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<document id="4FADAA2241FE73045B11AFC54BC59A4F" ID-DOI="10.15407/zoo2022.02.091" ID-ISSN="2707-7268" ID-Zenodo-Dep="7175709" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665294511910" checkinUser="felipe" docAuthor="Glotov, S., Hushtan, K., Hushtan, H., Koval, N. &amp; Diedus, V." docDate="2022" docId="03E487DDFFBAFF8DFF3F0D94FF108747" docLanguage="en" docName="Zoodiversity.56.1.91–110.pdf" docOrigin="Zoodiversity 56 (1)" docSource="http://dx.doi.org/10.15407/zoo2022.02.091" docStyle="DocumentStyle:65E6F982F72656C606C1639D3996B172.2:Zoodiversity.2020-.journal_article" docStyleId="65E6F982F72656C606C1639D3996B172" docStyleName="Zoodiversity.2020-.journal_article" docStyleVersion="2" docTitle="Atheta mortuorum Thomson 1867" docType="treatment" docVersion="3" lastPageNumber="107" masterDocId="FFDDFFA5FFAAFF9DFF990E30FFE2805E" masterDocTitle="The Genus Atheta (Coleoptera, Staphylinidae, Aleocharinae) In The Ukrainian Carpathians" masterLastPageNumber="110" masterPageNumber="91" pageNumber="107" updateTime="1699381295323" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="CB07A17B94BF99924AAEB6B87CEE66C3">The Genus Atheta (Coleoptera, Staphylinidae, Aleocharinae) In The Ukrainian Carpathians</mods:title>
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<mods:namePart id="E6260685F3FC3BCE765CBBE5D1AD473B">Glotov, S.</mods:namePart>
<mods:affiliation id="5DF095C0AA8E46A11092AE6D52EF2E19">State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine</mods:affiliation>
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<mods:namePart id="4B9B8A08D7F056C44D0A7C98FF87392F">Hushtan, K.</mods:namePart>
<mods:affiliation id="EFE391AE87FD7B59148294076136667A">State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine &amp; Ecological College of Lviv National Agrarian University, Zamarstynivska st., 167, Lviv, 79068 Ukraine</mods:affiliation>
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<mods:namePart id="929C5C19FDCBA42DF295CCCE9D91BCB5">Hushtan, H.</mods:namePart>
<mods:affiliation id="800692705F039664ECC701B387C6BFC7">State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine</mods:affiliation>
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<mods:namePart id="F126ABC93C4383C32B8C52501BE633A8">Koval, N.</mods:namePart>
<mods:affiliation id="EE8E48AA7FECBF75C9A980B855BDB306">Uzhgorod National University, A. Voloshyn st., 32, Uzhgorod, 88000 Ukraine &amp; Uzhansky National Nature Park, Nezalezhnosti st., 7, village Velykyi Bereznyi, Ukraine</mods:affiliation>
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<mods:namePart id="FB9D4D9F51B900D0E1C68C922882792F">Diedus, V.</mods:namePart>
<mods:affiliation id="ACB01D0FBAD7BFFF877759749D712726">Uzhgorod National University, A. Voloshyn st., 32, Uzhgorod, 88000 Ukraine</mods:affiliation>
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<mods:date id="38DDEF04269B557368B8ABEE71188565">2022</mods:date>
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<subSubSection id="C3576540FFBAFF8DFF3F0D94FDA583E0" box="[166,583,932,959]" pageId="16" pageNumber="107" type="nomenclature">
<paragraph id="8BF236CBFFBAFF8DFF3F0D94FDA583E0" blockId="16.[166,583,932,959]" box="[166,583,932,959]" pageId="16" pageNumber="107">
<heading id="D0BA81A7FFBAFF8DFF3F0D94FDA583E0" box="[166,583,932,959]" fontSize="36" level="2" pageId="16" pageNumber="107" reason="3">
<taxonomicName id="4C4D4D48FFBAFF8DFF3F0D94FDA583E0" ID-CoL="JDKN" authority="Thomson, 1867" authorityName="Thomson" authorityYear="1867" box="[166,583,932,959]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="107" phylum="Arthropoda" rank="species" species="mortuorum">
<emphasis id="B939EAD9FFBAFF8DFF3F0D94FE6883E1" bold="true" box="[166,394,932,959]" italics="true" pageId="16" pageNumber="107">Atheta mortuorum</emphasis>
Thomson, 1867
</taxonomicName>
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</paragraph>
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<subSubSection id="C3576540FFBAFF8DFF420DE3FD8F84C7" pageId="16" pageNumber="107" type="materials_examined">
<paragraph id="8BF236CBFFBAFF8DFF420DE3FC3083B7" blockId="16.[219,978,979,1001]" box="[219,978,979,1001]" pageId="16" pageNumber="107">
Material.
<specimenCount id="9D4BFD42FFBAFF8DFED60DE3FE9E83B7" box="[335,380,979,1001]" pageId="16" pageNumber="107" type="generic">2 ex.</specimenCount>
<materialsCitation id="3B253C96FFBAFF8DFE180DE3FC2C83B7" box="[385,974,979,1001]" collectionCode="SMNH" pageId="16" pageNumber="107" specimenCount="2">
<emphasis id="B939EAD9FFBAFF8DFE180DE3FE4F83B6" bold="true" box="[385,429,979,1000]" pageId="16" pageNumber="107">IFR:</emphasis>
Vorokhta [Worochta], date not specified,
<specimenCount id="9D4BFD42FFBAFF8DFCDD0DE3FC9383B7" box="[836,881,979,1001]" pageId="16" pageNumber="107" type="generic">2 ex.</specimenCount>
(
<collectionCode id="ED5CAE0EFFBAFF8DFCE50DE3FC2583B7" box="[892,967,979,1001]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="16" pageNumber="107">SMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF420DCFFDE78467" blockId="16.[166,1225,1023,1114]" pageId="16" pageNumber="107">
Distribution. Europe, North Africa, Siberia, North Korea (
<bibRefCitation id="EFDC4B3AFFBAFF8DFC490DCFFF018464" author="Schulke, M. &amp; Smetana, A." pageId="16" pageNumber="107" pagination="1 - 1702" refId="ref15821" refString="Schulke, M., Smetana, A. 2015. Staphylinidae Latreille 1802, 304 - 1134. In: Lobl, I. &amp; Lobl, D., eds. Catalogue of Palaearctic Coleoptera vols. 1 &amp; 2, Hydrophiloidea - Staphylinoidea, revised and updated edition. Brill, Leiden &amp; Boston, I - XXV, 1 - 1702." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
);
<emphasis id="B939EAD9FFBAFF8DFF6F0A2FFDE78467" bold="true" box="[246,517,1055,1081]" pageId="16" pageNumber="107">
<collectingCountry id="F35A765BFFBAFF8DFF6F0A2FFEB88467" box="[246,346,1055,1081]" name="Ukraine" pageId="16" pageNumber="107">Ukraine</collectingCountry>
(first record).
</emphasis>
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF420A0FFB6D8407" blockId="16.[166,1225,1023,1114]" box="[219,1167,1087,1114]" pageId="16" pageNumber="107">Bionomics. The species occurs mainly in lowlands and foothills, in forest litter.</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF3F0A4FFD8F84C7" blockId="16.[166,621,1151,1210]" box="[166,621,1151,1178]" pageId="16" pageNumber="107">
<emphasis id="B939EAD9FFBAFF8DFF3F0A4FFEFE84C7" bold="true" box="[166,284,1151,1177]" pageId="16" pageNumber="107">Subgenus</emphasis>
<taxonomicName id="4C4D4D48FFBAFF8DFEBA0A4FFD8F84C7" authority="Munster, 1923" authorityName="Munster" authorityYear="1923" box="[291,621,1151,1178]" class="Insecta" family="Staphylinidae" genus="Rhagocneme" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B939EAD9FFBAFF8DFEBA0A4FFE5E84C4" bold="true" box="[291,444,1151,1178]" italics="true" pageId="16" pageNumber="107">Rhagocneme</emphasis>
Munster, 1923
</taxonomicName>
</paragraph>
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<subSubSection id="C3576540FFBAFF8DFF3F0AAFFF108747" pageId="16" pageNumber="107" type="description">
<paragraph id="8BF236CBFFBAFF8DFF3F0AAFFDDA84E7" blockId="16.[166,621,1151,1210]" box="[166,568,1183,1210]" pageId="16" pageNumber="107">
<taxonomicName id="4C4D4D48FFBAFF8DFF3F0AAFFDDA84E7" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[166,568,1183,1210]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="107" phylum="Arthropoda" rank="species" species="subsinuata">
<emphasis id="B939EAD9FFBAFF8DFF3F0AAFFE6084E4" bold="true" box="[166,386,1183,1210]" italics="true" pageId="16" pageNumber="107">Atheta subsinuata</emphasis>
Erichson, 1839
</taxonomicName>
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF420AE8FC3084B0" blockId="16.[219,978,1240,1262]" box="[219,978,1240,1262]" pageId="16" pageNumber="107">
Material.
<specimenCount id="9D4BFD42FFBAFF8DFED60AE8FE9E84B0" box="[335,380,1240,1262]" pageId="16" pageNumber="107" type="generic">1 ex.</specimenCount>
<materialsCitation id="3B253C96FFBAFF8DFE180AE8FC2C84B0" box="[385,974,1240,1262]" collectionCode="SMNH" pageId="16" pageNumber="107" specimenCount="1">
<emphasis id="B939EAD9FFBAFF8DFE180AE8FE4F84B3" bold="true" box="[385,429,1240,1261]" pageId="16" pageNumber="107">IFR:</emphasis>
Vorokhta [Worochta], date not specified,
<specimenCount id="9D4BFD42FFBAFF8DFCDD0AE8FC9384B0" box="[836,881,1240,1262]" pageId="16" pageNumber="107" type="generic">1 ex.</specimenCount>
(
<collectionCode id="ED5CAE0EFFBAFF8DFCE50AE8FC2584B0" box="[892,967,1240,1262]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="16" pageNumber="107">SMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF420B20FD2E8537" blockId="16.[166,1225,1295,1450]" pageId="16" pageNumber="107">
D i s t r i b u t i o n. Europe, North America (
<bibRefCitation id="EFDC4B3AFFBAFF8DFD580B3FFC138574" author="Schulke, M. &amp; Smetana, A." box="[705,1009,1295,1322]" pageId="16" pageNumber="107" pagination="1 - 1702" refId="ref15821" refString="Schulke, M., Smetana, A. 2015. Staphylinidae Latreille 1802, 304 - 1134. In: Lobl, I. &amp; Lobl, D., eds. Catalogue of Palaearctic Coleoptera vols. 1 &amp; 2, Hydrophiloidea - Staphylinoidea, revised and updated edition. Brill, Leiden &amp; Boston, I - XXV, 1 - 1702." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
). In the Ukrainian Carpathians known from Transcarpathian Region, Berehovo (
<bibRefCitation id="EFDC4B3AFFBAFF8DFC3A0B1FFBDC8517" author="Kuthy, D." box="[931,1086,1327,1353]" pageId="16" pageNumber="107" pagination="1 - 214" refId="ref15193" refString="Kuthy, D. 1896. Ordo Coleoptera. Fauna Regni Hungariae. III. Arthropoda, Insecta. Budapest, 1 - 214." type="book chapter" year="1896">Kuthy, 1896</bibRefCitation>
). “
<collectingCountry id="F35A765BFFBAFF8DFBFB0B1FFB5B8517" box="[1122,1209,1327,1353]" name="Poland" pageId="16" pageNumber="107">Poland</collectingCountry>
”, without indication of region (
<bibRefCitation id="EFDC4B3AFFBAFF8DFD980B7FFD5D8537" author="Lomnicki, M." box="[513,703,1359,1385]" pageId="16" pageNumber="107" pagination="21 - 155" refId="ref15390" refString="Lomnicki, M. 1913. Wykaz chrzaszczow czyli Tegopokrywych (Coleoptera) ziem polskich. (Catalogus coleopterorum Poloniae). Kosmos. Seria A Biologia, 21 - 155." type="book chapter" year="1913">Łomnicki, 1913</bibRefCitation>
).
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF420B5FFE0885F4" blockId="16.[166,1225,1295,1450]" pageId="16" pageNumber="107">Bionomics. The species occurs mainly in lowlands and foothills, in forest litter, in rotten wood and in mosses.</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF3F0BFFFD6585B7" blockId="16.[166,662,1487,1546]" box="[166,647,1487,1514]" pageId="16" pageNumber="107">
<emphasis id="B939EAD9FFBAFF8DFF3F0BFFFEFE85B7" bold="true" box="[166,284,1487,1513]" pageId="16" pageNumber="107">Subgenus</emphasis>
<taxonomicName id="4C4D4D48FFBAFF8DFEBA0BFFFD6585B7" authority="Mulsant &amp; Rey, 1873" authorityName="Mulsant &amp; Rey" authorityYear="1873" box="[291,647,1487,1514]" class="Insecta" family="Staphylinidae" genus="Tetropla" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B939EAD9FFBAFF8DFEBA0BFFFE6985B4" bold="true" box="[291,395,1487,1514]" italics="true" pageId="16" pageNumber="107">Tetropla</emphasis>
Mulsant &amp; Rey, 1873
</taxonomicName>
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF3F0BDFFD748657" blockId="16.[166,662,1487,1546]" box="[166,662,1519,1546]" pageId="16" pageNumber="107">
<taxonomicName id="4C4D4D48FFBAFF8DFF3F0BDFFDC48657" authority="(Stephens, 1832)" baseAuthorityName="Stephens" baseAuthorityYear="1832" box="[166,550,1519,1546]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="107" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B939EAD9FFBAFF8DFF3F0BDFFEB98654" bold="true" box="[166,347,1519,1546]" italics="true" pageId="16" pageNumber="107">Atheta liturata</emphasis>
(Stephens, 1832)
</taxonomicName>
(
<figureCitation id="13762A4EFFBAFF8DFDAD0BDFFD8D8657" box="[564,623,1519,1545]" captionStart="Fig" captionStartId="4.[170,203,1668,1690]" captionTargetBox="[229,1172,250,1618]" captionTargetId="figure-13@4.[166,1205,244,1630]" captionTargetPageId="4" captionText="Fig. 1. Habitus: a — Atheta coriaria (Kraatz, 1856); b — A. divisa (Märkel, 1844); c — A. graminicola (Gravenhorst, 1806); d — A. oblita (Erichson, 1839); e — A. ravilla (Erichson, 1839); f — A. macrocera (Thomson, 1856); g — A. occulta (Erichson, 1837); h — A. longicornis (Gravenhorst, 1802); i — A. melanaria (Mannerheim, 1830); j — A. zosterae (Thomson, 1856); k — A. atramentaria (Gyllenhal, 1810); l — A. marcida (Erichson, 1837); m — A. subtilis (Scriba, 1866); n — A. amicula (Stephens, 1832); o — A. laticollis Stephens, 1832; p — A. liturata (Stephens, 1832); q — A. nigritula (Gravenhorst, 1802)." figureDoi="http://doi.org/10.5281/zenodo.7175711" httpUri="https://zenodo.org/record/7175711/files/figure.png" pageId="16" pageNumber="107">fig. 1</figureCitation>
, p)
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF420818FB948660" blockId="16.[219,1142,1576,1598]" box="[219,1142,1576,1598]" pageId="16" pageNumber="107">
Material.
<specimenCount id="9D4BFD42FFBAFF8DFED60818FE9E8660" box="[335,380,1576,1598]" pageId="16" pageNumber="107" type="generic">3 ex.</specimenCount>
<materialsCitation id="3B253C96FFBAFF8DFE180818FB918660" box="[385,1139,1576,1598]" collectionCode="ZMTSNU" location="Chernivtsi" pageId="16" pageNumber="107" specimenCount="3" stateProvince="Chernivtsi">
<emphasis id="B939EAD9FFBAFF8DFE180818FE578663" bold="true" box="[385,437,1576,1598]" pageId="16" pageNumber="107">CER:</emphasis>
<collectingRegion id="4989F829FFBAFF8DFE230818FDFD8660" box="[442,543,1576,1598]" country="Ukraine" name="Chernivtsi" pageId="16" pageNumber="107">Chernivtsi</collectingRegion>
[Bucowina. Czernowits], data not specified,
<specimenCount id="9D4BFD42FFBAFF8DFC550818FC1B8660" box="[972,1017,1576,1598]" pageId="16" pageNumber="107" type="generic">3 ex.</specimenCount>
(
<collectionCode id="ED5CAE0EFFBAFF8DFB9D0818FB8E8660" box="[1028,1132,1576,1598]" pageId="16" pageNumber="107">ZMTSNU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF42086FFC0F86C7" blockId="16.[166,1225,1631,1817]" pageId="16" pageNumber="107">
Distribution. Europe (
<bibRefCitation id="EFDC4B3AFFBAFF8DFD93086FFCAA8624" author="Schulke, M. &amp; Smetana, A." box="[522,840,1631,1658]" pageId="16" pageNumber="107" pagination="1 - 1702" refId="ref15821" refString="Schulke, M., Smetana, A. 2015. Staphylinidae Latreille 1802, 304 - 1134. In: Lobl, I. &amp; Lobl, D., eds. Catalogue of Palaearctic Coleoptera vols. 1 &amp; 2, Hydrophiloidea - Staphylinoidea, revised and updated edition. Brill, Leiden &amp; Boston, I - XXV, 1 - 1702." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
). In the Ukrainian Carpathians known from Transcarpathian Region, mountain Hoverla (
<bibRefCitation id="EFDC4B3AFFBAFF8DFCCB084FFC0786C7" author="Kuthy, D." box="[850,997,1663,1689]" pageId="16" pageNumber="107" pagination="1 - 214" refId="ref15193" refString="Kuthy, D. 1896. Ordo Coleoptera. Fauna Regni Hungariae. III. Arthropoda, Insecta. Budapest, 1 - 214." type="book chapter" year="1896">Kuthy, 1896</bibRefCitation>
)
</paragraph>
<paragraph id="8BF236CBFFBAFF8DFF4208AFFF108747" blockId="16.[166,1225,1631,1817]" pageId="16" pageNumber="107">
Bionomics. The species occurs mainly in foothills and high in the mountains in forests and mountain meadows, mainly in young mature and rotten fungi, mainly in
<taxonomicName id="4C4D4D48FFBAFF8DFF3F08D0FD8386A4" authority="(Bull.) Murr." authorityName="Murr." baseAuthorityName="Bull." box="[166,609,1759,1786]" class="Agaricomycetes" family="Fomitopsidaceae" genus="Laetiporus" kingdom="Fungi" order="Polyporales" pageId="16" pageNumber="107" phylum="Basidiomycota" rank="species" species="sulphureus">
<emphasis id="B939EAD9FFBAFF8DFF3F08D0FE5186A7" box="[166,435,1759,1785]" italics="true" pageId="16" pageNumber="107">Laetiporus sulphureus</emphasis>
(Bull.) Murr.
</taxonomicName>
, as well as in forest litter (
<bibRefCitation id="EFDC4B3AFFBAFF8DFC2E08EFFBBC86A7" author="Roubal, J." box="[951,1118,1759,1785]" pageId="16" pageNumber="107" pagination="1 - 527" refId="ref15689" refString="Roubal, J. 1930. Katalog Coleopter (brouku) Slovenska a Podkarpatska. Praha, 1 - 527." type="book chapter" year="1930">Roubal, 1930</bibRefCitation>
; Glotov, 2020).
</paragraph>
</subSubSection>
</treatment>
</document>
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