337 lines
37 KiB
XML
337 lines
37 KiB
XML
<document id="1E1B3CC9283C7603688D0896A8C9B91C" ID-DOI="10.5281/zenodo.5160708" ID-ISSN="1942-1354" ID-Zenodo-Dep="5160708" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628109944637" checkinUser="felipe" docAuthor="Warner, William B." docDate="2011" docId="03D587FC172DE02EFF66124C3166897D" docLanguage="en" docName="InsectaMundi.0172.1-17.pdf" docOrigin="Insecta Mundi 2011 (172)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Acoma howdenorum Warner 2011, new species" docType="treatment" docVersion="4" lastPageNumber="4" masterDocId="FFECFF841729E02BFFA6106F32548D66" masterDocTitle="Four new species of Acoma Casey, with a key to species in the genus (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="3" updateTime="1699034678431" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="AF0B7D8934FD5B5F3749876218A9FC98" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="45BAE3EB1AF6B59F55D74FDBE586AEFE">
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<mods:title id="57FCC1C642C7D00C6307A18DF323078C">Four new species of Acoma Casey, with a key to species in the genus (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A76DC1FF77EEE18307D11D05733D0D4B" type="personal">
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<mods:role id="054E27D38E16B567502BE6689249505D">
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<mods:roleTerm id="A07C5A61D48B8F1507FC4B07CAADB63B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B6C0E19DEAE37D3EE261000FB8EC3AB5">Warner, William B.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="CA4446A24DACF3E98EB54A761B45D7A6">text</mods:typeOfResource>
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<mods:titleInfo id="DD22D6787A38D980A7128FDA86B17B6B">
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<mods:title id="3DA3E4E87B9D1A5439392C01E0D7A0D7">Insecta Mundi</mods:title>
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</mods:titleInfo>
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||
<mods:part id="5BD61C7D8D8E2F07CC1CF616C5A928CE">
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||
<mods:date id="DBE93851E28923F1A243A184B97230DD">2011</mods:date>
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||
<mods:detail id="1C1BE20D3C51D0E68279D63FE04FD3D8" type="pubDate">
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||
<mods:number id="4E5F9F4599C00610EDC8AA29FC340BE1">2011-04-22</mods:number>
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||
</mods:detail>
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||
<mods:detail id="616F934CA0807F02F38CF5C90730A3B7" type="volume">
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||
<mods:number id="836317BF04B21D27C44528AF73B47773">2011</mods:number>
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||
</mods:detail>
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||
<mods:detail id="7F13744D99E53AD0CF08AC2AD9C6592D" type="issue">
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<mods:number id="54D1CC7A09FE007B08CF5539FD684A39">172</mods:number>
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</mods:detail>
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<mods:extent id="363CB05F67D68626F7F8E0D0A7A05DF2" unit="page">
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<mods:start id="7C3C9BAFF4C432B641B1F7D5C0EF8739">1</mods:start>
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<mods:end id="457A147B79503BD2C237C2827912C62B">17</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="2AC6097A82C460BDAC03A4F40D6EA7AF">journal article</mods:classification>
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<mods:identifier id="DE5285DBB05EFAF8C565592AEC7CCCDB" type="DOI">10.5281/zenodo.5160708</mods:identifier>
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<mods:identifier id="99B6CEDDFC5147D29F2E3CBC8A063C5E" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="F60F85B095DA5E288A4752F93528945A" type="Zenodo-Dep">5160708</mods:identifier>
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</mods:mods>
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<treatment id="03D587FC172DE02EFF66124C3166897D" ID-DOI="http://doi.org/10.5281/zenodo.5164481" ID-Zenodo-Dep="5164481" LSID="urn:lsid:plazi:treatment:03D587FC172DE02EFF66124C3166897D" httpUri="http://treatment.plazi.org/id/03D587FC172DE02EFF66124C3166897D" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
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<subSubSection id="C3666561172DE02FFF66124C302E8F5D" box="[192,634,546,571]" pageId="4" pageNumber="3" type="nomenclature">
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<paragraph id="8BC336EA172DE02FFF66124C302E8F5D" blockId="4.[192,634,546,603]" box="[192,634,546,571]" pageId="4" pageNumber="3">
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<heading id="D08B8186172DE02FFF66124C302E8F5D" bold="true" box="[192,634,546,571]" fontSize="10" level="2" pageId="4" pageNumber="3" reason="3">
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<emphasis id="B908EAF8172DE02FFF66124C302E8F5D" bold="true" box="[192,634,546,571]" pageId="4" pageNumber="3">
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||
<taxonomicName id="4C7C4D69172DE02FFF66124C33878F5D" authority="Warner, 2011" authorityName="Warner" authorityYear="2011" box="[192,467,546,571]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="howdenorum" status="sp. nov.">
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||
<emphasis id="B908EAF8172DE02FFF66124C33878F5D" bold="true" box="[192,467,546,571]" italics="true" pageId="4" pageNumber="3">Acoma howdenorum</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A23B5783172DE02FFE7E124C302E8F5D" box="[472,634,547,571]" pageId="4" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3666561172DE02FFF66122C33048F3D" box="[192,336,579,603]" pageId="4" pageNumber="3" type="description">
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||
<paragraph id="8BC336EA172DE02FFF66122C33048F3D" blockId="4.[192,634,546,603]" box="[192,336,579,603]" pageId="4" pageNumber="3">
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<figureCitation id="13472A6F172DE02FFF66122C33048F3D" box="[192,336,579,603]" captionStart="Figure 6-9" captionStartId="12.[192,276,1864,1886]" captionTargetBox="[246,1383,192,1841]" captionTargetId="figure-19@12.[246,1383,192,1841]" captionTargetPageId="12" captionText="Figure 6-9. Acoma spp. 6-7) Acoma howdenorum Warner: 6) Dorsal habitus. 7) Antenna. 8-9) Acoma glabrata Cazier (specimen from Algodones Dunes): 8) Dorsal habitus. 9) Antenna." figureDoi="http://doi.org/10.5281/zenodo.5160714" httpUri="https://zenodo.org/record/5160714/files/figure.png" pageId="4" pageNumber="3">Figures 6, 7</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3666561172DE02FFF6612EC31E98E1D" pageId="4" pageNumber="3" type="materials_examined">
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||
<paragraph id="8BC336EA172DE02FFF6612EC33A78FDD" blockId="4.[192,1440,643,891]" pageId="4" pageNumber="3">
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||
<emphasis id="B908EAF8172DE02FFF6612EC33D88FFD" bold="true" box="[192,396,643,667]" pageId="4" pageNumber="3">Type Material.</emphasis>
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||
<typeStatus id="54C78848172DE02FFE3512EC30578FFD" box="[403,515,643,667]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED6DAE2F172DE02FFDF012EC30F18FFD" box="[598,677,643,667]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
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||
): “Ariz: Yuma Co., Mohawk Dunes, ca.
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<quantity id="4C849B0F172DE02FFB2E12EC36928FFD" box="[1160,1222,643,667]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="4" pageNumber="3" unit="mi" value="1.0">1 mi.</quantity>
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||
S. I-8;
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<date id="FFC2102A172DE02FFAB812EC37CF8FFD" box="[1310,1435,643,667]" pageId="4" pageNumber="3" value="1988-05-14">v-14-1988</date>
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; U.V. light; W.B.Warner.”
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||
</paragraph>
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||
<paragraph id="8BC336EA172DE02FFF5612AB31D88E5D" blockId="4.[192,1440,643,891]" pageId="4" pageNumber="3">
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||
<typeStatus id="54C78848172DE02FFF5612AB33258FBA" box="[240,369,708,732]" pageId="4" pageNumber="3" type="paratype">Paratypes</typeStatus>
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||
, (
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<specimenCount id="9D7AFD63172DE02FFE2112AC30538FBD" box="[391,519,707,731]" pageId="4" pageNumber="3" type="male">193 males</specimenCount>
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) with data: same as
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<typeStatus id="54C78848172DE02FFCAB12AC31238FBD" box="[781,887,707,731]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
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||
(33);
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<materialsCitation id="3B143CB7172DE02FFC1912AC317F8F9D" collectingDate="1992-07-23" collectorName="H. & A. Howden" country="United States of America" county="Yuma Co." location="Tacna" municipality="Mohawk Dunes" pageId="4" pageNumber="3" specimenCount="1" typeStatus="paratype">
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“
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<collectingCountry id="F36B767A172DE02FFC6C12AC365C8FBD" box="[970,1032,707,731]" name="United States of America" pageId="4" pageNumber="3">USA</collectingCountry>
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. AZ:
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<collectingCounty id="62A24E66172DE02FFBE112AB36918FBD" box="[1095,1221,707,731]" pageId="4" pageNumber="3">Yuma Co.</collectingCounty>
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||
,
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||
<quantity id="4C849B0F172DE02FFB7412AC37758FBD" box="[1234,1313,707,731]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="3" unit="km" value="10.0">10 km</quantity>
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||
E.
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<location id="8EA36031172DE02FFAEA12AB37CF8FBD" LSID="urn:lsid:plazi:treatment:03D587FC172DE02EFF66124C3166897D:8EA36031172DE02FFAEA12AB37CF8FBD" box="[1356,1435,708,731]" country="United States of America" county="Yuma Co." municipality="Mohawk Dunes" name="Tacna" pageId="4" pageNumber="3">Tacna</location>
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||
,
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<collectingMunicipality id="6BA7AC90172DE02FFF66128C33D28F9D" box="[192,390,739,763]" pageId="4" pageNumber="3">Mohawk Dunes</collectingMunicipality>
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||
,
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||
<date id="FFC2102A172DE02FFE34128C30498F9D" box="[402,541,739,763]" pageId="4" pageNumber="3" value="1992-07-23">
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<collectingDate id="EF86E9C2172DE02FFE34128C30498F9D" box="[402,541,739,763]" pageId="4" pageNumber="3" value="1992-07-23">23 vii.1992</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="2689533C172DE02FFD8F128B30A18F9D" box="[553,757,739,763]" pageId="4" pageNumber="3">H. & A. Howden</collectorName>
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||
” (6)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B143CB7172DE02FFC91128B30138E7D" collectorName="W. Warner" country="United States of America" latitude="32.683334" location="Tacna" pageId="4" pageNumber="3" specimenCount="1" typeStatus="paratype">
|
||
same except: “
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||
<quantity id="4C849B0F172DE02FFC4F128C36628F9D" box="[1001,1078,739,763]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="4" pageNumber="3" unit="mi" value="10.0">10 mi.</quantity>
|
||
E of
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||
<location id="8EA36031172DE02FFBD1128B36928F9D" LSID="urn:lsid:plazi:treatment:03D587FC172DE02EFF66124C3166897D:8EA36031172DE02FFBD1128B36928F9D" box="[1143,1222,740,763]" country="United States of America" latitude="32.683334" name="Tacna" pageId="4" pageNumber="3">Tacna</location>
|
||
;
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||
<geoCoordinate id="EE48502D172DE02FFB74128C37678F9D" box="[1234,1331,737,763]" degrees="32" direction="north" minutes="41" orientation="latitude" pageId="4" pageNumber="3" precision="925" value="32.683334">
|
||
32
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||
<superScript id="7C099BA2172DE02FFB56128E36AC8F89" attach="none" box="[1264,1272,737,751]" fontSize="6" pageId="4" pageNumber="3">o</superScript>
|
||
41’N
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||
</geoCoordinate>
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||
, 113
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||
<superScript id="7C099BA2172DE02FFACB128E37218F89" attach="none" box="[1389,1397,737,751]" fontSize="6" pageId="4" pageNumber="3">o</superScript>
|
||
47’; vii.23.1992;UV,
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||
<collectorName id="2689533C172DE02FFE23136C30518E7D" box="[389,517,771,795]" pageId="4" pageNumber="3">W.Warner</collectorName>
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||
” (11)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B143CB7172DE02FFDF4136B36EE8E7D" box="[594,1210,769,795]" collectingDate="2010-06-04" country="United States of America" latitude="32.695835" pageId="4" pageNumber="3" specimenCount="1" typeStatus="paratype">
|
||
same except: “
|
||
<geoCoordinate id="EE48502D172DE02FFD58136C31D38E7D" box="[766,903,769,795]" degrees="32" direction="north" minutes="41" orientation="latitude" pageId="4" pageNumber="3" precision="15" seconds="45" value="32.695835">
|
||
32
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||
<superScript id="7C099BA2172DE02FFCBA136E31708E69" attach="right" box="[796,804,769,783]" fontSize="6" pageId="4" pageNumber="3">o</superScript>
|
||
41’45”N
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||
</geoCoordinate>
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||
, 113
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||
<superScript id="7C099BA2172DE02FFC18136E31928E69" attach="none" box="[958,966,769,783]" fontSize="6" pageId="4" pageNumber="3">o</superScript>
|
||
47’22”,
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||
<date id="FFC2102A172DE02FFBB8136B36EE8E7D" box="[1054,1210,771,795]" pageId="4" pageNumber="3" value="2010-06-04">
|
||
<collectingDate id="EF86E9C2172DE02FFBB8136B36EE8E7D" box="[1054,1210,771,795]" pageId="4" pageNumber="3" value="2010-06-04">June 4, 2010</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
; UV;
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||
<materialsCitation id="3B143CB7172DE02FFB5D136C31D88E5D" collectingDate="2010-06-04" collectorName="W. B. Warner & All" country="United States of America" latitude="32.695835" pageId="4" pageNumber="3" specimenCount="1" typeStatus="paratype">
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<collectorName id="2689533C172DE02FFB5D136C37C18E7D" box="[1275,1429,771,795]" pageId="4" pageNumber="3">W.B.Warner</collectorName>
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||
” (143). (
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||
<collectorName id="2689533C172DE02FFEB5134C336C8E5D" box="[275,312,803,827]" pageId="4" pageNumber="3">All</collectorName>
|
||
<typeStatus id="54C78848172DE02FFE9B134B33FF8E5A" box="[317,427,804,828]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
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||
locality data refer to the same location.)
|
||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BC336EA172DE02FFF56132B31E98E1D" blockId="4.[192,1440,643,891]" pageId="4" pageNumber="3">
|
||
<typeStatus id="54C78848172DE02FFF56132B333E8E3A" box="[240,362,836,860]" pageId="4" pageNumber="3" type="paratype">Paratypes</typeStatus>
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||
are deposited at
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||
<collectionCode id="ED6DAE2F172DE02FFD88132C302F8E3D" box="[558,635,835,859]" country="USA" httpUri="http://grbio.org/cool/jxy6-3sw7" name="Frank M. Hasbrouck Insect Collection" pageId="4" pageNumber="3">ASUT</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFD22132C30878E3D" box="[644,723,835,859]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="4" pageNumber="3">BCRC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFD7A132C317A8E3D" box="[732,814,835,859]" pageId="4" pageNumber="3">BDGC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFC90132C31D68E3D" box="[822,898,835,859]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="4" pageNumber="3" type="Museum">CSCA</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFC2D132C318B8E3D" box="[907,991,835,859]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="4" pageNumber="3" type="University or college">EMEC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFC4E132C36678E3D" box="[1000,1075,835,859]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFB9A132C36C78E3D" box="[1084,1171,835,859]" pageId="4" pageNumber="3">CMNC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFB3A132C36BE8E3D" box="[1180,1258,835,859]" pageId="4" pageNumber="3">MJPC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFB55132C37118E3D" box="[1267,1349,835,859]" country="USA" httpUri="http://grbio.org/cool/ktis-ep89" name="Northern Arizona University" pageId="4" pageNumber="3" type="University or college">NAUF</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFAE8132C37CF8E3D" box="[1358,1435,835,859]" pageId="4" pageNumber="3">PHSC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFF66130C33458E1D" box="[192,273,867,891]" pageId="4" pageNumber="3">RACC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFEBD130B33258E1D" box="[283,369,868,891]" pageId="4" pageNumber="3">RLWE</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFEDD130C33928E1D" box="[379,454,867,891]" country="0" httpUri="http://grbio.org/cool/tizb-nd41" name="University of Alabama, Ichthyological Collection" pageId="4" pageNumber="3">UAIC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFE76130C30778E1D" box="[464,547,867,891]" country="USA" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="4" pageNumber="3" type="University or college">UCRC</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFD8B130C30DD8E1D" box="[557,649,867,891]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="4" pageNumber="3">UNAM</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFD35130C30B88E1D" box="[659,748,867,891]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="4" pageNumber="3">UNSM</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFD50130C311B8E1D" box="[758,847,867,891]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="3" type="Museum">USNM</collectionCode>
|
||
,
|
||
<collectionCode id="ED6DAE2F172DE02FFCFF130C31EC8E1D" box="[857,952,867,891]" pageId="4" pageNumber="3">WBWC</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3666561172DE02FFF6613CC31C28E9D" pageId="4" pageNumber="3" type="diagnosis">
|
||
<paragraph id="8BC336EA172DE02FFF6613CC31C28E9D" blockId="4.[192,1440,931,1019]" pageId="4" pageNumber="3">
|
||
<emphasis id="B908EAF8172DE02FFF6613CC33048EDD" bold="true" box="[192,336,931,955]" pageId="4" pageNumber="3">Diagnosis.</emphasis>
|
||
Pale testaceous, pronotum and elytra subtransparent (when fresh) and glabrous except for marginal fimbrae and scattered long hairs at elytral base; clypeal disc nearly impunctate; antenna 9- segmented, with 3-segmented club; anterior tibia tridentate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3666561172DE02FFF66144C36148BBA" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="8BC336EA172DE02FFF66144C3714895D" blockId="4.[192,1441,1059,1627]" box="[192,1344,1059,1083]" pageId="4" pageNumber="3">
|
||
<emphasis id="B908EAF8172DE02FFF66144C3338895D" bold="true" box="[192,364,1059,1083]" pageId="4" pageNumber="3">Description.</emphasis>
|
||
<typeStatus id="54C78848172DE02FFED4144C33B6895D" box="[370,482,1059,1083]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
|
||
male, length:
|
||
<quantity id="4C849B0F172DE02FFD29144C30B9895D" box="[655,749,1059,1083]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="3" unit="mm" value="5.7">5.7 mm</quantity>
|
||
, width:
|
||
<quantity id="4C849B0F172DE02FFCEA144C31FE895D" box="[844,938,1059,1083]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="3" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, widest at about middle of elytra.
|
||
</paragraph>
|
||
<paragraph id="8BC336EA172DE02FFF56142C370A8B3D" blockId="4.[192,1441,1059,1627]" pageId="4" pageNumber="3">BODY light testaceous, integument translucent to subtransparent, shiny on head, slightly sericeous on pronotum and elytra. HEAD with front and clypeal base scabrously, reticulately punctate; clypeus subhemihexagonal, margins moderately reflexed, more strongly so apically, disc sparsely, shallowly, finely punctate, punctures separated by about 2 to 6 times their own diameters; antenna 9-segmented, club 3-segmented, segment immediately proximal to club anteriorly prolonged into short lamina. PRONOTUM glabrous except for marginal bead, length about 0.4 times elytral length, widest at about middle; anterior margin weakly bisinuate; posterior margin convex, lateral margins subparallel in about basal half, in apical half straight and converging to slightly obtuse anterior angles which are not drawn forward next to eyes; posterior angles rounded; disk weakly, but moderately punctate, punctures mostly separated by 2 to 4 times their own widths, longitudinal midline very slightly impressed and impunctate. SCUTELLUM lobiform, with apex evenly arcuate. ELYTRON glabrous, weakly striate, striae obsolescent basally, weakly contiguously punctate on disc; intervals more coarsely and distinctly punctate than pronotal disc, but punctures similarly separated. PYGIDIUM with apex broadly arcuate, disc shallowly subcontiguously punctate, punctures setigerous. LEGS: Anterior tibia narrow, gracile, tridentate, basal tooth weak; metatibia rather narrow, widest corbel diameter distinctly less than half pygidial length, posterior tarsal claws about 1/3 length of segment from which they arise. GENITALIA: paramera only weakly and arcuately deflexed in apical third, ventrobasal emargination shallow, moderately long.</paragraph>
|
||
<paragraph id="8BC336EA172DE02FFF6616EC36148BBA" blockId="4.[192,1440,1667,1756]" pageId="4" pageNumber="3">
|
||
<emphasis id="B908EAF8172DE02FFF6616EC33068BFD" bold="true" box="[192,338,1667,1691]" pageId="4" pageNumber="3">Variation.</emphasis>
|
||
Length: 4.2 – 6.0 mm, width:
|
||
<quantity id="4C849B0F172DE02FFD5016EC31F98BFD" box="[758,941,1667,1691]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.9" metricValueMin="1.8" pageId="4" pageNumber="3" unit="mm" value="2.35" valueMax="2.9" valueMin="1.8">1.8 – 2.9 mm</quantity>
|
||
. Clypeal shape varies occasionally to subsemicircular, and occasional
|
||
<typeStatus id="54C78848172DE02FFD9E16CB30FA8BDA" box="[568,686,1700,1724]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
have the apical clypeal margin vaguely concave. A few
|
||
<typeStatus id="54C78848172DE02FFA8C16CB37F48BDA" box="[1322,1440,1700,1724]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
had one antenna 8-segmented because of fusion in the funicular segments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3666561172DE02FFF66176C37108A5D" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8BC336EA172DE02FFF66176C37108A5D" blockId="4.[192,1440,1795,1852]" pageId="4" pageNumber="3">
|
||
<emphasis id="B908EAF8172DE02FFF66176C33098A7D" bold="true" box="[192,349,1795,1819]" pageId="4" pageNumber="3">Etymology.</emphasis>
|
||
I take pleasure in naming this species for Henry and Anne Howden, both in recognition of their many contributions to coleopterology and for their help in collecting part of the
|
||
<typeStatus id="54C78848172DE02FFB1C174B36BB8A5A" box="[1210,1263,1828,1852]" pageId="4" pageNumber="3">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3666561172CE02EFF6610AC3166897D" pageId="5" pageNumber="4" type="discussion">
|
||
<paragraph id="8BC336EA172CE02EFF6610AC336C8F9D" blockId="5.[192,1441,195,1051]" pageId="5" pageNumber="4">
|
||
<emphasis id="B908EAF8172CE02EFF6610AC33108DBD" bold="true" box="[192,324,195,219]" pageId="5" pageNumber="4">Remarks.</emphasis>
|
||
This species was treated as “variation” within
|
||
<taxonomicName id="4C7C4D69172CE02EFCDD10AB36C08DBD" authority="Cazier" authorityName="Cazier" authorityYear="1953" box="[891,1172,195,219]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFCDD10AB366A8DBD" box="[891,1086,195,219]" italics="true" pageId="5" pageNumber="4">Acoma glabrata</emphasis>
|
||
Cazier
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFED4B1B172CE02EFB1B10AC37378DBD" author="Cazier, M. A." box="[1213,1379,195,219]" pageId="5" pageNumber="4" pagination="1 - 13" refId="ref7057" refString="Cazier, M. A. 1953. A review of the scarab genus Acoma (Coleoptera: Scarabaeidae). American Museum Novitates, No. 1624: 1 - 13." type="book chapter" year="1953">Cazier (1953)</bibRefCitation>
|
||
, who listed
|
||
<specimenCount id="9D7AFD63172CE02EFEAE108C33EF8D9D" box="[264,443,227,251]" pageId="5" pageNumber="4" type="generic">four specimens</specimenCount>
|
||
from Welton and Dome,
|
||
<collectingRegion id="49B8F808172CE02EFD79108C31158D9D" box="[735,833,227,251]" country="United States of America" name="Arizona" pageId="5" pageNumber="4">Arizona</collectingRegion>
|
||
; but he purposely excluded them from the
|
||
<typeStatus id="54C78848172CE02EFA92108B37F48D9A" box="[1332,1440,228,252]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
|
||
series.
|
||
<bibRefCitation id="EFED4B1B172CE02EFEBA116C33888C7D" author="Howden, H. F." box="[284,476,259,283]" pageId="5" pageNumber="4" pagination="377 - 401" refId="ref7090" refString="Howden, H. F. 1958. Species of Acoma Casey having a three-segmented antennal club (Coleoptera: Scarabaeidae). Canadian Entomologist 90 (7): 377 - 401." type="journal article" year="1958">Howden (1958)</bibRefCitation>
|
||
followed that lead, quoting
|
||
<bibRefCitation id="EFED4B1B172CE02EFCED116C365D8C7D" author="Cazier, M. A." box="[843,1033,259,283]" pageId="5" pageNumber="4" pagination="1 - 13" refId="ref7057" refString="Cazier, M. A. 1953. A review of the scarab genus Acoma (Coleoptera: Scarabaeidae). American Museum Novitates, No. 1624: 1 - 13." type="book chapter" year="1953">Cazier’s (1953)</bibRefCitation>
|
||
comments on variation.
|
||
<taxonomicName id="4C7C4D69172CE02EFAEA116B330B8C5D" authorityName="Warner" authorityYear="2011" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="howdenorum">
|
||
<emphasis id="B908EAF8172CE02EFAEA116B330B8C5D" italics="true" pageId="5" pageNumber="4">Acoma howdenorum</emphasis>
|
||
</taxonomicName>
|
||
is indeed similar to
|
||
<taxonomicName id="4C7C4D69172CE02EFDE9114B30828C5D" authorityName="Cazier" authorityYear="1953" box="[591,726,291,315]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFDE9114B30828C5D" box="[591,726,291,315]" italics="true" pageId="5" pageNumber="4">A. glabrata</emphasis>
|
||
</taxonomicName>
|
||
; however,
|
||
<taxonomicName id="4C7C4D69172CE02EFCF0114B36418C5D" authorityName="Warner" authorityYear="2011" box="[854,1045,291,315]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="howdenorum">
|
||
<emphasis id="B908EAF8172CE02EFCF0114B36418C5D" box="[854,1045,291,315]" italics="true" pageId="5" pageNumber="4">A. howdenorum</emphasis>
|
||
</taxonomicName>
|
||
is separated both geographically and phenotypically from that species. It can immediately be differentiated from
|
||
<taxonomicName id="4C7C4D69172CE02EFB01112B37608C3D" authorityName="Cazier" authorityYear="1953" box="[1191,1332,323,347]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFB01112B37608C3D" box="[1191,1332,323,347]" italics="true" pageId="5" pageNumber="4">A. glabrata</emphasis>
|
||
</taxonomicName>
|
||
by its 9- segmented antennae, generally smaller size, less robust tarsi, more gracile tarsi and anterior tibia, differently shaped pronotum and clypeus, more rounded scutellar apex, and thinner integument.
|
||
<taxonomicName id="4C7C4D69172CE02EFAEB11EB337E8CDD" authorityName="Cazier" authorityYear="1953" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFAEB11EB337E8CDD" italics="true" pageId="5" pageNumber="4">Acoma glabrata</emphasis>
|
||
</taxonomicName>
|
||
has 10-segmented antennae (
|
||
<figureCitation id="13472A6F172CE02EFD3011CC30AD8CDD" box="[662,761,419,443]" captionStart="Figure 6-9" captionStartId="12.[192,276,1864,1886]" captionTargetBox="[246,1383,192,1841]" captionTargetId="figure-19@12.[246,1383,192,1841]" captionTargetPageId="12" captionText="Figure 6-9. Acoma spp. 6-7) Acoma howdenorum Warner: 6) Dorsal habitus. 7) Antenna. 8-9) Acoma glabrata Cazier (specimen from Algodones Dunes): 8) Dorsal habitus. 9) Antenna." figureDoi="http://doi.org/10.5281/zenodo.5160714" httpUri="https://zenodo.org/record/5160714/files/figure.png" pageId="5" pageNumber="4">Fig. 8, 9</figureCitation>
|
||
), although occasional specimens may have one antenna with one or more segments partially to completely fused. In all specimens examined (more than 100), however, at least one of the antennae is 10-segmented.
|
||
<taxonomicName id="4C7C4D69172CE02EFCFB118B36748C9D" authorityName="Cazier" authorityYear="1953" box="[861,1056,483,507]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFCFB118B36748C9D" box="[861,1056,483,507]" italics="true" pageId="5" pageNumber="4">Acoma glabrata</emphasis>
|
||
</taxonomicName>
|
||
ranges from the type locality at San Felipe, northward into the Algodones Dunes in SE
|
||
<collectingRegion id="49B8F808172CE02EFCCB126C31BB8F7D" box="[877,1007,515,539]" country="United States of America" name="California" pageId="5" pageNumber="4">California</collectingRegion>
|
||
. It also occurs in dunes of the Gran Desierto de Altar in northwest
|
||
<collectingRegion id="49B8F808172CE02EFD9B124C30C38F5D" box="[573,663,547,571]" country="Mexico" name="Sonora" pageId="5" pageNumber="4">Sonora</collectingRegion>
|
||
, with its northern terminus in the Yuma Desert in extreme SW
|
||
<collectingRegion id="49B8F808172CE02EFF66122C33748F3D" box="[192,288,579,603]" country="United States of America" name="Arizona" pageId="5" pageNumber="4">Arizona</collectingRegion>
|
||
west of the Tinajas Altas and Gila mountain ranges.
|
||
<taxonomicName id="4C7C4D69172CE02EFC3D122B36C68F3D" authorityName="Warner" authorityYear="2011" box="[923,1170,579,603]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="howdenorum">
|
||
<emphasis id="B908EAF8172CE02EFC3D122B36C68F3D" box="[923,1170,579,603]" italics="true" pageId="5" pageNumber="4">Acoma howdenorum</emphasis>
|
||
</taxonomicName>
|
||
occurs only to the east of those two mountain ranges in the Mohawk Dune field just west of the Mohawk Mountains, and associated small dunes in the Mohawk Valley near Welton and Dome, where it is sympatric with the southwestern form of
|
||
<taxonomicName id="4C7C4D69172CE02EFEE912CB31448FDD" authority="Brown. Apparently" authorityName="Brown. Apparently" authorityYear="1929" box="[335,784,675,699]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="arizonica">
|
||
<emphasis id="B908EAF8172CE02EFEE912CB30488FDD" box="[335,540,676,699]" italics="true" pageId="5" pageNumber="4">Acoma arizonica</emphasis>
|
||
Brown. Apparently
|
||
</taxonomicName>
|
||
<taxonomicName id="4C7C4D69172CE02EFCB012CB318C8FDD" authorityName="Warner" authorityYear="2011" box="[790,984,675,699]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="howdenorum">
|
||
<emphasis id="B908EAF8172CE02EFCB012CB318C8FDD" box="[790,984,675,699]" italics="true" pageId="5" pageNumber="4">A. howdenorum</emphasis>
|
||
</taxonomicName>
|
||
does not occur on the east side of the Mohawk Mountains as concurrent collecting on a dune field just south of Dateland, AZ, only produced
|
||
<taxonomicName id="4C7C4D69172CE02EFA2012AB33658F9D" authorityName="Brown. Apparently" authorityYear="1929" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="arizonica">
|
||
<emphasis id="B908EAF8172CE02EFA2012AB33658F9D" italics="true" pageId="5" pageNumber="4">A. arizonica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC336EA172CE02EFF56136C3166897D" blockId="5.[192,1441,195,1051]" pageId="5" pageNumber="4">
|
||
Both
|
||
<taxonomicName id="4C7C4D69172CE02EFE96136B33E38E7D" authorityName="Cazier" authorityYear="1953" box="[304,439,771,795]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFE96136B33E38E7D" box="[304,439,771,795]" italics="true" pageId="5" pageNumber="4">A. glabrata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7C4D69172CE02EFE49136B30F98E7D" authorityName="Warner" authorityYear="2011" box="[495,685,771,795]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="howdenorum">
|
||
<emphasis id="B908EAF8172CE02EFE49136B30F98E7D" box="[495,685,771,795]" italics="true" pageId="5" pageNumber="4">A. howdenorum</emphasis>
|
||
</taxonomicName>
|
||
appear to be restricted to deep eolian sand dunes. Specimens of
|
||
<taxonomicName id="4C7C4D69172CE02EFF66134B33188E5D" authorityName="Cazier" authorityYear="1953" box="[192,332,803,827]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFF66134B33188E5D" box="[192,332,803,827]" italics="true" pageId="5" pageNumber="4">A. glabrata</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingCountry id="F36B767A172CE02EFE65134C33A98E5D" box="[451,509,803,827]" name="United States of America" pageId="5" pageNumber="4">USA</collectingCountry>
|
||
tend to be rather uniform in size (mostly
|
||
<quantity id="4C849B0F172CE02EFBA6134C360A8E5D" box="[1024,1118,803,827]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="5" pageNumber="4" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6-7 mm</quantity>
|
||
long), whereas specimens from the
|
||
<typeStatus id="54C78848172CE02EFE89132B33318E3A" box="[303,357,836,860]" pageId="5" pageNumber="4">type</typeStatus>
|
||
locality vary widely in size, with some specimens much larger than those from the
|
||
<collectingCountry id="F36B767A172CE02EFAF8132C37C88E3D" box="[1374,1436,835,859]" name="United States of America" pageId="5" pageNumber="4">USA</collectingCountry>
|
||
. The
|
||
<collectingRegion id="49B8F808172CE02EFF53130C33018E1D" box="[245,341,867,891]" country="United States of America" name="Arizona" pageId="5" pageNumber="4">Arizona</collectingRegion>
|
||
specimens of
|
||
<taxonomicName id="4C7C4D69172CE02EFE5C130B30D68E1D" authorityName="Cazier" authorityYear="1953" box="[506,642,867,891]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFE5C130B30D68E1D" box="[506,642,867,891]" italics="true" pageId="5" pageNumber="4">A. glabrata</emphasis>
|
||
</taxonomicName>
|
||
from south of Yuma differ slightly from specimens collected on the Algodones Dunes (across the
|
||
<collectingRegion id="49B8F808172CE02EFD8613EC30DA8EFD" box="[544,654,899,923]" country="United States of America" name="Colorado" pageId="5" pageNumber="4">Colorado</collectingRegion>
|
||
River) as well, but do have 10 segmented antennae and the other differences from
|
||
<taxonomicName id="4C7C4D69172CE02EFE2D13CB301F8EDD" authorityName="Warner" authorityYear="2011" box="[395,587,931,955]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="howdenorum">
|
||
<emphasis id="B908EAF8172CE02EFE2D13CB301F8EDD" box="[395,587,931,955]" italics="true" pageId="5" pageNumber="4">A. howdenorum</emphasis>
|
||
</taxonomicName>
|
||
noted above.
|
||
<taxonomicName id="4C7C4D69172CE02EFD4813CB31FB8EDD" authorityName="Cazier" authorityYear="1953" box="[750,943,931,955]" class="Insecta" family="Melolonthidae" genus="Acoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="glabrata">
|
||
<emphasis id="B908EAF8172CE02EFD4813CB31FB8EDD" box="[750,943,931,955]" italics="true" pageId="5" pageNumber="4">Acoma glabrata</emphasis>
|
||
</taxonomicName>
|
||
’s
|
||
<typeStatus id="54C78848172CE02EFC6113CB31A88EDA" box="[967,1020,932,956]" pageId="5" pageNumber="4">type</typeStatus>
|
||
locality dunes are remote from the Sonoran and
|
||
<collectingCountry id="F36B767A172CE02EFEC113AC33DA8EBD" box="[359,398,963,987]" name="United States of America" pageId="5" pageNumber="4">US</collectingCountry>
|
||
localities, and further study may indicate that the three populations (
|
||
<typeStatus id="54C78848172CE02EFB5913AC37618EBD" box="[1279,1333,963,987]" pageId="5" pageNumber="4">type</typeStatus>
|
||
locality, Algodones Dunes, Gran Desierto) represent separate species or subspecies. A specimen from the Algodones Dunes in
|
||
<collectingRegion id="49B8F808172CE02EFE93146C33E7897D" box="[309,435,1027,1051]" country="United States of America" name="California" pageId="5" pageNumber="4">California</collectingRegion>
|
||
is shown in
|
||
<figureCitation id="13472A6F172CE02EFDEE146C30E9897D" box="[584,701,1027,1051]" captionStart="Figure 1-5" captionStartId="11.[192,276,1864,1886]" captionTargetBox="[200,1433,192,1843]" captionTargetId="figure-15@11.[200,1433,192,1843]" captionTargetPageId="11" captionText="Figure 1-5. Acoma spp. 1) Acoma glabrata Cazier, head and pronotum. 2-5) Acoma westcotti Warner: 2) Head and pronotum, 3) Dorsal habitus. 4) Left anterior tibia. 5) Pygidium." figureDoi="http://doi.org/10.5281/zenodo.5160710" httpUri="https://zenodo.org/record/5160710/files/figure.png" pageId="5" pageNumber="4">Figures 1</figureCitation>
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,
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<figureCitation id="13472A6F172CE02EFD6C146C308E897D" box="[714,730,1027,1051]" captionStart="Figure 6-9" captionStartId="12.[192,276,1864,1886]" captionTargetBox="[246,1383,192,1841]" captionTargetId="figure-19@12.[246,1383,192,1841]" captionTargetPageId="12" captionText="Figure 6-9. Acoma spp. 6-7) Acoma howdenorum Warner: 6) Dorsal habitus. 7) Antenna. 8-9) Acoma glabrata Cazier (specimen from Algodones Dunes): 8) Dorsal habitus. 9) Antenna." figureDoi="http://doi.org/10.5281/zenodo.5160714" httpUri="https://zenodo.org/record/5160714/files/figure.png" pageId="5" pageNumber="4">8</figureCitation>
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, and 9.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |