treatments-xml/data/03/B7/B5/03B7B5405E2AC173FF4BFA6AFE1FF8E0.xml

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<mods:title id="2F4B04591D07D82A051CD3AF6CD9E2FF">A new species of Varronia P. Browne (Cordiaceae) from central Colombia</mods:title>
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<mods:namePart id="C0521ADF56D6C0514F712574CC7B6136">Fernández-Alonso, Jose Luis</mods:namePart>
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<mods:affiliation id="1CC0A3650FBF7DEBCCF98F7A3388ED62">Departamento de Biodiversidad y Conservación, Real Jardín Botánico CSIC, Claudio Moyano 1, 28014 - Madrid, Spain jlfernandeza @ rjb. csic. es; https: // orcid. org / 0000 - 0002 - 1701 - 480 X</mods:affiliation>
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<mods:namePart id="FF4AC87796DDD43DE1C329CACD4480D8">Melo, José Iranildo Miranda De</mods:namePart>
<mods:affiliation id="DD737E48FD2AC9C2EBF0E67A99BA330A">Centro de Ciências Biológicas e da Saúde, Departamento de Biologia, Universidade Estadual da Paraíba, Rua das Baraúnas 351, Bairro Universitário, CEP 58429 - 500 - Campina Grande, Paraíba, Brazil</mods:affiliation>
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<paragraph id="8BA104565E2AC174FF4BFA6AFC8BFA74" blockId="1.[136,778,1468,1495]" box="[136,778,1468,1495]" pageId="1" pageNumber="122">
<heading id="D0E9B33A5E2AC174FF4BFA6AFC8BFA74" box="[136,778,1468,1495]" fontSize="11" level="2" pageId="1" pageNumber="122" reason="3">
<taxonomicName id="4C1E7FD55E2AC174FF4BFA6AFD29FA74" ID-CoL="8RTBC" authority="Fern. Alonso &amp; J.I.M. Melo" authorityName="Fern. Alonso &amp; J. I. M. Melo" authorityYear="2021" box="[136,680,1468,1495]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="1" pageNumber="122" phylum="Tracheophyta" rank="species" species="teguorum" status="sp. nov.">
<emphasis id="B96AD8445E2AC174FF4BFA6AFEE7FA74" bold="true" box="[136,358,1468,1494]" italics="true" pageId="1" pageNumber="122">Varronia teguorum</emphasis>
Fern. Alonso &amp; J.I.M. Melo
</taxonomicName>
, 
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<taxonomicNameLabel id="A259653F5E2AC174FD70FA6AFC8BFA74" box="[691,778,1468,1494]" pageId="1" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C30457DD5E2AC174FF4BFA36FF7DFA58" box="[136,252,1504,1530]" pageId="1" pageNumber="122" type="description">
<paragraph id="8BA104565E2AC174FF4BFA36FF7DFA58" blockId="1.[136,252,1504,1530]" box="[136,252,1504,1530]" pageId="1" pageNumber="122">
(
<figureCitation id="132518D35E2AC174FF53FA36FF72FA58" box="[144,243,1504,1530]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[136,229,1953,1975]" captionStartId-1="4.[136,229,1919,1941]" captionStartId-2="5.[136,229,1802,1824]" captionTargetBox-0="[226,1365,198,1921]" captionTargetBox-1="[243,1344,190,1895]" captionTargetBox-2="[270,1317,190,1778]" captionTargetId-0="figure-18@3.[218,1369,190,1929]" captionTargetId-1="figure-26@4.[243,1344,190,1895]" captionTargetId-2="figure-18@5.[270,1317,190,1778]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Image of the isotype of Varronia teguorum Fern.Alonso &amp; J.I.M. Melo, Fernández-Alonso 24079 (MA 888445)." captionText-1="FIGURE 2. Field images of Varronia teguorum. A.Apex of a young branch, showing the rusty indumentum and the tertiary and quaternary veins on the leaf adaxial surface. B. Branch with developed axillary inflorescences, showing floral buds. C. Branch with an inflorescence. D. Inflorescences with some flowers at anthesis (white corolla) and others at post-anthesis (senescent corolla), and immature fruits (red). E. Inflorescences with an open flower at a basal position, showing the senescent flowers with accrescent calyx and incurved lobes obscuring the calyx throat. Scale bar in all cases = 15 mm long. Photos by J.L. Fernández-Alonso taken in the type locality, Santa María, Boyacá." captionText-2="FIGURE 3. Light micrographs of a dried specimen of Varronia teguorum. A. Insertion of the petiole in the branch, the base of the leaf blade, and the axillary bud. B. Adaxial surface of leaf blade: impressed venation and indumentum with conspicuous long hairs along the veins and cystolithic pustules. C. Abaxial surface of leaf blade showing prominent reticulate venation (2º, 3º, and 4º orders) and hairs with cystolithic pustules. D. Slightly adnate region of the petiole (right) with the inflorescence axis (left). E. Young inflorescence, showing the indumentum of the peduncle and the incurved appendages of the calyx lobes. F. Inflorescence rachis, showing its dense indumentum and the flower arrangement. G. Inflorescence with flowers at anthesis: G1. Two open flowers showing the corolla lobes and some dehisced anthers; G2. Open flower, showing the glabrous anthers covering the corolla throat. Scale bar in all cases = 5 mm long. (Voucher: Fernández-Alonso 24079, COL). Photos by J.L. Fernández-Alonso." figureDoi-0="http://doi.org/10.5281/zenodo.5425569" figureDoi-1="http://doi.org/10.5281/zenodo.5425571" figureDoi-2="http://doi.org/10.5281/zenodo.5425573" httpUri-0="https://zenodo.org/record/5425569/files/figure.png" httpUri-1="https://zenodo.org/record/5425571/files/figure.png" httpUri-2="https://zenodo.org/record/5425573/files/figure.png" pageId="1" pageNumber="122">Figs 1–3</figureCitation>
)
</paragraph>
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<emphasis id="B96AD8445E2AC174FF4BF9F3FF49F998" bold="true" box="[136,200,1573,1594]" pageId="1" pageNumber="122">TYPE</emphasis>
: 
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<collectingCountry id="F30944C65E2AC174FF14F9F3FEE0F999" box="[215,353,1573,1595]" name="Colombia" pageId="1" pageNumber="122">COLOMBIA</collectingCountry>
. 
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. 
<collectingRegion id="49DACAB45E2AC174FE6EF9F2FE79F998" box="[429,504,1572,1594]" country="Colombia" name="Boyaca" pageId="1" pageNumber="122">Boyacá</collectingRegion>
</emphasis>
. 
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<collectingMunicipality id="6BC59E2C5E2AC174FDC6F9F3FCD5F999" box="[517,852,1573,1595]" pageId="1" pageNumber="122">Vertiente Oriental de la Cordillera</collectingMunicipality>
</collectorName>
oriental. Mpio. de 
<taxonomicName id="4C1E7FD55E2AC174FBD7F9F3FA9CF998" authority="Maria. Canon Negro" authorityName="Maria. Canon Negro" box="[1044,1309,1573,1595]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="122" phylum="Arthropoda" rank="genus">Santa María. Cañón Negro</taxonomicName>
, vía 
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, 
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, 
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, fl., fr., 
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<collectorName id="26EB61805E2AC174FD68F99CFC0BF9FC" box="[683,906,1609,1631]" pageId="1" pageNumber="122">J.L. Fernández-Alonso</collectorName>
&amp; al. 24079
</emphasis>
[
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: COL 537711!; isotypes (to be distributed): COL 557483!, G!, K!, MA (3 sheets: 888443!, 888444!, 888445!), MEDEL!, MO!, NY!, P!, 
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!,
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!].
</paragraph>
<paragraph id="8BA104565E2AC174FF4BF97CFDE7F88C" blockId="1.[136,1452,1706,2088]" pageId="1" pageNumber="122">
This new species is similar to 
<taxonomicName id="4C1E7FD55E2AC174FE26F97DFD56F966" authorityName="Friesen" authorityYear="1933" box="[485,727,1706,1732]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="1" pageNumber="122" phylum="Tracheophyta" rank="species" species="subtruncata">
<emphasis id="B96AD8445E2AC174FE26F97DFD56F966" box="[485,727,1706,1732]" italics="true" pageId="1" pageNumber="122">Varronia subtruncata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1E7FD55E2AC174FCD2F97DFCFCF966" authorityName="Linnaeus" authorityYear="1759" box="[785,893,1706,1732]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="1" pageNumber="122" phylum="Tracheophyta" rank="species" species="bullata">
<emphasis id="B96AD8445E2AC174FCD2F97DFCFCF966" box="[785,893,1706,1732]" italics="true" pageId="1" pageNumber="122">V. bullata</emphasis>
</taxonomicName>
, but it differs from these two by the presence of a rusty (
<emphasis id="B96AD8445E2AC174FF25F918FF7CF945" box="[230,253,1742,1767]" italics="true" pageId="1" pageNumber="122">vs</emphasis>
. white to yellow) indumentum covering the stems, leaves and inflorescences, longer (
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<emphasis id="B96AD8445E2AC174FA89F918FAE0F945" box="[1354,1377,1742,1767]" italics="true" pageId="1" pageNumber="122">vs</emphasis>
. &lt;
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long) and patent (
<emphasis id="B96AD8445E2AC174FE43F924FE16F8A9" box="[384,407,1778,1803]" italics="true" pageId="1" pageNumber="122">vs</emphasis>
. patent or adpressed) hairs; oblong to ovate-oblong (vs. lanceolate to ovate), larger (7.5–13 × 
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<emphasis id="B96AD8445E2AC174FECFF8C3FEA2F88C" box="[268,291,1813,1838]" italics="true" pageId="1" pageNumber="122">vs</emphasis>
. &lt;8.0 × 
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) leaf blades.
</paragraph>
<paragraph id="8BA104565E2AC177FF7EF8EEFEEAFCB1" blockId="1.[136,1452,1706,2088]" lastBlockId="2.[136,1452,158,2065]" lastPageId="2" lastPageNumber="123" pageId="1" pageNumber="122">
<emphasis id="B96AD8445E2AC174FF7EF8EEFE8EF8F1" bold="true" box="[189,271,1848,1875]" italics="true" pageId="1" pageNumber="122">Shrubs</emphasis>
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high, densely branched; branches sub-cylindric in cross-section, self-supporting, sometimes scrambling or vining, indumentum with eglandular hairs, densely hirsute, becoming denser towards the apex, hairs 
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long, simple, coarse, patent, rusty. 
<emphasis id="B96AD8445E2AC174FD44F8A9FD57F83B" bold="true" box="[647,726,1919,1945]" italics="true" pageId="1" pageNumber="122">Leaves</emphasis>
sparsely distributed along the branches, 
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apart from each other; petioles 
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long, slightly adnate at base, 
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long, hispid, hairs 
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long, rusty, enlarged at base; blades 7.5–13.0 × 
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, oblong to ovate-oblong (most rarely narrowly ovate), membranous to sub-chartaceous, asperous, drying blackish-brown, base cuneate to truncate, margins thick, irregularly dentate in upper 2/3, rarely dentate from base to apex, teeth short, 
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long, 
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apart from each other, apex acute to slightly acuminate, adaxially with long hairs, more or less adpressed, evenly distributed, with numerous cystolithic pustules; abaxially with hairs congested on primary veins, hispid, with numerous strigose hairs, sometimes presenting sessile papillae; midvein thick, 
<quantity id="4CE6A9B35E29C177FEF3FF33FE20FEA2" box="[304,417,229,256]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="123" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
wide at base, secondary veins 5–6 pairs, adaxially impressed-excavate, abaxially prominent; tertiary veins reticulate, generally conspicuous. 
<emphasis id="B96AD8445E29C177FD5AFEDEFCC0FE81" bold="true" box="[665,833,264,291]" italics="true" pageId="2" pageNumber="123">Inflorescences</emphasis>
axillary, pedunculated glomerules, peduncles cylindric, adnate at the base of the leaf 
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; 
<quantity id="4CE6A9B35E29C177FDF7FEFAFD31FEE5" box="[564,688,300,327]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="8.5" metricValueMin="2.5" pageId="2" pageNumber="123" unit="cm" value="5.5" valueMax="8.5" valueMin="2.5">2.5–8.5 cm</quantity>
long, ca. 
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wide, gradually thickened towards the apex; rarely inflorescences not associated with leaves; yellowish-green, with the same indumentum as the young branches, hairs long, drying dark brown; glomerules globose to narrowly ellipsoid, 
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diam., presenting ca. 60 flowers sessile, arranged in radiated series from base to apex; the residual inflorescence rachis (flowerless), cylindric, ca. 
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long. 
<emphasis id="B96AD8445E29C177FF4BFE6FFF64FE76" bold="true" box="[136,229,441,468]" italics="true" pageId="2" pageNumber="123">Flowers</emphasis>
bisexual, intensely fragrant, sessile; calyx slightly accrescent, 
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long (at the end of anthesis), urceolate, yellow, tube 
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long, turbinate-saccate, narrowing at apex, inconspicuously costate, externally densely pilose in the upper half, 5-toothed, teeth 
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long, incurved, covering the calyx throat, with a short appendage, up to 
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long, linear, with long, stiff, erect or patent hairs, internally glabrous; corolla 3.5–5 × 
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, slightly conspicuous, subcylindric, slightly campanulate, white to cream, externally pruinose, nitid, internally glabrous towards the apex, with short hairs at the throat, lobes scarcely distended, scarcely apart from each other; stamens included, shorter than the corolla tube, filaments short, densely pubescent at their insertion, anthers broadly cylindric; style bifurcated, branches bifurcated, stigmatic lobes filiform to clavate. 
<emphasis id="B96AD8445E29C177FCBFFD65FC51FD6F" bold="true" box="[892,976,691,717]" italics="true" pageId="2" pageNumber="123">Drupes</emphasis>
red, included in the persistent calyx when immature, 5–6 × 
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, ventricose, teeth 
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long, appendages 
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long; immature pyrenes ca. 2 × 
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, ovoid, bilobed.
</paragraph>
<paragraph id="8BA104565E29C177FF7EFCCBFDA3FC03" blockId="2.[136,1452,158,2065]" pageId="2" pageNumber="123">
<emphasis id="B96AD8445E29C177FF7EFCCBFEDEFC95" bold="true" box="[189,351,797,823]" pageId="2" pageNumber="123">Phenology:—</emphasis>
This new species was observed in flower during almost the whole year, except during January and February. This is most likely associated with the elevated rainfall levels on this side of the Eastern 
<collectingRegion id="49DACAB45E29C177FAF9FC96FA2DFCF8" box="[1338,1452,832,858]" country="Paraguay" name="Cordillera" pageId="2" pageNumber="123">Cordillera</collectingRegion>
throughout the year, which can surpass 
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of annual rainfall, on the edge of tropical rainforests and their transition to submontane rainforests.
</paragraph>
</subSubSection>
<subSubSection id="C30457DD5E29C177FF7EFC7DFD3FFB8D" pageId="2" pageNumber="123" type="etymology">
<paragraph id="8BA104565E29C177FF7EFC7DFD3FFB8D" blockId="2.[136,1452,158,2065]" pageId="2" pageNumber="123">
<emphasis id="B96AD8445E29C177FF7EFC7DFEE5FC67" bold="true" box="[189,356,939,965]" pageId="2" pageNumber="123">Etymology:—</emphasis>
The epithet refers to the indigenous Tegua people, who inhabited the base of the Boyacá-Casanare Mountain, bordering the Muiscas people territory (
<bibRefCitation id="EF8F79A75E29C177FD01FC18FB97FC4A" author="Fernandez de Piedrahita, L." box="[706,1046,974,1000]" pageId="2" pageNumber="123" refId="ref4724" refString="Fernandez de Piedrahita, L. (1973) Noticia historial de las conquistas del Nuevo Reino de Granada. Vol. 1. Editorial Kelly, Bogota, 432 pp." type="book" year="1973">Fernández de Piedrahita 1973</bibRefCitation>
). The chroniclers Fray Pedro Simón and B. Fernández de Oviedo highlighted the Tegua’s longevity and their knowledge and use of medicinal plants, which was also acknowledged by the Muiscas neighbors.
</paragraph>
</subSubSection>
<subSubSection id="C30457DD5E29C173FF7EFBEEFE1FF8E0" lastPageId="6" lastPageNumber="127" pageId="2" pageNumber="123" type="distribution">
<paragraph id="8BA104565E29C177FF7EFBEEFE9DFA8A" blockId="2.[136,1452,158,2065]" pageId="2" pageNumber="123">
<emphasis id="B96AD8445E29C177FF7EFBEEFD8FFBF0" bold="true" box="[189,526,1080,1106]" pageId="2" pageNumber="123">Distribution and Habitat:—</emphasis>
Currently this species is known exclusively from a small sector in the border area of 
<collectingRegion id="49DACAB45E29C177FF64FB8AFF7AFBD4" box="[167,251,1116,1142]" country="Colombia" name="Boyaca" pageId="2" pageNumber="123">Boyacá</collectingRegion>
and 
<collectingRegion id="49DACAB45E29C177FEF2FB8AFE55FBD4" box="[305,468,1116,1142]" country="Colombia" name="Cundinamarca" pageId="2" pageNumber="123">Cundinamarca</collectingRegion>
Departments, at the base of Colombia’s 
<collectingRegion id="49DACAB45E29C177FC5DFB8AFB91FBD4" box="[926,1040,1116,1142]" country="Paraguay" name="Cordillera" pageId="2" pageNumber="123">Cordillera</collectingRegion>
Oriental (
<figureCitation id="132518D35E29C177FB42FB8AFB65FBD4" box="[1153,1252,1116,1142]" captionStart="FIGURE 4" captionStartId="6.[136,229,1114,1136]" captionTargetBox="[179,1420,212,1068]" captionTargetId="figure-26@6.[151,1436,190,1090]" captionTargetPageId="6" captionText="FIGURE 4. Map showing the type locality of Varronia teguorum." figureDoi="http://doi.org/10.5281/zenodo.5425577" httpUri="https://zenodo.org/record/5425577/files/figure.png" pageId="2" pageNumber="123">Figure 4</figureCitation>
). It grows in very wet tropical forest and in its transition to pluvial submontane forest (5000 and 
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of annual rainfall). It has been exclusively collected at the edge of forests, in exposed areas such as creek margins and road slopes on forest edges, between 500 and 
<quantity id="4CE6A9B35E29C177FE97FB10FE2AFB43" box="[340,427,1222,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="123" unit="m" value="1150.0">1150 m</quantity>
elevation. Among the species restricted to this region of the mountain base are 
<taxonomicName id="4C1E7FD55E29C177FA84FB11FDC6FAA7" authority="Quinones (1995: 17)" authorityName="Quinones" authorityPageNumber="17" authorityYear="1995" class="Magnoliopsida" family="Fabaceae" genus="Brownea" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="enricii">
<emphasis id="B96AD8445E29C177FA84FB11FF50FAA1" italics="true" pageId="2" pageNumber="123">Brownea enricii</emphasis>
<bibRefCitation id="EF8F79A75E29C177FF1AFB3CFE45FAA6" author="Quinones, L. M." box="[217,452,1257,1284]" pageId="2" pageNumber="123" pagination="17 - 22" refId="ref6147" refString="Quinones, L. M. (1995) Dos especies nuevas del genero Brownea (Leguminosae-Caesalpinioideae). Caldasia 18 (86): 17 - 22." type="journal article" year="1995">Quiñones (1995: 17)</bibRefCitation>
(Fabaceae)
</taxonomicName>
and 
<taxonomicName id="4C1E7FD55E29C177FD43FB3CFE99FA8A" authority="Fern. Alonso &amp; Jaramillo-Mejia (1999: 125)" authorityName="Fern. Alonso &amp; Jaramillo-Mejia" authorityPageNumber="125" authorityYear="1999" class="Magnoliopsida" family="Malvaceae" genus="Phragmotheca" kingdom="Plantae" order="Malvales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="mambitana">
<emphasis id="B96AD8445E29C177FD43FB3CFC2AFAA6" box="[640,939,1258,1284]" italics="true" pageId="2" pageNumber="123">Phragmotheca mambitana</emphasis>
Fern. Alonso &amp; Jaramillo-Mejía (1999: 125) (Malvaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BA104565E29C177FF7EFAE6FADBFA7A" blockId="2.[136,1452,158,2065]" pageId="2" pageNumber="123">
<emphasis id="B96AD8445E29C177FF7EFAE6FE4EFAE9" bold="true" box="[189,463,1328,1355]" pageId="2" pageNumber="123">Conservation status:—</emphasis>
Based on fieldwork carried out in the regions of de Mámbita, 
<taxonomicName id="4C1E7FD55E29C177FBBFFAE7FA2AFAE9" authority="Maria, and Guateque" authorityName="Maria, and Guateque" box="[1148,1451,1329,1355]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="genus">Santa María, and Guateque</taxonomicName>
conducted by one of us (JLFA), it seems to represent a locally common species associated with unstable environments, presenting some degree of human influence. Thus, it seems likely that this species is not under any current threat. However, the species has never been formally assessed in 
<collectingCountry id="F30944C65E29C177FCE1FA4DFC12FA17" box="[802,915,1435,1461]" name="Colombia" pageId="2" pageNumber="123">Colombia</collectingCountry>
, based on the IUCN criteria (2019). Therefore, since it represents a locally used medicinal plant, it would be essential for this species to be properly assessed.
</paragraph>
<paragraph id="8BA104565E29C177FF7EFA34FACBF820" blockId="2.[136,1452,158,2065]" pageId="2" pageNumber="123">
<emphasis id="B96AD8445E29C177FF7EFA34FEA7FA5E" bold="true" box="[189,294,1506,1532]" pageId="2" pageNumber="123">Notes:—</emphasis>
<taxonomicName id="4C1E7FD55E29C177FEE5FA35FD81FA5E" box="[294,512,1507,1532]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="teguorum">
<emphasis id="B96AD8445E29C177FEE5FA35FD81FA5E" box="[294,512,1507,1532]" italics="true" pageId="2" pageNumber="123">Varronia teguorum</emphasis>
</taxonomicName>
is closely related morphologically to other species of the genus with globose inflorescences from northern South America, but it can be easily differentiated from 
<taxonomicName id="4C1E7FD55E29C177FB9AF9D0FF3DF9E6" authority="Friesen (1933: 151)" authorityName="Friesen" authorityPageNumber="151" authorityYear="1933" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="subtruncata">
<emphasis id="B96AD8445E29C177FB9AF9D0FB7CF9BD" box="[1113,1277,1541,1567]" italics="true" pageId="2" pageNumber="123">V. subtruncata</emphasis>
<bibRefCitation id="EF8F79A75E29C177FAC5F9D3FF3DF9E6" author="Friesen, C. V." pageId="2" pageNumber="123" pagination="117 - 199" refId="ref4952" refString="Friesen, C. V. (1933) Les caracteres essentiels de la famille des Sebestenaceae &amp; revision du genre Varronia. Bulletin de la Societe Botanique de Geneve 24 (2): 117 - 199." type="journal article" year="1933">Friesen (1933: 151)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C1E7FD55E29C177FF31F9FCFDD6F9E6" authority="Linnaeus (1759: 916)" authorityName="Linnaeus" authorityPageNumber="916" authorityYear="1759" box="[242,599,1577,1604]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="bullata">
<emphasis id="B96AD8445E29C177FF31F9FCFEDCF9E1" box="[242,349,1577,1603]" italics="true" pageId="2" pageNumber="123">V. bullata</emphasis>
<bibRefCitation id="EF8F79A75E29C177FEA6F9FFFDD6F9E6" author="Linnaeus, C." box="[357,599,1577,1604]" pageId="2" pageNumber="123" pagination="825 - 1384" refId="ref5457" refString="Linnaeus, C. (1759) Systema Naturae (ed. 10), 2. pp. 825 - 1384." type="book chapter" year="1759">Linnaeus (1759: 916)</bibRefCitation>
</taxonomicName>
due to its long and rusty indumentum with straight hairs 
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long 
<emphasis id="B96AD8445E29C177FA4EF9FCFA25F9E1" box="[1421,1444,1578,1603]" italics="true" pageId="2" pageNumber="123">vs</emphasis>
. white to yellow indumentum, with patent to adpressed hairs &lt;
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long, and oblong to ovate-oblong, 
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leaf blades 
<emphasis id="B96AD8445E29C177FECAF9A7FEA1F928" box="[265,288,1649,1674]" italics="true" pageId="2" pageNumber="123">vs</emphasis>
. lanceolate-ovate, 2.5–8.0 cm leaf blades. Moreover, it differs from 
<taxonomicName id="4C1E7FD55E29C177FBE0F9A7FD93F90F" authority="(Killip ex J. Estrada &amp; Garcia-Barr.) J. S. Mill. (2007: 373)" authorityName="J. S. Mill." authorityPageNumber="373" authorityYear="2007" baseAuthorityName="J. Estrada &amp; Garcia-Barr." class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="bullulata">
<emphasis id="B96AD8445E29C177FBE0F9A7FB25F928" box="[1059,1188,1648,1674]" italics="true" pageId="2" pageNumber="123">V. bullulata</emphasis>
(Killip ex J. Estrada &amp; García-Barr.) J.S. Mill. (2007: 373)
</taxonomicName>
because this one presents finely bullate (
<emphasis id="B96AD8445E29C177FC1AF942FC71F90F" box="[985,1008,1684,1709]" italics="true" pageId="2" pageNumber="123">vs</emphasis>
. flat) leaf blades, recurved (
<emphasis id="B96AD8445E29C177FAEAF942FAC1F90F" box="[1321,1344,1684,1709]" italics="true" pageId="2" pageNumber="123">vs</emphasis>
. straight) inflorescence peduncles, and appendages of the calyx lobes 
<quantity id="4CE6A9B35E29C177FCF3F961FC22F973" box="[816,931,1719,1746]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="5.0" metricValueMin="1.7" pageId="2" pageNumber="123" unit="mm" value="3.35" valueMax="5.0" valueMin="1.7">1.7–5 mm</quantity>
long (
<emphasis id="B96AD8445E29C177FC2DF96EFB84F973" box="[1006,1029,1720,1745]" italics="true" pageId="2" pageNumber="123">vs</emphasis>
. ≤ 
<quantity id="4CE6A9B35E29C177FBEFF961FB02F973" box="[1068,1155,1719,1746]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="123" unit="mm" value="1.5">1.5 mm</quantity>
long). Other species from this group, namely 
<taxonomicName id="4C1E7FD55E29C177FEA4F90DFC9FF956" authority="Desvaux (1808: 173)" authorityName="Desvaux" authorityPageNumber="173" authorityYear="1808" box="[359,798,1754,1781]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="macrocephala">
<emphasis id="B96AD8445E29C177FEA4F90DFDA5F956" box="[359,548,1754,1780]" italics="true" pageId="2" pageNumber="123">V. macrocephala</emphasis>
<bibRefCitation id="EF8F79A75E29C177FDEEF90DFC9FF956" author="Desvaux, N. A." box="[557,798,1754,1781]" pageId="2" pageNumber="123" pagination="129 - 165" refId="ref4570" refString="Desvaux, N. A. (1808) Memorie sur une monographie du genre Luzula. Journal de Botanique, Redige par une Societe di Botanistes 1: 129 - 165." type="journal article" year="1808">Desvaux (1808: 173)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C1E7FD55E29C177FC9AF90DFAF1F956" authority="(J. Estrada) J. S. Mill. (2007: 372)" authorityName="J. S. Mill." authorityPageNumber="372" authorityYear="2007" baseAuthorityName="J. Estrada" box="[857,1392,1754,1781]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="andreana">
<emphasis id="B96AD8445E29C177FC9AF90DFC60F956" box="[857,993,1754,1780]" italics="true" pageId="2" pageNumber="123">V. andreana</emphasis>
(J. Estrada) J.S. Mill. (2007: 372)
</taxonomicName>
, also significantly differ from 
<taxonomicName id="4C1E7FD55E29C177FE5CF928FDA7F8B5" box="[415,550,1790,1815]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="teguorum">
<emphasis id="B96AD8445E29C177FE5CF928FDA7F8B5" box="[415,550,1790,1815]" italics="true" pageId="2" pageNumber="123">V. teguorum</emphasis>
</taxonomicName>
. The first due to its unique indumentum with stellate hairs, and the latter due to its indumentum with short glandular hairs and flowers with larger (
<quantity id="4CE6A9B35E29C177FC46F8F7FB81F89E" box="[901,1024,1825,1852]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="2" pageNumber="123" unit="mm" value="20.5" valueMax="23.0" valueMin="18.0">18–23 mm</quantity>
long) corollas. Finally, 
<taxonomicName id="4C1E7FD55E29C177FAD3F8F4FE60F8FD" authority="(Kunth) T. S. Silva (2019: 297)" authorityName="T. S. Silva" authorityPageNumber="297" authorityYear="2019" baseAuthorityName="Kunth" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="serratifolia">
<emphasis id="B96AD8445E29C177FAD3F8F4FA2AF899" box="[1296,1451,1825,1851]" italics="true" pageId="2" pageNumber="123">V. serratifolia</emphasis>
(Kunth) T.S. 
<bibRefCitation id="EF8F79A75E29C177FED8F893FE60F8FD" author="Silva, T. S. &amp; Silva, L. P. &amp; Melo, J. I. M." box="[283,481,1861,1887]" pageId="2" pageNumber="123" pagination="25 - 26" refId="ref6441" refString="Silva, T. S., Silva, L. P. &amp; Melo, J. I. M. (2019) Nomenclatural updates in Varronia (Cordiaceae, Boraginales) from South America. Harvard Papers in Botany 24 (1): 25 - 26. https: // doi. org / 10.3100 / hpib. v 24 iss 1.2019. n 4" type="journal article" year="2019">Silva (2019: 297)</bibRefCitation>
</taxonomicName>
, which shares with 
<taxonomicName id="4C1E7FD55E29C177FD7DF893FCC4F8FC" box="[702,837,1861,1886]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="teguorum">
<emphasis id="B96AD8445E29C177FD7DF893FCC4F8FC" box="[702,837,1861,1886]" italics="true" pageId="2" pageNumber="123">V. teguorum</emphasis>
</taxonomicName>
the short calyx appendages, can be differentiated from the new species by its smaller leaf blades (usually 
<quantity id="4CE6A9B35E29C177FD7EF8BEFCBBF821" box="[701,826,1896,1923]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" metricValueMax="7.5" metricValueMin="2.1" pageId="2" pageNumber="123" unit="cm" value="4.8" valueMax="7.5" valueMin="2.1">2.1–7.5 cm</quantity>
long) and inflorescences less than 
<quantity id="4CE6A9B35E29C177FB00F8BEFA85F821" box="[1219,1284,1896,1923]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="123" unit="mm" value="8.0">8 mm</quantity>
diam.
</paragraph>
<paragraph id="8BA104565E29C177FF7EF85AFDAAF7B3" blockId="2.[136,1452,158,2065]" pageId="2" pageNumber="123">
<emphasis id="B96AD8445E29C177FF7EF85AFE3DF804" bold="true" box="[189,444,1932,1958]" pageId="2" pageNumber="123">Vernacular names:—</emphasis>
It is locally known as “ 
<emphasis id="B96AD8445E29C177FD7DF85DFC80F807" box="[702,769,1931,1957]" italics="true" pageId="2" pageNumber="123">salvia</emphasis>
” in the localities in which it is regarded as a medicinal plant. Other species of 
<taxonomicName id="4C1E7FD55E29C177FE80F866FE27F86B" box="[323,422,1968,1993]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AD8445E29C177FE80F866FE27F86B" box="[323,422,1968,1993]" italics="true" pageId="2" pageNumber="123">Varronia</emphasis>
</taxonomicName>
, such as 
<taxonomicName id="4C1E7FD55E29C177FDC9F866FBA6F86B" authority="Ruiz &amp; Pav. (1799: 23)" authorityName="Ruiz &amp; Pav." authorityPageNumber="23" authorityYear="1799" box="[522,1063,1967,1994]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="cylindristachya">
<emphasis id="B96AD8445E29C177FDC9F866FCA1F86B" box="[522,800,1967,1993]" italics="true" pageId="2" pageNumber="123">Varronia cylindristachya</emphasis>
Ruiz &amp; Pav. (1799: 23)
</taxonomicName>
and 
<taxonomicName id="4C1E7FD55E29C177FB9FF866FE27F84F" authority="(Linnaeus 1771: 206) Borhidi (1988: 393)" authorityName="Borhidi" authorityPageNumber="393" authorityYear="1988" baseAuthorityName="Linnaeus" baseAuthorityPageNumber="206" baseAuthorityYear="1771" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="123" phylum="Tracheophyta" rank="species" species="spinescens">
<emphasis id="B96AD8445E29C177FB9FF866FB6EF86B" box="[1116,1263,1968,1993]" italics="true" pageId="2" pageNumber="123">V. spinescens</emphasis>
(
<bibRefCitation id="EF8F79A75E29C177FB3EF879FF35F84F" author="Linnaeus, C." pageId="2" pageNumber="123" refId="ref5617" refString="Linnaeus, C. (1771) Mantissa Plantarum Altera. Impresis Direct Laurentii Salvii, Holmiae, 588 pp." type="book" year="1771">Linnaeus 1771: 206</bibRefCitation>
) 
<bibRefCitation id="EF8F79A75E29C177FF07F804FE27F84F" author="Borhidi, A. &amp; Gondar, E. &amp; Orosz-Kovacs, Z. S." box="[196,422,2002,2029]" pageId="2" pageNumber="123" pagination="375 - 423" refId="ref4429" refString="Borhidi, A., Gondar, E. &amp; Orosz-Kovacs, Z. S. (1988) The reconsideration of the genus Cordia. Acta Botanica Hungarica 34: 375 - 423." type="journal article" year="1988">Borhidi (1988: 393)</bibRefCitation>
</taxonomicName>
are also known by the same name, or small variations thereof (“ 
<emphasis id="B96AD8445E29C177FB45F804FA93F84E" box="[1158,1298,2002,2028]" italics="true" pageId="2" pageNumber="123">salvia negra</emphasis>
” or “ 
<emphasis id="B96AD8445E29C177FA90F804FA17F84E" box="[1363,1430,2002,2028]" italics="true" pageId="2" pageNumber="123">salvio</emphasis>
”) (
<bibRefCitation id="EF8F79A75E29C177FF4CF820FEFAF7B3" author="Garcia-Barriga, H." box="[143,379,2038,2065]" pageId="2" pageNumber="123" refId="ref4991" refString="Garcia-Barriga, H. (1975) Flora medicinal de Colombia. Vol. 2. Universidad Nacional de Colombia, Bogota, 345 pp." type="book" year="1975">García-Barriga 1975</bibRefCitation>
; 
<bibRefCitation id="EF8F79A75E29C177FE44F820FD9FF7B3" author="Estrada, J." box="[391,542,2038,2065]" pageId="2" pageNumber="123" refId="ref4603" refString="Estrada, J. (1995) Flora de Colombia. Cordia. Universidad Nacional de Colombia, Instituto de Ciencias Naturales, 176 pp." type="book" year="1995">Estrada 1995</bibRefCitation>
).
</paragraph>
<caption id="DF6154DE5E28C176FF4BF877FAC1F815" ID-DOI="http://doi.org/10.5281/zenodo.5425569" ID-Zenodo-Dep="5425569" box="[136,1344,1953,1975]" httpUri="https://zenodo.org/record/5425569/files/figure.png" pageId="3" pageNumber="124" startId="3.[136,229,1953,1975]" targetBox="[226,1365,198,1921]" targetPageId="3">
<paragraph id="8BA104565E28C176FF4BF877FAC1F815" blockId="3.[136,1344,1953,1975]" box="[136,1344,1953,1975]" pageId="3" pageNumber="124">
<emphasis id="B96AD8445E28C176FF4BF877FF7FF814" bold="true" box="[136,254,1953,1975]" pageId="3" pageNumber="124">FIGURE 1.</emphasis>
Image of the isotype of 
<taxonomicName id="4C1E7FD55E28C176FE2BF874FBECF814" authority="Fern. Alonso &amp; J. I. M. Melo, Fernandez-Alonso" authorityName="Fern. Alonso &amp; J. I. M. Melo, Fernandez-Alonso" box="[488,1133,1953,1975]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="3" pageNumber="124" phylum="Tracheophyta" rank="species" species="teguorum">
<emphasis id="B96AD8445E28C176FE2BF874FD1FF815" box="[488,670,1954,1975]" italics="true" pageId="3" pageNumber="124">Varronia teguorum</emphasis>
Fern. Alonso &amp; J.I.M. Melo, 
<emphasis id="B96AD8445E28C176FC7FF877FBECF814" box="[956,1133,1953,1974]" italics="true" pageId="3" pageNumber="124">Fernández-Alonso</emphasis>
</taxonomicName>
<emphasis id="B96AD8445E28C176FBB7F877FB31F814" box="[1140,1200,1953,1974]" italics="true" pageId="3" pageNumber="124">24079</emphasis>
(MA 888445).
</paragraph>
</caption>
<caption id="DF6154DE5E2FC171FF4BF8A9FA18F784" ID-DOI="http://doi.org/10.5281/zenodo.5425571" ID-Zenodo-Dep="5425571" httpUri="https://zenodo.org/record/5425571/files/figure.png" pageId="4" pageNumber="125" startId="4.[136,229,1919,1941]" targetBox="[243,1344,190,1895]" targetPageId="4">
<paragraph id="8BA104565E2FC171FF4BF8A9FA18F784" blockId="4.[136,1453,1919,2086]" pageId="4" pageNumber="125">
<emphasis id="B96AD8445E2FC171FF4BF8A9FF7DF837" bold="true" box="[136,252,1919,1941]" pageId="4" pageNumber="125">FIGURE 2.</emphasis>
Field images of 
<taxonomicName id="4C1E7FD55E2FC171FE5BF856FDCCF837" box="[408,589,1920,1941]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="4" pageNumber="125" phylum="Tracheophyta" rank="species" species="teguorum">
<emphasis id="B96AD8445E2FC171FE5BF856FDCCF837" box="[408,589,1920,1941]" italics="true" pageId="4" pageNumber="125">Varronia teguorum</emphasis>
</taxonomicName>
. A. Apex of a young branch, showing the rusty indumentum and the tertiary and quaternary veins on the leaf adaxial surface. B. Branch with developed axillary inflorescences, showing floral buds. C. Branch with an inflorescence. D. Inflorescences with some flowers at anthesis (white corolla) and others at post-anthesis (senescent corolla), and immature fruits (red). E. Inflorescences with an open flower at a basal position, showing the senescent flowers with accrescent calyx and incurved lobes obscuring the calyx throat. Scale bar in all cases = 15 mm long. Photos by J.L. Fernández-Alonso taken in the type locality, 
<taxonomicName id="4C1E7FD55E2FC171FB0AF7C6FAC1F784" authority="Maria" authorityName="Maria" box="[1225,1344,2064,2086]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="genus">Santa María</taxonomicName>
, Boyacá.
</paragraph>
</caption>
<caption id="DF6154DE5E2EC170FF4BF8DCFC92F7BE" ID-DOI="http://doi.org/10.5281/zenodo.5425573" ID-Zenodo-Dep="5425573" httpUri="https://zenodo.org/record/5425573/files/figure.png" pageId="5" pageNumber="126" startId="5.[136,229,1802,1824]" targetBox="[270,1317,190,1778]" targetPageId="5">
<paragraph id="8BA104565E2EC170FF4BF8DCFC92F7BE" blockId="5.[136,1453,1802,2076]" pageId="5" pageNumber="126">
<emphasis id="B96AD8445E2EC170FF4BF8DCFF7EF8BD" bold="true" box="[136,255,1802,1824]" pageId="5" pageNumber="126">FIGURE 3.</emphasis>
Light micrographs of a dried specimen of 
<taxonomicName id="4C1E7FD55E2EC170FD62F8DDFCD9F882" box="[673,856,1803,1824]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="teguorum">
<emphasis id="B96AD8445E2EC170FD62F8DDFCD9F882" box="[673,856,1803,1824]" italics="true" pageId="5" pageNumber="126">Varronia teguorum</emphasis>
</taxonomicName>
. A. Insertion of the petiole in the branch, the base of the leaf blade, and the axillary bud. B. Adaxial surface of leaf blade: impressed venation and indumentum with conspicuous long hairs along the veins and cystolithic pustules. C. Abaxial surface of leaf blade showing prominent reticulate venation (2º, 3º, and 4º orders) and hairs with cystolithic pustules. D. Slightly adnate region of the petiole (right) with the inflorescence axis (left). E. Young inflorescence, showing the indumentum of the peduncle and the incurved appendages of the calyx lobes. F. Inflorescence rachis, showing its dense indumentum and the flower arrangement. G. Inflorescence with flowers at anthesis: G1. Two open flowers showing the corolla lobes and some dehisced anthers; G2. Open flower, showing the glabrous anthers covering the corolla throat. Scale bar in all cases = 5 mm long. (Voucher: 
<emphasis id="B96AD8445E2EC170FF4BF7D0FEFAF7B9" box="[136,379,2054,2075]" italics="true" pageId="5" pageNumber="126">Fernández-Alonso 24079</emphasis>
, COL). Photos by J.L. Fernández-Alonso.
</paragraph>
</caption>
<caption id="DF6154DE5E2DC173FF4BFB8CFC8BFBD2" ID-DOI="http://doi.org/10.5281/zenodo.5425577" ID-Zenodo-Dep="5425577" box="[136,778,1114,1136]" httpUri="https://zenodo.org/record/5425577/files/figure.png" pageId="6" pageNumber="127" startId="6.[136,229,1114,1136]" targetBox="[179,1420,212,1068]" targetPageId="6">
<paragraph id="8BA104565E2DC173FF4BFB8CFC8BFBD2" blockId="6.[136,778,1114,1136]" box="[136,778,1114,1136]" pageId="6" pageNumber="127">
<emphasis id="B96AD8445E2DC173FF4BFB8CFF7FFBCD" bold="true" box="[136,254,1114,1136]" pageId="6" pageNumber="127">FIGURE 4.</emphasis>
Map showing the type locality of 
<taxonomicName id="4C1E7FD55E2DC173FD8EFB8DFC85FBD2" box="[589,772,1115,1136]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="species" species="teguorum">
<emphasis id="B96AD8445E2DC173FD8EFB8DFC85FBD2" box="[589,772,1115,1136]" italics="true" pageId="6" pageNumber="127">Varronia teguorum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA104565E2DC173FF7EFB49FCE0FB7F" blockId="6.[136,1452,1183,1858]" pageId="6" pageNumber="127">
<emphasis id="B96AD8445E2DC173FF7EFB49FEF1FB1B" bold="true" box="[189,368,1183,1209]" pageId="6" pageNumber="127">Known uses:—</emphasis>
In the region of 
<taxonomicName id="4C1E7FD55E2DC173FDDAFB49FD20FB1B" authority="Maria" authorityName="Maria" box="[537,673,1183,1209]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Santa María</taxonomicName>
, it has been recently cited as medicinal under the tentative identification of 
<taxonomicName id="4C1E7FD55E2DC173FF65FB15FCDCFB7F" authority="(Estrada) J. S. Mill. (Rodriguez-Mora et al. 2020)" box="[166,861,1219,1246]" class="Magnoliopsida" family="Cordiaceae" genus="Varronia" kingdom="Plantae" order="Boraginales" pageId="6" pageNumber="127" phylum="Tracheophyta" rank="species" species="andreana">
<emphasis id="B96AD8445E2DC173FF65FB15FEADFB7F" box="[166,300,1219,1245]" italics="true" pageId="6" pageNumber="127">V. andreana</emphasis>
(Estrada) J.S. Mill. (
<bibRefCitation id="EF8F79A75E2DC173FDD5FB15FCD4FB7F" author="Rodriguez-Mora, D. F. &amp; Velasquez-Avila, H. A. &amp; Fernandez-Alonso, J. L. &amp; Raz, L." box="[534,853,1219,1245]" pageId="6" pageNumber="127" refId="ref6176" refString="Rodriguez-Mora, D. F., Velasquez-Avila, H. A., Fernandez-Alonso, J. L. &amp; Raz, L. (2020) Guia de campo de los usos tradicionales no maderables de las plantas de Santa Maria, Boyaca (Andes colombianos). Serie Guias de Campo del Instituto de Ciencias Naturales de la Universidad Nacional de Colombia N ° 22. Universidad Nacional de Colombia, Bogota, 381 pp." type="book" year="2020">
Rodríguez-Mora 
<emphasis id="B96AD8445E2DC173FD1AFB15FC8AFB7F" box="[729,779,1219,1245]" italics="true" pageId="6" pageNumber="127">et al</emphasis>
. 2020
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BA104565E2DC173FF7EFB30FE1FF8E0" blockId="6.[136,1452,1183,1858]" pageId="6" pageNumber="127">
<emphasis id="B96AD8445E2DC173FF7EFB30FBA0FAA3" bold="true" box="[189,1057,1254,1281]" pageId="6" pageNumber="127">
Additional examined specimens (
<typeStatus id="54A5BAF45E2DC173FD85FB30FD40FAA2" box="[582,705,1254,1280]" pageId="6" pageNumber="127" type="paratype">paratypes</typeStatus>
). 
<collectingCountry id="F30944C65E2DC173FD16FB30FCF7FAA2" box="[725,886,1254,1280]" name="Colombia" pageId="6" pageNumber="127">COLOMBIA</collectingCountry>
. 
<collectingRegion id="49DACAB45E2DC173FC42FB31FC3FFAA3" box="[897,958,1255,1281]" country="Colombia" name="Dpto" pageId="6" pageNumber="127">Dpto</collectingRegion>
. 
<collectingRegion id="49DACAB45E2DC173FC09FB30FBA0FAA3" box="[970,1057,1254,1281]" country="Colombia" name="Boyaca" pageId="6" pageNumber="127">Boyacá</collectingRegion>
</emphasis>
. Vertiente Oriental de la Cordillera Oriental. 
<taxonomicName id="4C1E7FD55E2DC173FF30FADDFCBAFA87" authority="Maria, Vereda La Carbonera y Charco Largo" authorityName="Maria, Vereda La Carbonera y Charco Largo" box="[243,827,1291,1317]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Santa María, Vereda La Carbonera y Charco Largo</taxonomicName>
, 
<quantity id="4CE6A9B35E2DC173FC8BFADCFC47FA87" box="[840,966,1290,1317]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="6" pageNumber="127" unit="m" value="775.0" valueMax="900.0" valueMin="650.0">650-900 m</quantity>
, 
<date id="FFA022965E2DC173FC17FADDFBE3FA87" box="[980,1122,1291,1317]" pageId="6" pageNumber="127" value="2008-06-07">7 June 2008</date>
, fl., 
<emphasis id="B96AD8445E2DC173FB55FADDFA2AFA87" box="[1174,1451,1291,1317]" italics="true" pageId="6" pageNumber="127">J. Betancur &amp; al. 13020</emphasis>
(COL); Ibidem, en zonas de huertas y caminos en los márgenes del casco urbano, 
<quantity id="4CE6A9B35E2DC173FB82FAF9FB02FAEB" box="[1089,1155,1327,1353]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="127" unit="m" value="800.0">800m</quantity>
, arbusto scandens 
<quantity id="4CE6A9B35E2DC173FA98FAF9FA27FAEB" box="[1371,1446,1327,1353]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="127" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2–4 m</quantity>
, indumento ferrugíneo café, inflorescencias verdosas, corola crema, 
<date id="FFA022965E2DC173FC4FFA85FBAEFACF" box="[908,1071,1362,1389]" pageId="6" pageNumber="127" value="2005-04-22">22 April 2005</date>
, fl., 
<emphasis id="B96AD8445E2DC173FBA7FA85FF4FFA33" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 22914</emphasis>
(COL, MA); Ibidem, bajada de S. María a S. Luis de Gaceno, Zona del río Lengupá, Quebrada Mollas y Vereda Aguas Calientes, 
<quantity id="4CE6A9B35E2DC173FE92FA4CFE1DFA17" box="[337,412,1434,1461]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="127" unit="m" value="500.0">500 m</quantity>
, 
<date id="FFA022965E2DC173FE69FA4CFDCCFA17" box="[426,589,1434,1461]" pageId="6" pageNumber="127" value="2005-04-25">25 April 2005</date>
, fl., 
<emphasis id="B96AD8445E2DC173FD41FA4DFB99FA17" box="[642,1048,1435,1461]" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 23000</emphasis>
(COL 535465, 535816; HUA, MA 888438, 888439); Vía 
<taxonomicName id="4C1E7FD55E2DC173FE4CFA69FD9BFA7B" authority="Maria" authorityName="Maria" box="[399,538,1471,1497]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Santa María</taxonomicName>
a La Cristalina, bocatoma del acueducto, rio Batá, 
<quantity id="4CE6A9B35E2DC173FBB7FA68FB75FA7B" box="[1140,1268,1470,1497]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="6" pageNumber="127" unit="m" value="525.0" valueMax="550.0" valueMin="500.0">500-550 m</quantity>
, 
<date id="FFA022965E2DC173FAC0FA68FA26FA7B" box="[1283,1447,1470,1497]" pageId="6" pageNumber="127" value="2005-04-25">25 April 2005</date>
, fl., 
<emphasis id="B96AD8445E2DC173FF71FA35FDD1FA5F" box="[178,592,1507,1533]" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 23020</emphasis>
(COL 535820, 535826; HUA, MA 888440, 888441, MEDEL, MO, US); Ibidem, remanente de bmhT en el tramo al Acueducto, 
<quantity id="4CE6A9B35E2DC173FD28F9D1FCB2F983" box="[747,819,1543,1569]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="127" unit="m" value="800.0">800 m</quantity>
, 
<date id="FFA022965E2DC173FCFDF9D1FC70F983" box="[830,1009,1542,1569]" pageId="6" pageNumber="127" value="2005-08-24">24 August 2005</date>
, fl., 
<emphasis id="B96AD8445E2DC173FBE2F9D1FA2AF983" box="[1057,1451,1543,1569]" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 23579</emphasis>
(COL 533668; HUA, MA 888442, UPTC); Vía S. María a Mámbita, Trayecto desde el río Garagoa a 
<taxonomicName id="4C1E7FD55E2DC173FAD9F9FDFA26F9E7" authority="Maria" authorityName="Maria" box="[1306,1447,1579,1605]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Santa María</taxonomicName>
, 
<quantity id="4CE6A9B35E2DC173FF4BF999FE87F9CB" box="[136,262,1615,1641]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="6" pageNumber="127" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700-800 m</quantity>
, bosque intervenido y potreros, 
<date id="FFA022965E2DC173FDB0F999FC94F9CB" box="[627,789,1615,1641]" pageId="6" pageNumber="127" value="2007-03-06">6 March 2007</date>
, fr., 
<emphasis id="B96AD8445E2DC173FC8AF999FB5AF9CB" box="[841,1243,1615,1641]" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 24775</emphasis>
(COL, HUA, MA 888446); Ibidem, 
<quantity id="4CE6A9B35E2DC173FE93F9A5FE19F92F" box="[336,408,1651,1677]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="127" unit="m" value="800.0">800 m</quantity>
, 
<date id="FFA022965E2DC173FE61F9A5FDC0F92F" box="[418,577,1651,1677]" pageId="6" pageNumber="127" value="2007-03-04">4 March 2007</date>
, arbusto 
<quantity id="4CE6A9B35E2DC173FD60F9A5FD66F92F" box="[675,743,1651,1677]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="6" pageNumber="127" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">2-4 m</quantity>
, flores blancas con cáliz amarillento, fr., 
<emphasis id="B96AD8445E2DC173FB6FF9A5FE95F913" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 24933</emphasis>
(COL, MA 888447, 888448, MEDEL); Vía de 
<taxonomicName id="4C1E7FD55E2DC173FC83F941FC4AF913" authority="Maria" authorityName="Maria" box="[832,971,1687,1713]" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">Santa María</taxonomicName>
a Mámbita, 
<quantity id="4CE6A9B35E2DC173FBA2F940FB61F913" box="[1121,1248,1686,1713]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="6" pageNumber="127" unit="m" value="725.0" valueMax="750.0" valueMin="700.0">700-750 m</quantity>
, arbusto 1,5- 
<quantity id="4CE6A9B35E2DC173FAB5F940FA26F913" box="[1398,1447,1686,1713]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="127" unit="m" value="2.0">2 m</quantity>
, capítulos muy aromáticos, 
<date id="FFA022965E2DC173FE7FF96DFDF0F977" box="[444,625,1723,1749]" pageId="6" pageNumber="127" value="2007-10-02">2 October 2007</date>
, fl., 
<emphasis id="B96AD8445E2DC173FD64F96DFBBCF977" box="[679,1085,1723,1749]" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 25768</emphasis>
(COL, MA, UPTC, US). 
<emphasis id="B96AD8445E2DC173FAAAF96DFEBDF95A" bold="true" pageId="6" pageNumber="127">
<collectingRegion id="49DACAB45E2DC173FAAAF96DFA27F977" box="[1385,1446,1723,1749]" country="Colombia" name="Dpto" pageId="6" pageNumber="127">Dpto</collectingRegion>
. 
<collectingRegion id="49DACAB45E2DC173FF4BF908FEBDF95A" box="[136,316,1758,1784]" country="Colombia" name="Cundinamarca" pageId="6" pageNumber="127">Cundinamarca</collectingRegion>
</emphasis>
. Ubalá B, Vereda San Roque, Camino a Campo Hermoso, 
<geoCoordinate id="EE2A62915E2DC173FC13F908FBCEF95B" box="[976,1103,1758,1785]" degrees="4" direction="north" minutes="45" orientation="latitude" pageId="6" pageNumber="127" precision="15" seconds="50" value="4.763889">4º45’50’’N</geoCoordinate>
, 
<geoCoordinate id="EE2A62915E2DC173FB98F908FB6DF95B" box="[1115,1260,1758,1785]" degrees="73" direction="west" minutes="20" orientation="longitude" pageId="6" pageNumber="127" precision="15" seconds="56" value="-73.34889">73º20’56’’W</geoCoordinate>
, 
<quantity id="4CE6A9B35E2DC173FB34F908FACCF95B" box="[1271,1357,1758,1785]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="127" unit="m" value="1150.0">1150 m</quantity>
, arbusto 
<quantity id="4CE6A9B35E2DC173FF4BF8D5FF36F8BF" box="[136,183,1795,1821]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="127" unit="m" value="2.0">2 m</quantity>
, fl., 
<date id="FFA022965E2DC173FF28F8D5FE06F8BF" box="[235,391,1795,1821]" pageId="6" pageNumber="127" value="1998-06-30">30 June 1998</date>
, fl., fr., 
<emphasis id="B96AD8445E2DC173FE21F8D5FCF4F8BF" box="[482,885,1795,1821]" italics="true" pageId="6" pageNumber="127">J.L. Fernández-Alonso &amp; al. 16210</emphasis>
(COAH, COL 479190, 479208, 479209, 479210; HECASA, MA 888436).
</paragraph>
</subSubSection>
</treatment>
</document>