530 lines
69 KiB
XML
530 lines
69 KiB
XML
<document id="BCDCF2C200429AE8B4EB1BABBD5E651B" ID-DOI="10.11606/1807-0205/2020.60.special-issue.19" ID-ISSN="1807-0205" ID-Zenodo-Dep="4985175" ID-ZooBank="B723DFDD-1F03-4110-903A-A2B8A4CA7083" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624028473964" checkinUser="carolina" docAuthor="Bevilaqua, Marcus & Fonseca, Claudio Ruy Vasconcelos da" docDate="2020" docId="039E3733FFA47D0DECC49DE0FAB9F78A" docLanguage="en" docName="PapAvZool.60.special-issue.e202060.19.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 60 (19)" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.special-issue.19" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Passalus (Pertinax) deuterocerus Bevilaqua & Fonseca 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="FFA74F4BFFA57D0FEC6F9F5AFFF5FFD9" masterDocTitle="Two new species of Passalus Fabricius (Coleoptera: Passalidae) from the western Brazilian Amazon with comments on the taxonomic limits of the subgenera" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1698969348260" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:title id="762DF43E4C3696C7DE26CEB5F937FECE">Two new species of Passalus Fabricius (Coleoptera: Passalidae) from the western Brazilian Amazon with comments on the taxonomic limits of the subgenera</mods:title>
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<mods:namePart id="DE7FA6ECD0DDAA787934BCC053A63CE7">Bevilaqua, Marcus</mods:namePart>
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<mods:namePart id="C4732E78E1701F3B5965ED242C9B1C60">Fonseca, Claudio Ruy Vasconcelos da</mods:namePart>
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<mods:title id="71330B7F13335ED807B3EAAA49693667">Papéis Avulsos de Zoologia</mods:title>
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<mods:date id="BE2C08C33BC178F1FC9F60A5F92DABD4">2020</mods:date>
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<mods:title id="42044228DCD5BE61118B160576B95CF6">Pap. Avulsos Zool., S. Paulo</mods:title>
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<mods:number id="DE70252740516A9A0F2590B8680AC945">2020-03-04</mods:number>
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<mods:url id="588D855A79A25193A6A3544185A849F5">http://dx.doi.org/10.11606/1807-0205/2020.60.special-issue.19</mods:url>
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<heading id="D0C03149FFA47D0EECC49DE0FD74FD0A" box="[171,641,698,723]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
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<emphasis id="B9435A37FFA47D0EECC49DE0FD74FD0A" bold="true" box="[171,641,698,723]" pageId="1" pageNumber="2">
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<taxonomicName id="4C37FDA6FFA47D0EECC49DE0FDD7FD0A" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[171,546,698,723]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov." subGenus="Pertinax">
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<emphasis id="B9435A37FFA47D0EECC49DE0FDD7FD0A" bold="true" box="[171,546,698,723]" italics="true" pageId="1" pageNumber="2">Passalus (Pertinax) deuterocerus</emphasis>
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<emphasis id="B9435A37FFA47D0EED479D80FDF1FD2A" bold="true" box="[296,516,730,755]" pageId="1" pageNumber="2">
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(
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,
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<figureCitation id="130C9AA0FFA47D0EEDE79D80FE6CFD2A" box="[392,409,730,755]" captionStart="Figure 2" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-113@5.[94,1417,494,2044]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) deuterocerus sp.nov.(A) headand prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985183" httpUri="https://zenodo.org/record/4985183/files/figure.png" pageId="1" pageNumber="2">2</figureCitation>
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,
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)
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<emphasis id="B9435A37FFA47D0EEC319C40FEE7FCEA" bold="true" box="[94,274,794,819]" pageId="1" pageNumber="2">Type material:</emphasis>
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/ Fernando Pinto,
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col”
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/ Fernando Pinto, Gleic Sá col. (
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). Cruzeiro do Sul /
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364 Sítio Rosalvo /
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/ Fernando Pinto, Gleic Sá col. (
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). Cruzeiro do Sul /
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). Feijó / [Ramal] Deusa – Interflúvio Envira/Purús /
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/ Fernando Pinto, Gleic Sá col. (
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<specimenCount id="9D314DACFFA47D0EED0E9B20FE76FB4B" box="[353,387,1143,1171]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED261EE0FFA47D0EEDE19B20FE32FB4A" box="[398,455,1146,1171]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="2">INPA</collectionCode>
|
||
). Feijó / [Ramal] Deusa – Interflúvio Envira/Purús /
|
||
<geoCoordinate id="EE03E0E2FFA47D0EEDEC9BC0FDEAFB6A" box="[387,543,1178,1204]" degrees="08" direction="south" minutes="30" orientation="latitude" pageId="1" pageNumber="2" precision="1" seconds="32.41" value="-8.509003">08°30′32.41″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE03E0E2FFA47D0EEE449BC0FD3AFB6A" box="[555,719,1178,1204]" degrees="70" direction="west" minutes="02" orientation="longitude" pageId="1" pageNumber="2" precision="1" seconds="47.77" value="-70.0466">70°02′47.77″W</geoCoordinate>
|
||
/
|
||
<date id="FF89A0E5FFA47D0EEC1F9BE0FEF6FB0A" box="[112,259,1210,1235]" pageId="1" pageNumber="2" value="2011-09-05" valueMax="2011-09-11" valueMin="2011-09-05">05-11.ix.2011</date>
|
||
/ Fernando Pinto, Gleic Sá col. (
|
||
<specimenCount id="9D314DACFFA47D0EEE1E9BE0FD62FB0B" box="[625,663,1207,1235]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D314DACFFA47D0EEECB9BE0FD3CFB0B" box="[676,713,1207,1235]" pageId="1" pageNumber="2" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED261EE0FFA47D0EEC319B80FF63FB2A" box="[94,150,1242,1267]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="1" pageNumber="2">INPA</collectionCode>
|
||
).”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA47D0EEC319A40FF39F78A" pageId="1" pageNumber="2" type="diagnosis">
|
||
<paragraph id="8B888625FFA47D0EEC319A40FF39F78A" blockId="1.[94,719,1306,2131]" pageId="1" pageNumber="2">
|
||
<emphasis id="B9435A37FFA47D0EEC319A40FF29FAEA" bold="true" box="[94,220,1306,1331]" pageId="1" pageNumber="2">Diagnosis:</emphasis>
|
||
medium sized specimens (
|
||
<quantity id="4CCF2BC0FFA47D0EEE769A40FD64FAEA" box="[537,657,1306,1331]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="1" pageNumber="2" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">22-25 mm</quantity>
|
||
total length); body convex; anterior frontal edge straight or slightly convex coarsely punctated; posterior mediofrontal area with large and rounded cephalic nodule; latero + mediofrontal tubercles large and conspicuous, with acute apices attached to inner tubercles (which are larger and with rounded apices) by weak anterofrontal ridges; small secondary mediofrontal tubercles, small- er than the latero + mediofrontal tubercles, with acute apices, near to each other; secondary laterofrontal tubercles nearly inconspicuous with slightly obtuse apices; posterofrontal ridges strong, elevated and straight, attached to the central tubercle, which is small, flat, with only apex pronounced; lateroposterior tubercles conspicuous and transverse; mentum with a dilated mediobasal area and large, deep lateral scars; anterior angles of pronotum acute and protruded; mesosternal scars shallow, triangular-shaped, with opaque surface and presence of micro bristles; metasternal disc barely delimited, with group of punctures in posterior region; metasternal lateral grooves narrow and glabrous; aedeagus elongated and narrow, median lobe elongated, larger than basal piece and parameres together, containing two sclerotized plates (ventral view); parameres separated from basal piece by conspicuous suture (ventral view).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA47D0DEF759F80FE28FA2A" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
|
||
<paragraph id="8B888625FFA47D0EEF759F80FC51FF2A" blockId="1.[794,932,218,243]" box="[794,932,218,243]" pageId="1" pageNumber="2">
|
||
<emphasis id="B9435A37FFA47D0EEF759F80FC51FF2A" bold="true" box="[794,932,218,243]" pageId="1" pageNumber="2">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA47D0EEF759E40FC84FE8A" blockId="1.[794,1417,282,339]" pageId="1" pageNumber="2">
|
||
<emphasis id="B9435A37FFA47D0EEF759E40FC2DFEEA" bold="true" box="[794,984,282,307]" pageId="1" pageNumber="2">
|
||
Body (
|
||
<figureCitation id="130C9AA0FFA47D0EEF0B9E40FC3FFEEA" box="[868,970,282,307]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="4.[170,229,2106,2131]" captionStartId-1="5.[94,153,2074,2099]" captionStartId-2="6.[170,229,2074,2099]" captionTargetBox-0="[170,1493,338,2077]" captionTargetBox-1="[94,1417,494,2044]" captionTargetBox-2="[177,1482,696,2032]" captionTargetId-0="figure-44@4.[170,1493,338,2077]" captionTargetId-1="figure-113@5.[94,1417,494,2044]" captionTargetId-2="figure-199@6.[170,1493,683,2044]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." captionText-1="Figure 2. Passalus (Pertinax) deuterocerus sp.nov.(A) headand prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." captionText-2="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi-0="http://doi.org/10.5281/zenodo.4985179" figureDoi-1="http://doi.org/10.5281/zenodo.4985183" figureDoi-2="http://doi.org/10.5281/zenodo.4985187" httpUri-0="https://zenodo.org/record/4985179/files/figure.png" httpUri-1="https://zenodo.org/record/4985183/files/figure.png" httpUri-2="https://zenodo.org/record/4985187/files/figure.png" pageId="1" pageNumber="2">Figs. 1-3</figureCitation>
|
||
):
|
||
</emphasis>
|
||
convex;
|
||
<emphasis id="B9435A37FFA47D0EE8569E40FB93FEEA" box="[1081,1126,282,307]" italics="true" pageId="1" pageNumber="2">size:</emphasis>
|
||
medium (
|
||
<quantity id="4CCF2BC0FFA47D0EE8B89E40FAB8FEEA" box="[1239,1357,282,307]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="1" pageNumber="2" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">22-25 mm</quantity>
|
||
, total length).
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA47D0EEF759E20FB70FA2A" blockId="1.[794,1418,378,1523]" pageId="1" pageNumber="2">
|
||
<emphasis id="B9435A37FFA47D0EEF759E20FB76FE4A" bold="true" box="[794,1155,378,403]" pageId="1" pageNumber="2">
|
||
Head (
|
||
<figureCitation id="130C9AA0FFA47D0EEF079E20FC47FE4A" box="[872,946,378,403]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
|
||
A-C, 2A-B, 3A-B):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA47D0EE8E39E20FB13FE4A" box="[1164,1254,378,403]" italics="true" pageId="1" pageNumber="2">Labrum:</emphasis>
|
||
straight anterior border.
|
||
<emphasis id="B9435A37FFA47D0EEFCB9EC0FC08FE6A" box="[932,1021,410,435]" italics="true" pageId="1" pageNumber="2">Clypeus:</emphasis>
|
||
hidden under frons, with anterior angles large, acute and positioned between latero + mediofrontal and secondary laterofrontal tubercles, visible in dorsal view; lateroclypeal pits deep.
|
||
<emphasis id="B9435A37FFA47D0EE95D9EA0FC5DFDEA" italics="true" pageId="1" pageNumber="2">Anterior frontal edge:</emphasis>
|
||
straight, sometimes slightly convex and narrow.
|
||
<emphasis id="B9435A37FFA47D0EEF169D60FB04FD8A" box="[889,1265,570,595]" italics="true" pageId="1" pageNumber="2">Secondary mediofrontal tubercles:</emphasis>
|
||
small, small- er than the latero + mediofrontal tubercles, with acute apices, near to each other.
|
||
<emphasis id="B9435A37FFA47D0EE83F9D20FCB2FD6A" italics="true" pageId="1" pageNumber="2">Latero + mediofrontal tubercles:</emphasis>
|
||
large, conspicuous, with acute apices, projected forward.
|
||
<emphasis id="B9435A37FFA47D0EEF109DE0FB1BFD0A" box="[895,1262,698,723]" italics="true" pageId="1" pageNumber="2">Secondary laterofrontal tubercles:</emphasis>
|
||
small, almost inconspicuous, with obtuse apices.
|
||
<emphasis id="B9435A37FFA47D0EE8D39D80FA7FFD2A" box="[1212,1418,730,755]" italics="true" pageId="1" pageNumber="2">Mediofrontal area:</emphasis>
|
||
caliciform 2.1× wider than long, shallow; with coarse punctures, scattered over anterior region, and an inconspicuous longitudinal groove between secondary mediofrontal tubercles.
|
||
<emphasis id="B9435A37FFA47D0EE8569C00FB00FCAA" box="[1081,1269,858,883]" italics="true" pageId="1" pageNumber="2">Cephalic nodule:</emphasis>
|
||
large, wellmarked, rounded.
|
||
<emphasis id="B9435A37FFA47D0EEF9A9C20FB6AFC4A" box="[1013,1183,890,915]" italics="true" pageId="1" pageNumber="2">Inner tubercles:</emphasis>
|
||
large, conspicuous, with rounded apices, slightly larger than latero + mediofrontal tubercles, slightly near to each other than to the central tubercle.
|
||
<emphasis id="B9435A37FFA47D0EE8539C80FAD5FC2A" box="[1084,1312,986,1011]" italics="true" pageId="1" pageNumber="2">Anterofrontal ridges:</emphasis>
|
||
straight, small, weak, sometimes inconspicuous.
|
||
<emphasis id="B9435A37FFA47D0EE89F9CA0FC94FBEA" italics="true" pageId="1" pageNumber="2">Posterofrontal ridges:</emphasis>
|
||
elevated, strong, nearly straight, starting at apex of central tubercle.
|
||
<emphasis id="B9435A37FFA47D0EE86D9B60FB42FB8A" box="[1026,1207,1082,1107]" italics="true" pageId="1" pageNumber="2">Central tubercle:</emphasis>
|
||
conical, flat, with apex not free and dilated posteriorly.
|
||
<emphasis id="B9435A37FFA47D0EE88B9B00FC74FB4A" italics="true" pageId="1" pageNumber="2">Lateroposterior tubercles:</emphasis>
|
||
conspicuous, transverse, slightly distant from central tubercle, with evident apex.
|
||
<emphasis id="B9435A37FFA47D0EE8BE9BC0FCB6FB0A" italics="true" pageId="1" pageNumber="2">Laterofrontal areas:</emphasis>
|
||
slightly deep and rough, surface with a few stretch marks.
|
||
<emphasis id="B9435A37FFA47D0EEF079B80FB9CFB2A" box="[872,1129,1242,1267]" italics="true" pageId="1" pageNumber="2">Lateropostfrontal areas:</emphasis>
|
||
deep, with rough surface.
|
||
<emphasis id="B9435A37FFA47D0EEF759BA0FC26FACA" box="[794,979,1274,1299]" italics="true" pageId="1" pageNumber="2">Postfrontal area:</emphasis>
|
||
shallow, smooth surface.
|
||
<emphasis id="B9435A37FFA47D0EE97C9BA0FC9FFAEA" italics="true" pageId="1" pageNumber="2">Postfrontal groove:</emphasis>
|
||
well-marked, lacking medial notch.
|
||
<emphasis id="B9435A37FFA47D0EE9719A40FC98FA8A" italics="true" pageId="1" pageNumber="2">Epicranial sutures:</emphasis>
|
||
well-marked.
|
||
<emphasis id="B9435A37FFA47D0EE87D9A60FB47FA8A" box="[1042,1202,1338,1363]" italics="true" pageId="1" pageNumber="2">Epicranial pits:</emphasis>
|
||
deep.
|
||
<emphasis id="B9435A37FFA47D0EE96B9A60FC51FAAA" italics="true" pageId="1" pageNumber="2">Anterior angles of head:</emphasis>
|
||
well-developed, with acute apices, slightly smaller than latero + mediofrontal tubercles.
|
||
<emphasis id="B9435A37FFA47D0EE9409A20FC91FA6A" italics="true" pageId="1" pageNumber="2">Canthus ocular:</emphasis>
|
||
apices almost straight, not reaching half of eye.
|
||
<emphasis id="B9435A37FFA47D0EEF759AE0FC7CFA0A" box="[794,905,1466,1491]" italics="true" pageId="1" pageNumber="2">Antennae:</emphasis>
|
||
trilamellated, with small, robust lamellae, distal lamella wider than other two.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA47D0EEF759940FB63F8AA" blockId="1.[794,1418,1562,1907]" pageId="1" pageNumber="2">
|
||
<emphasis id="B9435A37FFA47D0EEF759940FBACF9EA" bold="true" box="[794,1113,1562,1587]" pageId="1" pageNumber="2">
|
||
Mouthparts (
|
||
<figureCitation id="130C9AA0FFA47D0EEFD79940FBEEF9EA" box="[952,1051,1562,1587]" captionStart="Figure 2" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-113@5.[94,1417,494,2044]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) deuterocerus sp.nov.(A) headand prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985183" httpUri="https://zenodo.org/record/4985183/files/figure.png" pageId="1" pageNumber="2">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA47D0EE8459940FBB9F9EA" box="[1066,1100,1562,1587]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="1" pageNumber="2">3B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA47D0EE80B9940FB5EF9EA" box="[1124,1195,1562,1587]" italics="true" pageId="1" pageNumber="2">
|
||
<taxonomicName id="4C37FDA6FFA47D0EE80B9940FB53F9EA" box="[1124,1190,1562,1587]" class="Cestoda" family="Diphyllobothriidae" genus="Ligula" kingdom="Animalia" order="Diphyllobothriidea" pageId="1" pageNumber="2" phylum="Platyhelminthes" rank="genus">Ligula</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
tridentate, median tooth larger, narrower than lateral teeth.
|
||
<emphasis id="B9435A37FFA47D0EE8B99960FCBFF9AA" italics="true" pageId="1" pageNumber="2">Hypostomal process:</emphasis>
|
||
wide, glabrous, slightly close to mentum. Mentum:
|
||
<emphasis id="B9435A37FFA47D0EEF759920FC26F94A" box="[794,979,1658,1683]" italics="true" pageId="1" pageNumber="2">mediobasal area:</emphasis>
|
||
dilated, smooth, glabrous, with slightly protruded anterior region lacking median notch;
|
||
<emphasis id="B9435A37FFA47D0EE92A99C0FCA7F90A" italics="true" pageId="1" pageNumber="2">lateral lobes</emphasis>
|
||
with apices externally rounded and slightly straight at the inner face;
|
||
<emphasis id="B9435A37FFA47D0EEF8E9980FB97F92A" box="[993,1122,1754,1779]" italics="true" pageId="1" pageNumber="2">lateral scars</emphasis>
|
||
large, deep, glabrous, unpunctate, rounded, sometimes slightly oval.
|
||
<emphasis id="B9435A37FFA47D0EE97B99A0FA7FF8CA" box="[1300,1418,1786,1811]" italics="true" pageId="1" pageNumber="2">Mandibles:</emphasis>
|
||
incisor lobe with three well-formed apical teeth; suprainternal teeth robust; infrabasal pits inconspicuous.
|
||
<emphasis id="B9435A37FFA47D0EEF759800FC8FF8AA" box="[794,890,1882,1907]" italics="true" pageId="1" pageNumber="2">Maxillae:</emphasis>
|
||
lacinia bidentate at apex.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA47D0EEF7598C0FAA0F78A" blockId="1.[794,1418,1946,2131]" pageId="1" pageNumber="2">
|
||
<emphasis id="B9435A37FFA47D0EEF7598C0FB1BF86A" bold="true" box="[794,1262,1946,1971]" pageId="1" pageNumber="2">
|
||
Prothorax: Pronotum (
|
||
<figureCitation id="130C9AA0FFA47D0EE84F98C0FB6FF86A" box="[1056,1178,1946,1971]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="1" pageNumber="2">Figs. 1A, C</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA47D0EE8CB98C0FB16F86A" box="[1188,1251,1946,1971]" captionStart="Figure 2" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-113@5.[94,1417,494,2044]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) deuterocerus sp.nov.(A) headand prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985183" httpUri="https://zenodo.org/record/4985183/files/figure.png" pageId="1" pageNumber="2">2A, C</figureCitation>
|
||
):
|
||
</emphasis>
|
||
anterior edge slightly straight; anterior angles acute and protracted; marginal groove well-marked, deep, narrow, and punctate throughout, with apex dilated, reaching more than ⅓ of pronotal width; lateral fossae large, well-marked, deep and rounded, with group of coarse punctures.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59F80FF11FE8A" blockId="2.[170,794,218,339]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59F80FE5AFF2A" bold="true" box="[170,431,218,243]" pageId="2" pageNumber="3">
|
||
Prosternum (
|
||
<figureCitation id="130C9AA0FFA77D0DED269F80FE54FF2A" box="[329,417,218,243]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="2" pageNumber="3">Fig. 1B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA77D0DEDD49F80FDBFFF2A" box="[443,586,218,243]" italics="true" pageId="2" pageNumber="3">prepisternum</emphasis>
|
||
with a few long, sparse setae on outer face;
|
||
<emphasis id="B9435A37FFA77D0DEDBC9FA0FDA2FECA" box="[467,599,250,275]" italics="true" pageId="2" pageNumber="3">prepimerum</emphasis>
|
||
with a few sparse bristled punctures;
|
||
<emphasis id="B9435A37FFA77D0DEDEE9E40FDE5FEEA" box="[385,528,282,307]" italics="true" pageId="2" pageNumber="3">prosternelum</emphasis>
|
||
rhomboidal, with acute base.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59E20FEC2FE2A" blockId="2.[170,794,378,499]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59E20FDE8FE4A" bold="true" box="[170,541,378,403]" pageId="2" pageNumber="3">
|
||
Mesosternum (
|
||
<figureCitation id="130C9AA0FFA77D0DED359E20FE42FE4A" box="[346,439,378,403]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="2" pageNumber="3">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA77D0DEDAD9E20FE11FE4A" box="[450,484,378,403]" captionStart="Figure 2" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-113@5.[94,1417,494,2044]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) deuterocerus sp.nov.(A) headand prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985183" httpUri="https://zenodo.org/record/4985183/files/figure.png" pageId="2" pageNumber="3">2D</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA77D0DED819E20FDFAFE4A" box="[494,527,378,403]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">3B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
smooth, glabrous;
|
||
<emphasis id="B9435A37FFA77D0DEE9F9E20FEBDFE6A" italics="true" pageId="2" pageNumber="3">mesosternal scars</emphasis>
|
||
triangular-shaped, shallow, with opaque surface containing micro bristles, without punctures or pubescence.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59D40FD8BFD6A" blockId="2.[170,795,538,691]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59D40FDD7FDEA" bold="true" box="[170,546,538,563]" pageId="2" pageNumber="3">
|
||
Metasternum (
|
||
<figureCitation id="130C9AA0FFA77D0DED369D40FE4FFDEA" box="[345,442,538,563]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="2" pageNumber="3">Figs. 1B</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA77D0DEDA79D40FE12FDEA" box="[456,487,538,563]" captionStart="Figure 2" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-113@5.[94,1417,494,2044]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) deuterocerus sp.nov.(A) headand prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985183" httpUri="https://zenodo.org/record/4985183/files/figure.png" pageId="2" pageNumber="3">2E</figureCitation>
|
||
,
|
||
<figureCitation id="130C9AA0FFA77D0DED9B9D40FDE0FDEA" box="[500,533,538,563]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">3B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA77D0DEE449D40FDA1FDEA" box="[555,596,538,563]" italics="true" pageId="2" pageNumber="3">disc</emphasis>
|
||
barely delimited, more dilated than lateral regions;
|
||
<emphasis id="B9435A37FFA77D0DEE519D60FD40FD8A" box="[574,693,570,595]" italics="true" pageId="2" pageNumber="3">punctation</emphasis>
|
||
forming group of punctures on lateroposterior region;
|
||
<emphasis id="B9435A37FFA77D0DEEBD9D00FF13FD4A" italics="true" pageId="2" pageNumber="3">pubescence</emphasis>
|
||
absent;
|
||
<emphasis id="B9435A37FFA77D0DED289D20FE29FD4A" box="[327,476,634,659]" italics="true" pageId="2" pageNumber="3">lateral groove</emphasis>
|
||
narrow, thinner than mesotibiae, deep, without punctures or bristles.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59D80FEBFFCAA" blockId="2.[170,795,730,883]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59D80FE73FD2A" bold="true" box="[170,390,730,755]" pageId="2" pageNumber="3">
|
||
Elytra (
|
||
<figureCitation id="130C9AA0FFA77D0DEC939D80FE8FFD2A" box="[252,378,730,755]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="2" pageNumber="3">Figs. 1A, C</figureCitation>
|
||
):
|
||
</emphasis>
|
||
approximately 2.5× longer and 1.1× wider than pronotum;
|
||
<emphasis id="B9435A37FFA77D0DEDDD9DA0FE18FCCA" box="[434,493,762,787]" italics="true" pageId="2" pageNumber="3">striae</emphasis>
|
||
narrower than interstriae, marked by circular and inconspicuous punctures on dorsal striae, well-defined in lateral striae;
|
||
<emphasis id="B9435A37FFA77D0DEE379C60FD49FC8A" box="[600,700,826,851]" italics="true" pageId="2" pageNumber="3">epipleura</emphasis>
|
||
and humeri glabrous.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59CC0FCEEFBEA" blockId="2.[170,795,922,1075]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59CC0FE17FC6A" bold="true" box="[170,482,922,947]" pageId="2" pageNumber="3">
|
||
Legs (
|
||
<figureCitation id="130C9AA0FFA77D0DEC809CC0FECDFC6A" box="[239,312,922,947]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
|
||
A-C, 2A, C, E):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA77D0DED859CC0FDBBFC6A" box="[490,590,922,947]" italics="true" pageId="2" pageNumber="3">profemur</emphasis>
|
||
with well-marked groove on anteroventral border not reaching apex; posteroventral border with a few setae near apex;
|
||
<emphasis id="B9435A37FFA77D0DEED69C80FCEFFC2A" box="[697,794,986,1011]" italics="true" pageId="2" pageNumber="3">protibiae</emphasis>
|
||
not dilated;
|
||
<emphasis id="B9435A37FFA77D0DED5D9CA0FE5DFBCA" box="[306,424,1018,1043]" italics="true" pageId="2" pageNumber="3">mesotibiae</emphasis>
|
||
with one or two small spines on outer face;
|
||
<emphasis id="B9435A37FFA77D0DED4C9B40FE62FBEA" box="[291,407,1050,1075]" italics="true" pageId="2" pageNumber="3">metatibiae</emphasis>
|
||
with one small spine on outer face.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59B00FDEEFB4A" blockId="2.[170,794,1114,1171]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59B00FE63FBAA" bold="true" box="[170,406,1114,1139]" pageId="2" pageNumber="3">
|
||
Abdomen (
|
||
<figureCitation id="130C9AA0FFA77D0DED5D9B00FE7DFBAA" box="[306,392,1114,1139]" captionStart="Figure 1" captionStartId="4.[170,229,2106,2131]" captionTargetBox="[170,1493,338,2077]" captionTargetId="figure-44@4.[170,1493,338,2077]" captionTargetPageId="4" captionText="Figure 1. Passalus (Pertinax) deuterocerussp.nov.,habitus.(A) dorsal;(B) ventral;(C) lateral." figureDoi="http://doi.org/10.5281/zenodo.4985179" httpUri="https://zenodo.org/record/4985179/files/figure.png" pageId="2" pageNumber="3">Fig. 1B</figureCitation>
|
||
):
|
||
</emphasis>
|
||
sternite VII with incomplete and well-marked groove; sides rough.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59BE0FE28FA2A" blockId="2.[170,795,1210,1523]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59BE0FE78FB0A" bold="true" box="[170,397,1210,1235]" pageId="2" pageNumber="3">
|
||
Aedeagus (
|
||
<figureCitation id="130C9AA0FFA77D0DED419BE0FE8AFB0A" box="[302,383,1210,1235]" captionStart="Figure 2" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[94,1417,494,2044]" captionTargetId="figure-113@5.[94,1417,494,2044]" captionTargetPageId="5" captionText="Figure 2. Passalus (Pertinax) deuterocerus sp.nov.(A) headand prothorax,dorsal;(B) head,ventral;(C) head and prothorax,dorsolateral;(D)posterior region of the prosternum and anterior region of the mesosternum;(E) metasternum (left side);(F) aedeagus (ventral,dorsal and lateral view)." figureDoi="http://doi.org/10.5281/zenodo.4985183" httpUri="https://zenodo.org/record/4985183/files/figure.png" pageId="2" pageNumber="3">Fig. 2F</figureCitation>
|
||
):
|
||
</emphasis>
|
||
<emphasis id="B9435A37FFA77D0DEDFC9BE0FDEBFB0A" box="[403,542,1210,1235]" italics="true" pageId="2" pageNumber="3">Median lobe:</emphasis>
|
||
narrower than parameres,almost as long as parameres and basal piece togeth- er; ventrally with two lateral sclerotized plates lacking at median region. Basal piece separated from parameres by distinct suture.
|
||
<emphasis id="B9435A37FFA77D0DED3C9A60FE31FA8A" box="[339,452,1338,1363]" italics="true" pageId="2" pageNumber="3">Parameres</emphasis>
|
||
ventrally, with median anterior region with central emargination; laterally, with rounded projections almost reaching half of median lobe length; dorsally, with unjoined projections.
|
||
<emphasis id="B9435A37FFA77D0DEE579AC0FD5BFA6A" box="[568,686,1434,1459]" italics="true" pageId="2" pageNumber="3">Basal piece</emphasis>
|
||
ventrally, short, with convexity in posteromedian region; dorsally lateral projections separate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA77D0DECC59940FCEEF8EA" pageId="2" pageNumber="3" type="etymology">
|
||
<paragraph id="8B888625FFA77D0DECC59940FDAAF96A" blockId="2.[170,794,1562,1715]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59940FEC7F9EA" bold="true" box="[170,306,1562,1587]" pageId="2" pageNumber="3">Etymology:</emphasis>
|
||
The specific epithet derives from the union of two Greek words, “deúteros” (“secondary” or “double”) and “keros” (“horn” or “any hornlike projection”), in reference to the two secondary mediofrontal tubercles, the most remarkable feature of the species.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DECC59980FCEEF8EA" blockId="2.[170,795,1754,1843]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59980FE9FF92A" bold="true" box="[170,362,1754,1779]" pageId="2" pageNumber="3">Polymorphisms:</emphasis>
|
||
The secondary mediofrontal tubercles may vary in size and distance from each other; the anterofrontal ridges may be almost absent in some specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32DD5AEFFA77D0DECC59800FAB9F78A" pageId="2" pageNumber="3" type="discussion">
|
||
<paragraph id="8B888625FFA77D0DECC59800FB08F94A" blockId="2.[170,795,1882,2131]" lastBlockId="2.[869,1495,218,2131]" pageId="2" pageNumber="3">
|
||
<emphasis id="B9435A37FFA77D0DECC59800FEE2F8AA" bold="true" box="[170,279,1882,1907]" pageId="2" pageNumber="3">Remarks:</emphasis>
|
||
Due to the convex body, well-developed inner tubercles,barely delimited metasternal disc,and metasternal pubescence absent,
|
||
<taxonomicName id="4C37FDA6FFA77D0DEDA898C0FD46F86A" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[455,691,1946,1971]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA77D0DEDA898C0FD46F86A" box="[455,691,1946,1971]" italics="true" pageId="2" pageNumber="3">Passalus deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA77D0DEEAF98C0FCEEF86A" box="[704,795,1946,1971]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is assigned to the subgenus
|
||
<emphasis id="B9435A37FFA77D0DED8798E0FD40F80A" box="[488,693,1978,2003]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C37FDA6FFA77D0DED8798E0FD44F80A" authority="" box="[488,689,1978,2003]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Pertinax">Passalus (Pertinax)</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It is very similar to
|
||
<taxonomicName id="4C37FDA6FFA77D0DED789880FDCBF82A" authority="Percheron, 1841" authorityName="Percheron" authorityYear="1841" box="[279,574,2010,2035]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="latifrons">
|
||
<emphasis id="B9435A37FFA77D0DED789880FE71F82A" box="[279,388,2010,2035]" italics="true" pageId="2" pageNumber="3">P. latifrons</emphasis>
|
||
Percheron, 1841
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C37FDA6FFA77D0DEE269880FF16F7CA" authority="Percheron, 1835" authorityName="Percheron" authorityYear="1835" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="morio">
|
||
<emphasis id="B9435A37FFA77D0DEE269880FD68F82A" box="[585,669,2010,2035]" italics="true" pageId="2" pageNumber="3">P. morio</emphasis>
|
||
Percheron, 1835
|
||
</taxonomicName>
|
||
,and especially to
|
||
<taxonomicName id="4C37FDA6FFA77D0DEDCB98A0FD1EF7CA" authority="(Kuwert, 1891)" baseAuthorityName="Kuwert" baseAuthorityYear="1891" box="[420,747,2042,2067]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="epiphanoides">
|
||
<emphasis id="B9435A37FFA77D0DEDCB98A0FDB2F7CA" box="[420,583,2042,2067]" italics="true" pageId="2" pageNumber="3">P.epiphanoides</emphasis>
|
||
(Kuwert, 1891)
|
||
</taxonomicName>
|
||
.The differences between
|
||
<taxonomicName id="4C37FDA6FFA77D0DEDE39740FDDDF7EA" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[396,552,2074,2099]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA77D0DEDE39740FDDDF7EA" box="[396,552,2074,2099]" italics="true" pageId="2" pageNumber="3">P.deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA77D0DEE439740FD8BF7EA" box="[556,638,2074,2099]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C37FDA6FFA77D0DEEC09740FCEFF7EA" authorityName="Percheron" authorityYear="1841" box="[687,794,2074,2099]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="latifrons">
|
||
<emphasis id="B9435A37FFA77D0DEEC09740FCEFF7EA" box="[687,794,2074,2099]" italics="true" pageId="2" pageNumber="3">P.latifrons</emphasis>
|
||
</taxonomicName>
|
||
are more pronounced because the new species has small- er size, two secondary mediofrontal tubercles, mesosternal scars, and absence of pubescence on posterior region of the pronotum ventrally,while
|
||
<taxonomicName id="4C37FDA6FFA77D0DE8AC9E40FAD8FEEA" authorityName="Percheron" authorityYear="1841" box="[1219,1325,282,307]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="latifrons">
|
||
<emphasis id="B9435A37FFA77D0DE8AC9E40FAD8FEEA" box="[1219,1325,282,307]" italics="true" pageId="2" pageNumber="3">P.latifrons</emphasis>
|
||
</taxonomicName>
|
||
is larger,has no secondary mediofrontal tubercles or mesosternal scars, and has pubescence on the posterior region of the pronotum ventrally (
|
||
<tableCitation id="C6B5B39EFFA77D0DE84A9E20FB83FE4A" box="[1061,1142,378,403]" captionStart="Table 1" captionStartId="3.[94,144,218,243]" captionText="Table 1. Differential characters of Passalus deuterocerussp.nov.,P.morio Percheron,1835, P.latifrons Percheron,1841 and P.epiphanoides (Kuwert,1891)." pageId="2" pageNumber="3">Table 1</tableCitation>
|
||
).
|
||
<taxonomicName id="4C37FDA6FFA77D0DE8E19E20FA82FE4A" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[1166,1399,378,403]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA77D0DE8E19E20FA82FE4A" box="[1166,1399,378,403]" italics="true" pageId="2" pageNumber="3">Passalus deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA77D0DE9EF9E20FA23FE4A" box="[1408,1494,378,403]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C37FDA6FFA77D0DEF849EC0FBCAFE6A" authorityName="Percheron" authorityYear="1835" box="[1003,1087,410,435]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="morio">
|
||
<emphasis id="B9435A37FFA77D0DEF849EC0FBCAFE6A" box="[1003,1087,410,435]" italics="true" pageId="2" pageNumber="3">P. morio</emphasis>
|
||
</taxonomicName>
|
||
by having acute latero + mediofrontal tubercles (obtuse in
|
||
<taxonomicName id="4C37FDA6FFA77D0DE8059EE0FB4AFE0A" authorityName="Percheron" authorityYear="1835" box="[1130,1215,442,467]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="morio">
|
||
<emphasis id="B9435A37FFA77D0DE8059EE0FB4AFE0A" box="[1130,1215,442,467]" italics="true" pageId="2" pageNumber="3">P. morio</emphasis>
|
||
</taxonomicName>
|
||
), anterior frontal edge of the head thinner (which is thicker in
|
||
<taxonomicName id="4C37FDA6FFA77D0DE89F9E80FAB1FE2A" authorityName="Percheron" authorityYear="1835" box="[1264,1348,474,499]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="morio">
|
||
<emphasis id="B9435A37FFA77D0DE89F9E80FAB1FE2A" box="[1264,1348,474,499]" italics="true" pageId="2" pageNumber="3">P.morio</emphasis>
|
||
</taxonomicName>
|
||
), presence of secondary mediofrontal tubercles (absent in
|
||
<taxonomicName id="4C37FDA6FFA77D0DE9289EA0FA6CFDCA" authorityName="Percheron" authorityYear="1835" box="[1351,1433,506,531]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="morio">
|
||
<emphasis id="B9435A37FFA77D0DE9289EA0FA6CFDCA" box="[1351,1433,506,531]" italics="true" pageId="2" pageNumber="3">P.morio</emphasis>
|
||
</taxonomicName>
|
||
), and by the general shape of aedeagus (
|
||
<tableCitation id="C6B5B39EFFA77D0DE8819D40FACBFDEA" box="[1262,1342,538,563]" captionStart="Table 1" captionStartId="3.[94,144,218,243]" captionText="Table 1. Differential characters of Passalus deuterocerussp.nov.,P.morio Percheron,1835, P.latifrons Percheron,1841 and P.epiphanoides (Kuwert,1891)." pageId="2" pageNumber="3">Table 1</tableCitation>
|
||
). When comparing the new species with
|
||
<emphasis id="B9435A37FFA77D0DE8F89D60FACAFD8A" box="[1175,1343,570,595]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C37FDA6FFA77D0DE8F89D60FACEFD8A" baseAuthorityName="Kuwert" baseAuthorityYear="1891" box="[1175,1339,570,595]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="epiphanoides">P.epiphanoides</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the differences are more subtle and require attention.
|
||
<bibRefCitation id="EFA6FBD4FFA77D0DE9519D00FC5FFD4A" author="Reyes-Castillo, P." pageId="2" pageNumber="3" pagination="1541 - 1587" refId="ref8490" refString="Reyes-Castillo, P. 1973. Passalidae de la Guayana Francesa (Coleptera, Lamellicornia). Bulletin du Museum National d'Histoire Naturelle, Serie Zoologie, 129 (197): 1541 - 1587." type="journal article" year="1973">Reyes-Castillo (1973)</bibRefCitation>
|
||
redescribed
|
||
<taxonomicName id="4C37FDA6FFA77D0DE8549D20FB17FD4A" baseAuthorityName="Kuwert" baseAuthorityYear="1891" box="[1083,1250,634,659]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="epiphanoides">
|
||
<emphasis id="B9435A37FFA77D0DE8549D20FB17FD4A" box="[1083,1250,634,659]" italics="true" pageId="2" pageNumber="3">P. epiphanoides</emphasis>
|
||
</taxonomicName>
|
||
adding some characters as interspecific variations, such as protruding edges of the median notch of the anterior frontal edge of the head forming two secondary mediofrontal tubercles and mesosternal scars, which may be presented as two triangular-shaped matte areas covered with submicroscopic spicules. However, a reexamination the specimens studied by
|
||
<bibRefCitation id="EFA6FBD4FFA77D0DEFC19C00FB22FCAA" author="Bevilaqua, M. & Fonseca, C. R. V." box="[942,1239,858,883]" pageId="2" pageNumber="3" pagination="449 - 466" refId="ref7425" refString="Bevilaqua, M. & Fonseca, C. R. V. 2019. Passalidae (Coleoptera: Scarabaeoidea) from the West-Most Brazilian Amazon Region: Checklist, New Records, and Identification Key. Neotropical Entomology, 48 (3): 449 - 466. DOI" type="journal article" year="2019">Bevilaqua & Fonseca (2019)</bibRefCitation>
|
||
showed that these features are constant in series of specimens. A better analysis showed that these so-called polymorphic characters are diagnostic features that distinguish the two species.Thus,
|
||
<taxonomicName id="4C37FDA6FFA77D0DEF0A9C80FBF7FC2A" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[869,1026,986,1011]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA77D0DEF0A9C80FBF7FC2A" box="[869,1026,986,1011]" italics="true" pageId="2" pageNumber="3">P.deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA77D0DE8689C80FBACFC2A" box="[1031,1113,986,1011]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be differentiated from
|
||
<taxonomicName id="4C37FDA6FFA77D0DE9129C80FC4DFBCA" baseAuthorityName="Kuwert" baseAuthorityYear="1891" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="epiphanoides">
|
||
<emphasis id="B9435A37FFA77D0DE9129C80FC4DFBCA" italics="true" pageId="2" pageNumber="3">P.epiphanoides</emphasis>
|
||
</taxonomicName>
|
||
by the narrower mediofrontal area (
|
||
<figureCitation id="130C9AA0FFA77D0DE9249CA0FA6FFBCA" box="[1355,1434,1018,1043]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3A</figureCitation>
|
||
); two secondary mediofrontal tubercles (
|
||
<figureCitation id="130C9AA0FFA77D0DE89F9B40FAB7FBEA" box="[1264,1346,1050,1075]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3A</figureCitation>
|
||
); mediobasal area of the mentum with rounded anterior region, without punctures or bristles (
|
||
<figureCitation id="130C9AA0FFA77D0DE8E19B00FB28FBAA" box="[1166,1245,1114,1139]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3B</figureCitation>
|
||
); shallow mesosternal scars with slightly triangular shape with matte surface and presence of micro bristles (
|
||
<figureCitation id="130C9AA0FFA77D0DE8AE9BC0FAFAFB6A" box="[1217,1295,1178,1203]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3C</figureCitation>
|
||
); metasternal disc with a group of punctures on the lateroposterior region (
|
||
<figureCitation id="130C9AA0FFA77D0DEF039B80FC4CFB2A" box="[876,953,1242,1267]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3C</figureCitation>
|
||
); narrower aedeagus, with median lobe with narrower sclerotized plates, parameres with median anterior emargination and basal piece with concavity at posterior edge.
|
||
<taxonomicName id="4C37FDA6FFA77D0DEFC69A60FB62FA8A" baseAuthorityName="Kuwert" baseAuthorityYear="1891" box="[937,1175,1338,1363]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="epiphanoides">
|
||
<emphasis id="B9435A37FFA77D0DEFC69A60FB62FA8A" box="[937,1175,1338,1363]" italics="true" pageId="2" pageNumber="3">Passalus epiphanoides</emphasis>
|
||
</taxonomicName>
|
||
presents wider mediofrontal area (
|
||
<figureCitation id="130C9AA0FFA77D0DEFCD9A00FC04FAAA" box="[930,1009,1370,1395]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3D</figureCitation>
|
||
), absence of secondary mediofrontal tubercles (
|
||
<figureCitation id="130C9AA0FFA77D0DEFF39A20FC19FA4A" box="[924,1004,1402,1427]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3D</figureCitation>
|
||
), mediobasal area of the mentum may have punctures or bristles and a notch in the anterior region (
|
||
<figureCitation id="130C9AA0FFA77D0DEF039AE0FC4DFA0A" box="[876,952,1466,1491]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3E</figureCitation>
|
||
), deep and oval or drop shape mesosternal scars with matt surface and presence of micro bristles (
|
||
<figureCitation id="130C9AA0FFA77D0DE9EE9A80FA3EFA2A" box="[1409,1483,1498,1523]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3F</figureCitation>
|
||
), metasternal disc with a small group of punctures in the lateroposterior region (
|
||
<figureCitation id="130C9AA0FFA77D0DE80F9940FB5FF9EA" box="[1120,1194,1562,1587]" captionStart="Figure 3" captionStartId="6.[170,229,2074,2099]" captionTargetBox="[177,1482,696,2032]" captionTargetId="figure-199@6.[170,1493,683,2044]" captionTargetPageId="6" captionText="Figure 3. Passalus(Pertinax) deuterocerus sp.nov.(A) cephaliccapsule,dorsal;(B)mentum;(C) meso- and metasternum.Passalus (Pertinax) epiphanoides (Kuwert, 1891):(D) cephalic capsule,dorsal.(E)mentum. (F)meso- and metasternum." figureDoi="http://doi.org/10.5281/zenodo.4985187" httpUri="https://zenodo.org/record/4985187/files/figure.png" pageId="2" pageNumber="3">Fig. 3F</figureCitation>
|
||
), wider aedeagus, with median lobe with wider sclerotized plates, parameres with median convexity on the anterior edge and basal piece with convexity on the posterior edge.
|
||
</paragraph>
|
||
<paragraph id="8B888625FFA77D0DEFE499C0FAB9F78A" blockId="2.[869,1495,218,2131]" pageId="2" pageNumber="3">
|
||
The main difference between
|
||
<taxonomicName id="4C37FDA6FFA77D0DE8B599C0FA8FF96A" authority="Bevilaqua & Fonseca, 2020" authorityName="Bevilaqua & Fonseca" authorityYear="2020" box="[1242,1402,1690,1715]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="deuterocerus" status="sp. nov.">
|
||
<emphasis id="B9435A37FFA77D0DE8B599C0FA8FF96A" box="[1242,1402,1690,1715]" italics="true" pageId="2" pageNumber="3">P. deuterocerus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A270E74CFFA77D0DE9EE99C0FA23F96A" box="[1409,1494,1690,1715]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and other species currently allocated in
|
||
<emphasis id="B9435A37FFA77D0DE97699E0FA60F90A" box="[1305,1429,1722,1747]" italics="true" pageId="2" pageNumber="3">P.(Pertinax)</emphasis>
|
||
is the presence of secondary mediofrontal tubercles, a characteristic that apparently belongs to the Rhodocanthopus group. However, other characters such as eyes reduction, presence of secondary inner tubercles and presence of large spines on the external face of the meso- and metatibia, besides the general shape of the aedeagus (Jiménez- Ferbans
|
||
<emphasis id="B9435A37FFA77D0DEFAD98C0FC0EF86A" box="[962,1019,1946,1971]" italics="true" pageId="2" pageNumber="3">et al.,</emphasis>
|
||
2016a), do not allow the new species to be allocate in this group. Furthermore, it seems to be more related to species belonging to
|
||
<emphasis id="B9435A37FFA77D0DE8BA9880FAA0F82A" box="[1237,1365,2010,2035]" italics="true" pageId="2" pageNumber="3">P. (Pertinax)</emphasis>
|
||
s. str. such as
|
||
<emphasis id="B9435A37FFA77D0DEFE898A0FAECF7CA" box="[903,1305,2042,2067]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C37FDA6FFA77D0DEFE898A0FC17F7CA" authorityName="Percheron" authorityYear="1835" box="[903,994,2042,2067]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="morio">P. morio</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C37FDA6FFA77D0DEF8098A0FB97F7CA" authorityName="Percheron" authorityYear="1841" box="[1007,1122,2042,2067]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="latifrons">P. latifrons</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C37FDA6FFA77D0DE80098A0FAECF7CA" baseAuthorityName="Kuwert" baseAuthorityYear="1891" box="[1135,1305,2042,2067]" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="epiphanoides">P. epiphanoides</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C37FDA6FFA77D0DE93698A0FBF7F7EA" authority="Dalman, 1817" authorityName="Dalman" authorityYear="1817" class="Insecta" family="Passallidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="convexus">
|
||
<emphasis id="B9435A37FFA77D0DE93698A0FA20F7CA" box="[1369,1493,2042,2067]" italics="true" pageId="2" pageNumber="3">P. convexus</emphasis>
|
||
Dalman, 1817
|
||
</taxonomicName>
|
||
. However, phylogenetic analysis that is in process may have clarified this issue further.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |