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<document ID-DOI="10.11646/zootaxa.5230.4.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7564103" ID-ZooBank="FA55F137-EF78-4ADB-B0CF-B9F84FFC86FD" approvalRequired="4" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="3" checkinTime="1674544415157" checkinUser="plazi" docAuthor="Lee, Seung-Gyu &amp; Ahn, Kee-Jeong" docDate="2023" docId="03C1E0117C0AFF8E14E7F9469B900155" docLanguage="en" docName="zootaxa.5230.4.5.pdf" docOrigin="Zootaxa 5230 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5230.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amischa bifoveolata" docType="treatment" docVersion="2" lastPageNumber="482" masterDocId="FFF898697C09FF8A1470FFE59E000504" masterDocTitle="Korean species of the genus Amischa Thomson (Coleoptera: Staphylinidae Aleocharinae), with a description of new species" masterLastPageNumber="488" masterPageNumber="478" pageNumber="481" updateTime="1674544840578" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Korean species of the genus Amischa Thomson (Coleoptera: Staphylinidae Aleocharinae), with a description of new species</mods:title>
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<mods:namePart>Lee, Seung-Gyu</mods:namePart>
<mods:affiliation>Animal Resources Division, National Institute of Biological Resources, Incheon 22689, South Korea.</mods:affiliation>
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<mods:namePart>Ahn, Kee-Jeong</mods:namePart>
<mods:affiliation>Department of Biology, Chungnam National University, Daejeon 34134, South Korea.</mods:affiliation>
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<taxonomicName ID-CoL="66BW8" authority="(Mannerheim, 1830)" baseAuthorityName="Mannerheim" baseAuthorityYear="1830" box="[151,661,1699,1726]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="bifoveolata">
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<emphasis bold="true" box="[151,398,1699,1726]" italics="true" pageId="3" pageNumber="481">Amischa bifoveolata</emphasis>
(
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<paragraph blockId="3.[151,1015,1804,2043]" box="[151,637,1804,1828]" pageId="3" pageNumber="481">
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<emphasis box="[151,393,1804,1828]" italics="true" pageId="3" pageNumber="481">Bolitochara bifoveolata</emphasis>
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<treatmentCitation author="Sharp, D. S." box="[151,535,1835,1859]" page="177" pageId="3" pageNumber="481" year="1869">
<taxonomicName ID-CoL="6LZR4" authority="Sharp, 1869: 177" authorityName="Sharp" authorityPageNumber="177" authorityYear="1869" box="[151,535,1835,1859]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis box="[151,352,1835,1859]" italics="true" pageId="3" pageNumber="481">Homalota cavifrons</emphasis>
<bibRefCitation author="Sharp, D. S." box="[359,535,1835,1859]" pageId="3" pageNumber="481" pagination="91 - 272" refId="ref5570" refString="Sharp, D. S. (1869) A revision of the British species of Homalota. The Transactions of the Entomological Society of London, 1869, 91 - 272. https: // doi. org / 10.1111 / j. 1365 - 2311.1869. tb 01102. x" type="journal article" year="1869">Sharp, 1869: 177</bibRefCitation>
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<treatmentCitation author="Mulsant, E. &amp; Rey, C." box="[151,590,1865,1890]" page="105" pageId="3" pageNumber="481" year="1870">
<taxonomicName ID-CoL="6LZFS" authority="Mulsant &amp; Rey, 1870: 105" authorityName="Mulsant &amp; Rey" authorityPageNumber="105" authorityYear="1870" box="[151,590,1865,1890]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="filum">
<emphasis box="[151,311,1865,1889]" italics="true" pageId="3" pageNumber="481">Homalota filum</emphasis>
<bibRefCitation author="Mulsant, E. &amp; Rey, C." box="[317,590,1865,1890]" pageId="3" pageNumber="481" pagination="105 - 122" refId="ref5212" refString="Mulsant, E. &amp; Rey, C. (1870) Description de diverses especes nouvelles de coleopteres. Opuscules Entomologiques, 14, 105 - 122." type="journal article" year="1870">Mulsant &amp; Rey, 1870: 105</bibRefCitation>
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<paragraph blockId="3.[151,1015,1804,2043]" box="[151,603,1896,1921]" pageId="3" pageNumber="481">
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<treatmentCitation author="Mulsant, E. &amp; Rey, C." box="[151,599,1896,1921]" page="179" pageId="3" pageNumber="481" year="1873">
<taxonomicName ID-CoL="CSN6" authority="Mulsant &amp; Rey, 1873: 179" authorityName="Mulsant &amp; Rey" authorityPageNumber="179" authorityYear="1873" box="[151,599,1896,1921]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="minima">
<emphasis box="[151,321,1896,1920]" italics="true" pageId="3" pageNumber="481">Amischa minima</emphasis>
<bibRefCitation author="Mulsant, E. &amp; Rey, C." box="[327,599,1896,1921]" pageId="3" pageNumber="481" pagination="147 - 189" refId="ref5241" refString="Mulsant, E. &amp; Rey, C. (1873) Description de divers coleopteres brevipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147 - 189." type="journal article" year="1873">Mulsant &amp; Rey, 1873: 179</bibRefCitation>
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<treatmentCitation author="Scheerpeltz, O." box="[151,540,1927,1951]" page="13" pageId="3" pageNumber="481" year="1967">
<taxonomicName ID-CoL="CSP6" authority="Scheerpeltz, 1967: 13" authorityName="Scheerpeltz" authorityPageNumber="13" authorityYear="1967" box="[151,540,1927,1951]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="strupii">
<emphasis box="[151,311,1927,1951]" italics="true" pageId="3" pageNumber="481">Amischa strupii</emphasis>
<bibRefCitation author="Scheerpeltz, O." box="[317,540,1927,1951]" pageId="3" pageNumber="481" pagination="13 - 15" refId="ref5468" refString="Scheerpeltz, O. (1967) Eine neue alpine Art der Gattung Amischa C. G. Thomson (Col. Staphylinidae). Entomologische Blatter, 63 (1), 13 - 15." type="journal article" year="1967">Scheerpeltz, 1967: 13</bibRefCitation>
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<paragraph blockId="3.[151,1015,1804,2043]" box="[151,514,1957,1982]" pageId="3" pageNumber="481">
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<treatmentCitation author="Benick, G." box="[151,509,1957,1982]" page="24" pageId="3" pageNumber="481" year="1967">
<taxonomicName ID-CoL="CSPB" authority="G. Benick, 1967: 24" authorityName="G. Benick" authorityPageNumber="24" authorityYear="1967" box="[151,509,1957,1982]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="vogti">
<emphasis box="[151,296,1957,1981]" italics="true" pageId="3" pageNumber="481">Amischa vogti</emphasis>
G.
<bibRefCitation author="Benick, G." box="[333,509,1957,1982]" pageId="3" pageNumber="481" pagination="16 - 29" refId="ref4705" refString="Benick, G. (1967) Die palaearktischen Arten der Gattung Amischa C. G. Thomson (Col. Staph.). Entomologische Blatter, 63, 16 - 29." type="journal article" year="1967">Benick, 1967: 24</bibRefCitation>
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<paragraph blockId="3.[151,1015,1804,2043]" box="[151,1015,1988,2012]" pageId="3" pageNumber="481">
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<treatmentCitation author="Palm, T." box="[151,521,1988,2012]" page="101" pageId="3" pageNumber="481" year="1968">
<taxonomicName ID-CoL="6LZR4" authority=": Palm, 1968: 101" authorityName="Palm" authorityPageNumber="101" authorityYear="1968" box="[151,521,1988,2012]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis box="[151,338,1988,2012]" italics="true" pageId="3" pageNumber="481">Amischa cavifrons</emphasis>
:
<bibRefCitation author="Palm, T." box="[351,521,1988,2012]" pageId="3" pageNumber="481" refId="ref5340" refString="Palm, T. (1968) Svensk Insektfauna: Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Underfam. Aleocharinae (De nopsis-Trichomicra). Hafte 5. Entomologiska Foreningen I, Stockholm, 9 + 112 pp." type="book" year="1968">Palm, 1968: 101</bibRefCitation>
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<bibRefCitation author="Benick, G. &amp; Lohse, G. A." box="[532,815,1988,2012]" pageId="3" pageNumber="481" pagination="72 - 220" refId="ref4739" refString="Benick, G. &amp; Lohse, G. A. (1974) [Tribus 14 (Callicerini)]. In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5. Staphylinidae II (Hypocyptinae und Aleocharinae), Pselaphidae. Goecke and Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse, 1974: 101</bibRefCitation>
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;
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<bibRefCitation author="Pasnik, G." box="[826,1011,1988,2012]" pageId="3" pageNumber="481" pagination="185 - 234" refId="ref5382" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoological Cracoviensia, 44, 185 - 234." type="journal article" year="2001">Paśnik, 2001: 198</bibRefCitation>
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<treatmentCitationGroup box="[151,480,2019,2043]" pageId="3" pageNumber="481">
<treatmentCitation author="Pace, R." box="[151,476,2019,2043]" page="280" pageId="3" pageNumber="481" year="1987">
<taxonomicName authority="Pace, 1987: 280" authorityName="Pace" authorityPageNumber="280" authorityYear="1987" box="[151,476,2019,2043]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="481" phylum="Arthropoda" rank="species" species="uhligi">
<emphasis box="[151,304,2019,2043]" italics="true" pageId="3" pageNumber="481">Amischa uhligi</emphasis>
<bibRefCitation author="Pace, R." box="[310,476,2019,2043]" pageId="3" pageNumber="481" pagination="279 - 283" refId="ref5309" refString="Pace, R. (1987) Nuove specie di Aleocharinae orofile della Bulgaria (Coleoptera, Staphylinidae). Giornale Italiano di Entomologia, 3, 279 - 283." type="journal article" year="1987">Pace, 1987: 280</bibRefCitation>
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<treatmentCitation author="Smetana, A." box="[151,688,151,175]" page="370" pageId="4" pageNumber="482" year="2004">
<taxonomicName ID-CoL="66BW8" authority=": Smetana, 2004: 370" authorityName="Smetana" authorityPageNumber="370" authorityYear="2004" box="[151,688,151,175]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="species" species="bifoveolata" subGenus="Amischa">
<emphasis box="[151,238,151,175]" italics="true" pageId="4" pageNumber="482">Amischa</emphasis>
(
<emphasis box="[253,340,151,175]" italics="true" pageId="4" pageNumber="482">Amischa</emphasis>
)
<emphasis box="[355,469,151,175]" italics="true" pageId="4" pageNumber="482">bifoveolata</emphasis>
:
<bibRefCitation author="Smetana, A." box="[483,688,151,175]" pageId="4" pageNumber="482" pagination="504 - 698" refId="ref5622" refString="Smetana, A. (2004) Family Saphylinidae: Omaliinae, Proteininae, Micropeplinae, Dasycerinae, Phleocharinae, Olisthaerinae, Tachyporinae, Trichophyinae, Habrocerinae, Aleocharinae, Trigonurinae, Apaticinae, Piestinae, Osoriinae, Oxytelinae, Oxyporinae, Megalopsiinae, Steninae, Euaesthetinae, Leptotyphlinae, Pseudopsinae, Paederinae, Staphylininae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, pp. 237 - 272 + 329 - 495 + 504 - 698." type="book chapter" year="2004">Smetana, 2004: 370</bibRefCitation>
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<bibRefCitation author="Schulke, M. &amp; Smetana, A." box="[698,1017,150,175]" pageId="4" pageNumber="482" pagination="304 - 1134" refId="ref5505" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae Latreille, 1802. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2 / 1 - 2. Revised and updated edition. Hydrophiloidea - Staphylinoidea. Brill, Leiden, pp. 304 - 1134." type="book chapter" year="2015">Schülke &amp; Smetana, 2015: 512</bibRefCitation>
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<treatmentCitation author="Assing, V." box="[151,539,181,205]" page="33" pageId="4" pageNumber="482" year="2021">
<taxonomicName ID-CoL="66BW8" authority=": Assing 2021: 33" authorityName=": Assing" authorityPageNumber="33" authorityYear="2021" box="[151,539,181,205]" class="Insecta" family="Staphylinidae" genus="Amischa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="482" phylum="Arthropoda" rank="species" species="bifoveolata">
<emphasis box="[151,359,181,205]" italics="true" pageId="4" pageNumber="482">Amischa bifoveolata</emphasis>
:
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<emphasis bold="true" box="[151,289,251,277]" pageId="4" pageNumber="482">Description</emphasis>
. Length about 2.0 mm. Body (
<figureCitation box="[640,726,251,278]" captionStart="FIGURE 1" captionStartId="2.[152,255,2016,2040]" captionTargetBox="[307,1285,1116,1961]" captionTargetId="figure-453@2.[223,1365,1101,1993]" captionTargetPageId="2" captionText="FIGURE 1. Habitus. A: Amischa analis (1.9 mm); B: A. bifoveolata (2.0 mm); C: A. koreana sp. nov. (1.8 mm)." figureDoi="http://doi.org/10.5281/zenodo.7564107" httpUri="https://zenodo.org/record/7564107/files/figure.png" pageId="4" pageNumber="482">Fig. 1B</figureCitation>
) subparallel-sided; surface glossy and densely pubescent, with fine microsculpture. Body reddish brown; head and abdomen darker than other parts.
<emphasis box="[1112,1173,287,313]" italics="true" pageId="4" pageNumber="482">Head</emphasis>
. Subtriangular, slightly transverse, about 1.2 times as wide as long, widest behind eyes, slightly narrower than pronotum; vertex depressed; eyes relatively small and slightly prominent, about 0.8 times as long as temples; gular sutures moderately separated, diverged basally; infraorbital carina absent. Antennae relatively short and more or less dilated apically; antennomeres 13 elongate, 1 about as long as 11, 2 longer than 3, 45 slightly transverse, 610 transverse, 11 longer than wide and about as long as preceding two combined.
<emphasis box="[732,813,467,493]" italics="true" pageId="4" pageNumber="482">Thorax</emphasis>
. Pronotum slightly transverse and depressed in posteromedian region, approximately 1.2 times as wide as long, widest in about middle; hypomeron partially visible in lateral aspect. Elytra slightly transverse, slightly wider than pronotum, approximately 1.71.8 times as long as wide, pubescence directed posteriorly and postero-laterally, postero-lateral margin sinuate; hind wings fully developed. Legs slender and long, with pubescence and macrosetae; tibiae with two spurs at apex; tarsal formula 4-5-5; hind tarsomere 1 about as long as 2.
<emphasis box="[512,618,647,673]" italics="true" pageId="4" pageNumber="482">Abdomen</emphasis>
. Parallel-sided, widest at middle; surface glossy and densely pubescent; tergites IIIV impressed in basal region; male tergite VIII with 4 long macrosetae on each side of midline, posterior margin subtriangularly emarginate; male sternite VIII with about 4 long macrosetae on each side of midline, posterior margin truncate in median region, with 78 marginal setae conspicuously; female tergite VIII similar to male; posterior margin of female sternite VIII truncate in median region with conspicuous and long marginal setae.
<emphasis italics="true" pageId="4" pageNumber="482">Aedeagus</emphasis>
. Median lobe ovate; apical process long and slender, pointed apically in lateral aspect; triangular and pointed apically in ventral aspect.
<emphasis box="[444,590,863,889]" italics="true" pageId="4" pageNumber="482">Spermatheca</emphasis>
.
<collectingRegion box="[604,669,864,890]" country="Turkey" name="Bursa" pageId="4" pageNumber="482">Bursa</collectingRegion>
elongate, extended in apical half; duct coiled apically.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="482" type="materials_examined">
<paragraph blockId="4.[151,1437,251,1106]" pageId="4" pageNumber="482">
<emphasis bold="true" box="[199,436,899,925]" pageId="4" pageNumber="482">Material examined.</emphasis>
<materialsCitation collectionCode="ISEA" collectorName="Myohyangsan Mts. Exp. Inst. Zool. Cr. &amp; Thesongho A. Szeptycki &amp; Taeseongho-lake &amp; A. Szeptycki" country="North Korea" location="Myohyangsan Mts." municipality="Mts. Exp. Inst. Zool. Cr." pageId="4" pageNumber="482" specimenCount="1">
4 exx., Corea spet. 1983
<collectorName box="[741,1176,899,926]" pageId="4" pageNumber="482">
<location LSID="urn:lsid:plazi:treatment:03C1E0117C0AFF8E14E7F9469B900155:8EB707DC7C0DFF8E1695FC669DC1069A" box="[741,961,899,926]" country="North Korea" municipality="Mts. Exp. Inst. Zool. Cr." name="Myohyangsan Mts." pageId="4" pageNumber="482">
Myohyangsan
<collectingMunicipality box="[910,961,900,926]" pageId="4" pageNumber="482">Mts.</collectingMunicipality>
</location>
Exp. Inst.Zool.Cr.
</collectorName>
[
<collectingCountry box="[1192,1342,899,926]" name="North Korea" pageId="4" pageNumber="482">North Korea</collectingCountry>
, Pyeon- ganbuk Prov., Mt. Myohyangsan, 1983 [
<collectionCode box="[605,675,936,962]" pageId="4" pageNumber="482">ISEA</collectionCode>
];
<specimenCount box="[694,751,936,962]" pageId="4" pageNumber="482" type="generic">1 ex.</specimenCount>
, Korea 1981
<collectorName box="[902,1177,935,962]" pageId="4" pageNumber="482">Thesongho A. Szeptycki</collectorName>
[
<collectingCountry box="[1192,1266,936,962]" name="South Korea" pageId="4" pageNumber="482">Korea</collectingCountry>
, Pyeongannam Prov.,
<collectorName box="[223,415,971,997]" pageId="4" pageNumber="482">Taeseongho-lake</collectorName>
, 12 vi l981,
<collectorName box="[552,703,971,998]" pageId="4" pageNumber="482">A. Szeptycki</collectorName>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[199,1132,1007,1034]" pageId="4" pageNumber="482" type="distribution">
<paragraph blockId="4.[151,1437,251,1106]" box="[199,1132,1007,1034]" pageId="4" pageNumber="482">
<emphasis bold="true" box="[199,353,1007,1033]" pageId="4" pageNumber="482">Distribution.</emphasis>
<collectingCountry box="[359,427,1008,1034]" name="South Korea" pageId="4" pageNumber="482">Korea</collectingCountry>
(North),
<collectingCountry box="[533,608,1007,1033]" name="Russia" pageId="4" pageNumber="482">Russia</collectingCountry>
(East Siberia, Far East, West Siberia), Europe.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="482" type="discussion">
<paragraph blockId="4.[151,1437,251,1106]" pageId="4" pageNumber="482">
<emphasis bold="true" box="[199,315,1044,1070]" pageId="4" pageNumber="482">Remarks.</emphasis>
This species was recorded by
<bibRefCitation author="Pasnik, G." box="[659,815,1043,1070]" pageId="4" pageNumber="482" pagination="185 - 234" refId="ref5382" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta Zoological Cracoviensia, 44, 185 - 234." type="journal article" year="2001">Paśnik (2001)</bibRefCitation>
from
<collectingCountry box="[884,1028,1043,1070]" name="North Korea" pageId="4" pageNumber="482">North Korea</collectingCountry>
, but no specimen has been collected in
<collectingCountry box="[180,324,1079,1106]" name="South Korea" pageId="4" pageNumber="482">South Korea</collectingCountry>
. We could not describe the mouthparts in detail because no specimen was available for dissection.
</paragraph>
</subSubSection>
</treatment>
</document>