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<document id="7C76DFF97D720B1271804B0458EDC3EA" ID-DOI="10.11646/zootaxa.5432.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10898520" ID-ZooBank="102FEF9D-51B0-49AF-A226-37B2D6E55925" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711817688943" checkinUser="plazi" docAuthor="Heiss, Ernst, Shimamoto, Shusuke &amp; Nagashima, Seidai" docDate="2024" docId="03CDA902FF89FFBF778CFA25B3DAFED5" docLanguage="en" docName="zootaxa.5432.1.4.pdf" docOrigin="Zootaxa 5432 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5432.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aradosyrtis shonoi Heiss &amp; Shimamoto &amp; Nagashima 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="66" masterDocId="FFF4D17AFF80FFB2771BFFEFB20BFFC8" masterDocTitle="Review of the flat bug subfamily Calisiinae from Japan, with description of a new genus and four new species (Hemiptera: Heteroptera: Aradidae)" masterLastPageNumber="68" masterPageNumber="54" pageNumber="63" updateTime="1711818425965" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8CD0A19D51DCAB9D5BF60129D66EB8E8">Review of the flat bug subfamily Calisiinae from Japan, with description of a new genus and four new species (Hemiptera: Heteroptera: Aradidae)</mods:title>
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<mods:namePart id="F17D6CDC24AD34F6C988A07B24EDFD47">Heiss, Ernst</mods:namePart>
<mods:affiliation id="2DE3755F2998CA6214992CE919BDD5B8">Entomology Research Associate, Tiroler Landesmuseum, 2 a Josef Schraffl Strasse, A- 6020 Innsbruck, Austria</mods:affiliation>
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<mods:namePart id="47B62641617D1C0A177F4A1B4F0EF123">Shimamoto, Shusuke</mods:namePart>
<mods:affiliation id="40ABF704A9D221D7C67EAB1731A83747">Laboratory of Entomology, Faculty of Agriculture, Tokyo University of Agriculture, 1737 Funako, Atsugi-shi, Kanagawa, Japan</mods:affiliation>
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<mods:namePart id="B46A67C71E05852FF4EABCC8F7DA97D0">Nagashima, Seidai</mods:namePart>
<mods:affiliation id="895D19B11320CF0EAA1A40E8A44B8F97">Itami City Museum of Insects, 3 - 1 Koyaike, Itami-shi, Hyogo, Japan</mods:affiliation>
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<mods:title id="12F6B6EB2BE8CFD92ED0C982370737D7">Zootaxa</mods:title>
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<mods:date id="3D9D7129AF926D67D0C7B7EAC7E85212">2024</mods:date>
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<mods:number id="6BF5ADDA5EEEA129CD37C144D8982E8D">2024-03-27</mods:number>
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<mods:number id="2E7C3B0ED5ACF46C426039044BFE45AA">5432</mods:number>
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<treatment id="03CDA902FF89FFBF778CFA25B3DAFED5" ID-DOI="http://doi.org/10.5281/zenodo.10898458" ID-Zenodo-Dep="10898458" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5" httpUri="http://treatment.plazi.org/id/03CDA902FF89FFBF778CFA25B3DAFED5" lastPageId="13" lastPageNumber="66" pageId="9" pageNumber="63">
<subSubSection id="C37E4B9FFF89FFBB778CFA25B3D4FA2D" box="[151,479,1482,1509]" pageId="9" pageNumber="63" type="nomenclature">
<paragraph id="8BDB1814FF89FFBB778CFA25B3D4FA2D" blockId="9.[151,518,1482,1545]" box="[151,479,1482,1509]" pageId="9" pageNumber="63">
<heading id="D093AF78FF89FFBB778CFA25B3D4FA2D" bold="true" box="[151,479,1482,1509]" fontSize="11" level="1" pageId="9" pageNumber="63" reason="1">
<emphasis id="B910C406FF89FFBB778CFA25B3D4FA2D" bold="true" box="[151,479,1482,1509]" pageId="9" pageNumber="63">
<taxonomicName id="4C646397FF89FFBB778CFA25B370FA2D" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[151,379,1482,1509]" class="Insecta" family="Aradidae" genus="Aradosyrtis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="63" phylum="Arthropoda" rank="species" species="shonoi" status="sp. nov.">
<emphasis id="B910C406FF89FFBB778CFA25B370FA2D" bold="true" box="[151,379,1482,1509]" italics="true" pageId="9" pageNumber="63">Aradosyrtis shonoi</emphasis>
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<taxonomicNameLabel id="A223797DFF89FFBB7699FA24B3D4FA2D" box="[386,479,1483,1509]" pageId="9" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C37E4B9FFF89FFBB778CFA00B00DF9C1" box="[151,518,1518,1545]" pageId="9" pageNumber="63" type="description">
<paragraph id="8BDB1814FF89FFBB778CFA00B00DF9C1" blockId="9.[151,518,1482,1545]" box="[151,518,1518,1545]" pageId="9" pageNumber="63">
(
<figureCitation id="135F0491FF89FFBB7784FA00B312F9C1" box="[159,281,1519,1545]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[170,1419,193,1895]" captionTargetId="figure-17@2.[158,1429,181,1918]" captionTargetPageId="2" captionText="FIGURES 1–10. Japanese Nipponocalisius and Aradosyrtis species, dorsal habitus. 1–2, N. ishikawanus sp. nov., (1) ♂, (2) ♀; 3–5, N. tomokunii sp. nov., (3–4) ♂, (5) ♀; 6–8, N. dimorphus sp. nov., (6) brachypterous ♂, (7) brachypterous ♀, (8) macropterous ♀; 9–10, A. shonoi sp. nov., (9) ♂, (10) ♀" figureDoi="http://doi.org/10.5281/zenodo.10898522" httpUri="https://zenodo.org/record/10898522/files/figure.png" pageId="9" pageNumber="63">Figs. 9–10</figureCitation>
,
<figureCitation id="135F0491FF89FFBB763EFA00B348F9C1" box="[293,323,1519,1545]" captionStart="FIGURES 11–16" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[192,1395,188,1913]" captionTargetId="figure-17@6.[186,1401,181,1919]" captionTargetPageId="6" captionText="FIGURES 11–16. Japanese Nipponocalisius and Aradosyrtis species, dorsal habitus and (a) ventral head. 11, N. ishikawanus sp. nov., ♂; 12, N. tomokunii sp. nov., ♂; 13–15, N. dimorphus sp. nov., (13) brachypterous ♂, (14) brachypterous ♀, (15) macropterous ♀; 16, A. shonoi sp. nov., ♂." figureDoi="http://doi.org/10.5281/zenodo.10898526" httpUri="https://zenodo.org/record/10898526/files/figure.png" pageId="9" pageNumber="63">16</figureCitation>
,
<figureCitation id="135F0491FF89FFBB7654FA01B39CF9C1" box="[335,407,1518,1545]" captionStart="FIGURES 17–25" captionStartId="7.[151,264,1581,1605]" captionTargetBox="[165,1422,195,1543]" captionTargetId="figure-18@7.[151,1435,181,1557]" captionTargetPageId="7" captionText="FIGURES 17–25. Japanese Nipponocalisius and Aradosyrtis species, right antenna, (a) dorsal view and (b) ventral view. 17–18, N. ishikawanus sp. nov.; 19–20, N. tomokunii sp. nov.; 21–23, N. dimorphus sp. nov., (21–22) brachypterous, (23) macropterous; 24–25, A. shonoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10898528" httpUri="https://zenodo.org/record/10898528/files/figure.png" pageId="9" pageNumber="63">24–25</figureCitation>
,
<figureCitation id="135F0491FF89FFBB76B8FA00B3CAF9C1" box="[419,449,1519,1545]" captionStart="FIGURES 26–29" captionStartId="8.[151,264,1624,1648]" captionTargetBox="[154,1429,745,1597]" captionTargetId="figure-291@8.[151,1436,741,1601]" captionTargetPageId="8" captionText="FIGURES 26–29. Japanese Nipponocalisius and Aradosyrtis species, left paramere, (a) dorsal view and (b) ventral view. 26, N. ishikawanus sp. nov.; 27, N. tomokunii sp. nov., 28, N. dimorphus sp. nov., 29, A. shonoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10898530" httpUri="https://zenodo.org/record/10898530/files/figure.png" pageId="9" pageNumber="63">29</figureCitation>
,
<figureCitation id="135F0491FF89FFBB76D6FA00B3F5F9C1" box="[461,510,1519,1545]" captionStart="FIGURE 33" captionStartId="11.[151,250,1648,1672]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-18@11.[151,1436,181,1623]" captionTargetPageId="11" captionText="FIGURE 33. Living individual and habitat of Japanese Nipponocalisius and Aradosyrtis species.A–C, N. ishikawanus sp. nov., (A) ♂, (B) ♀, (C) nymph; D–I, N. tomokunii sp. nov., (D–E) ♂, (F) egg and nymph, (G–H) ♀, (I) nymph; J–N, N. dimorphus sp. nov., (J) macropterous ♀, (K) brachypterous ♀, (L) nymph, (M) brachypterous ♂ and nymph, (N) habitat; O, A. shonoi sp. nov., ♂ (photographed by Jun Souma)." figureDoi="http://doi.org/10.5281/zenodo.10898534" httpUri="https://zenodo.org/record/10898534/files/figure.png" pageId="9" pageNumber="63">33O</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37E4B9FFF89FFBB778CF9DCB0EFF983" box="[151,740,1587,1611]" pageId="9" pageNumber="63" type="reference_group">
<paragraph id="8BDB1814FF89FFBB778CF9DCB0EFF983" blockId="9.[151,740,1587,1611]" box="[151,740,1587,1611]" pageId="9" pageNumber="63">
<treatmentCitationGroup id="AB743F3AFF89FFBB778CF9DCB0EFF983" box="[151,740,1587,1611]" pageId="9" pageNumber="63">
<taxonomicName id="4C646397FF89FFBB778CF9DCB353F983" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[151,344,1587,1611]" class="Insecta" family="Aradidae" genus="Aradosyrtis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="63" phylum="Arthropoda" rank="species" species="shonoi">
<emphasis id="B910C406FF89FFBB778CF9DCB353F983" box="[151,344,1587,1611]" italics="true" pageId="9" pageNumber="63">Aradosyrtis shonoi</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AC53E05FF89FFBB767EF9DCB010F983" author="Heiss, E." box="[357,539,1587,1611]" page="111" pageId="9" pageNumber="63" year="2023">
<bibRefCitation id="EFF565E5FF89FFBB767EF9DCB010F983" author="Heiss, E." box="[357,539,1587,1611]" pageId="9" pageNumber="63" pagination="110 - 120" refId="ref8284" refString="Heiss, E. (2023) New taxonomic assignments for two Palaearctic Calisiinae genera (Heteroptera, Aradidae) with description of a new species from China. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 75, 110 - 120." type="journal article" year="2023">Heiss (2023: 111)</bibRefCitation>
</treatmentCitation>
. Unavailable name.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37E4B9FFF89FFBB778CF99CB3EEF83D" pageId="9" pageNumber="63" type="materials_examined">
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<emphasis id="B910C406FF89FFBB778CF99CB342F944" bold="true" box="[151,329,1650,1677]" pageId="9" pageNumber="63">Type material.</emphasis>
<materialsCitation id="3B0C1249FF89FFBB764FF99CB3EEF83D" collectingDate="1998-07-13" collectingDateMax="2022-09-08" collectingDateMin="1998-07-13" collectionCode="TUA, CEHI" collectorName="Ishigaki Is. &amp; Tadafumi Nakata &amp; Jun Souma &amp; Okinawa-ken &amp; Yaeyama-gun &amp; Taketomi-cho &amp; Aira-gawa &amp; Fumitaka Nakano &amp; Iriomote Is. &amp; Tedwu-san &amp; Reo Ito &amp; All" country="Japan" county="Ryukyus" latitude="24.3475" location="Tadashi" longLatPrecision="21" longitude="123.90305" municipality="Itona" pageId="9" pageNumber="63" specimenCount="22" specimenCount-female="12" specimenCount-male="10" stateProvince="Ishikawa" typeStatus="holotype">
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<typeStatus id="54DFA6B6FF89FFBB764FF99CB3CAF945" box="[340,449,1651,1677]" pageId="9" pageNumber="63" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D62D39DFF89FFBB76CDF99CB3E5F945" box="[470,494,1651,1677]" count="1" pageId="9" pageNumber="63" type="male">♂</specimenCount>
):
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,
<collectingCounty id="62BA6098FF89FFBB7543F99CB0B1F945" box="[600,698,1651,1677]" pageId="9" pageNumber="63">Ryukyus</collectingCounty>
/
<collectorName id="26917DC2FF89FFBB75C2F99CB150F945" box="[729,859,1651,1677]" pageId="9" pageNumber="63">Ishigaki Is.</collectorName>
,
<collectingMunicipality id="6BBF826EFF89FFBB7477F99CB1ADF945" box="[876,934,1651,1677]" pageId="9" pageNumber="63">Itona</collectingMunicipality>
/
<date id="FFDA3ED4FF89FFBB74D8F99CB635F945" box="[963,1086,1651,1677]" pageId="9" pageNumber="63" value="2003-06-03">
<collectingDate id="EF9EC73CFF89FFBB74D8F99CB635F945" box="[963,1086,1651,1677]" pageId="9" pageNumber="63" value="2003-06-03">3.Jun.2003</collectingDate>
</date>
/
<location id="8EBB4ECFFF89FFBB7347F99CB6B8F945" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7347F99CB6B8F945" box="[1116,1203,1651,1677]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Tadashi" pageId="9" pageNumber="63" stateProvince="Ishikawa">Tadashi</location>
<collectingRegion id="49A0D6F6FF89FFBB73A6F99CB729F945" box="[1213,1314,1651,1677]" country="Japan" name="Ishikawa" pageId="9" pageNumber="63">Ishikawa</collectingRegion>
// (
<collectionCode id="ED7580D1FF89FFBB724BF99CB784F945" box="[1360,1423,1651,1677]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<emphasis id="B910C406FF89FFBB778CF978B306F979" bold="true" box="[151,269,1687,1713]" pageId="9" pageNumber="63">
<typeStatus id="54DFA6B6FF89FFBB778CF978B306F979" box="[151,269,1687,1713]" pageId="9" pageNumber="63" type="paratype">paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D62D39DFF89FFBB7605F978B341F979" box="[286,330,1687,1713]" count="1" pageId="9" pageNumber="63" type="female">1 ♀</specimenCount>
collected with holotype (
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);
<specimenCount id="9D62D39DFF89FFBB75D6F978B0F0F979" box="[717,763,1687,1713]" count="1" pageId="9" pageNumber="63" type="female">1 ♀</specimenCount>
:
<collectingCountry id="F3735884FF89FFBB7411F978B146F979" box="[778,845,1687,1713]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
,
<location id="8EBB4ECFFF89FFBB7441F978B1B7F979" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7441F978B1B7F979" box="[858,956,1687,1713]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Ryukyus" pageId="9" pageNumber="63" stateProvince="Ishikawa">Ryukyus</location>
/
<collectorName id="26917DC2FF89FFBB74CDF978B65DF979" box="[982,1110,1687,1713]" pageId="9" pageNumber="63">Ishigaki Is.</collectorName>
,
<location id="8EBB4ECFFF89FFBB737EF978B6EEF979" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB737EF978B6EEF979" box="[1125,1253,1687,1713]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Mt. Yarabu" pageId="9" pageNumber="63" stateProvince="Ishikawa">Mt. Yarabu</location>
/
<date id="FFDA3ED4FF89FFBB73E5F978B780F979" box="[1278,1419,1687,1713]" pageId="9" pageNumber="63" value="2003-06-10">
<collectingDate id="EF9EC73CFF89FFBB73E5F978B780F979" box="[1278,1419,1687,1713]" pageId="9" pageNumber="63" value="2003-06-10">10 Jun 2003</collectingDate>
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<location id="8EBB4ECFFF89FFBB778CF954B368F91D" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB778CF954B368F91D" box="[151,355,1723,1749]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Kazutaka Yamada" pageId="9" pageNumber="63" stateProvince="Ishikawa">Kazutaka Yamada</location>
// (
<collectionCode id="ED7580D1FF89FFBB7695F954B3C6F91D" box="[398,461,1723,1749]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<specimenCount id="9D62D39DFF89FFBB76F8F954B019F91D" box="[483,530,1723,1749]" count="1" pageId="9" pageNumber="63" type="female">1 ♀</specimenCount>
:
<collectingCountry id="F3735884FF89FFBB753AF954B06FF91D" box="[545,612,1723,1749]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
,
<location id="8EBB4ECFFF89FFBB756AF954B0D8F91D" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB756AF954B0D8F91D" box="[625,723,1723,1749]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Ryukyus" pageId="9" pageNumber="63" stateProvince="Ishikawa">Ryukyus</location>
/
<collectorName id="26917DC2FF89FFBB75F5F954B166F91D" box="[750,877,1723,1749]" pageId="9" pageNumber="63">Ishigaki Is.</collectorName>
/
<location id="8EBB4ECFFF89FFBB7493F954B64DF91D" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7493F954B64DF91D" box="[904,1094,1723,1749]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Mt. Yarabu-dake" pageId="9" pageNumber="63" stateProvince="Ishikawa">Mt. Yarabu-dake</location>
/ 27.January.2003 /
<collectorName id="26917DC2FF89FFBB722AF954B2EDF931" pageId="9" pageNumber="63">Tadafumi Nakata</collectorName>
leg. // (
<collectionCode id="ED7580D1FF89FFBB765BF930B374F931" box="[320,383,1759,1785]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<specimenCount id="9D62D39DFF89FFBB768FF930B3CAF931" box="[404,449,1759,1785]" count="1" pageId="9" pageNumber="63" type="female">1 ♀</specimenCount>
:
<collectingCountry id="F3735884FF89FFBB76D4F930B019F931" box="[463,530,1759,1785]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
,
<location id="8EBB4ECFFF89FFBB7505F930B08BF931" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7505F930B08BF931" box="[542,640,1759,1785]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Ryukyus" pageId="9" pageNumber="63" stateProvince="Ishikawa">Ryukyus</location>
/
<collectorName id="26917DC2FF89FFBB7583F930B11CF931" box="[664,791,1759,1785]" pageId="9" pageNumber="63">Ishigaki Is.</collectorName>
,
<location id="8EBB4ECFFF89FFBB743FF930B17FF931" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB743FF930B17FF931" box="[804,884,1759,1785]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Takeda" pageId="9" pageNumber="63" stateProvince="Ishikawa">Takeda</location>
/
<date id="FFDA3ED4FF89FFBB7496F930B62BF931" box="[909,1056,1759,1785]" pageId="9" pageNumber="63" value="1998-07-16">
<collectingDate id="EF9EC73CFF89FFBB7496F930B62BF931" box="[909,1056,1759,1785]" pageId="9" pageNumber="63" value="1998-07-16">16 July 1998</collectingDate>
</date>
/
<location id="8EBB4ECFFF89FFBB7323F930B708F931" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7323F930B708F931" box="[1080,1283,1759,1785]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Keiichi Takahashi" pageId="9" pageNumber="63" stateProvince="Ishikawa">Keiichi Takahashi</location>
//[same label written in Japanese] // (
<collectionCode id="ED7580D1FF89FFBB7682F8ECB3D3F8D5" box="[409,472,1795,1821]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<specimenCount id="9D62D39DFF89FFBB76F0F8ECB01DF8D5" box="[491,534,1795,1821]" count="1" pageId="9" pageNumber="63" type="female">1 ♀</specimenCount>
:
<collectingCountry id="F3735884FF89FFBB7539F8ECB06EF8D5" box="[546,613,1795,1821]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
,
<location id="8EBB4ECFFF89FFBB7574F8ECB0DAF8D5" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7574F8ECB0DAF8D5" box="[623,721,1795,1821]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Ryukyus" pageId="9" pageNumber="63" stateProvince="Ishikawa">Ryukyus</location>
/
<collectorName id="26917DC2FF89FFBB75FEF8ECB169F8D5" box="[741,866,1795,1821]" pageId="9" pageNumber="63">Ishigaki Is.</collectorName>
,
<location id="8EBB4ECFFF89FFBB7476F8ECB1B6F8D5" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7476F8ECB1B6F8D5" box="[877,957,1795,1821]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Takeda" pageId="9" pageNumber="63" stateProvince="Ishikawa">Takeda</location>
/
<date id="FFDA3ED4FF89FFBB74C9F8ECB66BF8D5" box="[978,1120,1795,1821]" pageId="9" pageNumber="63" value="1998-07-30">
<collectingDate id="EF9EC73CFF89FFBB74C9F8ECB66BF8D5" box="[978,1120,1795,1821]" pageId="9" pageNumber="63" value="1998-07-30">30 July 1998</collectingDate>
</date>
/
<location id="8EBB4ECFFF89FFBB736FF8ECB736F8D5" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB736FF8ECB736F8D5" box="[1140,1341,1795,1821]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Keiichi Takahashi" pageId="9" pageNumber="63" stateProvince="Ishikawa">Keiichi Takahashi</location>
// [same label written in Japanese] // (
<collectionCode id="ED7580D1FF89FFBB76C5F8C8B016F889" box="[478,541,1831,1857]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<specimenCount id="9D62D39DFF89FFBB7529F8C8B054F889" box="[562,607,1831,1857]" count="1" pageId="9" pageNumber="63" type="female">1 ♀</specimenCount>
:
<collectingCountry id="F3735884FF89FFBB7577F8C8B0A4F889" box="[620,687,1831,1857]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
,
<location id="8EBB4ECFFF89FFBB75A0F8C8B116F889" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB75A0F8C8B116F889" box="[699,797,1831,1857]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Ryukyus" pageId="9" pageNumber="63" stateProvince="Ishikawa">Ryukyus</location>
/
<collectorName id="26917DC2FF89FFBB742FF8C8B1B8F889" box="[820,947,1831,1857]" pageId="9" pageNumber="63">Ishigaki Is.</collectorName>
,
<location id="8EBB4ECFFF89FFBB74DBF8C8B61BF889" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB74DBF8C8B61BF889" box="[960,1040,1831,1857]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Takeda" pageId="9" pageNumber="63" stateProvince="Ishikawa">Takeda</location>
/
<date id="FFDA3ED4FF89FFBB7333F8C8B6B1F889" box="[1064,1210,1831,1857]" pageId="9" pageNumber="63" value="1998-07-13">
<collectingDate id="EF9EC73CFF89FFBB7333F8C8B6B1F889" box="[1064,1210,1831,1857]" pageId="9" pageNumber="63" value="1998-07-13">13 July 1998</collectingDate>
</date>
/
<location id="8EBB4ECFFF89FFBB73CAF8C8B797F889" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB73CAF8C8B797F889" box="[1233,1436,1831,1857]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Keiichi Takahashi" pageId="9" pageNumber="63" stateProvince="Ishikawa">Keiichi Takahashi</location>
// [same label written in Japanese] // (
<collectionCode id="ED7580D1FF89FFBB7555F8A4B086F8AD" box="[590,653,1867,1893]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<specimenCount id="9D62D39DFF89FFBB75B8F8A4B0D8F8AD" box="[675,723,1867,1893]" count="1" pageId="9" pageNumber="63" type="male">1 ♂</specimenCount>
:
<collectingCountry id="F3735884FF89FFBB75F9F8A4B12EF8AD" box="[738,805,1867,1893]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
, the
<location id="8EBB4ECFFF89FFBB7445F8A4B1CBF8AD" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB7445F8A4B1CBF8AD" box="[862,960,1867,1893]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Ryukyus" pageId="9" pageNumber="63" stateProvince="Ishikawa">Ryukyus</location>
/ Yaeyama Isls.,
<collectorName id="26917DC2FF89FFBB7391F8A4B702F8AD" box="[1162,1289,1867,1893]" pageId="9" pageNumber="63">Ishigaki Is.</collectorName>
/
<location id="8EBB4ECFFF89FFBB723FF8A4B793F8AD" LSID="urn:lsid:plazi:treatment:03CDA902FF89FFBF778CFA25B3DAFED5:8EBB4ECFFF89FFBB723FF8A4B793F8AD" box="[1316,1432,1867,1893]" country="Japan" county="Ryukyus" latitude="24.3475" longLatPrecision="21" longitude="123.90305" municipality="Itona" name="Tonoshiro" pageId="9" pageNumber="63" stateProvince="Ishikawa">Tonoshiro</location>
,
<taxonomicName id="4C646397FF89FFBB778CF880B376F841" box="[151,381,1903,1929]" class="Magnoliopsida" family="Oleaceae" genus="Osmanthus" kingdom="Plantae" order="Lamiales" pageId="9" pageNumber="63" phylum="Tracheophyta" rank="species" species="insularis">
<emphasis id="B910C406FF89FFBB778CF880B376F841" box="[151,381,1903,1929]" italics="true" pageId="9" pageNumber="63">Osmanthus insularis</emphasis>
</taxonomicName>
/
<geoCoordinate id="EE507ED3FF89FFBB7689F880B039F841" box="[402,562,1903,1929]" degrees="24" direction="north" minutes="22" orientation="latitude" pageId="9" pageNumber="63" precision="1" seconds="40.0" value="24.377777">24°22’40.0”N</geoCoordinate>
,
<geoCoordinate id="EE507ED3FF89FFBB7526F880B0EDF841" box="[573,742,1903,1929]" degrees="124" direction="east" minutes="10" orientation="longitude" pageId="9" pageNumber="63" precision="1" seconds="24.0" value="124.17333">124°10’24.0”E</geoCoordinate>
/
<date id="FFDA3ED4FF89FFBB75E1F880B160F841" box="[762,875,1903,1929]" pageId="9" pageNumber="63" value="2022-09-08">
<collectingDate id="EF9EC73CFF89FFBB75E1F880B160F841" box="[762,875,1903,1929]" pageId="9" pageNumber="63" value="2022-09-08">8 IX 2022</collectingDate>
</date>
/
<collectorName id="26917DC2FF89FFBB7464F880B1F1F841" box="[895,1018,1903,1929]" pageId="9" pageNumber="63">Jun Souma</collectorName>
// (
<collectionCode id="ED7580D1FF89FFBB7305F880B656F841" box="[1054,1117,1903,1929]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<specimenCount id="9D62D39DFF89FFBB736BF880B6A5F841" box="[1136,1198,1903,1929]" count="2" pageId="9" pageNumber="63" type="male">2 ♂♂</specimenCount>
<specimenCount id="9D62D39DFF89FFBB73AFF880B6FEF841" box="[1204,1269,1903,1929]" count="2" pageId="9" pageNumber="63" type="female">2 ♀♀</specimenCount>
: Iriomotejima-Is.,
<collectingCountry id="F3735884FF89FFBB77DFF87CB30FF865" box="[196,260,1939,1965]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
/
<collectorName id="26917DC2FF89FFBB763BF87CB3B1F865" box="[288,442,1939,1965]" pageId="9" pageNumber="63">Okinawa-ken</collectorName>
,
<collectorName id="26917DC2FF89FFBB76DCF87CB06AF865" box="[455,609,1939,1965]" pageId="9" pageNumber="63">Yaeyama-gun</collectorName>
/
<collectorName id="26917DC2FF89FFBB7566F87CB110F865" box="[637,795,1939,1965]" pageId="9" pageNumber="63">Taketomi-chō</collectorName>
,
<collectorName id="26917DC2FF89FFBB7433F87CB195F865" box="[808,926,1939,1965]" pageId="9" pageNumber="63">Aira-gawa</collectorName>
/
<geoCoordinate id="EE507ED3FF89FFBB74A1F87DB64DF865" box="[954,1094,1938,1965]" degrees="24" direction="north" minutes="20" orientation="latitude" pageId="9" pageNumber="63" precision="15" seconds="51" value="24.3475">24°20’51”N</geoCoordinate>
,
<geoCoordinate id="EE507ED3FF89FFBB734EF87DB6E3F865" box="[1109,1256,1938,1965]" degrees="123" direction="east" minutes="54" orientation="longitude" pageId="9" pageNumber="63" precision="15" seconds="11" value="123.90305">123°54’11”E</geoCoordinate>
/
<date id="FFDA3ED4FF89FFBB721FF87CB781F865" box="[1284,1418,1939,1965]" pageId="9" pageNumber="63" value="2018-11-24">
<collectingDate id="EF9EC73CFF89FFBB721FF87CB781F865" box="[1284,1418,1939,1965]" pageId="9" pageNumber="63" value="2018-11-24">24 XI 2018</collectingDate>
</date>
/
<collectorName id="26917DC2FF89FFBB778CF858B36BF819" box="[151,352,1975,2001]" pageId="9" pageNumber="63">Fumitaka Nakano</collectorName>
// (
<collectionCode id="ED7580D1FF89FFBB7693F858B3CCF819" box="[392,455,1975,2001]" pageId="9" pageNumber="63">TUA</collectionCode>
);
<specimenCount id="9D62D39DFF89FFBB76C7F858B017F819" box="[476,540,1975,2001]" count="6" pageId="9" pageNumber="63" type="male">6 ♂♂</specimenCount>
<specimenCount id="9D62D39DFF89FFBB753FF858B06DF819" box="[548,614,1975,2001]" count="4" pageId="9" pageNumber="63" type="female">4 ♀♀</specimenCount>
:
<collectingCountry id="F3735884FF89FFBB756FF858B0BCF819" box="[628,695,1975,2001]" name="Japan" pageId="9" pageNumber="63">Japan</collectingCountry>
,
<collectorName id="26917DC2FF89FFBB75D9F858B14DF819" box="[706,838,1975,2001]" pageId="9" pageNumber="63">Iriomote Is.</collectorName>
/
<collectorName id="26917DC2FF89FFBB7446F858B1DCF819" box="[861,983,1975,2001]" pageId="9" pageNumber="63">Tedwu-san</collectorName>
/
<date id="FFDA3ED4FF89FFBB74EBF858B687F819" box="[1008,1164,1975,2001]" pageId="9" pageNumber="63" value="2020-02-23">
<collectingDate id="EF9EC73CFF89FFBB74EBF858B687F819" box="[1008,1164,1975,2001]" pageId="9" pageNumber="63" value="2020-02-23">23. Feb. 2020</collectingDate>
</date>
/
<collectorName id="26917DC2FF89FFBB73BFF858B6F3F819" box="[1188,1272,1975,2001]" pageId="9" pageNumber="63">Reo Ito</collectorName>
leg. // (
<collectionCode id="ED7580D1FF89FFBB724AF858B79AF819" box="[1361,1425,1975,2001]" pageId="9" pageNumber="63">TUA</collectionCode>
).
<collectorName id="26917DC2FF89FFBB778CF834B2B0F83D" box="[151,187,2011,2037]" pageId="9" pageNumber="63">All</collectorName>
provided with type labels.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF1B489CFF8AFFB8778CF8E1B639F882" ID-DOI="http://doi.org/10.5281/zenodo.10898532" ID-Zenodo-Dep="10898532" httpUri="https://zenodo.org/record/10898532/files/figure.png" pageId="10" pageNumber="64" startId="10.[151,264,1806,1830]" targetBox="[151,1436,209,1767]" targetPageId="10" targetType="figure">
<paragraph id="8BDB1814FF8AFFB8778CF8E1B639F882" blockId="10.[151,1436,1806,1866]" pageId="10" pageNumber="64">
<emphasis id="B910C406FF8AFFB8778CF8E1B352F8EE" bold="true" box="[151,345,1806,1830]" pageId="10" pageNumber="64">FIGURES 30–32.</emphasis>
Japanese
<taxonomicName id="4C646397FF8AFFB876DCF8E1B0CBF8EE" box="[455,704,1806,1830]" pageId="10" pageNumber="64">
<emphasis id="B910C406FF8AFFB876DCF8E1B067F8EE" box="[455,620,1806,1830]" italics="true" pageId="10" pageNumber="64">Nipponocalisius</emphasis>
species
</taxonomicName>
examined in previous studies, (a) figure from original literature, (b) relevant specimen, and (c) label. 30–31,
<taxonomicName id="4C646397FF8AFFB87528F8DCB0C6F882" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[563,717,1842,1866]" class="Insecta" family="Aradidae" genus="Nipponocalisius" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="64" phylum="Arthropoda" rank="species" species="ishikawanus" status="sp. nov.">
<emphasis id="B910C406FF8AFFB87528F8DCB04FF882" box="[563,580,1843,1866]" italics="true" pageId="10" pageNumber="64">N</emphasis>
.
<emphasis id="B910C406FF8AFFB8754BF8DDB0C6F882" box="[592,717,1842,1866]" italics="true" pageId="10" pageNumber="64">ishikawanus</emphasis>
</taxonomicName>
<emphasis id="B910C406FF8AFFB875CFF8DDB12FF882" bold="true" box="[724,804,1842,1866]" pageId="10" pageNumber="64">
<taxonomicNameLabel id="A223797DFF8AFFB875CFF8DDB12FF882" box="[724,804,1842,1866]" pageId="10" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 32,
<taxonomicName id="4C646397FF8AFFB87443F8DCB1D0F882" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[856,987,1842,1866]" class="Insecta" family="Aradidae" genus="Nipponocalisius" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="64" phylum="Arthropoda" rank="species" species="tomokunii" status="sp. nov.">
<emphasis id="B910C406FF8AFFB87443F8DCB162F882" box="[856,873,1843,1866]" italics="true" pageId="10" pageNumber="64">N</emphasis>
.
<emphasis id="B910C406FF8AFFB8746DF8DDB1D0F882" box="[886,987,1842,1866]" italics="true" pageId="10" pageNumber="64">tomokunii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A223797DFF8AFFB874FAF8DDB639F882" box="[993,1074,1842,1866]" pageId="10" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF1B489CFF8BFFB9778CF99FB028F93C" ID-DOI="http://doi.org/10.5281/zenodo.10898534" ID-Zenodo-Dep="10898534" httpUri="https://zenodo.org/record/10898534/files/figure.png" pageId="11" pageNumber="65" startId="11.[151,250,1648,1672]" targetBox="[151,1436,181,1623]" targetPageId="11" targetType="figure">
<paragraph id="8BDB1814FF8BFFB9778CF99FB028F93C" blockId="11.[151,1436,1648,1780]" pageId="11" pageNumber="65">
<emphasis id="B910C406FF8BFFB9778CF99FB314F940" bold="true" box="[151,287,1648,1672]" pageId="11" pageNumber="65">FIGURE 33.</emphasis>
Living individual and habitat of Japanese
<taxonomicName id="4C646397FF8BFFB975D3F99FB166F940" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[712,877,1648,1672]" class="Insecta" family="Aradidae" genus="Nipponocalisius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="65" phylum="Arthropoda" rank="genus">
<emphasis id="B910C406FF8BFFB975D3F99FB166F940" box="[712,877,1648,1672]" italics="true" pageId="11" pageNumber="65">Nipponocalisius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C646397FF8BFFB97487F99FB675F940" box="[924,1150,1648,1672]" class="Insecta" family="Aradidae" genus="Aradosyrtis" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="65" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B910C406FF8BFFB97487F99FB619F940" box="[924,1042,1648,1672]" italics="true" pageId="11" pageNumber="65">Aradosyrtis</emphasis>
species. A
</taxonomicName>
–C,
<taxonomicName id="4C646397FF8BFFB973BDF99EB74BF940" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[1190,1344,1648,1672]" class="Insecta" family="Aradidae" genus="Nipponocalisius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="65" phylum="Arthropoda" rank="species" species="ishikawanus" status="sp. nov.">
<emphasis id="B910C406FF8BFFB973BDF99EB6BCF940" box="[1190,1207,1649,1672]" italics="true" pageId="11" pageNumber="65">N</emphasis>
.
<emphasis id="B910C406FF8BFFB973D8F99FB74BF940" box="[1219,1344,1648,1672]" italics="true" pageId="11" pageNumber="65">ishikawanus</emphasis>
</taxonomicName>
<emphasis id="B910C406FF8BFFB9725DF99FB79EF940" bold="true" box="[1350,1429,1648,1672]" pageId="11" pageNumber="65">
<taxonomicNameLabel id="A223797DFF8BFFB9725DF99FB79EF940" box="[1350,1429,1648,1672]" pageId="11" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (A) ♂, (B) ♀, (C) nymph; D–I,
<taxonomicName id="4C646397FF8BFFB976C6F97AB06BF964" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[477,608,1684,1708]" class="Insecta" family="Aradidae" genus="Nipponocalisius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="65" phylum="Arthropoda" rank="species" species="tomokunii" status="sp. nov.">
<emphasis id="B910C406FF8BFFB976C6F97AB3E5F964" box="[477,494,1685,1708]" italics="true" pageId="11" pageNumber="65">N</emphasis>
.
<emphasis id="B910C406FF8BFFB976E0F97BB06BF964" box="[507,608,1684,1708]" italics="true" pageId="11" pageNumber="65">tomokunii</emphasis>
</taxonomicName>
<emphasis id="B910C406FF8BFFB9757CF97BB0B3F964" bold="true" box="[615,696,1684,1708]" pageId="11" pageNumber="65">
<taxonomicNameLabel id="A223797DFF8BFFB9757CF97BB0B3F964" box="[615,696,1684,1708]" pageId="11" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (D–E) ♂, (F) egg and nymph, (G–H) ♀, (I) nymph; J–N,
<taxonomicName id="4C646397FF8BFFB97209F97AB797F964" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[1298,1436,1684,1708]" class="Insecta" family="Aradidae" genus="Nipponocalisius" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="65" phylum="Arthropoda" rank="species" species="dimorphus" status="sp. nov.">
<emphasis id="B910C406FF8BFFB97209F97AB728F964" box="[1298,1315,1685,1708]" italics="true" pageId="11" pageNumber="65">N</emphasis>
.
<emphasis id="B910C406FF8BFFB9722BF97BB797F964" box="[1328,1436,1684,1708]" italics="true" pageId="11" pageNumber="65">dimorphus</emphasis>
</taxonomicName>
<emphasis id="B910C406FF8BFFB9778CF957B2E3F918" bold="true" box="[151,232,1720,1744]" pageId="11" pageNumber="65">
<taxonomicNameLabel id="A223797DFF8BFFB9778CF957B2E3F918" box="[151,232,1720,1744]" pageId="11" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (J) macropterous ♀, (K) brachypterous ♀, (L) nymph, (M) brachypterous ♂ and nymph, (N) habitat; O,
<taxonomicName id="4C646397FF8BFFB97203F957B773F918" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[1304,1400,1720,1744]" class="Insecta" family="Aradidae" genus="Aradosyrtis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="65" phylum="Arthropoda" rank="species" species="shonoi" status="sp. nov.">
<emphasis id="B910C406FF8BFFB97203F957B72CF907" box="[1304,1319,1720,1743]" italics="true" pageId="11" pageNumber="65">A</emphasis>
.
<emphasis id="B910C406FF8BFFB9722FF957B773F918" box="[1332,1400,1720,1744]" italics="true" pageId="11" pageNumber="65">shonoi</emphasis>
</taxonomicName>
<emphasis id="B910C406FF8BFFB97265F957B2C8F93C" bold="true" pageId="11" pageNumber="65">
<taxonomicNameLabel id="A223797DFF8BFFB97265F957B2C8F93C" pageId="11" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ (photographed by Jun Souma).
</paragraph>
</caption>
<subSubSection id="C37E4B9FFF8BFFB977DCF8A9B65FF804" pageId="11" pageNumber="65" type="diagnosis">
<paragraph id="8BDB1814FF8BFFB977DCF8A9B65FF804" blockId="11.[151,1436,1862,1997]" pageId="11" pageNumber="65">
<emphasis id="B910C406FF8BFFB977DCF8A9B349F8A8" bold="true" box="[199,322,1862,1888]" pageId="11" pageNumber="65">Diagnosis.</emphasis>
Recognized and distinguished from Palaearctic congeners and
<taxonomicName id="4C646397FF8BFFB97315F8A9B716F8A8" box="[1038,1309,1862,1888]" pageId="11" pageNumber="65">
<emphasis id="B910C406FF8BFFB97315F8A9B6CFF8A8" box="[1038,1220,1862,1888]" italics="true" pageId="11" pageNumber="65">Nipponocalisius</emphasis>
species
</taxonomicName>
by a set of the following characters: clypeus apically rounded without dilated genae, antennal segments I–III of equal length, carinae of pronotum, lateral margins of scutellum and median ridge granulate without distinct tubercles, lateral margins of deltg II–VII with flat nodules and the specific colour pattern of scutellum.
</paragraph>
</subSubSection>
<caption id="DF1B489CFF8CFFBE778CFBCCB367FBA8" ID-DOI="http://doi.org/10.5281/zenodo.10898536" ID-Zenodo-Dep="10898536" httpUri="https://zenodo.org/record/10898536/files/figure.png" pageId="12" pageNumber="66" startId="12.[151,250,1059,1083]" targetBox="[155,1430,185,1012]" targetPageId="12" targetType="figure">
<paragraph id="8BDB1814FF8CFFBE778CFBCCB367FBA8" blockId="12.[151,1436,1059,1120]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE778CFBCCB314FBF4" bold="true" box="[151,287,1059,1084]" pageId="12" pageNumber="66">FIGURE 34.</emphasis>
Distribution map of Japanese
<emphasis id="B910C406FF8CFFBE7554FBCBB0FFFBF4" box="[591,756,1060,1084]" italics="true" pageId="12" pageNumber="66">Nippponcalisius</emphasis>
and
<taxonomicName id="4C646397FF8CFFBE7438FBCBB1E4FBF4" box="[803,1007,1060,1084]" class="Insecta" family="Aradidae" genus="Aradosyrtis" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="66" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B910C406FF8CFFBE7438FBCBB192FBF4" box="[803,921,1060,1084]" italics="true" pageId="12" pageNumber="66">Aradosyrtis</emphasis>
species.
</taxonomicName>
Red icons = holotype localities; blue icons = paratype localities.
</paragraph>
</caption>
<subSubSection id="C37E4B9FFF8CFFBE77DCFB5DB119F968" pageId="12" pageNumber="66" type="description">
<paragraph id="8BDB1814FF8CFFBE77DCFB5DB3BDFB38" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCFB5DB352FB04" bold="true" box="[199,345,1202,1228]" pageId="12" pageNumber="66">Description.</emphasis>
Macropterous male. Colour stramineous with brown markings on deltg II–VII, color pattern of scutellum as in
<figureCitation id="135F0491FF8CFFBE765DFB39B3BAFB38" box="[326,433,1238,1264]" captionStart="FIGURES 1–10" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[170,1419,193,1895]" captionTargetId="figure-17@2.[158,1429,181,1918]" captionTargetPageId="2" captionText="FIGURES 1–10. Japanese Nipponocalisius and Aradosyrtis species, dorsal habitus. 1–2, N. ishikawanus sp. nov., (1) ♂, (2) ♀; 3–5, N. tomokunii sp. nov., (3–4) ♂, (5) ♀; 6–8, N. dimorphus sp. nov., (6) brachypterous ♂, (7) brachypterous ♀, (8) macropterous ♀; 9–10, A. shonoi sp. nov., (9) ♂, (10) ♀" figureDoi="http://doi.org/10.5281/zenodo.10898522" httpUri="https://zenodo.org/record/10898522/files/figure.png" pageId="12" pageNumber="66">Figs. 6–7</figureCitation>
.
</paragraph>
<paragraph id="8BDB1814FF8CFFBE77DCFB15B773FAF0" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCFB15B300FADC" box="[199,267,1274,1300]" italics="true" pageId="12" pageNumber="66">Head.</emphasis>
Slightly longer than wide (22:19), clypeus granulate and rounded anteriorly; antennae 0.89 times as long as width of head, length of antennal segments I:II:III:IV = 4:4:4:7; vertex with a double row of round tubercles.
</paragraph>
<paragraph id="8BDB1814FF8CFFBE77DCFAACB716FA48" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCFAACB335FA94" box="[199,318,1347,1372]" italics="true" pageId="12" pageNumber="66">Pronotum.</emphasis>
About 1.5 times as wide as long (34:19); lateral margins sinuate, beset with 4 larger tubercles on anterior lobe and a few smaller ones on posterior lobe; disk with two pairs of carinae without tubercles.
</paragraph>
<paragraph id="8BDB1814FF8CFFBE77DCFA65B1B0FA00" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCFA65B336FA6C" box="[199,317,1418,1444]" italics="true" pageId="12" pageNumber="66">Scutellum.</emphasis>
Basal triangular elevation with three pairs of tubercles, four of them overlapping pronotum; median carina and lateral margins granulate, surface of disk with flat punctures.
</paragraph>
<paragraph id="8BDB1814FF8CFFBE77DCFA3DB61CF9D8" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCFA3DB332FA24" box="[199,313,1490,1516]" italics="true" pageId="12" pageNumber="66">Abdomen.</emphasis>
Lateral margins of deltg II–VII with a double row of three flat, round nodules, posterior ones stramineous, anterior ones dark brown; spiracles II–VI ventral, VII–VIII lateral.
</paragraph>
<paragraph id="8BDB1814FF8CFFBE77DCF9F5B08EF9FC" blockId="12.[151,1437,1202,2021]" box="[199,645,1562,1588]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCF9F5B332F9FC" box="[199,313,1562,1588]" italics="true" pageId="12" pageNumber="66">Genitalia.</emphasis>
Paramere blade as in
<figureCitation id="135F0491FF8CFFBE7535F9F5B08AF9FC" box="[558,641,1562,1588]" captionStart="FIGURES 26–29" captionStartId="8.[151,264,1624,1648]" captionTargetBox="[154,1429,745,1597]" captionTargetId="figure-291@8.[151,1436,741,1601]" captionTargetPageId="8" captionText="FIGURES 26–29. Japanese Nipponocalisius and Aradosyrtis species, left paramere, (a) dorsal view and (b) ventral view. 26, N. ishikawanus sp. nov.; 27, N. tomokunii sp. nov., 28, N. dimorphus sp. nov., 29, A. shonoi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.10898530" httpUri="https://zenodo.org/record/10898530/files/figure.png" pageId="12" pageNumber="66">Fig. 29</figureCitation>
.
</paragraph>
<paragraph id="8BDB1814FF8CFFBE77DCF9D1B161F990" blockId="12.[151,1437,1202,2021]" box="[199,874,1598,1624]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCF9D1B328F990" box="[199,291,1598,1624]" italics="true" pageId="12" pageNumber="66">Female.</emphasis>
As male but of larger size and different tergite VIII.
</paragraph>
<paragraph id="8BDB1814FF8CFFBE77DCF98CB119F968" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCF98CB37AF9B4" box="[199,369,1635,1660]" italics="true" pageId="12" pageNumber="66">Measurements.</emphasis>
<specimenCount id="9D62D39DFF8CFFBE7662F98DB3A8F9B4" box="[377,419,1634,1660]" count="1" pageId="12" pageNumber="66" type="male">♂♂</specimenCount>
(n = 5): body length
<quantity id="4C9CB5F1FF8CFFBE758CF98DB136F9B4" box="[663,829,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="12" pageNumber="66" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.40–2.50 mm</quantity>
, width of abdomen 1.00–
<quantity id="4C9CB5F1FF8CFFBE737BF98DB6CCF9B4" box="[1120,1223,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="66" unit="mm" value="1.15">1.15 mm</quantity>
;
<specimenCount id="9D62D39DFF8CFFBE73CFF98DB6F5F9B4" box="[1236,1278,1634,1660]" count="1" pageId="12" pageNumber="66" type="female">♀♀</specimenCount>
(n = 5): body length
<quantity id="4C9CB5F1FF8CFFBE77FFF969B382F968" box="[228,393,1670,1697]" metricMagnitude="-3" metricUnit="m" metricValue="2.725" metricValueMax="2.85" metricValueMin="2.6" pageId="12" pageNumber="66" unit="mm" value="2.725" valueMax="2.85" valueMin="2.6">2.60–2.85 mm</quantity>
, width of abdomen
<quantity id="4C9CB5F1FF8CFFBE7573F969B105F968" box="[616,782,1670,1697]" metricMagnitude="-3" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" pageId="12" pageNumber="66" unit="mm" value="1.175" valueMax="1.25" valueMin="1.1">1.10–1.25 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E4B9FFF8CFFBE77DCF945B021F920" pageId="12" pageNumber="66" type="etymology">
<paragraph id="8BDB1814FF8CFFBE77DCF945B021F920" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCF945B345F90C" bold="true" box="[199,334,1706,1732]" pageId="12" pageNumber="66">Etymology.</emphasis>
The specific epithet is dedicated to our friend Yoshinori Shono, recognizing his valuable advice and activities concerning this paper.
</paragraph>
</subSubSection>
<subSubSection id="C37E4B9FFF8CFFBE77DCF91DB31EF8F8" pageId="12" pageNumber="66" type="biology_ecology">
<paragraph id="8BDB1814FF8CFFBE77DCF91DB31EF8F8" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCF91DB327F8C4" bold="true" box="[199,300,1778,1804]" pageId="12" pageNumber="66">Ecology.</emphasis>
A male specimen was collected on
<taxonomicName id="4C646397FF8CFFBE75DDF91DB608F8C4" authority="Koidz." authorityName="Koidz." box="[710,1027,1778,1804]" class="Magnoliopsida" family="Oleaceae" genus="Osmanthus" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="66" phylum="Tracheophyta" rank="species" species="insularis">
<emphasis id="B910C406FF8CFFBE75DDF91DB1A4F8C4" box="[710,943,1778,1804]" italics="true" pageId="12" pageNumber="66">Osmanthus insularis</emphasis>
Koidz.
</taxonomicName>
(
<taxonomicName id="4C646397FF8CFFBE7308F91DB676F8C4" authorityName="von Hoffmannsegg &amp; Link" authorityYear="1809" box="[1043,1149,1778,1804]" class="Magnoliopsida" family="Oleaceae" kingdom="Plantae" order="Lamiales" pageId="12" pageNumber="66" phylum="Tracheophyta" rank="family">Oleaceae</taxonomicName>
) (pers. comm. J. Souma) (
<figureCitation id="135F0491FF8CFFBE7784F8F9B303F8F8" box="[159,264,1814,1840]" captionStart="FIGURE 33" captionStartId="11.[151,250,1648,1672]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-18@11.[151,1436,181,1623]" captionTargetPageId="11" captionText="FIGURE 33. Living individual and habitat of Japanese Nipponocalisius and Aradosyrtis species.A–C, N. ishikawanus sp. nov., (A) ♂, (B) ♀, (C) nymph; D–I, N. tomokunii sp. nov., (D–E) ♂, (F) egg and nymph, (G–H) ♀, (I) nymph; J–N, N. dimorphus sp. nov., (J) macropterous ♀, (K) brachypterous ♀, (L) nymph, (M) brachypterous ♂ and nymph, (N) habitat; O, A. shonoi sp. nov., ♂ (photographed by Jun Souma)." figureDoi="http://doi.org/10.5281/zenodo.10898534" httpUri="https://zenodo.org/record/10898534/files/figure.png" pageId="12" pageNumber="66">Fig. 33O</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E4B9FFF8CFFBE77DCF8D5B30BF8B0" pageId="12" pageNumber="66" type="distribution">
<paragraph id="8BDB1814FF8CFFBE77DCF8D5B30BF8B0" blockId="12.[151,1437,1202,2021]" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCF8D5B36AF89C" bold="true" box="[199,353,1850,1876]" pageId="12" pageNumber="66">Distribution.</emphasis>
Recorded from Ishigaki and Iriomote Islands in the Ryukyu Archipelago, supposed to be endemic (
<figureCitation id="135F0491FF8CFFBE7784F8B1B2FFF8B0" box="[159,244,1886,1912]" captionStart="FIGURE 34" captionStartId="12.[151,250,1059,1083]" captionTargetBox="[155,1430,185,1012]" captionTargetId="figure-17@12.[151,1435,181,1036]" captionTargetPageId="12" captionText="FIGURE 34. Distribution map of Japanese Nippponcalisius and Aradosyrtis species. Red icons = holotype localities; blue icons = paratype localities." figureDoi="http://doi.org/10.5281/zenodo.10898536" httpUri="https://zenodo.org/record/10898536/files/figure.png" pageId="12" pageNumber="66">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37E4B9FFF8CFFBF77DCF86DB3DAFED5" lastPageId="13" lastPageNumber="67" pageId="12" pageNumber="66" type="discussion">
<paragraph id="8BDB1814FF8CFFBF77DCF86DB3DAFED5" blockId="12.[151,1437,1202,2021]" lastBlockId="13.[151,1436,151,285]" lastPageId="13" lastPageNumber="67" pageId="12" pageNumber="66">
<emphasis id="B910C406FF8CFFBE77DCF86DB330F854" bold="true" box="[199,315,1922,1948]" pageId="12" pageNumber="66">Remarks.</emphasis>
It is remarkable two sympatric species of
<taxonomicName id="4C646397FF8CFFBE7404F86DB184F854" authorityName="Stal" authorityYear="1873" box="[799,911,1922,1948]" class="Insecta" family="Aradidae" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="66" phylum="Arthropoda" rank="subFamily" subFamily="Calisiinae">Calisiinae</taxonomicName>
occur on such rather small islands as Ishigaki and Iriomote of the Yaeyama group, about
<quantity id="4C9CB5F1FF8CFFBE759BF849B0DDF808" box="[640,726,1958,1984]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="66" unit="km" value="400.0">400 km</quantity>
southwest of the Okinawa Island. A female of
<taxonomicName id="4C646397FF8CFFBE73F4F849B3C3F82D" authority="Matsuda &amp; Usinger, 1957" authorityName="Matsuda &amp; Usinger" authorityYear="1957" class="Insecta" family="Aradidae" genus="Calisius" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="66" phylum="Arthropoda" rank="species" species="acutus">
<emphasis id="B910C406FF8CFFBE73F4F849B797F808" box="[1263,1436,1958,1984]" italics="true" pageId="12" pageNumber="66">Calisius acutus</emphasis>
Matsuda &amp; Usinger, 1957
</taxonomicName>
, a species described from the
<collectingCountry id="F3735884FF8CFFBE7434F825B1E7F82C" box="[815,1004,1994,2020]" name="Micronesia" pageId="12" pageNumber="66">Caroline Islands</collectingCountry>
, has been reported from Iriomote by Miyamoto &amp; Hidaka (1963). This record is likely a misidentification, pertaining either to this new species or
<taxonomicName id="4C646397FF8DFFBF778CFF54B3DBFF1D" authority="Heiss &amp; Shimamoto &amp; Nagashima, 2024" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[151,464,187,213]" class="Insecta" family="Aradidae" genus="Nipponocalisius" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="67" phylum="Arthropoda" rank="species" species="dimorphus" status="sp. nov.">
<emphasis id="B910C406FF8DFFBF778CFF54B3DBFF1D" box="[151,464,187,213]" italics="true" pageId="13" pageNumber="67">Nipponocalisius dimorphus</emphasis>
</taxonomicName>
<emphasis id="B910C406FF8DFFBF76C1FF54B03CFF1D" bold="true" box="[474,567,187,213]" pageId="13" pageNumber="67">
<taxonomicNameLabel id="A223797DFF8DFFBF76C1FF54B03CFF1D" box="[474,567,187,213]" pageId="13" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but in lack of the voucher specimens of Miyamoto &amp; Hidaka (1963) this question cannot be settled at the present. The unavailable name
<taxonomicName id="4C646397FF8DFFBF749AFF30B652FF31" authorityName="Heiss &amp; Shimamoto &amp; Nagashima" authorityYear="2024" box="[897,1113,223,249]" class="Insecta" family="Aradidae" genus="Aradosyrtis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="67" phylum="Arthropoda" rank="species" species="shonoi">
<emphasis id="B910C406FF8DFFBF749AFF30B652FF31" box="[897,1113,223,249]" italics="true" pageId="13" pageNumber="67">Aradosyrtis shonoi</emphasis>
</taxonomicName>
mentioned by
<bibRefCitation id="EFF565E5FF8DFFBF7212FF30B797FF31" author="Heiss, E." box="[1289,1436,223,249]" pageId="13" pageNumber="67" pagination="110 - 120" refId="ref8284" refString="Heiss, E. (2023) New taxonomic assignments for two Palaearctic Calisiinae genera (Heteroptera, Aradidae) with description of a new species from China. Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 75, 110 - 120." type="journal article" year="2023">Heiss (2023)</bibRefCitation>
pertains to this new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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