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<document id="465A45B0124B78579EA6E2A6E588AD72" ID-DOI="10.11646/zootaxa.3616.1.1" ID-GBIF-Dataset="ba683f8e-57d5-4573-bf93-b1bbb53be92e" ID-ISSN="1175-5326" ID-Zenodo-Dep="283352" ID-ZooBank="0D52FFF5-0099-42D4-9298-7942A03C5057" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460575469493" checkinUser="plazi" docAuthor="Viraktamath, C. A. &amp; Gonçalves, Ana Clara" docDate="2013" docId="03E487C6FFE41735FF498C48FE545A1E" docLanguage="en" docName="zt03616p021.pdf" docOrigin="Zootaxa 3616 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Igerna neosa Webb, comb. nov." docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FFDDFFBEFFED173FFFDE8D27FFFF5E4E" masterDocTitle="Review of Madagascaran Agalliini (Hemiptera: Cicadellidae: Megophthalminae) with descriptions of a new genus and six new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="10" updateTime="1698323014752" updateUser="plazi">
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<mods:title id="C751BF35B05CE09A3AF7C3B71BAA5F82">Review of Madagascaran Agalliini (Hemiptera: Cicadellidae: Megophthalminae) with descriptions of a new genus and six new species</mods:title>
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<mods:namePart id="A2BB042F15593556172B4BAE2C34E03C">Viraktamath, C. A.</mods:namePart>
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<mods:namePart id="0A96DB055AF569D5F70050DA2374735B">Gonçalves, Ana Clara</mods:namePart>
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<mods:date id="CBC8D7E5795EC88C9B5C55F505A5C678">2013</mods:date>
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<mods:number id="15A5DC6EB7CC09376CF306F10F0258BF">3616</mods:number>
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<treatment id="03E487C6FFE41735FF498C48FE545A1E" ID-DOI="http://doi.org/10.5281/zenodo.5694139" ID-GBIF-Taxon="127663278" ID-Zenodo-Dep="5694139" LSID="urn:lsid:plazi:treatment:03E487C6FFE41735FF498C48FE545A1E" httpUri="http://treatment.plazi.org/id/03E487C6FFE41735FF498C48FE545A1E" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C357655BFFE41736FF498C48FB965FBE" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BF236D0FFE41736FF498C48FDD85FC7" blockId="9.[151,551,367,426]" box="[151,551,367,393]" pageId="9" pageNumber="10">
<heading id="D0BA81BCFFE41736FF498C48FDD85FC7" bold="true" box="[151,551,367,393]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B939EAC2FFE41736FF498C48FDD85FC7" bold="true" box="[151,551,367,393]" pageId="9" pageNumber="10">
<taxonomicName id="4C4D4D53FFE41736FF498C48FE655FC7" ID-CoL="8JTWY" authority="Webb" authorityName="Webb" box="[151,410,367,393]" class="Insecta" family="Cicadellidae" genus="Igerna" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="neosa" status="comb. nov.">
<emphasis id="B939EAC2FFE41736FF498C48FECA5FC7" bold="true" box="[151,309,367,393]" italics="true" pageId="9" pageNumber="10">Igerna neosa</emphasis>
(Webb)
</taxonomicName>
<taxonomicNameLabel id="A20A57B9FFE41736FE7F8C48FDD85FC7" box="[417,551,367,393]" pageId="9" pageNumber="10">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF236D0FFE41736FF498CB5FDDD5FE4" blockId="9.[151,551,367,426]" box="[151,546,402,426]" pageId="9" pageNumber="10">
(
<figureCitation id="13762A55FFE41736FF418CB5FEDF5FE4" box="[159,288,402,426]" captionStart="FIGURES 15 22" captionStartId="13.[151,264,1834,1857]" captionTargetBox="[192,1408,241,1769]" captionTargetId="figure@13.[160,1427,193,1813]" captionTargetPageId="13" captionText="FIGURES 15 22. Species of Madagascaran Agalliini. Body, lateral and dorsal view: (15 16) Igerna delta sp. nov.; (17 18) I. neosa (Webb); (19 20) I. malagasica sp. nov., male; (21 22) same, female." httpUri="https://zenodo.org/record/283355/files/figure.png" pageId="9" pageNumber="10">Figs 1718</figureCitation>
,
<figureCitation id="13762A55FFE41736FEF58CB5FEB55FE4" box="[299,330,402,426]" captionStart="FIGURES 23 33" captionStartId="14.[151,264,1852,1875]" captionTargetBox="[215,1378,239,1801]" captionTargetId="figure@14.[192,1396,193,1831]" captionTargetPageId="14" captionText="FIGURES 23 33. Species of Agalliini. Face: (23) Agallidwipa biramosa sp. nov.; (24) A. bispinosa sp. nov., male; (25) same, female; (26) A. pauliana (Evans); (27) A. webbi sp. nov., male; (28) same, female; (29) Igerna delta sp. nov.; (30) I. malagasica sp. nov., male; (31) same, female; (32) I. neosa (Webb); (33) Agalliana ensigera Oman." httpUri="https://zenodo.org/record/283356/files/figure.png" pageId="9" pageNumber="10">32</figureCitation>
,
<figureCitation id="13762A55FFE41736FE8B8CB5FE8C5FE4" box="[341,371,402,426]" captionStart="FIGURES 34 41" captionStartId="15.[151,264,1583,1606]" captionTargetBox="[169,1422,225,1525]" captionTargetId="figure@15.[151,1436,193,1562]" captionTargetPageId="15" captionText="FIGURES 34 41. Species of Madagascaran Agalliini. Genital capsules, lateral view: (34) Igerna delta sp. nov.; (35) I. neosa (Webb); (36) I. malagasica sp. nov.; (37) I. flavocosta sp. nov.; (38) Agallidwipa biramosa sp. nov.; (39) A. webbi sp. nov.; (40) A. bispinosa sp. nov.; (41) A. pauliana (Evans), Arrows point out posterior margin of pygofer." httpUri="https://zenodo.org/record/283357/files/figure.png" pageId="9" pageNumber="10">35</figureCitation>
,
<figureCitation id="13762A55FFE41736FEA18CB5FE625FE4" box="[383,413,402,426]" captionStart="FIGURES 42 53" captionStartId="16.[151,264,1569,1592]" captionTargetBox="[168,1420,205,1523]" captionTargetId="figure@16.[151,1436,193,1548]" captionTargetPageId="16" captionText="FIGURES 42 53. Species of Madagascaran Agalliini. 42 49. Styles, dorsal views (except 46), respectively: (42) Igerna malagasica sp. nov.; (43) I. flavocosta sp. nov.; (44) I. delta sp. nov.; (45) I. neosa (Webb); (46 47) Agallidwipa bispinosa sp. nov., lateral and dorsal view respectively; (48) A. biramosa sp. nov.; (49) A. pauliana (Evans). 50 53. Subgenital plates, ventral view: (50) I. flavocosta sp. nov.; (51) A. biramosa sp. nov.; (52) I. delta sp. nov.; (53) A. bispinosa sp. nov." httpUri="https://zenodo.org/record/283358/files/figure.png" pageId="9" pageNumber="10">45</figureCitation>
,
<figureCitation id="13762A55FFE41736FE778CB5FE385FE4" box="[425,455,402,426]" captionStart="FIGURES 54 61" captionStartId="17.[151,264,1440,1463]" captionTargetBox="[157,1416,229,1364]" captionTargetId="figure@17.[151,1436,193,1419]" captionTargetPageId="17" captionText="FIGURES 54 61. Species of Madagascaran Agalliini. Aedeagus (except 57), lateral views: (54) Agallidwipa pauliana (Evans); (55) A. bispinosa sp. nov.; (56) A. biramosa sp. nov.; (57) A. webbi sp. nov., style, connective and aedeagus; (58) Igerna delta sp. nov.; (59) I. neosa (Webb); (60) Igerna malagasica sp. nov.; (61) I. flavocosta sp. nov., Arrows point out the ventral projection prominence." httpUri="https://zenodo.org/record/283359/files/figure.png" pageId="9" pageNumber="10">59</figureCitation>
,
<figureCitation id="13762A55FFE41736FE0D8CB5FE0D5FE4" box="[467,498,402,426]" captionStart="FIGURES 62 69" captionStartId="18.[151,264,1614,1637]" captionTargetBox="[188,1407,217,1575]" captionTargetId="figure@18.[172,1414,193,1593]" captionTargetPageId="18" captionText="FIGURES 62 69. Species of Madagascaran Agalliini. Aedeagus, ventral views (except 65 &amp; 66): (62) Agallidwipa bispinosa sp. nov.; (63) A. biramosa sp. nov.; (64) A. webbi sp. nov.; (65) A. pauliana (Evans) aedeagus and connective, ventral view;; (66) Igerna delta sp. nov., dorsal view; (67) I. neosa (Webb); (68) I. flavocosta sp. nov.; (69) I. malagasica sp. nov." httpUri="https://zenodo.org/record/283360/files/figure.png" pageId="9" pageNumber="10">67</figureCitation>
,
<figureCitation id="13762A55FFE41736FE238CB5FDE55FE4" box="[509,538,402,426]" captionStart="FIGURES 70 85" captionStartId="19.[151,264,1860,1883]" captionTargetBox="[239,1366,222,1797]" captionTargetId="figure@19.[207,1380,193,1839]" captionTargetPageId="19" captionText="FIGURES 70 85. Morphological aspects of Agallidwipa and Igerna species. (70 73) A. biramosa sp. nov.: (70) profemur; (71) apodemes of second abdominal sternite; (72) connective; (73) X segment processes, lateral view; (74 76) A. bispinosa sp. nov.: (74) forewing, lateral view; (75) connective; (76) X segment processes, lateral view; (77 79) A. pauliana (Evans): (77) apodemes of second abdominal sternite; (78) pygofer posterior margin; (79) X segment processes, lateral view; (80) A. webbi sp. nov., style and connective, dorsal view; (81 82) I. delta sp. nov.: (81) pygofer process, caudal view; (82) connective; (83) I. neosa (Webb), apex of pygofer process; (84 85) I. malagasica sp. nov.: (84) pygofer processes, caudal view; (85) connective." httpUri="https://zenodo.org/record/283361/files/figure.png" pageId="9" pageNumber="10">83</figureCitation>
)
</paragraph>
<paragraph id="8BF236D0FFE41736FF498CFDFB965FBE" blockId="9.[151,1129,474,496]" box="[151,1129,474,496]" pageId="9" pageNumber="10">
<taxonomicName id="4C4D4D53FFE41736FF498CFDFE235FBE" authority="Webb 1980: 837" authorityName="Webb" authorityPageNumber="837" authorityYear="1980" box="[151,476,474,496]" class="Insecta" family="Cicadellidae" genus="Stonasla" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="neosa">
<emphasis id="B939EAC2FFE41736FF498CFDFED15FBE" box="[151,302,474,496]" italics="true" pageId="9" pageNumber="10">Stonasla neosa</emphasis>
Webb 1980: 837
</taxonomicName>
838, figs 2937.
<typeStatus id="54F68872FFE41736FD4D8CFDFD0E5FBE" box="[659,753,474,496]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
3, Aldabra [BMNH, not examined].
</paragraph>
</subSubSection>
<subSubSection id="C357655BFFE41736FF498F3AFA735BF7" pageId="9" pageNumber="10" type="description">
<paragraph id="8BF236D0FFE41736FF498F3AFDFB5C14" blockId="9.[151,1437,541,1684]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF498F3AFEB25C7B" bold="true" box="[151,333,541,565]" pageId="9" pageNumber="10">Measurements.</emphasis>
Male
<quantity id="4CB59B35FFE41736FE478F3AFDC15C7B" box="[409,574,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.55" metricValueMin="3.45" pageId="9" pageNumber="10" unit="mm" value="3.5" valueMax="3.55" valueMin="3.45">3.453.55 mm</quantity>
long and
<quantity id="4CB59B35FFE41736FD6A8F3AFCA65C7B" box="[692,857,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="9" pageNumber="10" unit="mm" value="1.2" valueMax="1.25" valueMin="1.15">1.151.25 mm</quantity>
wide across eyes; female
<quantity id="4CB59B35FFE41736FB548F3AFACF5C7B" box="[1162,1328,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="3.825" metricValueMax="3.85" metricValueMin="3.8" pageId="9" pageNumber="10" unit="mm" value="3.825" valueMax="3.85" valueMin="3.8">3.803.85 mm</quantity>
long and
<quantity id="4CB59B35FFE41736FF498F66FEC55C17" box="[151,314,577,601]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" metricValueMax="1.3" metricValueMin="1.12" pageId="9" pageNumber="10" unit="mm" value="1.21" valueMax="1.3" valueMin="1.12">1.121.30 mm</quantity>
wide across eyes.
</paragraph>
<paragraph id="8BF236D0FFE41736FF178F42FF0D5A63" blockId="9.[151,1437,541,1684]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF178F42FE015C30" bold="true" box="[201,510,613,638]" pageId="9" pageNumber="10">Head and thorax (color).</emphasis>
Ground color ochraceous with brown markings. Crown (
<figureCitation id="13762A55FFE41736FB788F42FB035C33" box="[1190,1276,613,637]" captionStart="FIGURES 15 22" captionStartId="13.[151,264,1834,1857]" captionTargetBox="[192,1408,241,1769]" captionTargetId="figure@13.[160,1427,193,1813]" captionTargetPageId="13" captionText="FIGURES 15 22. Species of Madagascaran Agalliini. Body, lateral and dorsal view: (15 16) Igerna delta sp. nov.; (17 18) I. neosa (Webb); (19 20) I. malagasica sp. nov., male; (21 22) same, female." httpUri="https://zenodo.org/record/283355/files/figure.png" pageId="9" pageNumber="10">Fig. 18</figureCitation>
) with pair of elliptical dark brown spots equally distant between eyes and median line, and with median line bearing brown macula, all of them contiguous to facial ones. Face (
<figureCitation id="13762A55FFE41736FD338F8AFCBF5C8B" box="[749,832,685,709]" captionStart="FIGURES 23 33" captionStartId="14.[151,264,1852,1875]" captionTargetBox="[215,1378,239,1801]" captionTargetId="figure@14.[192,1396,193,1831]" captionTargetPageId="14" captionText="FIGURES 23 33. Species of Agalliini. Face: (23) Agallidwipa biramosa sp. nov.; (24) A. bispinosa sp. nov., male; (25) same, female; (26) A. pauliana (Evans); (27) A. webbi sp. nov., male; (28) same, female; (29) Igerna delta sp. nov.; (30) I. malagasica sp. nov., male; (31) same, female; (32) I. neosa (Webb); (33) Agalliana ensigera Oman." httpUri="https://zenodo.org/record/283356/files/figure.png" pageId="9" pageNumber="10">Fig. 32</figureCitation>
) with conspicuous inverted brown Y-shaped macula; area adjacent to ocelli dark brown followed by brown maculae; pair of brown maculae adjacent to upper margin of compound eyes. Frontoclypeus (
<figureCitation id="13762A55FFE41736FDC68FD2FD9D5D43" box="[536,610,757,781]" captionStart="FIGURES 23 33" captionStartId="14.[151,264,1852,1875]" captionTargetBox="[215,1378,239,1801]" captionTargetId="figure@14.[192,1396,193,1831]" captionTargetPageId="14" captionText="FIGURES 23 33. Species of Agalliini. Face: (23) Agallidwipa biramosa sp. nov.; (24) A. bispinosa sp. nov., male; (25) same, female; (26) A. pauliana (Evans); (27) A. webbi sp. nov., male; (28) same, female; (29) Igerna delta sp. nov.; (30) I. malagasica sp. nov., male; (31) same, female; (32) I. neosa (Webb); (33) Agalliana ensigera Oman." httpUri="https://zenodo.org/record/283356/files/figure.png" pageId="9" pageNumber="10">Fig.32</figureCitation>
) with several irregular brown maculae forming pair of lateral rows; clypellus pale brown with brown median stripe; genae ochraceous with dark brown, elongate maculae on adjacent antennal ledges; lora pale brown with inner margin dark brown. Pronotum (
<figureCitation id="13762A55FFE41736FBD48E1AFB725D1B" box="[1034,1165,829,853]" captionStart="FIGURES 15 22" captionStartId="13.[151,264,1834,1857]" captionTargetBox="[192,1408,241,1769]" captionTargetId="figure@13.[160,1427,193,1813]" captionTargetPageId="13" captionText="FIGURES 15 22. Species of Madagascaran Agalliini. Body, lateral and dorsal view: (15 16) Igerna delta sp. nov.; (17 18) I. neosa (Webb); (19 20) I. malagasica sp. nov., male; (21 22) same, female." httpUri="https://zenodo.org/record/283355/files/figure.png" pageId="9" pageNumber="10">Figs 1718</figureCitation>
) with five markings as follows: one brown line contiguous to coronal median macula, and attaining the pronotal posterior margin; one pair of small maculae at anterior area, and one pair of large spots at lateral area of disc, dark brown. Mesoscutum (
<figureCitation id="13762A55FFE41736FAB08EA2FF4A5D8F" captionStart="FIGURES 15 22" captionStartId="13.[151,264,1834,1857]" captionTargetBox="[192,1408,241,1769]" captionTargetId="figure@13.[160,1427,193,1813]" captionTargetPageId="13" captionText="FIGURES 15 22. Species of Madagascaran Agalliini. Body, lateral and dorsal view: (15 16) Igerna delta sp. nov.; (17 18) I. neosa (Webb); (19 20) I. malagasica sp. nov., male; (21 22) same, female." httpUri="https://zenodo.org/record/283355/files/figure.png" pageId="9" pageNumber="10">Fig. 18</figureCitation>
), pale brown with pair of brown laterobasal triangular markings, scuto-scutellar suture dark brown; mesoscutellum pale brown. Forewings (
<figureCitation id="13762A55FFE41736FD878EEAFD265DAB" box="[601,729,973,997]" captionStart="FIGURES 15 22" captionStartId="13.[151,264,1834,1857]" captionTargetBox="[192,1408,241,1769]" captionTargetId="figure@13.[160,1427,193,1813]" captionTargetPageId="13" captionText="FIGURES 15 22. Species of Madagascaran Agalliini. Body, lateral and dorsal view: (15 16) Igerna delta sp. nov.; (17 18) I. neosa (Webb); (19 20) I. malagasica sp. nov., male; (21 22) same, female." httpUri="https://zenodo.org/record/283355/files/figure.png" pageId="9" pageNumber="10">Figs 1718</figureCitation>
) pale brown with veins brown, and some longitudinal areas of cells paler. Legs (
<figureCitation id="13762A55FFE41736FE818ED6FE4D5A47" box="[351,434,1009,1033]" captionStart="FIGURES 15 22" captionStartId="13.[151,264,1834,1857]" captionTargetBox="[192,1408,241,1769]" captionTargetId="figure@13.[160,1427,193,1813]" captionTargetPageId="13" captionText="FIGURES 15 22. Species of Madagascaran Agalliini. Body, lateral and dorsal view: (15 16) Igerna delta sp. nov.; (17 18) I. neosa (Webb); (19 20) I. malagasica sp. nov., male; (21 22) same, female." httpUri="https://zenodo.org/record/283355/files/figure.png" pageId="9" pageNumber="10">Fig. 17</figureCitation>
) with longitudinal pale brown to brown patches. Thoracic pleura dark brown with paler borders.
</paragraph>
<paragraph id="8BF236D0FFE41736FF17891DFDFD5A1F" blockId="9.[151,1437,541,1684]" box="[201,514,1081,1106]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF17891DFE9D5A1C" bold="true" box="[201,354,1082,1106]" pageId="9" pageNumber="10">Morphology.</emphasis>
As in
<taxonomicName id="4C4D4D53FFE41736FE72891DFE045A1F" box="[428,507,1081,1105]" class="Insecta" family="Cicadellidae" genus="Igerna" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="delta">
<emphasis id="B939EAC2FFE41736FE72891DFE045A1F" box="[428,507,1081,1105]" italics="true" pageId="9" pageNumber="10">I. delta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF236D0FFE41736FF17897AFE3F5BDB" blockId="9.[151,1437,541,1684]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF17897AFE825A38" bold="true" box="[201,381,1117,1142]" pageId="9" pageNumber="10">Male genitalia.</emphasis>
Pygofer (
<figureCitation id="13762A55FFE41736FE2E897AFDB95A3B" box="[496,582,1117,1141]" captionStart="FIGURES 34 41" captionStartId="15.[151,264,1583,1606]" captionTargetBox="[169,1422,225,1525]" captionTargetId="figure@15.[151,1436,193,1562]" captionTargetPageId="15" captionText="FIGURES 34 41. Species of Madagascaran Agalliini. Genital capsules, lateral view: (34) Igerna delta sp. nov.; (35) I. neosa (Webb); (36) I. malagasica sp. nov.; (37) I. flavocosta sp. nov.; (38) Agallidwipa biramosa sp. nov.; (39) A. webbi sp. nov.; (40) A. bispinosa sp. nov.; (41) A. pauliana (Evans), Arrows point out posterior margin of pygofer." httpUri="https://zenodo.org/record/283357/files/figure.png" pageId="9" pageNumber="10">Fig. 35</figureCitation>
), in lateral view, with posterior margin narrowing towards bifid apex, and directed inward; in caudal view (
<figureCitation id="13762A55FFE41736FDC889A6FD955AD7" box="[534,618,1153,1177]" captionStart="FIGURES 70 85" captionStartId="19.[151,264,1860,1883]" captionTargetBox="[239,1366,222,1797]" captionTargetId="figure@19.[207,1380,193,1839]" captionTargetPageId="19" captionText="FIGURES 70 85. Morphological aspects of Agallidwipa and Igerna species. (70 73) A. biramosa sp. nov.: (70) profemur; (71) apodemes of second abdominal sternite; (72) connective; (73) X segment processes, lateral view; (74 76) A. bispinosa sp. nov.: (74) forewing, lateral view; (75) connective; (76) X segment processes, lateral view; (77 79) A. pauliana (Evans): (77) apodemes of second abdominal sternite; (78) pygofer posterior margin; (79) X segment processes, lateral view; (80) A. webbi sp. nov., style and connective, dorsal view; (81 82) I. delta sp. nov.: (81) pygofer process, caudal view; (82) connective; (83) I. neosa (Webb), apex of pygofer process; (84 85) I. malagasica sp. nov.: (84) pygofer processes, caudal view; (85) connective." httpUri="https://zenodo.org/record/283361/files/figure.png" pageId="9" pageNumber="10">Fig. 83</figureCitation>
) projection with crenulate dorsal margin, caudoventral margin sinuate. Styles (
<figureCitation id="13762A55FFE41736FF338982FEBC5AF3" box="[237,323,1189,1213]" captionStart="FIGURES 42 53" captionStartId="16.[151,264,1569,1592]" captionTargetBox="[168,1420,205,1523]" captionTargetId="figure@16.[151,1436,193,1548]" captionTargetPageId="16" captionText="FIGURES 42 53. Species of Madagascaran Agalliini. 42 49. Styles, dorsal views (except 46), respectively: (42) Igerna malagasica sp. nov.; (43) I. flavocosta sp. nov.; (44) I. delta sp. nov.; (45) I. neosa (Webb); (46 47) Agallidwipa bispinosa sp. nov., lateral and dorsal view respectively; (48) A. biramosa sp. nov.; (49) A. pauliana (Evans). 50 53. Subgenital plates, ventral view: (50) I. flavocosta sp. nov.; (51) A. biramosa sp. nov.; (52) I. delta sp. nov.; (53) A. bispinosa sp. nov." httpUri="https://zenodo.org/record/283358/files/figure.png" pageId="9" pageNumber="10">Fig. 45</figureCitation>
), in dorsal view, well developed, outer fork very large, subtruncate, longer than inner fork, with subapical digitiform process, bearing scattered setae; inner fork with truncate apex, bearing digitiform process directed outwards. Connective as in
<taxonomicName id="4C4D4D53FFE41736FDE089C9FD6F5B4B" box="[574,656,1261,1285]" class="Insecta" family="Cicadellidae" genus="Igerna" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="delta">
<emphasis id="B939EAC2FFE41736FDE089C9FD6F5B4B" box="[574,656,1261,1285]" italics="true" pageId="9" pageNumber="10">I. delta</emphasis>
</taxonomicName>
. Aedeagus (
<figureCitation id="13762A55FFE41736FCFC89CAFC835B4B" box="[802,892,1261,1285]" captionStart="FIGURES 54 61" captionStartId="17.[151,264,1440,1463]" captionTargetBox="[157,1416,229,1364]" captionTargetId="figure@17.[151,1436,193,1419]" captionTargetPageId="17" captionText="FIGURES 54 61. Species of Madagascaran Agalliini. Aedeagus (except 57), lateral views: (54) Agallidwipa pauliana (Evans); (55) A. bispinosa sp. nov.; (56) A. biramosa sp. nov.; (57) A. webbi sp. nov., style, connective and aedeagus; (58) Igerna delta sp. nov.; (59) I. neosa (Webb); (60) Igerna malagasica sp. nov.; (61) I. flavocosta sp. nov., Arrows point out the ventral projection prominence." httpUri="https://zenodo.org/record/283359/files/figure.png" pageId="9" pageNumber="10">Figs 59</figureCitation>
,
<figureCitation id="13762A55FFE41736FC5589CAFC575B4B" box="[907,936,1261,1285]" captionStart="FIGURES 62 69" captionStartId="18.[151,264,1614,1637]" captionTargetBox="[188,1407,217,1575]" captionTargetId="figure@18.[172,1414,193,1593]" captionTargetPageId="18" captionText="FIGURES 62 69. Species of Madagascaran Agalliini. Aedeagus, ventral views (except 65 &amp; 66): (62) Agallidwipa bispinosa sp. nov.; (63) A. biramosa sp. nov.; (64) A. webbi sp. nov.; (65) A. pauliana (Evans) aedeagus and connective, ventral view;; (66) Igerna delta sp. nov., dorsal view; (67) I. neosa (Webb); (68) I. flavocosta sp. nov.; (69) I. malagasica sp. nov." httpUri="https://zenodo.org/record/283360/files/figure.png" pageId="9" pageNumber="10">67</figureCitation>
) with well developed dorsal apodeme 0.33 times as long as shaft in lateral view, and prominent ventral projection; shaft in ventral view, almost straight; bearing anterolateral foliaceous expansion at left side with crenulate margin; gonopore apical, on ventral margin; apex rounded. Anal tube (
<figureCitation id="13762A55FFE41736FE63887EFDEE5B3F" box="[445,529,1369,1393]" captionStart="FIGURES 34 41" captionStartId="15.[151,264,1583,1606]" captionTargetBox="[169,1422,225,1525]" captionTargetId="figure@15.[151,1436,193,1562]" captionTargetPageId="15" captionText="FIGURES 34 41. Species of Madagascaran Agalliini. Genital capsules, lateral view: (34) Igerna delta sp. nov.; (35) I. neosa (Webb); (36) I. malagasica sp. nov.; (37) I. flavocosta sp. nov.; (38) Agallidwipa biramosa sp. nov.; (39) A. webbi sp. nov.; (40) A. bispinosa sp. nov.; (41) A. pauliana (Evans), Arrows point out posterior margin of pygofer." httpUri="https://zenodo.org/record/283357/files/figure.png" pageId="9" pageNumber="10">Fig. 35</figureCitation>
), with segment X simple, without any process; segment XI with bifid spiniform process on ventral margin.
</paragraph>
<paragraph id="8BF236D0FFE41736FF178886FA735BF7" blockId="9.[151,1437,541,1684]" box="[201,1420,1441,1466]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF178886FE245BF4" bold="true" box="[201,475,1441,1466]" pageId="9" pageNumber="10">Intraspecific variation.</emphasis>
Some individuals have paler ground color with facial and pronotal markings darker.
</paragraph>
</subSubSection>
<subSubSection id="C357655BFFE41736FF1788E2FC185802" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BF236D0FFE41736FF1788E2FC185802" blockId="9.[151,1437,541,1684]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF1788E2FE4D5B90" bold="true" box="[201,434,1477,1502]" pageId="9" pageNumber="10">Material examined.</emphasis>
<collectingCountry id="F35A7640FFE41736FE6688E2FD7B5B93" box="[440,644,1477,1501]" name="Madagascar" pageId="9" pageNumber="10">MADAGASCAR</collectingCountry>
: 1 3, 2 Ƥ, Province dAntsiranana, Parc National Montagne, dAmbre,
<quantity id="4CB59B35FFE41736FF4988CBFF0D584A" box="[151,242,1516,1540]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="9" pageNumber="10" unit="m" value="1125.0">1125 m</quantity>
,
<date id="FFF31010FFE41736FEDD88CBFD7D584A" box="[259,642,1516,1541]" pageId="9" pageNumber="10" value="2001-01-29" valueMax="2001-02-11" valueMin="2001-01-29">29 January to 11 February 2001</date>
,
<geoCoordinate id="EE795017FFE41736FD4D88CBFCDD584A" box="[659,802,1513,1540]" direction="south" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="-12.520278">12o31 13”S</geoCoordinate>
,
<geoCoordinate id="EE795017FFE41736FCEA88CBFC47584A" box="[820,952,1513,1540]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="49.17917">49o1045”E</geoCoordinate>
, California Acad Science, R. HarinHala collector, Malaise trap, MA-01-01D-04; 1 3 1 Ƥ, same data except
<date id="FFF31010FFE41736FC478B37FBA25866" box="[921,1117,1552,1577]" pageId="9" pageNumber="10" value="2001-05-14" valueMax="2001-05-30" valueMin="2001-05-14">1430 May 2001</date>
, MA-01-01D-11; 2 Ƥ, same data except
<date id="FFF31010FFE41736FEC28B13FD7C5802" box="[284,643,1588,1613]" pageId="9" pageNumber="10" value="2001-01-29" valueMax="2001-02-11" valueMin="2001-01-29">29 January to 11 February 2001</date>
, MA-01-01D-03 (CAS, INHS).
</paragraph>
</subSubSection>
<subSubSection id="C357655BFFE41736FF178B7FFAA95983" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BF236D0FFE41736FF178B7FFD0D58DA" blockId="9.[151,1437,541,1684]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF178B7FFEC9583E" bold="true" box="[201,310,1624,1648]" pageId="9" pageNumber="10">Remarks</emphasis>
. This species differs from other species of
<taxonomicName id="4C4D4D53FFE41736FCC58B7EFC99583E" box="[795,870,1625,1648]" class="Insecta" family="Cicadellidae" genus="Igerna" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B939EAC2FFE41736FCC58B7EFC99583E" box="[795,870,1625,1648]" italics="true" pageId="9" pageNumber="10">Igerna</emphasis>
</taxonomicName>
known from the island in having a less developed anterolateral foliaceous process on the aedeagal shaft.
</paragraph>
<paragraph id="8BF236D0FFE41736FF498BFAFE8058B8" blockId="9.[151,383,1757,1782]" box="[151,383,1757,1782]" pageId="9" pageNumber="10">
<emphasis id="B939EAC2FFE41736FF498BFAFE8058B8" bold="true" box="[151,383,1757,1782]" pageId="9" pageNumber="10">Acknowledgements</emphasis>
</paragraph>
<paragraph id="8BF236D0FFE41736FF498A01FAA95983" blockId="9.[151,1436,1829,1998]" pageId="9" pageNumber="10">
We are grateful to Dr. C.H. Dietrich (INHS), California Academy of Sciences and Dr. M. Boulard, Paris Museum,
<collectingCountry id="F35A7640FFE41736FF498A6DFF19592F" box="[151,230,1866,1889]" name="France" pageId="9" pageNumber="10">France</collectingCountry>
, for making available the material used in this study, to Dr. J. Poorani, National Bureau of Agriculturally Important Insects, Bangalore, and Mr. Yeshwanth, H.M. University of Agricultural Sciences, Bangalore for help in preparing the illustrations. A fellowship from Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) to A. C. G. is acknowledged. We also thank the anonymous reviewer for his constructive comments.
</paragraph>
</subSubSection>
<subSubSection id="C357655BFFE71735FF498DB0FE545A1E" pageId="10" pageNumber="11" type="reference_group">
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Quelques nouveuax Cicadellides (Homopteres) de
<collectingCountry id="F35A7640FFE71735FCBB8C91FC1C5F82" box="[869,995,438,460]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
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<emphasis id="B939EAC2FFE71735FDD98CF2FD7D5FA5" box="[519,642,469,491]" italics="true" pageId="10" pageNumber="11">Coloborrhis</emphasis>
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