treatments-xml/data/03/96/87/039687F1FFC9FF85FF06FEB4FB4D9ED7.xml

<document id="D88432A919B345BCDE69EE2F64EFD259" ID-CLB-Dataset="27679" ID-DOI="10.11646/zootaxa.4544.1.8" ID-GBIF-Dataset="ae29b145-77e4-4ead-85f2-4f5ceabb94fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="2618205" ID-ZooBank="853D2455-55B3-4695-8AE1-89355E87D0B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553929234917" checkinUser="plazi" docAuthor="Reyes, Scarleth Margarita Raudez, Espinoza, Roberto Antonio Cano &amp; Girón, Jennifer C." docDate="2019" docId="039687F1FFC9FF85FF06FEB4FB4D9ED7" docLanguage="en" docName="zootaxa.4544.1.8.pdf" docOrigin="Zootaxa 4544 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yara marmontsedu Reyes &amp; Espinoza &amp; Girón 2019, n. sp." docType="treatment" docUuid="7824E895-F234-4DC6-A6DE-BA416852E819" docUuidSource="ZooBank" docVersion="9" lastPageNumber="133" masterDocId="FFAFFF89FFCBFF8DFF91FFF8FF849F7B" masterDocTitle="A new species and immature stages of the skiff beetle genus Yara Reichardt and Hinton (Coleoptera: Myxophaga: Hydroscaphidae) from Nicaragua" masterLastPageNumber="134" masterPageNumber="125" pageNumber="127" updateTime="1698715618769" updateUser="ExternalLinkService">
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<mods:title id="AD8A55384F145C45CC027248660EDA00">A new species and immature stages of the skiff beetle genus Yara Reichardt and Hinton (Coleoptera: Myxophaga: Hydroscaphidae) from Nicaragua</mods:title>
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<mods:namePart id="BC977D0FA7C9102975F96A2791DE082D">Reyes, Scarleth Margarita Raudez</mods:namePart>
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<mods:namePart id="FAAE4A820DF5C1D170FD3E595156DCAB">Espinoza, Roberto Antonio Cano</mods:namePart>
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<taxonomicName id="4C3F4D64FFC9FF8FFF06FEB4FEF89E1D" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[151,380,332,358]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC9FF8FFF06FEB4FEF89E1D" bold="true" box="[151,380,332,358]" italics="true" pageId="2" pageNumber="127">Yara marmontsedu</emphasis>
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<paragraph id="8B8036E7FFC9FF8FFF06FE96FE909EFC" blockId="2.[151,454,332,391]" box="[151,276,366,391]" pageId="2" pageNumber="127">
<figureCitation id="13042A62FFC9FF8FFF06FE96FE949EFC" box="[151,272,366,391]" captionStart-0="FIGURES 1–6" captionStart-1="FIGURES 7–12" captionStart-2="FIGURES 13–16" captionStart-3="FIGURES 17–25" captionStart-4="FIGURES 26–38" captionStart-5="FIGURES 39–42" captionStart-6="FIGURES 43–47" captionStartId-0="2.[151,264,1415,1438]" captionStartId-1="3.[151,264,729,752]" captionStartId-2="3.[151,264,1345,1368]" captionStartId-3="5.[151,264,1857,1880]" captionStartId-4="6.[151,264,1866,1889]" captionStartId-5="7.[151,264,950,973]" captionStartId-6="7.[151,264,1843,1866]" captionTargetBox-0="[263,1324,696,1394]" captionTargetBox-1="[199,1381,193,708]" captionTargetBox-2="[198,1411,878,1308]" captionTargetBox-3="[278,1298,193,1823]" captionTargetBox-4="[285,1290,242,1815]" captionTargetBox-5="[247,1320,208,922]" captionTargetBox-6="[197,1389,1092,1822]" captionTargetId-0="figure@2.[263,1324,695,1394]" captionTargetId-1="figure@3.[199,1387,193,708]" captionTargetId-2="figure@3.[167,1420,863,1324]" captionTargetId-3="figure@5.[278,1298,193,1826]" captionTargetId-4="figure@6.[271,1306,223,1824]" captionTargetId-5="figure@7.[219,1356,193,933]" captionTargetId-6="figure@7.[196,1390,1092,1822]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="7" captionText-0="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." captionText-1="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." captionText-2="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." captionText-3="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." captionText-4="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." captionText-5="FIGURES 39–42. Pupa of Yara marmontsedu: (39) dorso-lateral view; (40) detail of head and antenna; (41) detail of pretarsus; (42) detail of apex of abdomen." captionText-6="FIGURES 43–47. Distribution and collecting: (43) map for species of Yara; (44) map for Central American species of Yara; (45) sampling locality at La Libertad, Chontales, Nicaragua; (46) Pijibay Creek; (47) collecting macroinvertebrates using a Dnet at Pijibay Creek." httpUri-0="https://zenodo.org/record/2618207/files/figure.png" httpUri-1="https://zenodo.org/record/2618209/files/figure.png" httpUri-2="https://zenodo.org/record/2618211/files/figure.png" httpUri-3="https://zenodo.org/record/2618213/files/figure.png" httpUri-4="https://zenodo.org/record/2618215/files/figure.png" httpUri-5="https://zenodo.org/record/2618217/files/figure.png" httpUri-6="https://zenodo.org/record/2618219/files/figure.png" pageId="2" pageNumber="127">Figs. 1–47</figureCitation>
.
</paragraph>
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<paragraph id="8B8036E7FFC9FF8FFF06FE4EFD829D6C" blockId="2.[151,1436,438,643]" pageId="2" pageNumber="127">
<emphasis id="B94BEAF5FFC9FF8FFF06FE4EFDA39EB4" bold="true" box="[151,551,438,463]" pageId="2" pageNumber="127">
Type material. 
<materialsCitation id="3B573CBAFFC9FF8FFEC9FE4EFDA39EB4" ID-GBIF-Occurrence="2147525269" box="[344,551,438,463]" collectingDate="2012" collectingDateMax="2013" collectingDateMin="2012" collectionCode="CIRA, UNAN" collectorName="Col. Scarleth Raudez" country="Nicaragua" location="Municipality of La Libertad" municipality="Pijibay Creek" pageId="2" pageNumber="127" specimenCount="1" stateProvince="Chontales" typeStatus="holotype">
<typeStatus id="54848845FFC9FF8FFEC9FE4EFE419EB4" box="[344,453,438,463]" pageId="2" pageNumber="127" type="holotype">Holotype</typeStatus>
(male):
</materialsCitation>
</emphasis>
“ 
<collectingCountry id="F3287677FFC9FF8FFDAFFE4FFD6D9EB4" box="[574,745,439,463]" name="Nicaragua" pageId="2" pageNumber="127">NICARAGUA</collectingCountry>
/ 
<collectingMunicipality id="6BE4AC9DFFC9FF8FFC9BFE4EFC2E9EB4" box="[778,938,438,463]" pageId="2" pageNumber="127">Pijibay Creek</collectingMunicipality>
/ 
<location id="8EE0603CFFC9FF8FFC5DFE4EFA929EB4" LSID="urn:lsid:plazi:treatment:039687F1FFC9FF85FF06FEB4FB4D9ED7:8EE0603CFFC9FF8FFC5DFE4EFA929EB4" box="[972,1302,438,463]" country="Nicaragua" municipality="Pijibay Creek" name="Municipality of La Libertad" pageId="2" pageNumber="127" stateProvince="Chontales">Municipality of La Libertad</location>
/ 
<collectingRegion id="49FBF805FFC9FF8FFABDFE4EFE9F9E89" country="Nicaragua" name="Chontales" pageId="2" pageNumber="127">Chontales Department</collectingRegion>
/ 
<collectingDate id="EFC5E9CFFFC9FF8FFEA2FE23FE179E88" box="[307,403,475,499]" pageId="2" pageNumber="127" value="2012" valueMax="2013">
<date id="FF811027FFC9FF8FFEA2FE23FE179E88" box="[307,403,475,499]" pageId="2" pageNumber="127" value="2012" valueMax="2013">2012-13</date>
</collectingDate>
. In a shallow environment with grasses and clay. 
<collectorName id="26CA5331FFC9FF8FFC56FE22FB329E88" box="[967,1206,474,499]" pageId="2" pageNumber="127">Col. Scarleth Ráudez</collectorName>
/ Roberto Cano and Rafael Varela”. (CIRA/ UNAN).
</paragraph>
<paragraph id="8B8036E7FFC9FF8FFF56FDDAFDDC9DF8" blockId="2.[151,1436,438,643]" pageId="2" pageNumber="127">
<materialsCitation id="3B573CBAFFC9FF8FFF56FDDAFDD09DF8" ID-GBIF-Occurrence="2147525336" collectingDate="2012" collectingDateMax="2013" collectingDateMin="2012" collectionCode="CIRA, UNAN, SEMC, USNM" collectorName="Col. Scarleth Raudez" country="Nicaragua" location="Municipality of La Libertad" municipality="Pijibay Creek" pageId="2" pageNumber="127" specimenCount="39" specimenCount-female="19" specimenCount-male="20" stateProvince="La Libertad" typeStatus="paratype">
<emphasis id="B94BEAF5FFC9FF8FFF56FDDAFD6E9D40" bold="true" box="[199,746,546,571]" pageId="2" pageNumber="127">
<typeStatus id="54848845FFC9FF8FFF56FDDAFEBB9D40" box="[199,319,546,571]" pageId="2" pageNumber="127" type="paratype">Paratypes</typeStatus>
(39): 
<collectingCountry id="F3287677FFC9FF8FFE1AFDDAFDBA9D40" box="[395,574,546,571]" name="Nicaragua" pageId="2" pageNumber="127">NICARAGUA</collectingCountry>
: 
<collectingRegion id="49FBF805FFC9FF8FFDDDFDDAFD619D40" box="[588,741,546,571]" pageId="2" pageNumber="127">La Libertad</collectingRegion>
:
</emphasis>
same data as holotype (
<specimenCount id="9D39FD6EFFC9FF8FFB96FDDBFBE79D40" box="[1031,1123,546,571]" pageId="2" pageNumber="127" type="male">7 males</specimenCount>
, 
<specimenCount id="9D39FD6EFFC9FF8FFBE3FDDAFB619D40" box="[1138,1253,546,571]" pageId="2" pageNumber="127" type="female">6 females</specimenCount>
; all cleared and mounted in microscope slides; 
<collectionCode id="ED2EAE22FFC9FF8FFE6DFDBFFDBA9D24" box="[508,574,583,607]" pageId="2" pageNumber="127">CIRA</collectionCode>
/ 
<collectionCode id="ED2EAE22FFC9FF8FFDC1FDBFFD229D24" box="[592,678,583,607]" country="Nicaragua" httpUri="http://grbio.org/cool/k7i5-im6v" name="Universidad Nacional Autonoma de Nicaragua" pageId="2" pageNumber="127">UNAN</collectionCode>
); (
<specimenCount id="9D39FD6EFFC9FF8FFD52FDBFFCA99D24" box="[707,813,582,607]" pageId="2" pageNumber="127" type="male">10 males</specimenCount>
, 
<specimenCount id="9D39FD6EFFC9FF8FFCAAFDBFFC3E9D24" box="[827,954,582,607]" pageId="2" pageNumber="127" type="female">10 females</specimenCount>
, preserved in glycerol, 
<collectionCode id="ED2EAE22FFC9FF8FFB59FDBFFA9C9D24" box="[1224,1304,583,607]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="2" pageNumber="127" type="Museum">SEMC</collectionCode>
; 
<specimenCount id="9D39FD6EFFC9FF8FFAB4FDBFFA059D24" box="[1317,1409,582,607]" pageId="2" pageNumber="127" type="male">3 males</specimenCount>
, 
<specimenCount id="9D39FD6EFFC9FF8FFA1FFDBFFF759DF8" pageId="2" pageNumber="127" type="female">3 females</specimenCount>
, preserved in glycerol, 
<collectionCode id="ED2EAE22FFC9FF8FFE67FD93FDC89DF8" box="[502,588,619,643]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="127" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF40666FFFC9FF8FFF06FA7FFF6F9AC7" httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="2" pageNumber="127" startId="2.[151,264,1415,1438]" targetBox="[263,1324,696,1394]" targetPageId="2">
<paragraph id="8B8036E7FFC9FF8FFF06FA7FFF6F9AC7" blockId="2.[151,1435,1415,1468]" pageId="2" pageNumber="127">
<emphasis id="B94BEAF5FFC9FF8FFF06FA7FFEBF9AE6" bold="true" box="[151,315,1415,1438]" pageId="2" pageNumber="127">FIGURES 1–6.</emphasis>
Habitus of 
<taxonomicName id="4C3F4D64FFC9FF8FFE24FA71FDF19AE5" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[437,629,1416,1438]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu">
<emphasis id="B94BEAF5FFC9FF8FFE24FA71FDF19AE5" box="[437,629,1416,1438]" italics="true" pageId="2" pageNumber="127">Yara marmontsedu</emphasis>
</taxonomicName>
: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C325656CFFC9FF8FFF56FA1CFC2899F6" pageId="2" pageNumber="127" type="diagnosis">
<paragraph id="8B8036E7FFC9FF8FFF56FA1CFC2899F6" blockId="2.[151,1436,1508,2001]" pageId="2" pageNumber="127">
<emphasis id="B94BEAF5FFC9FF8FFF56FA1CFE559A86" bold="true" box="[199,465,1508,1533]" pageId="2" pageNumber="127">Differential diagnosis.</emphasis>
Body length, females (
<quantity id="4CC79B02FFC9FF8FFD70FA1DFC0F9A86" box="[737,907,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="1.285" metricValueMax="1.37" metricValueMin="1.2" pageId="2" pageNumber="127" unit="mm" value="1.285" valueMax="1.37" valueMin="1.2">1.20–1.37 mm</quantity>
) and males (
<quantity id="4CC79B02FFC9FF8FFBB3FA1DFB499A86" box="[1058,1229,1509,1533]" metricMagnitude="-3" metricUnit="m" metricValue="1.335" metricValueMax="1.47" metricValueMin="1.2" pageId="2" pageNumber="127" unit="mm" value="1.335" valueMax="1.47" valueMin="1.2">1.20–1.47 mm</quantity>
); hindwings welldeveloped; femoral plate of metacoxae triangular, extending beyond area of articulation with metatrochanter, almost covering the metatrochanter completely; metatrochanter with three thick spines along inner margin; female abdominal tergite and sternite VIII with a median apical emargination; median lobe of aedeagus nearly 1.5× longer than basal apophysis; basal apophysis of aedeagus bifurcate near base.
</paragraph>
</subSubSection>
<subSubSection id="C325656CFFC9FF89FF56F961FDC19AF9" lastPageId="4" lastPageNumber="129" pageId="2" pageNumber="127" type="description">
<paragraph id="8B8036E7FFC9FF8FFF56F961FD6C98AB" blockId="2.[151,1436,1508,2001]" pageId="2" pageNumber="127">
<taxonomicName id="4C3F4D64FFC9FF8FFF56F961FE2599CB" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[199,417,1688,1712]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC9FF8FFF56F961FE2599CB" box="[199,417,1688,1712]" italics="true" pageId="2" pageNumber="127">Yara marmontsedu</emphasis>
</taxonomicName>
<emphasis id="B94BEAF5FFC9FF8FFE3CF960FE7699CA" bold="true" box="[429,498,1688,1713]" pageId="2" pageNumber="127">
<taxonomicNameLabel id="A278578EFFC9FF8FFE3CF960FE7699CA" box="[429,498,1688,1713]" pageId="2" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C3F4D64FFC9FF8FFDECF961FC6699CA" authority="Reichardt &amp; Hinton" authorityName="Reichardt &amp; Hinton" box="[637,994,1688,1713]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="dybasi">
<emphasis id="B94BEAF5FFC9FF8FFDECF961FD6399CB" box="[637,743,1688,1712]" italics="true" pageId="2" pageNumber="127">Y. dybasi</emphasis>
Reichardt &amp; Hinton
</taxonomicName>
, mostly because of characters of the abdominal apex of females. Both species can be distinguished by the development of the hindwings (welldeveloped in 
<taxonomicName id="4C3F4D64FFC9FF8FFEBFF919FE669983" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[302,482,1760,1784]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC9FF8FFEBFF919FE669983" box="[302,482,1760,1784]" italics="true" pageId="2" pageNumber="127">Y. marmontsedu</emphasis>
</taxonomicName>
<emphasis id="B94BEAF5FFC9FF8FFE7BF918FDAD9982" bold="true" box="[490,553,1760,1785]" pageId="2" pageNumber="127">
<taxonomicNameLabel id="A278578EFFC9FF8FFE7BF918FDAD9982" box="[490,553,1760,1785]" pageId="2" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; reduced in 
<taxonomicName id="4C3F4D64FFC9FF8FFD27F919FB829982" authority="Reichardt &amp; Hinton" authorityName="Reichardt &amp; Hinton" box="[694,1030,1760,1785]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="dybasi">
<emphasis id="B94BEAF5FFC9FF8FFD27F919FC9F9983" box="[694,795,1760,1784]" italics="true" pageId="2" pageNumber="127">Y. dybasi</emphasis>
Reichardt &amp; Hinton
</taxonomicName>
), the size of the femoral plate (large in 
<taxonomicName id="4C3F4D64FFC9FF8FFF26F8FDFEE99867" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[183,365,1796,1820]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC9FF8FFF26F8FDFEE99867" box="[183,365,1796,1820]" italics="true" pageId="2" pageNumber="127">Y. marmontsedu</emphasis>
</taxonomicName>
<emphasis id="B94BEAF5FFC9FF8FFEE9F8FCFE3E9866" bold="true" box="[376,442,1796,1821]" pageId="2" pageNumber="127">
<taxonomicNameLabel id="A278578EFFC9FF8FFEE9F8FCFE3E9866" box="[376,442,1796,1821]" pageId="2" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, see 
<figureCitation id="13042A62FFC9FF8FFE68F8FCFDCF9867" box="[505,587,1796,1821]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="2" pageNumber="127">Figs. 7</figureCitation>
, 
<figureCitation id="13042A62FFC9FF8FFDCBF8FCFDFD9866" box="[602,633,1796,1821]" captionStart="FIGURES 13–16" captionStartId="3.[151,264,1345,1368]" captionTargetBox="[198,1411,878,1308]" captionTargetId="figure@3.[167,1420,863,1324]" captionTargetPageId="3" captionText="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="2" pageNumber="127">16</figureCitation>
; short (as in the original generic diagnosis) in 
<taxonomicName id="4C3F4D64FFC9FF8FFB0CF8FDFF6D983A" authority="Reichardt &amp; Hinton" authorityName="Reichardt &amp; Hinton" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="dybasi">
<emphasis id="B94BEAF5FFC9FF8FFB0CF8FDFA819867" box="[1181,1285,1796,1820]" italics="true" pageId="2" pageNumber="127">Y. dybasi</emphasis>
Reichardt &amp; Hinton
</taxonomicName>
), and the presence of three distinct spines along the inner margin of the metatrochanter in 
<taxonomicName id="4C3F4D64FFC9FF8FFB79F8D1FA18983B" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[1256,1436,1832,1856]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC9FF8FFB79F8D1FA18983B" box="[1256,1436,1832,1856]" italics="true" pageId="2" pageNumber="127">Y. marmontsedu</emphasis>
</taxonomicName>
<emphasis id="B94BEAF5FFC9FF8FFF06F8B4FF51981E" bold="true" box="[151,213,1868,1893]" pageId="2" pageNumber="127">
<taxonomicNameLabel id="A278578EFFC9FF8FFF06F8B4FF51981E" box="[151,213,1868,1893]" pageId="2" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(with a row of long setae, as in the generic diagnosis in 
<taxonomicName id="4C3F4D64FFC9FF8FFCD4F8B5FB16981E" authority="Reichardt &amp; Hinton" authorityName="Reichardt &amp; Hinton" box="[837,1170,1868,1893]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="dybasi">
<emphasis id="B94BEAF5FFC9FF8FFCD4F8B5FC2D981F" box="[837,937,1868,1892]" italics="true" pageId="2" pageNumber="127">Y. dybasi</emphasis>
Reichardt &amp; Hinton
</taxonomicName>
). Furthermore, the form of the aedeagus of 
<taxonomicName id="4C3F4D64FFC9FF8FFEE0F889FDA298F3" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[369,550,1904,1928]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC9FF8FFEE0F889FDA298F3" box="[369,550,1904,1928]" italics="true" pageId="2" pageNumber="127">Y. marmontsedu</emphasis>
</taxonomicName>
<emphasis id="B94BEAF5FFC9FF8FFDBEF888FDEB98F2" bold="true" box="[559,623,1904,1929]" pageId="2" pageNumber="127">
<taxonomicNameLabel id="A278578EFFC9FF8FFDBEF888FDEB98F2" box="[559,623,1904,1929]" pageId="2" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is similar to the form in 
<taxonomicName id="4C3F4D64FFC9FF8FFC00F889FA8F98F3" authority="Short, Joly &amp; Garcia" authorityName="Short, Joly &amp; Garcia" box="[913,1291,1904,1929]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B94BEAF5FFC9FF8FFC00F889FB9298F3" box="[913,1046,1904,1928]" italics="true" pageId="2" pageNumber="127">Y. maculata</emphasis>
Short, Joly &amp; García
</taxonomicName>
(J-shaped at base), except by the basally bifurcated basal apophysis in 
<taxonomicName id="4C3F4D64FFC9FF8FFC8DF86DFC4998D7" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[796,973,1940,1964]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC9FF8FFC8DF86DFC4998D7" box="[796,973,1940,1964]" italics="true" pageId="2" pageNumber="127">Y. marmontsedu</emphasis>
</taxonomicName>
<emphasis id="B94BEAF5FFC9FF8FFC47F86CFB9098D6" bold="true" box="[982,1044,1940,1965]" pageId="2" pageNumber="127">
<taxonomicNameLabel id="A278578EFFC9FF8FFC47F86CFB9098D6" box="[982,1044,1940,1965]" pageId="2" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(apically bifurcated in 
<taxonomicName id="4C3F4D64FFC9FF8FFA88F86DFE0698AB" authority="Short, Joly &amp; Garcia" authorityName="Short, Joly &amp; Garcia" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="127" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B94BEAF5FFC9FF8FFA88F86DFA1898D7" box="[1305,1436,1940,1964]" italics="true" pageId="2" pageNumber="127">Y. maculata</emphasis>
Short, Joly &amp; García
</taxonomicName>
, see 
<figureCitation id="13042A62FFC9FF8FFE26F840FE7D98AA" box="[439,505,1976,2001]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="2" pageNumber="127">
Fig. 
<quantity id="4CC79B02FFC9FF8FFE7AF840FE7D98AA" box="[491,505,1976,2001]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="127" unit="in" value="6.0">6</quantity>
</figureCitation>
in 
<bibRefCitation id="EFAE4B16FFC9FF8FFD8CF840FD5898AB" author="Short, A. E. Z. &amp; Joly, L. J. &amp; Garcia, M." box="[541,732,1976,2001]" pageId="2" pageNumber="127" pagination="151 - 156" refId="ref4993" refString="Short, A. E. Z., Joly, L. J. &amp; Garcia, M. (2010) Discovery of the Skiff Beetle Genus Yara Reichardt and Hinton (Coleoptera: Myxophaga: Hydroscaphidae) in Venezuela with description of a new species. The Coleopterists Bulletin, 64 (2), 151 - 156. https: // www. jstor. org / stable / 40664383" type="journal article" year="2010">
Short 
<emphasis id="B94BEAF5FFC9FF8FFDF1F841FD1C98AB" box="[608,664,1976,2000]" italics="true" pageId="2" pageNumber="127">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
<caption id="DF40666FFFC8FF8EFF06FD21FF5B9C56" httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128" startId="3.[151,264,729,752]" targetBox="[199,1381,193,708]" targetPageId="3">
<paragraph id="8B8036E7FFC8FF8EFF06FD21FF5B9C56" blockId="3.[151,1436,729,813]" pageId="3" pageNumber="128">
<emphasis id="B94BEAF5FFC8FF8EFF06FD21FEC39D94" bold="true" box="[151,327,729,752]" pageId="3" pageNumber="128">FIGURES 7–12.</emphasis>
Ventral view and head appendages of 
<taxonomicName id="4C3F4D64FFC8FF8EFD5FFD23FC0A9D8B" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[718,910,730,752]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="marmontsedu">
<emphasis id="B94BEAF5FFC8FF8EFD5FFD23FC0A9D8B" box="[718,910,730,752]" italics="true" pageId="3" pageNumber="128">Yara marmontsedu</emphasis>
</taxonomicName>
: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm.
</paragraph>
</caption>
<caption id="DF40666FFFC8FF8EFF06FAB9FE379A0D" httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="3" pageNumber="128" startId="3.[151,264,1345,1368]" targetBox="[198,1411,878,1308]" targetPageId="3">
<paragraph id="8B8036E7FFC8FF8EFF06FAB9FE379A0D" blockId="3.[151,1436,1345,1398]" pageId="3" pageNumber="128">
<emphasis id="B94BEAF5FFC8FF8EFF06FAB9FED19A2C" bold="true" box="[151,341,1345,1368]" pageId="3" pageNumber="128">FIGURES 13–16.</emphasis>
Hindwings and adult legs of 
<taxonomicName id="4C3F4D64FFC8FF8EFD19FABBFCCC9A23" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[648,840,1346,1368]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="128" phylum="Arthropoda" rank="species" species="marmontsedu">
<emphasis id="B94BEAF5FFC8FF8EFD19FABBFCCC9A23" box="[648,840,1346,1368]" italics="true" pageId="3" pageNumber="128">Yara marmontsedu</emphasis>
</taxonomicName>
: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8036E7FFC8FF89FF56FA65FB099C73" blockId="3.[151,1437,1437,2029]" lastBlockId="4.[151,1437,151,1977]" lastPageId="4" lastPageNumber="129" pageId="3" pageNumber="128">
<emphasis id="B94BEAF5FFC8FF8EFF56FA65FEDD9ACD" bold="true" box="[199,345,1437,1462]" pageId="3" pageNumber="128">Description.</emphasis>
Body, elongated, fusiform, with dense setae all over the surface (
<figureCitation id="13042A62FFC8FF8EFBD7FA65FB379ACD" box="[1094,1203,1437,1462]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="3" pageNumber="128">Figs. 1–6</figureCitation>
). Body length 
<quantity id="4CC79B02FFC8FF8EFACCFA66FF7F9AAC" metricMagnitude="-3" metricUnit="m" metricValue="1.285" metricValueMax="1.37" metricValueMin="1.2" pageId="3" pageNumber="128" unit="mm" value="1.285" valueMax="1.37" valueMin="1.2">1.20– 1.37 mm</quantity>
in females, 
<quantity id="4CC79B02FFC8FF8EFE17FA47FDAE9AAC" box="[390,554,1471,1495]" metricMagnitude="-3" metricUnit="m" metricValue="1.335" metricValueMax="1.47" metricValueMin="1.2" pageId="3" pageNumber="128" unit="mm" value="1.335" valueMax="1.47" valueMin="1.2">1.20–1.47 mm</quantity>
in males. 
<emphasis id="B94BEAF5FFC8FF8EFD0FFA47FD5A9AA3" bold="true" box="[670,734,1471,1496]" pageId="3" pageNumber="128">Head</emphasis>
. Prognathous, transverse, nearly 2× wider than long. Eyes not protruding from outline of head (
<figureCitation id="13042A62FFC8FF8EFD82FA18FDE09A82" box="[531,612,1504,1529]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="3" pageNumber="128">Figs. 4</figureCitation>
, 
<figureCitation id="13042A62FFC8FF8EFDE2FA19FD079A82" box="[627,643,1505,1529]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">7</figureCitation>
); dorsal surface of head microreticulate, with sparse, rather long fine setae; frontoclypeal suture well defined, bell-shaped. Clypeus yellowish, reticulated, with abundant fine setae. Labrum with dorsal surface coarsely reticulated, bearing a transverse row of rather long and thick setae at midlength (
<figureCitation id="13042A62FFC8FF8EFE88F9BCFEDA9926" box="[281,350,1604,1629]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 7</figureCitation>
); ventral edge of labrum (
<figureCitation id="13042A62FFC8FF8EFD16F9BCFD5C9926" box="[647,728,1604,1629]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 11</figureCitation>
) fringed by stout setae, bearing lateral tufts of fine long setae; hypopharynx (
<figureCitation id="13042A62FFC8FF8EFEABF99DFE0D9905" box="[314,393,1637,1662]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 11</figureCitation>
) as a median longitudinal tuft of fine setae. Mandibles (
<figureCitation id="13042A62FFC8FF8EFC6CF99DFBCA9905" box="[1021,1102,1637,1662]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 10</figureCitation>
) with well-developed mola, a median tuft of fine setae, and two teeth (apparently three) on the incisor region, with proximal tooth bifurcated. Antennae with eight antennomeres (
<figureCitation id="13042A62FFC8FF8EFDBEF950FD1499BA" box="[559,656,1704,1729]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 7, 8</figureCitation>
); antennomere I 1.6× longer than antennomere II; antennomeres III– VII subquadrate, similar in size and shape to each other, nearly as long as wide, half as long as antennomere II; antennomere VIII somewhat wedge-shaped, 1.4× longer than antennomere I. Maxillary palps (
<figureCitation id="13042A62FFC8FF8EFB4BF913FAA4987F" box="[1242,1312,1771,1796]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 9</figureCitation>
) with four palpomeres; palpomere III thicker than palpomere II, palpomere IV short, digitiform, bearing a lateral and an apical sensilla. Mentum (
<figureCitation id="13042A62FFC8FF8EFEFBF8D5FE38983D" box="[362,444,1837,1862]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 12</figureCitation>
) well-developed, subquadrate, with a basal row of six long setae. Prementum (
<figureCitation id="13042A62FFC8FF8EFAD0F8D5FA10983D" box="[1345,1428,1837,1862]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 12</figureCitation>
) well-developed, with two transverse rows (preapical and apical) of subquadrate teeth, laterally limited by one spiniform tooth and a seta on each side. Labial palps (
<figureCitation id="13042A62FFC8FF8EFD67F888FCCC98F2" box="[758,840,1904,1929]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="3" pageNumber="128">Fig. 12</figureCitation>
) with two palpomeres; palpomere II digitiform, with an outer basal sensillum and a group of apical sensilla, central apical sensillum digitiform, largest. 
<emphasis id="B94BEAF5FFC8FF8EFAADF86AFA1898D0" bold="true" box="[1340,1436,1938,1963]" pageId="3" pageNumber="128">Thorax.</emphasis>
Pronotum trapezoidal in dorsal view, with posterior margin as wide as base of elytra (
<figureCitation id="13042A62FFC8FF8EFBC0F84BFB3998B7" box="[1105,1213,1971,1996]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="3" pageNumber="128">Figs. 1, 4</figureCitation>
). Elytra dark brown with yellow markings: an irregular area on basal third, a small median spot at 2/3, and an apical transverse band; elytra densely covered by long setae on dorsal surface; each elytron 1.5× longer than wide, with anterolateral corner rounded, truncated at apex, covering up to basal half of third abdominal tergite (
<figureCitation id="13042A62FFCFFF89FBE3FF41FB649FAA" box="[1138,1248,185,210]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="4" pageNumber="129">Figs. 1, 4</figureCitation>
); ventral surface with long setae along lateral and posterior margins. Hindwings well-developed (examined in 14 individuals; 
<figureCitation id="13042A62FFCFFF89FAE1FF22FF319E6F" captionStart="FIGURES 13–16" captionStartId="3.[151,264,1345,1368]" captionTargetBox="[198,1411,878,1308]" captionTargetId="figure@3.[167,1420,863,1324]" captionTargetPageId="3" captionText="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="4" pageNumber="129">Fig. 13</figureCitation>
), each 2.3× longer than wide, densely covered by short setae over entire surface, and fringed by long setae along margins, except antero-basal (costal) margin; anal lobe well-developed. Legs (
<figureCitation id="13042A62FFCFFF89FB97FEE5FB149E4D" box="[1030,1168,285,310]" captionStart="FIGURES 13–16" captionStartId="3.[151,264,1345,1368]" captionTargetBox="[198,1411,878,1308]" captionTargetId="figure@3.[167,1420,863,1324]" captionTargetPageId="3" captionText="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="4" pageNumber="129">Figs. 14–16</figureCitation>
). All femora with a row of setae along dorsal margin; all tibiae with a row of spines along ventral margin, mesotibia with additional row of well-developed spines along dorsal margin; metacoxae with long, triangular femoral plate (
<figureCitation id="13042A62FFCFFF89FB51FEA7FA979E0C" box="[1216,1299,351,376]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,729,752]" captionTargetBox="[199,1381,193,708]" captionTargetId="figure@3.[199,1387,193,708]" captionTargetPageId="3" captionText="FIGURES 7–12. Ventral view and head appendages of Yara marmontsedu: (7) ventral view of head and thorax, scale bar: 100 µm; (8) antenna, (9) maxilla, (10) left mandible, (11) ventral view of labrum including hypopharynx, (12) labium. Scale bars: 50 µm." httpUri="https://zenodo.org/record/2618209/files/figure.png" pageId="4" pageNumber="129">Figs. 7</figureCitation>
, 
<figureCitation id="13042A62FFCFFF89FAB2FEA7FAC59E03" box="[1315,1345,351,376]" captionStart="FIGURES 13–16" captionStartId="3.[151,264,1345,1368]" captionTargetBox="[198,1411,878,1308]" captionTargetId="figure@3.[167,1420,863,1324]" captionTargetPageId="3" captionText="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="4" pageNumber="129">16</figureCitation>
), partly covering metatrochanter, extending beyond area of articulation; metatrochanter (
<figureCitation id="13042A62FFCFFF89FB8EFE79FBF49EE1" box="[1055,1136,385,410]" captionStart="FIGURES 13–16" captionStartId="3.[151,264,1345,1368]" captionTargetBox="[198,1411,878,1308]" captionTargetId="figure@3.[167,1420,863,1324]" captionTargetPageId="3" captionText="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="4" pageNumber="129">Fig. 16</figureCitation>
) with three rigid and thick spines along posterior margin; all tarsi with three tarsomeres (
<figureCitation id="13042A62FFCFFF89FCCAFE5AFC629EC0" box="[859,998,418,443]" captionStart="FIGURES 13–16" captionStartId="3.[151,264,1345,1368]" captionTargetBox="[198,1411,878,1308]" captionTargetId="figure@3.[167,1420,863,1324]" captionTargetPageId="3" captionText="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="4" pageNumber="129">Figs. 14–16</figureCitation>
); tarsal claws long and thin (
<figureCitation id="13042A62FFCFFF89FAA0FE5AFF329EA6" captionStart="FIGURES 13–16" captionStartId="3.[151,264,1345,1368]" captionTargetBox="[198,1411,878,1308]" captionTargetId="figure@3.[167,1420,863,1324]" captionTargetPageId="3" captionText="FIGURES 13–16. Hindwings and adult legs of Yara marmontsedu: (13) extended hindwing; (14) proleg; (15) mesoleg; (16) metaleg. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2618211/files/figure.png" pageId="4" pageNumber="129">Figs. 14– 16</figureCitation>
). 
<emphasis id="B94BEAF5FFCFFF89FF5DFE3CFECC9EA6" bold="true" box="[204,328,452,477]" pageId="4" pageNumber="129">Abdomen.</emphasis>
Abdominal segments of both males and females moderate and rather uniformly covered by setae (
<figureCitation id="13042A62FFCFFF89FF0EFE1DFE8F9E85" box="[159,267,485,510]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="4" pageNumber="129">Figs. 1–6</figureCitation>
); tergite IV lacking dense cover of setae; tergite III light brown, tergites IV and V dark brown, tergite VI and VII yellowish; tergite VIII of females (
<figureCitation id="13042A62FFCFFF89FD17FDFEFD719D64" box="[646,757,518,543]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="4" pageNumber="129">Figs. 4, 5</figureCitation>
, 
<figureCitation id="13042A62FFCFFF89FC92FDFFFCD39D64" box="[771,855,519,543]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">17, 17a</figureCitation>
) conical, yellowish along basal and apical fourths, dark brown along mesal region, with apical margin mesally emarginate; tergites VIII and IX of males (
<figureCitation id="13042A62FFCFFF89FABBFDDFFA139D3B" box="[1322,1431,551,576]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1415,1438]" captionTargetBox="[263,1324,696,1394]" captionTargetId="figure@2.[263,1324,695,1394]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Yara marmontsedu: (1, 4) dorsal, (2, 5) lateral, (3, 6) ventral; male (1–3); female (4–6). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/2618207/files/figure.png" pageId="4" pageNumber="129">Figs. 1, 2</figureCitation>
, 
<figureCitation id="13042A62FFCFFF89FF06FDB1FF6E9D1A" box="[151,234,585,609]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">19, 19a</figureCitation>
) conical, tergite VIII with narrow yellowish bands along basal and apical margins, dark brown along mesal region; sternite VIII of males with apex asymmetrically projected (
<figureCitation id="13042A62FFCFFF89FC1CFD92FB9C9DF8" box="[909,1048,618,643]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Fig. 20, 20a</figureCitation>
), bearing three thick, curved setae along left side of projection. Female genitalia with two long lateral cylindrical projections extending beyond last tergite (
<figureCitation id="13042A62FFCFFF89FF7FFD55FE619DBD" box="[238,485,685,710]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Figs. 17, 17a, 18, 18a</figureCitation>
); both projections similar in length, covered by setae, with two long preapical setae. Aedeagus (
<figureCitation id="13042A62FFCFFF89FE89FD36FEE89D9C" box="[280,364,718,743]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Fig. 21</figureCitation>
) with median lobe slender, non-coiled, at most J-shaped at base (
<figureCitation id="13042A62FFCFFF89FBF8FD36FB7C9D9C" box="[1129,1272,718,743]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Figs. 21, 22</figureCitation>
); median lobe nearly 1.5× longer than basal apophysis; basal apophysis bifurcated near its base (
<figureCitation id="13042A62FFCFFF89FBBDFD17FB059C73" box="[1068,1153,751,776]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8B8036E7FFCFFF89FF56FCE9FDC19AF9" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="129">
<emphasis id="B94BEAF5FFCFFF89FF56FCE9FE939C51" bold="true" box="[199,279,785,810]" pageId="4" pageNumber="129">Larva.</emphasis>
Body fusiform in dorsal view (
<figureCitation id="13042A62FFCFFF89FDEFFCE9FD569C51" box="[638,722,785,810]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 26</figureCitation>
), slightly compressed dorsoventrally (
<figureCitation id="13042A62FFCFFF89FB13FCE9FB529C51" box="[1154,1238,785,810]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 27</figureCitation>
), with abdominal segments gradually decreasing in width towards apex (
<figureCitation id="13042A62FFCFFF89FC80FCCAFC1A9C30" box="[785,926,818,843]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Figs. 26–28</figureCitation>
); body 
<quantity id="4CC79B02FFCFFF89FC67FCCAFB0A9C31" box="[1014,1166,818,843]" metricMagnitude="-3" metricUnit="m" metricValue="1.1749999999999998" metricValueMax="1.4" metricValueMin="0.95" pageId="4" pageNumber="129" unit="mm" value="1.1749999999999998" valueMax="1.4" valueMin="0.95">0.95–1.4 mm</quantity>
in length, light to dark brown in coloration; surface of body tuberculate and rather densely covered by thick and rather long, curved setae (e.g. 
<figureCitation id="13042A62FFCFFF89FF44FC8DFEAB9CF5" box="[213,303,885,910]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Fig. 24</figureCitation>
). Head (
<figureCitation id="13042A62FFCFFF89FE08FC8DFE749CF5" box="[409,496,885,910]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 28</figureCitation>
) transverse, nearly 2× wider than long; postero-dorsal margin of head slightly emarginate, postero ventral margin strongly emarginate. Stemmata relatively large, each well defined, six on each side; each group of stemmata abruptly projecting from the surface of the head capsule, bearing a well-developed central seta. Antennae (
<figureCitation id="13042A62FFCFFF89FE34FC21FE7D9C89" box="[421,505,985,1010]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 30</figureCitation>
) nearly as long as stemmatal tubercle; antennal insertions projected as tubercles; basal antennomere cylindrical, nearly as wide as long; apical antenommere with two branches, one of them bifurcate at apex. Frontoclypeal suture indistinct. Labrum (
<figureCitation id="13042A62FFCFFF89FCF3FBE4FC389B4F" box="[866,956,1052,1077]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 29</figureCitation>
) semirectangular, with posterior margin rounded, and a row of tiny denticles along median region of ventral apical margin which is bordered by paired larger teeth. Ventral mouthparts exposed, well-developed (
<figureCitation id="13042A62FFCFFF89FCB6FBA6FCFF9B0C" box="[807,891,1118,1143]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 29</figureCitation>
). Maxillary palps well developed (
<figureCitation id="13042A62FFCFFF89FA90FBA6FAD09B0C" box="[1281,1364,1118,1143]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 33</figureCitation>
), each with two elongated palpomeres; palpomere II with an apical seta. Labium (
<figureCitation id="13042A62FFCFFF89FC76FB87FBBD9BE3" box="[999,1081,1151,1176]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 31</figureCitation>
) subquadrate, nearly as long as wide, with two rows of tiny irregular denticles along apical margin, bordered by paired larger teeth; each labial palp with one single palpomere (
<figureCitation id="13042A62FFCFFF89FE40FB3AFD029BA0" box="[465,646,1218,1243]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Figs. 29, 31, 32</figureCitation>
). Thorax about half as long as abdomen (
<figureCitation id="13042A62FFCFFF89FBCBFB3AFB2B9BA0" box="[1114,1199,1218,1243]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 26</figureCitation>
); posterior margin of each thoracic tergum fringed by rather long and thick setae. Legs (
<figureCitation id="13042A62FFCFFF89FC0FFB1CFBAC9B86" box="[926,1064,1252,1277]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Figs. 36–38</figureCitation>
) slender, each pair separated by nearly the width of a coxa (
<figureCitation id="13042A62FFCFFF89FE75FAFDFDB99A65" box="[484,573,1285,1310]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 28</figureCitation>
); femora slightly longer than tibiotarsus, pretarsus nearly 0.7× as long as tibiotarsus. Abdominal segments II to VII with lateral projections posteriorly directed; each projection with apical curved and thick setae (
<figureCitation id="13042A62FFCFFF89FE30FABFFE729A1B" box="[417,502,1351,1376]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 34</figureCitation>
); abdominal tergite VIII with apical margin fringed by setae (
<figureCitation id="13042A62FFCFFF89FB38FABFFB799A1B" box="[1193,1277,1351,1376]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 35</figureCitation>
); sternite VIII with paired hook-like spines (
<figureCitation id="13042A62FFCFFF89FE75FA91FDBC9AF9" box="[484,568,1385,1410]" captionStart="FIGURES 26–38" captionStartId="6.[151,264,1866,1889]" captionTargetBox="[285,1290,242,1815]" captionTargetId="figure@6.[271,1306,223,1824]" captionTargetPageId="6" captionText="FIGURES 26–38. Larva of Yara marmontsedu: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm." httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="4" pageNumber="129">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C325656CFFCFFF89FF56FA72FEA89ABF" pageId="4" pageNumber="129" type="distribution">
<paragraph id="8B8036E7FFCFFF89FF56FA72FEA89ABF" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="129">
<emphasis id="B94BEAF5FFCFFF89FF56FA72FEE49AD8" bold="true" box="[199,352,1418,1443]" pageId="4" pageNumber="129">Distribution.</emphasis>
Pijibay Creek, La Libertad Municipality, 
<collectingRegion id="49FBF805FFCFFF89FCA3FA72FBAB9AD9" box="[818,1071,1418,1443]" country="Nicaragua" name="Chontales" pageId="4" pageNumber="129">Chontales Department</collectingRegion>
, 
<collectingCountry id="F3287677FFCFFF89FBA8FA72FB289AD8" box="[1081,1196,1418,1443]" name="Nicaragua" pageId="4" pageNumber="129">Nicaragua</collectingCountry>
(
<geoCoordinate id="EE0B5020FFCFFF89FB2DFA73FAA69AD9" box="[1212,1314,1419,1443]" direction="north" orientation="latitude" pageId="4" pageNumber="129" precision="925" value="12.2">12°12' N</geoCoordinate>
, 
<geoCoordinate id="EE0B5020FFCFFF89FABDFA72FA1C9AD9" box="[1324,1432,1418,1443]" direction="west" orientation="longitude" pageId="4" pageNumber="129" precision="925" value="-85.166664">85°10' W</geoCoordinate>
; 
<figureCitation id="13042A62FFCFFF89FF06FA54FEA49ABF" box="[151,288,1452,1477]" captionStart="FIGURES 43–47" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[197,1389,1092,1822]" captionTargetId="figure@7.[196,1390,1092,1822]" captionTargetPageId="7" captionText="FIGURES 43–47. Distribution and collecting: (43) map for species of Yara; (44) map for Central American species of Yara; (45) sampling locality at La Libertad, Chontales, Nicaragua; (46) Pijibay Creek; (47) collecting macroinvertebrates using a Dnet at Pijibay Creek." httpUri="https://zenodo.org/record/2618219/files/figure.png" pageId="4" pageNumber="129">Figs. 43–47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C325656CFFCFFF85FF56FA35FB4D9ED7" lastPageId="8" lastPageNumber="133" pageId="4" pageNumber="129" type="etymology">
<paragraph id="8B8036E7FFCFFF89FF56FA35FADD997C" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="129">
<emphasis id="B94BEAF5FFCFFF89FF56FA35FECA9A9D" bold="true" box="[199,334,1485,1510]" pageId="4" pageNumber="129">Etymology.</emphasis>
The specific epithet 
<taxonomicName id="4C3F4D64FFCFFF89FDA6FA35FD4B9A9E" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[567,719,1485,1509]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="marmontsedu">
<emphasis id="B94BEAF5FFCFFF89FDA6FA35FD4B9A9E" box="[567,719,1485,1509]" italics="true" pageId="4" pageNumber="129">marmontsedu</emphasis>
</taxonomicName>
is formed by the abbreviation of the names of the three children of authors Raudez and Cano: Margarita, 
<collectingCountry id="F3287677FFCFFF89FDCEFA17FD5E997D" box="[607,730,1519,1542]" name="Montserrat" pageId="4" pageNumber="129">Montserrat</collectingCountry>
and Eduardo, to whom the species is lovingly dedicated.
</paragraph>
<paragraph id="8B8036E7FFCFFF89FF56F9F7FF58982E" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="129">
<emphasis id="B94BEAF5FFCFFF89FF56F9F7FE0D9953" bold="true" box="[199,393,1551,1576]" pageId="4" pageNumber="129">Natural history.</emphasis>
Adult specimens were collected during the wet season (October) in 2012 and in the dry season (April) of 2013 at the margins of Pijibay Creek. Ovigerous females were found during the dry season (
<date id="FF811027FFCFFF89FACEF9C9FF509910" pageId="4" pageNumber="129" value="2013-04">April 2013</date>
). Each female contained between 1 and 
<specimenCount id="9D39FD6EFFCFFF89FD23F9ABFD7B9910" box="[690,767,1619,1643]" pageId="4" pageNumber="129" type="egg">3 eggs</specimenCount>
(
<figureCitation id="13042A62FFCFFF89FC83F9AAFCEF9910" box="[786,875,1618,1643]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Fig. 24</figureCitation>
). The eggs are elliptical, 2.4× longer than wide (length: 199.8±9.5µm; width: 81.9±6.39µm; n = 8), light brown in color with smooth surface (
<figureCitation id="13042A62FFCFFF89FB26F98CFA8F99F6" box="[1207,1291,1652,1677]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Fig. 25</figureCitation>
); each egg is slightly longer than abdominal segments II and III of the female combined (
<figureCitation id="13042A62FFCFFF89FB82F96DFBEF99D5" box="[1043,1131,1685,1710]" captionStart="FIGURES 17–25" captionStartId="5.[151,264,1857,1880]" captionTargetBox="[278,1298,193,1823]" captionTargetId="figure@5.[278,1298,193,1826]" captionTargetPageId="5" captionText="FIGURES 17–25. Abdomen of Yara marmontsedu: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm." httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="4" pageNumber="129">Fig. 24</figureCitation>
). Larvae and a pupa were collected during the wet season. The larvae described here are likely on the last larval instar, given their strong resemblance to the adults. The head of the larva is also generally similar to the head of third instar 
<taxonomicName id="4C3F4D64FFCFFF89FA97F920FF66986A" baseAuthorityName="Reichardt &amp; Hinton" baseAuthorityYear="1976" class="Insecta" family="Hydroscaphidae" genus="Hydroscapha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="129" phylum="Arthropoda" rank="species" species="natans">
<emphasis id="B94BEAF5FFCFFF89FA97F920FF66986A" italics="true" pageId="4" pageNumber="129">Hydroscapha natans</emphasis>
</taxonomicName>
as described by 
<bibRefCitation id="EFAE4B16FFCFFF89FE38F901FD299869" author="Beutel, R. G. &amp; Haas, A." box="[425,685,1785,1810]" pageId="4" pageNumber="129" pagination="103 - 116" refId="ref4520" refString="Beutel, R. G. &amp; Haas, A. (1998) Larval head morphology of Hydroscapha natans (Coleoptera, Myxophaga) with reference to miniaturization and the systematic position of Hydroscaphidae. Zoomorphology, 118 (2), 103 - 116. https: // doi. org / 10.1007 / s 004350050061" type="journal article" year="1998">Beutel &amp; Haas (1998)</bibRefCitation>
. The pupa was brown in coloration and covered by a membrane (
<figureCitation id="13042A62FFCFFF89FF0EF8E2FEAD9848" box="[159,297,1818,1843]" captionStart="FIGURES 39–42" captionStartId="7.[151,264,950,973]" captionTargetBox="[247,1320,208,922]" captionTargetId="figure@7.[219,1356,193,933]" captionTargetPageId="7" captionText="FIGURES 39–42. Pupa of Yara marmontsedu: (39) dorso-lateral view; (40) detail of head and antenna; (41) detail of pretarsus; (42) detail of apex of abdomen." httpUri="https://zenodo.org/record/2618217/files/figure.png" pageId="4" pageNumber="129">Figs. 39–42</figureCitation>
); unfortunately, only one pupa was collected, and given its condition we are not able to describe it in detail.
</paragraph>
<paragraph id="8B8036E7FFCFFF89FF56F8A5FBDF98C3" blockId="4.[151,1437,151,1977]" pageId="4" pageNumber="129">
The site is characterized by shallow waters (less than 
<quantity id="4CC79B02FFCFFF89FC8EF8A6FCCE980E" box="[799,842,1886,1910]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="129" unit="m" value="1.0">1 m</quantity>
deep), with permanent flow. The dominant materials of the creek bed are silt and clay, resulting in a muddy substrate devoid of rocks or sand; the marginal areas of the creek are covered with grassland vegetation and limited riparian shrubs (
<figureCitation id="13042A62FFCFFF89FC54F858FBCB98C3" box="[965,1103,1952,1977]" captionStart="FIGURES 43–47" captionStartId="7.[151,264,1843,1866]" captionTargetBox="[197,1389,1092,1822]" captionTargetId="figure@7.[196,1390,1092,1822]" captionTargetPageId="7" captionText="FIGURES 43–47. Distribution and collecting: (43) map for species of Yara; (44) map for Central American species of Yara; (45) sampling locality at La Libertad, Chontales, Nicaragua; (46) Pijibay Creek; (47) collecting macroinvertebrates using a Dnet at Pijibay Creek." httpUri="https://zenodo.org/record/2618219/files/figure.png" pageId="4" pageNumber="129">Figs. 43–47</figureCitation>
).
</paragraph>
<caption id="DF40666FFFCEFF88FF06F8B9FA869897" httpUri="https://zenodo.org/record/2618213/files/figure.png" pageId="5" pageNumber="130" startId="5.[151,264,1857,1880]" targetBox="[278,1298,193,1823]" targetPageId="5">
<paragraph id="8B8036E7FFCEFF88FF06F8B9FA869897" blockId="5.[151,1436,1857,2028]" pageId="5" pageNumber="130">
<emphasis id="B94BEAF5FFCEFF88FF06F8B9FED0982C" bold="true" box="[151,340,1857,1880]" pageId="5" pageNumber="130">FIGURES 17–25.</emphasis>
Abdomen of 
<taxonomicName id="4C3F4D64FFCEFF88FE49F8BBFD0A9823" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[472,654,1858,1880]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="marmontsedu">
<emphasis id="B94BEAF5FFCEFF88FE49F8BBFD0A9823" box="[472,654,1858,1880]" italics="true" pageId="5" pageNumber="130">Yara marmontsedu</emphasis>
</taxonomicName>
: (17) female abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (17a) same, line drawing; (18) female abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII; (18a) same, line drawing; (19) male abdomen, dorsal view, black arrow indicates emargination on apical margin of tergite VIII; (19a) same, line drawing; (20) male abdomen, ventral view, black arrow indicates emargination on apical margin of sternite VIII, white arrow pointing curved setae; (20a) same, line drawing; (21) aedeagus; (22) basal apophysis in lateral view; (23) basal apophysis in dorsal view; (24) ventral view of ovigerous female; (25) egg. Scale bar: 17–20: 0.1 mm.
</paragraph>
</caption>
<caption id="DF40666FFFCDFF8BFF06F8B2FB7C98C6" httpUri="https://zenodo.org/record/2618215/files/figure.png" pageId="6" pageNumber="131" startId="6.[151,264,1866,1889]" targetBox="[285,1290,242,1815]" targetPageId="6">
<paragraph id="8B8036E7FFCDFF8BFF06F8B2FB7C98C6" blockId="6.[151,1436,1866,1981]" pageId="6" pageNumber="131">
<emphasis id="B94BEAF5FFCDFF8BFF06F8B2FED2981B" bold="true" box="[151,342,1866,1889]" pageId="6" pageNumber="131">FIGURES 26–38.</emphasis>
Larva of 
<taxonomicName id="4C3F4D64FFCDFF8BFE51F8B4FD06981A" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[448,642,1867,1889]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="131" phylum="Arthropoda" rank="species" species="marmontsedu">
<emphasis id="B94BEAF5FFCDFF8BFE51F8B4FD06981A" box="[448,642,1867,1889]" italics="true" pageId="6" pageNumber="131">Yara marmontsedu</emphasis>
</taxonomicName>
: (26) dorsal view; (27) lateral view; (28) ventral view; (29) ventral view of head; (30) antenna; (31) ventral view of mandible, prementum and mentum; (32) detail of labial palp; (33) detail of maxilla including maxillary palp; (34) detail of lateral projection of abdominal segments; (35) detail of lateral view of last abdominal segments; (36) proleg; (37) mesoleg; (38) metaleg. Scale bars: 26–28: 0.5 mm; 29–33: 50 µm; 34–37: 0.1 mm.
</paragraph>
</caption>
<caption id="DF40666FFFCCFF8AFF06FC4EFE539C90" httpUri="https://zenodo.org/record/2618217/files/figure.png" pageId="7" pageNumber="132" startId="7.[151,264,950,973]" targetBox="[247,1320,208,922]" targetPageId="7">
<paragraph id="8B8036E7FFCCFF8AFF06FC4EFE539C90" blockId="7.[151,1436,950,1003]" pageId="7" pageNumber="132">
<emphasis id="B94BEAF5FFCCFF8AFF06FC4EFED79CB7" bold="true" box="[151,339,950,973]" pageId="7" pageNumber="132">FIGURES 39–42.</emphasis>
Pupa of 
<taxonomicName id="4C3F4D64FFCCFF8AFE3DFC40FDEF9CB6" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[428,619,951,973]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="132" phylum="Arthropoda" rank="species" species="marmontsedu">
<emphasis id="B94BEAF5FFCCFF8AFE3DFC40FDEF9CB6" box="[428,619,951,973]" italics="true" pageId="7" pageNumber="132">Yara marmontsedu</emphasis>
</taxonomicName>
: (39) dorso-lateral view; (40) detail of head and antenna; (41) detail of pretarsus; (42) detail of apex of abdomen.
</paragraph>
</caption>
<caption id="DF40666FFFCCFF8AFF06F8CBFEE298FC" httpUri="https://zenodo.org/record/2618219/files/figure.png" pageId="7" pageNumber="132" startId="7.[151,264,1843,1866]" targetBox="[197,1389,1092,1822]" targetPageId="7">
<paragraph id="8B8036E7FFCCFF8AFF06F8CBFEE298FC" blockId="7.[151,1436,1843,1927]" pageId="7" pageNumber="132">
<emphasis id="B94BEAF5FFCCFF8AFF06F8CBFED19832" bold="true" box="[151,341,1843,1866]" pageId="7" pageNumber="132">FIGURES 43–47.</emphasis>
Distribution and collecting: (43) map for species of 
<taxonomicName id="4C3F4D64FFCCFF8AFCE5F8CDFC259831" authorityName="Reichardt &amp; Hinton" authorityYear="1976" box="[884,929,1845,1866]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAF5FFCCFF8AFCE5F8CDFC259831" box="[884,929,1845,1866]" italics="true" pageId="7" pageNumber="132">Yara</emphasis>
</taxonomicName>
; (44) map for Central American species of 
<taxonomicName id="4C3F4D64FFCCFF8AFAF7F8CDFA179831" authorityName="Reichardt &amp; Hinton" authorityYear="1976" box="[1382,1427,1845,1866]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="132" phylum="Arthropoda" rank="genus">
<emphasis id="B94BEAF5FFCCFF8AFAF7F8CDFA179831" box="[1382,1427,1845,1866]" italics="true" pageId="7" pageNumber="132">Yara</emphasis>
</taxonomicName>
; (45) sampling locality at La Libertad, Chontales, Nicaragua; (46) Pijibay Creek; (47) collecting macroinvertebrates using a Dnet at Pijibay Creek.
</paragraph>
</caption>
<paragraph id="8B8036E7FFC3FF85FF56FF6FFB4D9ED7" blockId="8.[151,1436,151,429]" pageId="8" pageNumber="133">
According to the physicochemical parameters measured, Pijibay Creek has permanent flow of well oxygenated water in both wet and dry seasons (5.3 ± 
<quantity id="4CC79B02FFC3FF85FD15FF44FD509FAF" box="[644,724,188,213]" metricMagnitude="-7" metricUnit="kg" metricValue="5.0" pageId="8" pageNumber="133" unit="mg" value="0.5">0.5 mg</quantity>
/L mean value; oxygen saturation: 64.2 ± 1.7%); the average temperature is 26 ± 0.6 °C; the electrical conductivity is low (51.3 ± 12.7 µS/cm), and the pH is slightly acidic (6.90 ± 0.57). All the parameters indicate that because of the moderate temperatures, slow flow of water, and moderate concentrations of oxygen, Pijibay Creek offers optimal conditions for the survival of aquatic life. Other groups of invertebrates sampled along with 
<taxonomicName id="4C3F4D64FFC3FF85FD18FEB5FCB99E1F" authority="Reyes &amp; Espinoza &amp; Girón, 2019" authorityName="Reyes &amp; Espinoza &amp; Girón" authorityYear="2019" box="[649,829,332,356]" class="Insecta" family="Hydroscaphidae" genus="Yara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="133" phylum="Arthropoda" rank="species" species="marmontsedu" status="sp. nov.">
<emphasis id="B94BEAF5FFC3FF85FD18FEB5FCB99E1F" box="[649,829,332,356]" italics="true" pageId="8" pageNumber="133">Y. marmontsedu</emphasis>
</taxonomicName>
<emphasis id="B94BEAF5FFC3FF85FCD4FEB4FC029E1E" bold="true" box="[837,902,332,357]" pageId="8" pageNumber="133">
<taxonomicNameLabel id="A278578EFFC3FF85FCD4FEB4FC029E1E" box="[837,902,332,357]" pageId="8" pageNumber="133" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
include, among others, certain members of the orders Ephemeroptera, Odonata, Plecoptera, Trichoptera, and other 
<taxonomicName id="4C3F4D64FFC3FF85FC05FE97FB969EF3" authorityName="" authorityYear="1758" box="[916,1042,367,392]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="133" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
, generally associated with clean to moderately contaminated (elevated contents of organic matter) waters (e.g. 
<bibRefCitation id="EFAE4B16FFC3FF85FC74FE6DFB399ED7" author="Hanson, P. &amp; Springer, M. &amp; Ramirez, A." box="[997,1213,404,428]" pageId="8" pageNumber="133" pagination="1 - 198" refId="ref4830" refString="Hanson, P., Springer, M. &amp; Ramirez, A. (2010) Macroinvertebrados de Agua Dulce de Costa Rica I. Revista de Biologia Tropical, 58 (4), 1 - 198. https: // doi. org / 10.15517 / rbt. v 58 i 4" type="journal article" year="2010">
Hanson 
<emphasis id="B94BEAF5FFC3FF85FBD0FE6DFBFD9ED7" box="[1089,1145,404,428]" italics="true" pageId="8" pageNumber="133">et al.</emphasis>
2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>