 189.5 ± 2.4 µm, width at level of coxae II: 156.2 ± 3.1 µm) (n = 8 individuals) with the usual 3 pairs of smooth setae and 2 pairs of pores. Median transverse line procurved, discrete and marked by a row of small punctures, the two oblique anterior lines joined anterioly, angular and arched lines partially hidden in the anterior half by the reticular ornamentation; punctuate zone anterior to coxae II extending from oblique anterior line. Posterior part of sternal shield with irregular punctate pattern, mostly large medially, extended backward behind the midlevel of coxae III. Metasternal plates close to the sternal shield, not extended behind the posterior edge of coxae III. Epigynial shield with a pair of long setae, broader than long (110–112 µm long, 145– 166 µm maximum width). Epigynial shield with sclerotization attenuated anteriorly, sclerotized lines forming arched pattern, marked by punctures, and a median zone with sub­polygonal pattern (figure 2). Ventri­anal shield as wide as long (ca. 260 µm long), with 6 main transverse procurved lines, longitudinal ornamentation masked, with the polygons marked by punctures and arranged as irregularly defined overlapping tiles, with 3 pairs of pre­anal setae and three para­anal setae; cribrum present. Metapodal sclerites elongate. The postcoxal pore is visible, included in the podal shield.
</paragraph>
<paragraph id="8BC0A37AFF83FFC7FF51FAA5FD5D907F" blockId="3.[151,1437,226,2012]" pageId="3" pageNumber="22">
<taxonomicName id="4C7FD8F9FF83FFC7FF51FAA5FE8F93DF" box="[198,348,1387,1412]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="3" pageNumber="22" phylum="Nematoda" rank="genus">
<emphasis id="B90B7F68FF83FFC7FF51FAA5FE8F93DF" box="[198,348,1387,1412]" italics="true" pageId="3" pageNumber="22">Gnathosoma</emphasis>
</taxonomicName>
: similar to that of 
<taxonomicName id="4C7FD8F9FF83FFC7FDA5FAA5FD7693DF" box="[562,677,1387,1412]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="B90B7F68FF83FFC7FDA5FAA5FD7693DF" box="[562,677,1387,1412]" italics="true" pageId="3" pageNumber="22">M. glaber</emphasis>
</taxonomicName>
group species. Chelicerae robust (figure 4). Fixed digit with well developed median tooth pyramidal in shape and a terminal hook, 
<emphasis id="B90B7F68FF83FFC7FC39FA5DFB9593F7" box="[942,1094,1427,1452]" italics="true" pageId="3" pageNumber="22">pilus dentilis</emphasis>
straight, movable digit with four teeth. Hypostomal groove with five transverse rows of denticles. Arthrodial brush reaching the median level of movable digit. Palpi as in 
<taxonomicName id="4C7FD8F9FF83FFC7FDADFA2DFD7C93A7" box="[570,687,1507,1532]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="B90B7F68FF83FFC7FDADFA2DFD7C93A7" box="[570,687,1507,1532]" italics="true" pageId="3" pageNumber="22">M. glaber</emphasis>
</taxonomicName>
group species. Epistome (figure 10) 
<taxonomicName id="4C7FD8F9FF83FFC7FBE7FA2DFB6F93A7" box="[1136,1212,1507,1532]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="B90B7F68FF83FFC7FBE7FA2DFB6F93A7" box="[1136,1212,1507,1532]" italics="true" pageId="3" pageNumber="22">glaber</emphasis>
</taxonomicName>
­like in shape with smooth lateral lobes, central branch pilose.
</paragraph>
<paragraph id="8BC0A37AFF83FFC7FF51F9FDFAB2902F" blockId="3.[151,1437,226,2012]" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF51F9FDFE739017" box="[198,416,1587,1612]" italics="true" pageId="3" pageNumber="22">Sacculus foeminus</emphasis>
(figure 5): sacculus wide, elongate (45 µm width), tubuli opening into spherical lateral vesicles that join a thickened median section which feeds into the sperm duct, corniculi long (ca 50 µm).
</paragraph>
<paragraph id="8BC0A37AFF83FFC7FF51F94DFB20909F" blockId="3.[151,1437,226,2012]" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF51F94DFF2E90C7" box="[198,253,1667,1692]" italics="true" pageId="3" pageNumber="22">Legs</emphasis>
: chaetotaxy normal for genus, setae simple or lightly pilose distally, except for the tibial posteroventral and distal basitarsal setae on leg IV, and a ventral seta on genu I. Tarsal setae thin and long.
</paragraph>
<paragraph id="8BC0A37AFF83FFC7FF51F91CFD129167" blockId="3.[151,1437,226,2012]" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF51F91CFEDE90B7" bold="true" box="[198,269,1746,1772]" pageId="3" pageNumber="22">Male.</emphasis>
<emphasis id="B90B7F68FF83FFC7FE8FF91DFE6990B7" box="[280,442,1747,1772]" italics="true" pageId="3" pageNumber="22">Dorsal shield</emphasis>
(figure 6): elongate (485 µm long, 285 µm wide) with rough ornamental pattern, reticulation weakened laterally, irregularly sclerotized laterally, with 28 pairs of setae, 
<emphasis id="B90B7F68FF83FFC7FB3CF935FB12914F" box="[1195,1217,1787,1812]" italics="true" pageId="3" pageNumber="22">j1</emphasis>
long and smooth, distally close together; 
<emphasis id="B90B7F68FF83FFC7FE30F8EDFE129167" box="[423,449,1827,1852]" italics="true" pageId="3" pageNumber="22">r2</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FE47F8EDFE389167" box="[464,491,1827,1852]" italics="true" pageId="3" pageNumber="22">z4</emphasis>
, 
<emphasis id="B90B7F68FF83FFC7FE6EF8EDFDCB9167" box="[505,536,1827,1852]" italics="true" pageId="3" pageNumber="22">Z5</emphasis>
and 
<emphasis id="B90B7F68FF83FFC7FDC6F8EDFDBE9167" box="[593,621,1827,1852]" italics="true" pageId="3" pageNumber="22">J5</emphasis>
pilose.
</paragraph>
<paragraph id="8BC0A37AFF83FFC0FF51F885FBCD9681" blockId="3.[151,1437,226,2012]" lastBlockId="4.[151,1437,152,538]" lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22">
<emphasis id="B90B7F68FF83FFC7FF51F885FEA7913F" box="[198,372,1867,1892]" italics="true" pageId="3" pageNumber="22">Ventral shields</emphasis>
: sterno­genital, ventri­anal, endopodal, metapodal and metasternal shields all fused into a holoventral shield; sterno­genital shield with 5 pairs of setae and 3 pairs of pores; length of the shield 395 µm, width at level of coxae II: 90 µm (n = 1). Ventri­anal shield longer than wide, with 3 pairs of pre­anal setae, a pair of para­anal setae and a post­anal seta, all setae simple. Surface ornamentation lacking, only weakly punctuate, stronger at lateral margins. Cribrum present, bordering posterior end of shield. Metapodal sclerites long, with characteristic shape. Pore visible on podal sclerite behind coxa IV.
</paragraph>
<paragraph id="8BC0A37AFF84FFC0FF51FF26FB6F9721" blockId="4.[151,1437,152,538]" pageId="4" pageNumber="23">
<taxonomicName id="4C7FD8F9FF84FFC0FF51FF26FE8F975A" box="[198,348,232,257]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="23" phylum="Nematoda" rank="genus">
<emphasis id="B90B7F68FF84FFC0FF51FF26FE8F975A" box="[198,348,232,257]" italics="true" pageId="4" pageNumber="23">Gnathosoma</emphasis>
</taxonomicName>
well sclerotized. Epistome as in female, 
<taxonomicName id="4C7FD8F9FF84FFC0FCC5FF26FC1A975A" box="[850,969,232,257]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="glaber">
<emphasis id="B90B7F68FF84FFC0FCC5FF26FC1A975A" box="[850,969,232,257]" italics="true" pageId="4" pageNumber="23">M. glaber</emphasis>
</taxonomicName>
­like. Chelicerae (figure 7) with robust digits, weakly dentate, 
<emphasis id="B90B7F68FF84FFC0FE3BFEDEFD969772" box="[428,581,272,297]" italics="true" pageId="4" pageNumber="23">pilus dentilis</emphasis>
straight; dorsal seta conspicuous; movable digit with three small teeth, arthrodial brush reaching only 1/3 length of the movable digit; spermatodactyl with large basal bulb and distally blunt, elbowed in the proximal third, strongly recurved backward, distally attenuated.
</paragraph>
<paragraph id="8BC0A37AFF84FFC0FF51FE46FEE59441" blockId="4.[151,1437,152,538]" pageId="4" pageNumber="23">
<emphasis id="B90B7F68FF84FFC0FF51FE46FF2E97FA" box="[198,253,392,417]" italics="true" pageId="4" pageNumber="23">Legs</emphasis>
: tarsus I without ambulacrum and claw; tarsi II–IV with developed ambulacra and claws; legs with a large spur on femur II bearing strong simple anterior seta, genu II with setigerous tubercle. Femur IV with a double spur bearing a pilose seta and a small, non setose tubercle. Dorsal and ventral setae pilose or simple (figures 8–9).
</paragraph>
<caption id="DF00F3F2FF84FFC0FF00F988FF69909D" httpUri="https://zenodo.org/record/175140/files/figure.png" pageId="4" pageNumber="23" targetBox="[168,1432,601,1568]" targetPageId="4">
<paragraph id="8BC0A37AFF84FFC0FF00F988FF69909D" blockId="4.[151,1437,1606,1734]" pageId="4" pageNumber="23">
<emphasis id="B90B7F68FF84FFC0FF00F988FEEE9005" bold="true" box="[151,317,1606,1630]" pageId="4" pageNumber="23">FIGURES 1–7</emphasis>
. 
<taxonomicName id="4C7FD8F9FF84FFC0FEDBF989FD939005" box="[332,576,1607,1630]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="seraphim" status="sp. nov.">
<emphasis id="B90B7F68FF84FFC0FEDBF989FD939005" box="[332,576,1607,1630]" italics="true" pageId="4" pageNumber="23">Macrocheles seraphim</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238C213FF84FFC0FDDEF989FD729005" box="[585,673,1607,1630]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
: 1. female, dorsal shield; 2. female, ventral view; 3a. female, detail of pilose dorsal setae. 3b. female, detail of distally pilose dorsal setae. 3c. female, detail of distally irregular dorsal setae. 4. female, distal end of chelicera. 5. female, sacculus foeminus. 6. male, dorsal shield. 7. male, chelicera and spermatodactyl.
</paragraph>
</caption>
<paragraph id="8BC0A37AFF84FFC1FF51F8DAFA9A9681" blockId="4.[151,1437,1812,2038]" lastBlockId="5.[151,1437,152,658]" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23">
<emphasis id="B90B7F68FF84FFC0FF51F8DAFEB09175" bold="true" box="[198,355,1812,1838]" pageId="4" pageNumber="23">Protonymph</emphasis>
(figures 11 &amp; 13). Length of idiosoma: 413 µm; width at level of 
<emphasis id="B90B7F68FF84FFC0FBFAF8DAFB549176" box="[1133,1159,1812,1837]" italics="true" pageId="4" pageNumber="23">r3</emphasis>
: 207 µm (n = 1). Anterior shield with distally pilose setae 
<emphasis id="B90B7F68FF84FFC0FDC6F8F2FDB8910E" box="[593,619,1852,1877]" italics="true" pageId="4" pageNumber="23">z4</emphasis>
, 
<emphasis id="B90B7F68FF84FFC0FDEAF8F2FD47910E" box="[637,660,1852,1877]" italics="true" pageId="4" pageNumber="23">j6</emphasis>
, 
<emphasis id="B90B7F68FF84FFC0FD31F8F2FD13910E" box="[678,704,1852,1877]" italics="true" pageId="4" pageNumber="23">s5</emphasis>
, 
<emphasis id="B90B7F68FF84FFC0FD45F8F2FD3B910E" box="[722,744,1852,1877]" italics="true" pageId="4" pageNumber="23">j4</emphasis>
, other setae simple. Opisthonotal shield reduced, weakly delimited and with distally pilose setae 
<emphasis id="B90B7F68FF84FFC0FDF1F8AAFD579126" box="[614,644,1892,1917]" italics="true" pageId="4" pageNumber="23">S4</emphasis>
, 
<emphasis id="B90B7F68FF84FFC0FD05F8AAFD7C9126" box="[658,687,1892,1917]" italics="true" pageId="4" pageNumber="23">S5</emphasis>
, 
<emphasis id="B90B7F68FF84FFC0FD29F8AAFD0E9126" box="[702,733,1892,1917]" italics="true" pageId="4" pageNumber="23">Z4</emphasis>
; concave posterior part of the shield delimited anteriorly by a curved line posterior to insertions of 
<emphasis id="B90B7F68FF84FFC0FDCDF842FDAD91FE" box="[602,638,1932,1957]" italics="true" pageId="4" pageNumber="23">J4.</emphasis>
Sterno­ventral shield rather soft, poorly sclerotized, with three pairs of setae and two pairs of pores. Anal shield distinct, with opisthogastric setae. 
<taxonomicName id="4C7FD8F9FF84FFC0FB13F87AFACB9195" box="[1156,1304,1972,1998]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="23" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
similar to deutonymph, but less developed. Palpal setation deficient, with a crown of only eight terminal setae, but the larger sensory setae of later stages, i.e. one large, hollow and curved, and the bent and hollow seta, are present. Tritosternal base short, with relatively long laciniae, epistome similar to the adult. Genu I with 6 setae.
</paragraph>
<paragraph id="8BC0A37AFF85FFC1FF51FF29FF1F9441" blockId="5.[151,1437,152,658]" pageId="5" pageNumber="24">
<emphasis id="B90B7F68FF85FFC1FF51FF29FEBA975A" bold="true" box="[198,361,231,257]" pageId="5" pageNumber="24">Deutonymph</emphasis>
(figure 12). Length of idiosoma: 546.5 ± 16.3 µm; width at level of setae 
<emphasis id="B90B7F68FF85FFC1FB44FF26FB3F975A" box="[1235,1260,232,257]" italics="true" pageId="5" pageNumber="24">r3</emphasis>
: 283 ± 8.5 µm (n = 2). Dorsal shield with 28 pairs of setae, weakly ornamented and incompletely fused, with opisthonotal portion marked posteriorly by depressed zone, punctate, delimited anteriorly by an arched line that isolates setae 
<emphasis id="B90B7F68FF85FFC1FF4EFEAEFF2E9722" box="[217,253,352,377]" italics="true" pageId="5" pageNumber="24">J5.</emphasis>
Dorsal setae pilose (i.e. 
<emphasis id="B90B7F68FF85FFC1FD88FEAEFDE59722" box="[543,566,352,377]" italics="true" pageId="5" pageNumber="24">j1</emphasis>
, 
<emphasis id="B90B7F68FF85FFC1FDD3FEAEFD889722" box="[580,603,352,377]" italics="true" pageId="5" pageNumber="24">j4</emphasis>
, 
<emphasis id="B90B7F68FF85FFC1FDFDFEAEFD579722" box="[618,644,352,377]" italics="true" pageId="5" pageNumber="24">z4</emphasis>
, 
<emphasis id="B90B7F68FF85FFC1FD04FEAEFD7E9722" box="[659,685,352,377]" italics="true" pageId="5" pageNumber="24">r3</emphasis>
, 
<emphasis id="B90B7F68FF85FFC1FD2BFEAEFD089722" box="[700,731,352,377]" italics="true" pageId="5" pageNumber="24">Z3</emphasis>
, 
<emphasis id="B90B7F68FF85FFC1FD7DFEAEFCFD9722" box="[746,814,352,377]" italics="true" pageId="5" pageNumber="24">Z4 S4</emphasis>
, 
<emphasis id="B90B7F68FF85FFC1FCAAFEAEFC899722" box="[829,858,352,377]" italics="true" pageId="5" pageNumber="24">S5</emphasis>
), 
<emphasis id="B90B7F68FF85FFC1FCE4FEAEFC5C9722" box="[883,911,352,377]" italics="true" pageId="5" pageNumber="24">J5</emphasis>
short and serrate, other dorsal setae smooth; vertical setae 
<emphasis id="B90B7F68FF85FFC1FEADFE46FE8297FA" box="[314,337,392,417]" italics="true" pageId="5" pageNumber="24">j1</emphasis>
widely separated. Tritosternum well developed, with a pair of pilose laciniae. Sterno­ventral shield with 4 pairs of simple setae and 3 pairs of pores. Four pairs of simple setae located between sterno­ventral and anal shields; anal shield longer than wide; with a pair of simple para­anal setae and a shorter post­anal seta.
</paragraph>
</subSubSection>
<subSubSection id="C365F0F1FF85FFC1FF51FDE9FE3894C9" pageId="5" pageNumber="24" type="etymology">
<paragraph id="8BC0A37AFF85FFC1FF51FDE9FE3894C9" blockId="5.[151,1437,152,658]" pageId="5" pageNumber="24">
<emphasis id="B90B7F68FF85FFC1FF51FDE9FE80941A" bold="true" box="[198,339,551,577]" pageId="5" pageNumber="24">Etymology.</emphasis>
<taxonomicName id="4C7FD8F9FF85FFC1FECBFDE6FE22941A" box="[348,497,552,577]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="seraphim">
<emphasis id="B90B7F68FF85FFC1FECBFDE6FE22941A" box="[348,497,552,577]" italics="true" pageId="5" pageNumber="24">M. seraphim</emphasis>
</taxonomicName>
is dedicated to Mr Seraphin Dhotel. 
<taxonomicName id="4C7FD8F9FF85FFC1FC3BFDE6FBCE941A" box="[940,1053,552,577]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="28" phylum="Arthropoda" rank="species" species="seraphim">
<emphasis id="B90B7F68FF85FFC1FC3BFDE6FBCE941A" box="[940,1053,552,577]" italics="true" pageId="5" pageNumber="24">Seraphim</emphasis>
</taxonomicName>
(Latin etym., from Hebrew) are Biblical creatures similar to angels but they are able to hide and to conceal their presence (Dead Sea Scrolls, &quot;Michael's words&quot; (4Q529)).
</paragraph>
</subSubSection>
</treatment>
</document>