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<document id="BE4BA79E4E3F3EEDE60EBEF9CF18CC74" ID-DOI="10.11646/zootaxa.5403.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10561558" ID-ZooBank="7458460D-2367-48C5-9ACE-D28457C27463" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1706103452799" checkinUser="plazi" docAuthor="Dondorp, Esther &amp; Denzer, Wolfgang" docDate="2024" docId="0387E3417C37FFC474C8CB25CB57ED13" docLanguage="en" docName="zootaxa.5403.1.4.pdf" docOrigin="Zootaxa 5403 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lophyrus kuhlii Schlegel 1851" docType="treatment" docVersion="3" lastPageNumber="74" masterDocId="FFBE9B397C3EFFCD745FCA6FC845E926" masterDocTitle="Annotated catalogue of acrodont lizard types in the collection of the Naturalis Biodiverstity Center (Squamata: Agamidae, Chamaeleonidae)" masterLastPageNumber="79" masterPageNumber="65" pageNumber="74" updateTime="1706278432058" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="679A18F344334712537365240C04134A">Annotated catalogue of acrodont lizard types in the collection of the Naturalis Biodiverstity Center (Squamata: Agamidae, Chamaeleonidae)</mods:title>
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<mods:name id="34CDFB8AB876A3078443073EEBB79DE3" type="personal">
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<mods:roleTerm id="D7BBF9491DAAF82CA08B852402D45A61">Author</mods:roleTerm>
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<mods:namePart id="7DC85FBB818019435DE450678E18ED51">Dondorp, Esther</mods:namePart>
<mods:affiliation id="0521CAD8F8CA3AC134E8D15CD6DB83A9">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR, Leiden, The Netherlands.</mods:affiliation>
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<mods:namePart id="BF2FE15414CD5A80F909B025653ECC77">Denzer, Wolfgang</mods:namePart>
<mods:nameIdentifier id="09765DA5CB77DF1D634CD256C4A4E48C" type="ORCID">0000-0003-2619-4555</mods:nameIdentifier>
<mods:affiliation id="C037891EA69D77C83C471852B94B09E9">Society for Southeast Asian Herpetology, Calle Rio Segura 26, 30600 Archena / Murcia, Spain.</mods:affiliation>
<mods:nameIdentifier id="368E51571090EFFD4621013C82159B80" type="email">lobo@herpetologica.org</mods:nameIdentifier>
</mods:name>
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<mods:title id="CF57913EDC371D53E7092CD6892148BF">Zootaxa</mods:title>
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<mods:part id="5F7EE5C7C3EBFBB65407688D9194C8BF">
<mods:date id="ADD6FD6A5C1762330B86E8075BBF7A84">2024</mods:date>
<mods:detail id="8586B02E33B3C9F95EDBDF6D33D9D267" type="pubDate">
<mods:number id="77D993A3631FAB5FC21F2150EDE18EEE">2024-01-18</mods:number>
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<mods:detail id="CE6FECBFB09DBC21146C0C6DC97349EB" type="volume">
<mods:number id="BD96446FA81AAE7F6F126F6C54F88D60">5403</mods:number>
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<mods:number id="AA08488F4F8EDB7D991FEFE788297721">1</mods:number>
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<mods:identifier id="A9EF74D960D18493C6406B15BC1B33C2" type="DOI">10.11646/zootaxa.5403.1.4</mods:identifier>
<mods:identifier id="CC245B7C868DE1CE4B993463D92FFEA3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="47A4047367E748515FD9788888921AAE" type="Zenodo-Dep">10561558</mods:identifier>
<mods:identifier id="11A5B6FD465764B3BA15D6E3E729B3E4" type="ZooBank">7458460D-2367-48C5-9ACE-D28457C27463</mods:identifier>
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<treatment id="0387E3417C37FFC474C8CB25CB57ED13" LSID="urn:lsid:plazi:treatment:0387E3417C37FFC474C8CB25CB57ED13" httpUri="http://treatment.plazi.org/id/0387E3417C37FFC474C8CB25CB57ED13" lastPageNumber="74" pageId="9" pageNumber="74">
<subSubSection id="C33401DC7C37FFC474C8CB25CA74E842" box="[151,561,330,357]" pageId="9" pageNumber="74" type="nomenclature">
<paragraph id="8B9152577C37FFC474C8CB25CA74E842" blockId="9.[151,1048,330,357]" box="[151,561,330,357]" pageId="9" pageNumber="74">
<heading id="D0D9E53B7C37FFC474C8CB25CA74E842" bold="true" box="[151,561,330,357]" fontSize="11" level="1" pageId="9" pageNumber="74" reason="1">
<treatmentCitationGroup id="AB3E75797C37FFC474C8CB25CA74E842" box="[151,561,330,357]" pageId="9" pageNumber="74">
<treatmentCitation id="0A8F74467C37FFC474C8CB25CA74E842" author="Schlegel, H." box="[151,561,330,357]" page="5" pageId="9" pageNumber="74" year="1851">
<taxonomicName id="4C2E29D47C37FFC474C8CB25CA74E842" ID-CoL="3W55S" authority="Schlegel, 1851: 5" authorityName="Schlegel" authorityPageNumber="5" authorityYear="1851" box="[151,561,330,357]" class="Squamata" family="Agamidae" genus="Lophyrus" kingdom="Animalia" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="B95A8E457C37FFC474C8CB25CA74E842" bold="true" box="[151,561,330,357]" pageId="9" pageNumber="74">
<emphasis id="B95A8E457C37FFC474C8CB25C91FE843" bold="true" box="[151,346,330,357]" italics="true" pageId="9" pageNumber="74">Lophyrus kuhlii</emphasis>
<bibRefCitation id="EFBF2FA67C37FFC4753ECB25CA74E842" author="Schlegel, H." box="[353,561,330,356]" pageId="9" pageNumber="74" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel, 1851: 5</bibRefCitation>
</emphasis>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33401DC7C37FFC47667CB25CC5DE842" box="[568,1048,330,357]" pageId="9" pageNumber="74" type="reference_group">
<paragraph id="8B9152577C37FFC47667CB25CC5DE842" blockId="9.[151,1048,330,357]" box="[568,1048,330,357]" pageId="9" pageNumber="74">
<emphasis id="B95A8E457C37FFC47667CB25CC5DE842" bold="true" box="[568,1048,330,357]" pageId="9" pageNumber="74">
=
<taxonomicName id="4C2E29D47C37FFC4760FCB25CC5DE842" authority="(Schlegel, 1851)" baseAuthorityName="Schlegel" baseAuthorityYear="1851" box="[592,1048,330,357]" class="Squamata" family="Agamidae" genus="Gonocephalus" kingdom="Animalia" order="Scorpaeniformes" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="B95A8E457C37FFC4760FCB25CB0BE843" bold="true" box="[592,846,330,357]" italics="true" pageId="9" pageNumber="74">Gonocephalus kuhlii</emphasis>
(
<bibRefCitation id="EFBF2FA67C37FFC47702CB25CC55E842" author="Schlegel, H." box="[861,1040,330,356]" pageId="9" pageNumber="74" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel, 1851</bibRefCitation>
)
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33401DC7C37FFC474C8CBFDCAEAEB1A" pageId="9" pageNumber="74" type="materials_examined">
<paragraph id="8B9152577C37FFC474C8CBFDC9A7EB3F" blockId="9.[151,1437,402,1077]" pageId="9" pageNumber="74">
<materialsCitation id="3B46580A7C37FFC474C8CBFDCC4DE88A" box="[151,1032,402,429]" collectionCode="RMNH, RENA" collectorName="Kuhl &amp; van Hasselt" location="Java" pageId="9" pageNumber="74" specimenCount="1" stateProvince="Jawa Timur" typeStatus="syntype">
<emphasis id="B95A8E457C37FFC474C8CBFDC947E88A" bold="true" box="[151,258,402,428]" pageId="9" pageNumber="74">
<typeStatus id="5495ECF57C37FFC474C8CBFDC947E88A" box="[151,258,402,428]" pageId="9" pageNumber="74" type="syntype">Syntypes</typeStatus>
</emphasis>
:
<collectionCode id="ED3FCA927C37FFC4754ACBFDC92BE88B" box="[277,366,402,429]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="74" type="Museum">RMNH</collectionCode>
.
<collectionCode id="ED3FCA927C37FFC4752ECBFDC987E88B" box="[369,450,402,429]" pageId="9" pageNumber="74">RENA</collectionCode>
.3045, male,
<collectingRegion id="49EA9CB57C37FFC4760ACBFCCACCE88B" box="[597,649,403,429]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="74">Java</collectingRegion>
, coll.
<collectorName id="26DB37817C37FFC4768CCBFDCB4EE88A" box="[723,779,402,428]" pageId="9" pageNumber="74">Kuhl</collectorName>
&amp;
<collectorName id="26DB37817C37FFC47769CBFCCBFBE88A" box="[822,958,402,429]" pageId="9" pageNumber="74">van Hasselt</collectorName>
, 1821
</materialsCitation>
;
<materialsCitation id="3B46580A7C37FFC47048CBFDCA4CE8F6" collectionCode="RMNH, RENA" collectorName="Kuhl &amp; van Hasselt" location="Java" pageId="9" pageNumber="74" specimenCount="1" stateProvince="Jawa Timur" typeStatus="syntype">
<collectionCode id="ED3FCA927C37FFC47048CBFDCC35E88A" box="[1047,1136,402,428]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="74" type="Museum">RMNH</collectionCode>
.
<collectionCode id="ED3FCA927C37FFC4702CCBFDCC81E88A" box="[1139,1220,402,428]" pageId="9" pageNumber="74">RENA</collectionCode>
.3047a, male,
<collectingRegion id="49EA9CB57C37FFC4713CCBFCCDD2E88B" box="[1379,1431,403,429]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="74">Java</collectingRegion>
, coll.
<collectorName id="26DB37817C37FFC4748CCBD9C94EE8F6" box="[211,267,438,464]" pageId="9" pageNumber="74">Kuhl</collectorName>
&amp;
<collectorName id="26DB37817C37FFC47569CBD8C9FAE8F6" box="[310,447,438,465]" pageId="9" pageNumber="74">van Hasselt</collectorName>
, 1821
</materialsCitation>
;
<materialsCitation id="3B46580A7C37FFC47647CBD9CD77E8F6" box="[536,1330,438,465]" collectionCode="RMNH, RENA" collectorName="Kuhl &amp; van Hasselt" location="Java" pageId="9" pageNumber="74" specimenCount="1" stateProvince="Jawa Timur" typeStatus="syntype">
<collectionCode id="ED3FCA927C37FFC47647CBD9CA34E8F6" box="[536,625,438,464]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="74" type="Museum">RMNH</collectionCode>
.
<collectionCode id="ED3FCA927C37FFC4762BCBD9CA80E8F6" box="[628,709,438,464]" pageId="9" pageNumber="74">RENA</collectionCode>
.3047b, female,
<collectingRegion id="49EA9CB57C37FFC47723CBD8CBF5E8F7" box="[892,944,439,465]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="74">Java</collectingRegion>
, coll.
<collectorName id="26DB37817C37FFC477A3CBD9CC71E8F6" box="[1020,1076,438,464]" pageId="9" pageNumber="74">Kuhl</collectorName>
&amp;
<collectorName id="26DB37817C37FFC47000CBD8CCA2E8F6" box="[1119,1255,438,465]" pageId="9" pageNumber="74">van Hasselt</collectorName>
, 1821
</materialsCitation>
;
<materialsCitation id="3B46580A7C37FFC4711ECBD8C999EB3E" collectionCode="RMNH, RENA" location="Sumatra" pageId="9" pageNumber="74" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="syntype">
<collectionCode id="ED3FCA927C37FFC4711ECBD8CDDCE8F7" box="[1345,1433,439,465]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="74" type="Museum">RMNH</collectionCode>
.
<collectionCode id="ED3FCA927C37FFC474C8CBB5C8ADE8D2" box="[151,232,474,500]" pageId="9" pageNumber="74">RENA</collectionCode>
.3046, possibly male (not dissected),
<collectingRegion id="49EA9CB57C37FFC476D4CBB5CAAEE8D2" box="[651,747,474,500]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="74">Sumatra</collectingRegion>
, 1848; all specimens originally registered in the handwritten catalogue as
<taxonomicName id="4C2E29D47C37FFC47577CB91C999EB3E" authorityName="Lepeletier" authorityYear="1823" box="[296,476,510,536]" class="Squamata" family="Agamidae" genus="Lophyrus" kingdom="Animalia" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="minor">
<emphasis id="B95A8E457C37FFC47577CB91C999EB3E" box="[296,476,510,536]" italics="true" pageId="9" pageNumber="74">Lophyrus minor</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9152577C37FFC47498C84DCAEAEB1A" blockId="9.[151,1437,402,1077]" box="[199,687,546,573]" pageId="9" pageNumber="74">
<emphasis id="B95A8E457C37FFC47498C84DC925EB1A" bold="true" box="[199,352,546,572]" pageId="9" pageNumber="74">Type locality</emphasis>
:
<materialsCitation id="3B46580A7C37FFC47530C84CCAEAEB1A" box="[367,687,546,573]" country="Indonesia" location="Java" pageId="9" pageNumber="74" specimenCount="1" stateProvince="Jawa Timur">
<collectingRegion id="49EA9CB57C37FFC47530C84CC9E4EB1B" box="[367,417,547,573]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="74">Java</collectingRegion>
and Sumatra,
<collectingCountry id="F33912C77C37FFC4761DC84DCAEAEB1A" box="[578,687,546,572]" name="Indonesia" pageId="9" pageNumber="74">Indonesia</collectingCountry>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33401DC7C37FFC47498C829CB57ED13" pageId="9" pageNumber="74" type="discussion">
<paragraph id="8B9152577C37FFC47498C829CC74EA33" blockId="9.[151,1437,402,1077]" pageId="9" pageNumber="74">
<emphasis id="B95A8E457C37FFC47498C829C971EB46" bold="true" box="[199,308,582,608]" pageId="9" pageNumber="74">Remarks</emphasis>
:
<bibRefCitation id="EFBF2FA67C37FFC4751BC829C9B3EB47" author="Schlegel, H." box="[324,502,582,609]" pageId="9" pageNumber="74" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel (1851)</bibRefCitation>
noted that the
<specimenCount id="9D2899DE7C37FFC476FFC828CB02EB46" box="[672,839,582,609]" count="2" pageId="9" pageNumber="74" type="generic">two specimens</specimenCount>
collected by Kuhl &amp; van Hasselt were in a state that ”would not allow a rigorous examination“ [our translation]. When we retrieved the specimens we were surprised that they were actually in an acceptable state of preservation. While their colours are faded, morphological traits necessary for species identification are well preserved. The total length (10 pouces, 1 pouce approx.
<quantity id="4CD6FFB27C37FFC470B3C8DDCD7FEBEB" box="[1260,1338,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="74" unit="mm" value="25.0">25 mm</quantity>
in 1851) given in
<bibRefCitation id="EFBF2FA67C37FFC474A4C8B9C9EBEBD7" author="Schlegel, H." box="[251,430,726,753]" pageId="9" pageNumber="74" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel (1851)</bibRefCitation>
agrees well with the size of RMNH.RENA.3046 (
<quantity id="4CD6FFB27C37FFC477B3C8B9CC09EBD6" box="[1004,1100,726,752]" metricMagnitude="-1" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="74" unit="mm" value="260.0">260 mm</quantity>
) from
<collectingRegion id="49EA9CB57C37FFC470C4C8B9CCB9EBD6" box="[1179,1276,726,752]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="74">Sumatra</collectingRegion>
. The
<typeStatus id="5495ECF57C37FFC47165C8B8CDD9EBD7" box="[1338,1436,727,753]" pageId="9" pageNumber="74" type="syntype">syntypes</typeStatus>
RMNH.RENA 3047a, b are probably the specimens illustrated in
<bibRefCitation id="EFBF2FA67C37FFC47727C895CC68EA33" author="Schlegel, H." box="[888,1069,762,789]" pageId="9" pageNumber="74" pagination="4 - 6" refId="ref11831" refString="Schlegel, H. (1851) Descriptions de plusieurs expeces nouvelles du genre Lophyrus. Bijdragen tot de Dierkunde, Amsterdam, 3 (1), 4 - 6. https: // doi. org / 10.1163 / 26660644 - 00301002" type="journal article" year="1851">Schlegel (1851)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9152577C37FFC47498C971CB57ED13" blockId="9.[151,1437,402,1077]" pageId="9" pageNumber="74">
Several publications (e.g.
<bibRefCitation id="EFBF2FA67C37FFC475AAC971CAD8EA1F" author="de Rooij, N." box="[501,669,798,825]" pageId="9" pageNumber="74" refId="ref9589" refString="de Rooij, N. (1915) The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. E. J. Brill, Leiden, xiv + 384 pp. https: // doi. org / 10.5962 / bhl. title. 10610" type="book" year="1915">de Rooij 1915</bibRefCitation>
,
<bibRefCitation id="EFBF2FA67C37FFC476F4C971CB8BEA1E" author="Manthey, U. &amp; Denzer, W." box="[683,974,798,825]" pageId="9" pageNumber="74" pagination="23 - 28" refId="ref11027" refString="Manthey, U. &amp; Denzer, W. (1993) Die echten Winkelkopfagamen der Gattung Gonocephalus Kaup (Sauria: Agamidae), Teil 4. Die chamaeleontinus - Gruppe. Sauria, 15 (2), 23 - 28." type="journal article" year="1993">Manthey &amp; Denzer 1993</bibRefCitation>
) gave 1848 as the publication date, but according to the publisher Brill the third volume of the Bijdragen tot de Dierkunde was published on
<date id="FF9074977C37FFC47171C92DC897EAA7" pageId="9" pageNumber="74" value="1851-06-14">14th June 1851</date>
. We would like to note that while the authorship of this taxon is typically assigned to Schlegel, who provided a detailed description and illustrations of
<specimenCount id="9D2899DE7C37FFC47617C9E4CAA9EA82" box="[584,748,906,933]" count="2" pageId="9" pageNumber="74" type="generic">two specimens</specimenCount>
taken in life, already
<bibRefCitation id="EFBF2FA67C37FFC4778AC9E5CC19EA82" author="Gray, J. E." box="[981,1116,906,932]" pageId="9" pageNumber="74" refId="ref10005" refString="Gray, J. E. (1831) A synopsis of the species of Class Reptilia. In: Griffith, E &amp; E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed. V. Whittaker, Treacher and Co., London, 481 + 110 pp. [1830]" type="book" year="1831">Gray (1831)</bibRefCitation>
gave a very short description of “
<taxonomicName id="4C2E29D47C37FFC4749BC9C1C9BCEAEE" authority=", Boie, MS" authorityName="Boie, MS" box="[196,505,942,969]" class="Squamata" family="Agamidae" genus="Lophyrus" kingdom="Animalia" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="kuhlii">
<emphasis id="B95A8E457C37FFC4749BC9C1C932EAEE" box="[196,375,942,968]" italics="true" pageId="9" pageNumber="74">Lophyrus kuhlii</emphasis>
, Boie, MS
</taxonomicName>
“. The manuscript
<bibRefCitation id="EFBF2FA67C37FFC47694C9C1CB12EAEE" author="Gray, J. E." box="[715,855,942,968]" pageId="9" pageNumber="74" refId="ref10005" refString="Gray, J. E. (1831) A synopsis of the species of Class Reptilia. In: Griffith, E &amp; E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed. V. Whittaker, Treacher and Co., London, 481 + 110 pp. [1830]" type="book" year="1831">Gray (1831)</bibRefCitation>
referred to was most probably Boie’s unpublished Erpetologie de
<collectingRegion id="49EA9CB57C37FFC4751FC9BCC930EACB" box="[320,373,979,1005]" country="Indonesia" name="Jawa Timur" pageId="9" pageNumber="74">Java</collectingRegion>
. If Gray’s description is accepted as the original publication RMNH.RENA.3046 does no longer constitute a
<typeStatus id="5495ECF57C37FFC4757DC998C93CED37" box="[290,377,1015,1041]" pageId="9" pageNumber="74" type="syntype">syntype</typeStatus>
owing to its late collection date, and only specimens that were available to Boie prior to his finishing the manuscript in 1825 would have type status.
</paragraph>
</subSubSection>
</treatment>
</document>
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