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<document id="4B3D54244F4A8084AB56A44EAB4EDB2F" ID-CLB-Dataset="300345" ID-DOI="10.1186/s13358-022-00257-z" ID-GBIF-Dataset="21b570bc-eafa-4448-b23b-61a89ddddd63" ID-ISSN="1664-2384" ID-PMC="PMC9499918" ID-PubMed="36164559" ID-Zenodo-Dep="11998918" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1718652706118" checkinUser="felipe" docAuthor="Ferrante, Christophe, Menkveld-Gfeller, Ursula &amp; Cavin, Lionel" docDate="2022" docId="03E487A4F34BFF86139F8B66BDCDFBFD" docLanguage="en" docName="SwissJPalaeontol.141.15.pdf" docOrigin="Swiss Journal of Palaeontology (15) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00257-z" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Libys callolepis Ferrante &amp; Menkveld-Gfeller &amp; Cavin 2022, sp. nov." docType="treatment" docUuid="876BF37D-30B0-4DBD-BEFA-FC66C579129A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="6" masterDocId="FFDDFFDCF34FFF83132A8D4DBF0DFFFE" masterDocTitle="The first Jurassic coelacanth from Switzerland" masterLastPageNumber="20" masterPageNumber="1" pageNumber="5" updateTime="1722275673157" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="23EE8E50BA3F288B37DCC08829F3DEEA">The first Jurassic coelacanth from Switzerland</mods:title>
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<mods:namePart id="4595A22BC4D2818341EB52D08219E574">Ferrante, Christophe</mods:namePart>
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<mods:namePart id="B338F74E12517065F1E7EED7DCF7726E">Menkveld-Gfeller, Ursula</mods:namePart>
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<mods:namePart id="D409E23DF4E091DC2B1A71C4635BCE54">Cavin, Lionel</mods:namePart>
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<mods:date id="3D7F7FE76DC9FAD1442A1A62126F5561">2022</mods:date>
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<treatment id="03E487A4F34BFF86139F8B66BDCDFBFD" ID-DOI="http://doi.org/10.5281/zenodo.13126254" ID-Zenodo-Dep="13126254" LSID="urn:lsid:zoobank.org:act:876BF37D-30B0-4DBD-BEFA-FC66C579129A" httpUri="http://treatment.plazi.org/id/03E487A4F34BFF86139F8B66BDCDFBFD" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C3576539F34BFF87139F8B66BE91F9BD" box="[181,412,1579,1603]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BF236B2F34BFF87139F8B66BE91F9BD" blockId="4.[181,412,1579,1603]" box="[181,412,1579,1603]" pageId="4" pageNumber="5">
<heading id="D0BA81DEF34BFF87139F8B66BE91F9BD" box="[181,412,1579,1603]" fontSize="10" level="2" pageId="4" pageNumber="5" reason="7">
<taxonomicName id="4C4D4D31F34BFF87139F8B66BE32F9BD" authority="Ferrante &amp; Menkveld-Gfeller &amp; Cavin, 2022" authorityName="Ferrante &amp; Menkveld-Gfeller &amp; Cavin" authorityYear="2022" box="[181,319,1579,1603]" class="Coelacanthi" family="Mawsoniidae" genus="Libys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coelacanthiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="callolepis" status="sp. nov.">L. callolepis</taxonomicName>
<taxonomicNameLabel id="A20A57DBF34BFF87126F8B66BE91F9BD" box="[325,412,1579,1603]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B939EAA0F34BFF87126F8B66BE98F9BD" bold="true" box="[325,405,1579,1603]" pageId="4" pageNumber="5">sp. nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8713BD8B26BCBCFEDD" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BF236B2F34BFF8713BD8B26BE07F97D" blockId="4.[151,775,1643,1891]" box="[151,266,1643,1667]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8713BD8B26BE07F97D" bold="true" box="[151,266,1643,1667]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8713BD8BC6BCBCFEDD" blockId="4.[151,775,1643,1891]" lastBlockId="4.[813,1437,235,291]" pageId="4" pageNumber="5">
<taxonomicName id="4C4D4D31F34BFF8713BD8BC6BE2BF95D" box="[151,294,1675,1699]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8713BD8BC6BFC2F95D" box="[151,207,1675,1699]" italics="true" pageId="4" pageNumber="5">Libys</emphasis>
species
</taxonomicName>
with the postparietal shield about half the length of the parietonasal shield (the parietonasal is then proportionally shorter than in the
<typeStatus id="54F68810F34BFF8711208B81BD34F91A" box="[522,569,1740,1764]" pageId="4" pageNumber="5">type</typeStatus>
species). Te teeth covering the prearticular are very small, and rounded and smooth. Between 4147 neural arches. Fin rays are slender than in the
<typeStatus id="54F68810F34BFF87126E8A61BE7EF8BA" box="[324,371,1836,1860]" pageId="4" pageNumber="5">type</typeStatus>
species and then not expanded. Te scales are strongly ornamented with irregularly sized and elongated round-to-ovoid ridges disposed along a longitudinal axis.
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8710078C06BBE6FDFD" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF236B2F34BFF8710078C06BB63FE9D" blockId="4.[813,1436,331,419]" box="[813,1134,331,355]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078C06BB63FE9D" bold="true" box="[813,1134,331,355]" pageId="4" pageNumber="5">Measurements and meristic</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078C26BB0FFE5D" blockId="4.[813,1436,331,419]" pageId="4" pageNumber="5">
(TB) Total body length
<quantity id="4CB59B57F34BFF87171A8C26BB86FE7D" box="[1072,1163,363,387]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="5" unit="mm" value="290.0">290 mm</quantity>
(estimation); (SL) standard length
<quantity id="4CB59B57F34BFF8710888CC6BCF3FE5D" box="[930,1022,395,419]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" pageId="4" pageNumber="5" unit="mm" value="255.0">255 mm</quantity>
.
</paragraph>
<paragraph id="8BF236B2F34BFF8710078C86BBE6FDFD" blockId="4.[813,1437,458,515]" pageId="4" pageNumber="5">d1.f= 10; d2.f =16; pect.f =1822; pelv.f ≥17; ana.f= 2023; cau.f= 15/1416; n.a =4447; h.a= 1820.</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8710078F66BB90FD3D" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BF236B2F34BFF8710078F66BCA4FDBD" blockId="4.[813,1436,555,708]" box="[813,937,555,579]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078F66BCA4FDBD" bold="true" box="[813,937,555,579]" pageId="4" pageNumber="5">Etymology</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078F01BB90FD3D" blockId="4.[813,1436,555,708]" pageId="4" pageNumber="5">
From the ancient Greek ΚΑλός, kalós, (beautiful, nice) and λεπίς, lepís, (scale) in reference to the nicely ornamented scales of the species, which differentiates it from the
<typeStatus id="54F68810F34BFF8717278FE1BB33FD3A" box="[1037,1086,684,708]" pageId="4" pageNumber="5">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF8710078FA6BC87FA3D" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BF236B2F34BFF8710078FA6BBC6FCFD" blockId="4.[813,1437,747,995]" box="[813,1227,747,771]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078FA6BBC6FCFD" bold="true" box="[813,1227,747,771]" pageId="4" pageNumber="5">
<typeStatus id="54F68810F34BFF8710078FA6BC94FCFD" box="[813,921,747,771]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
and only known specimen
</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078E41BB74FC1D" blockId="4.[813,1437,747,995]" pageId="4" pageNumber="5">
<materialsCitation id="3B253CEFF34BFF8710078E41BB74FC1D" collectionCode="NMBE" country="Switzerland" county="Natural History Museum" location="Most" municipality="Te" pageId="4" pageNumber="5" specimenCode="NMBE 5034072" specimenCount="1" stateProvince="Bern" typeStatus="holotype">
<specimenCode id="DBEB9EC9F34BFF8710078E41BCD0FCDD" box="[813,989,779,803]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="4" pageNumber="5">NMBE 5034072</specimenCode>
and 5034073, a sub-complete specimen preserved in right lateral view as part and counterpart. Most of the bones, including the scales on the body, are preserved in anatomical position and only the bones of the cheek and the jaw are missing. Te specimen is kept in the collections of the Natural History Museum Bern (Canton of Bern, Switzerland).
</materialsCitation>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078946BB5EFBDD" blockId="4.[813,1437,1035,1155]" box="[813,1107,1035,1059]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078946BB5EFBDD" bold="true" box="[813,1107,1035,1059]" pageId="4" pageNumber="5">
Horizon and
<typeStatus id="54F68810F34BFF8710EE8946BCFAFBDD" box="[964,1015,1035,1059]" pageId="4" pageNumber="5">type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078966BB25FB7D" blockId="4.[813,1437,1035,1155]" pageId="4" pageNumber="5">
Toarcian (Lower Jurassic), Creux de lOurs section, locality of Les Pueys near the Teysachaux summit (
<collectingRegion id="4989F850F34BFF8716078906BC82FB7D" country="Switzerland" name="Fribourg" pageId="4" pageNumber="5">Canton of Fribourg</collectingRegion>
,
<collectingCountry id="F35A7622F34BFF8710B28926BB11FB7D" box="[920,1052,1131,1155]" name="Switzerland" pageId="4" pageNumber="5">Switzerland</collectingCountry>
).
</paragraph>
<paragraph id="8BF236B2F34BFF87100789E6BB09FB3D" blockId="4.[813,1437,1195,1475]" box="[813,1028,1195,1219]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF87100789E6BB09FB3D" bold="true" box="[813,1028,1195,1219]" pageId="4" pageNumber="5">Nomenclatural act</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078981BC87FA3D" blockId="4.[813,1437,1195,1475]" pageId="4" pageNumber="5">
Te present work and its nomenclatural act are registered in ZooBank, the online registration system for the International Commission on Zoological Nomenclature. Te Life Science Identifiers for this publication is “
<uri id="FFDC3AB0F34BFF8710728806BB27FA7D" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:pub:
<uuid id="FFEB0C67F34BFF8717558806BB27FA7D" pageId="4" pageNumber="5">D03A2AC9-51F9-45E6- 8CAC-F06A1526AF2C</uuid>
</uri>
” and can be resolved appending the prefix “http://zoobank.org/” in any standard web browser.
</paragraph>
</subSubSection>
<subSubSection id="C3576539F34BFF86100788A7BDCDFBFD" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="8BF236B2F34BFF87100788A7BCB6F9FD" blockId="4.[813,1437,1514,1891]" box="[813,955,1514,1539]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF87100788A7BCB6F9FD" bold="true" box="[813,955,1514,1539]" pageId="4" pageNumber="5">Description</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078B40BCA1F9DA" blockId="4.[813,1437,1514,1891]" box="[813,940,1549,1572]" pageId="4" pageNumber="5">
<emphasis id="B939EAA0F34BFF8710078B40BCA1F9DA" bold="true" box="[813,940,1549,1572]" pageId="4" pageNumber="5">Generalities</emphasis>
</paragraph>
<paragraph id="8BF236B2F34BFF8710078B61BB84F89D" blockId="4.[813,1437,1514,1891]" pageId="4" pageNumber="5">
Most of the bones of the specimen are preserved on the part (
<collectionCode id="ED5CAE77F34BFF87104C8B06BCBCF99D" box="[870,945,1611,1635]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="4" pageNumber="5">NMBE</collectionCode>
5034073) (
<figureCitation id="13762A37F34BFF8717028B06BB6EF99D" box="[1064,1123,1611,1635]" captionStart="Fig" captionStartId="5.[168,199,1822,1842]" captionTargetBox="[151,1437,237,1950]" captionTargetId="figure-14@6.[192,1395,253,1913]" captionTargetPageId="6" captionText="Fig. 3 Skeleton of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073).A Photos with osteological details: 1, denticles on the proximal fin rays of the caudal fin. 2, Postparietal shield with the otic sensory canal opening as a deep groove crossed by pillars (white arrowhead). 3, Posterior parietal and the supraorbitals with their pillars (white arrowhead). 4, Consolidated snout with the anterior opening for the rostral organ (white arrowhead). 5,Teeth on the prearticular. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126231" httpUri="https://zenodo.org/record/13126231/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
). From the head, most of the dermal bones of the skull roof are preserved. Te bones of the cheek and the lower jaw are missing, revealing then the bones of the palatoquadrate and of the branchial apparatus. Te axial skeleton, the girdles and the fins are almost entirely preserved, except the posterior tip of the supplementary caudal fin lobe. Te scales are well preserved on the entire body in natural position, especially on the counterpart (
<collectionCode id="ED5CAE77F34BFF8710A88A06BCC0F89D" box="[898,973,1867,1891]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="4" pageNumber="5">NMBE</collectionCode>
5034072) (
<figureCitation id="13762A37F34BFF87176B8A06BB76F89D" box="[1089,1147,1867,1891]" captionStart="Fig" captionStartId="7.[168,199,1783,1803]" captionTargetBox="[152,1438,237,1949]" captionTargetId="figure-14@8.[198,1392,253,1911]" captionTargetPageId="8" captionText="Fig. 4 Skeleton of Libys callolepis sp. nov. on the counterpart (holotype, NMBE 5034072).A Photos with osteological details: 1, articular head of the scapulocoracoid. 2, Scales on the flank immediately beneath the first anterior dorsal fin. 3, Scales of the lateral line showing the ornamental pattern with the larger central tubercles (white arrowheads point, showed only on one scale).4, Scales on the ventral flank from the pelvic to the anal fin. 5, Axial mesomere (white arrowhead) surrounded by some fin rays of the anal fin. 6, Axial mesomeres (white arrowhead) partially covered by sediment in the pelvic fin. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126233" httpUri="https://zenodo.org/record/13126233/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BF236B2F34AFF8613878DA1BE0EFDBD" blockId="5.[151,776,235,579]" pageId="5" pageNumber="6">
Te specimen may represent an adult individual as all the basal plates are fully ossified, which is a feature observed in adult coelacanths (e.g.,
<bibRefCitation id="EFDC4B43F34AFF8612878C66BD4AFEBD" author="Schultze" box="[429,583,299,323]" firstAuthor="Schultze" pageId="5" pageNumber="6" pagination="101 - 108" refId="ref14584" refString="Schultze, H. - P. (1980). Eier legende und lebendgebarende Quastenflosser. Natur und Museum, 110, 101 - 108." type="journal article" year="1980">Schultze, 1980</bibRefCitation>
;
<bibRefCitation id="EFDC4B43F34AFF86117F8C66BFC7FE9D" author="Witzmann" etAl="et al." firstAuthor="Witzmann" pageId="5" pageNumber="6" pagination="309 - 316" refId="ref15023" refString="Witzmann, F., Dorka, M., &amp; Korn, D. (2010). A juvenile Early Carboniferous (Visean) coelacanth from Rosenbeck (Rhenish Mountains, Germany) with derived postcranial characters. Fossil Record, 13, 309 - 316. https: // doi. org / 10.1002 / mmng. 201000004" type="journal article" year="2010">Witzmann et al., 2010</bibRefCitation>
). Tis view is reinforced because the specimen shows some well ossified axial mesomere and the scapulocoracoid (
<figureCitation id="13762A37F34AFF8613B48CC6BFFDFE5D" box="[158,240,395,419]" captionStart="Fig" captionStartId="7.[168,199,1783,1803]" captionTargetBox="[152,1438,237,1949]" captionTargetId="figure-14@8.[198,1392,253,1911]" captionTargetPageId="8" captionText="Fig. 4 Skeleton of Libys callolepis sp. nov. on the counterpart (holotype, NMBE 5034072).A Photos with osteological details: 1, articular head of the scapulocoracoid. 2, Scales on the flank immediately beneath the first anterior dorsal fin. 3, Scales of the lateral line showing the ornamental pattern with the larger central tubercles (white arrowheads point, showed only on one scale).4, Scales on the ventral flank from the pelvic to the anal fin. 5, Axial mesomere (white arrowhead) surrounded by some fin rays of the anal fin. 6, Axial mesomeres (white arrowhead) partially covered by sediment in the pelvic fin. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126233" httpUri="https://zenodo.org/record/13126233/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
<figureCitation id="13762A37F34AFF8613DA8CC6BFF4FE5D" box="[240,249,395,419]" captionStart="Fig" captionStartId="1.[168,199,1175,1195]" captionTargetBox="[152,1437,237,1315]" captionTargetId="figure-295@1.[169,1421,253,1162]" captionTargetPageId="1" captionText="Fig. 1 Maps showing the localization of the fossiliferous outcrops. A Paleogeographical map showing the continent configuration during the Toarcian (modified from Scotese,2014, Map 39) with the distribution of (1) Diplurus longicaudatus, (2) Holophagus gulo, (3) Trachymetopon, (4) Undina (?) barroviensis, (5) Libys callolepis sp. nov. (6) Atacamaia solitaria and (7) Indocoelacanthus robustus. B Paleogeographic map of the NW Tethys Ocean (modified from Fantasia et al., 2019), enlargement of B (red square). C Map of Switzerland with the locality of Les Pueys near the Teysachaux summit (Canton of Fribourg, Switzerland)" figureDoi="http://doi.org/10.5281/zenodo.11998922" httpUri="https://zenodo.org/record/11998922/files/figure.png" pageId="5" pageNumber="6">1</figureCitation>
, A
<figureCitation id="13762A37F34AFF8612308CC6BE2BFE5D" box="[282,294,395,419]" captionStart="Fig" captionStartId="9.[168,199,1857,1877]" captionTargetBox="[152,1438,237,1889]" captionTargetId="figure-11@9.[284,1304,253,1844]" captionTargetPageId="9" captionText="Fig. 5 Skull of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073). A Photos and B outline of the skull" figureDoi="http://doi.org/10.5281/zenodo.11998937" httpUri="https://zenodo.org/record/11998937/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
, A
<figureCitation id="13762A37F34AFF86126F8CC6BE5FFE5D" box="[325,338,395,419]" captionStart="Fig" captionStartId="13.[168,199,1136,1156]" captionTargetBox="[152,1437,237,1136]" captionTargetId="figure-273@13.[168,1420,253,1124]" captionTargetPageId="13" captionText="Fig. 6 Comparison of scales between Libys callolepis sp. nov. and some other Latimeriidae. A Libys callolepis sp. nov. (holotype, NMBE 5034072). Scales of the flank below the neural arches between the basal plates of the anterior and posterior dorsal fin. B Libys polypterus (L. superbus) (BSM AS I 801a). Scales of the flank located between the neural arches and the basal plate of the second dorsal fin. C Undina penicillata (BSM 1870 XIV 22). Scales of the middle flank located on the neural arches below the basal plate of the second dorsal fin. D Trachymetopon (Macropoma) substriolatum (holotype, SMC J27415) from the Kimmeridgian of Cottenham, Cambridgeshire. Scales of the middle flank. It is worth noting that this species was first attributed by Huxley (1866) to the genus Macropoma and latter to Coccoderma by Reis (1888). Forey (1998) kept this species within Coccoderma, but remarked that it shows affinities with Holophagus. Recently,Cavin et al. (2021a) attributed this specimen to the genus of Trachymetopon regarding, among other characteristics, the coarse ornamentation on the dermal bones" figureDoi="http://doi.org/10.5281/zenodo.11998940" httpUri="https://zenodo.org/record/11998940/files/figure.png" pageId="5" pageNumber="6">6</figureCitation>
, B). Although a long supplementary lobe of the caudal fin is considered as a juvenile character (e.g., Forey, 1981;
<bibRefCitation id="EFDC4B43F34AFF8612368C86BEB8FE1D" author="Schultze" box="[284,437,459,483]" firstAuthor="Schultze" pageId="5" pageNumber="6" pagination="90 - 91" refId="ref14543" refString="Schultze, H. - P. (1972). Early growth stages in coelacanth fishes. Nature New Biology, 236, 90 - 91. https: // doi. org / 10.1038 / newbio 236090 a 0" type="journal article" year="1972">Schultze, 1972</bibRefCitation>
), the prominent supplementary lobe of the caudal fin (
<figureCitation id="13762A37F34AFF8612A08CA6BECBFDFD" box="[394,454,491,515]" captionStart="Fig" captionStartId="5.[168,199,1822,1842]" captionTargetBox="[151,1437,237,1950]" captionTargetId="figure-14@6.[192,1395,253,1913]" captionTargetPageId="6" captionText="Fig. 3 Skeleton of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073).A Photos with osteological details: 1, denticles on the proximal fin rays of the caudal fin. 2, Postparietal shield with the otic sensory canal opening as a deep groove crossed by pillars (white arrowhead). 3, Posterior parietal and the supraorbitals with their pillars (white arrowhead). 4, Consolidated snout with the anterior opening for the rostral organ (white arrowhead). 5,Teeth on the prearticular. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126231" httpUri="https://zenodo.org/record/13126231/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) observed in the specimen of Teysachaux represents rather a generic character of the specimen.
</paragraph>
<paragraph id="8BF236B2F34AFF8613BD8F20BEDBFD7A" blockId="5.[151,776,621,1027]" box="[151,470,621,644]" pageId="5" pageNumber="6">
<emphasis id="B939EAA0F34AFF8613BD8F20BEDBFD7A" bold="true" box="[151,470,621,644]" pageId="5" pageNumber="6">Dermal bones of the skull roof</emphasis>
</paragraph>
<paragraph id="8BF236B2F34AFF8613BD8FC1BDCDFBFD" blockId="5.[151,776,621,1027]" pageId="5" pageNumber="6">
Te bones of the skull are only preserved on the part (
<figureCitation id="13762A37F34AFF8613B48FE6BFE8FD3D" box="[158,229,683,707]" captionStart="Fig" captionStartId="5.[168,199,1822,1842]" captionTargetBox="[151,1437,237,1950]" captionTargetId="figure-14@6.[192,1395,253,1913]" captionTargetPageId="6" captionText="Fig. 3 Skeleton of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073).A Photos with osteological details: 1, denticles on the proximal fin rays of the caudal fin. 2, Postparietal shield with the otic sensory canal opening as a deep groove crossed by pillars (white arrowhead). 3, Posterior parietal and the supraorbitals with their pillars (white arrowhead). 4, Consolidated snout with the anterior opening for the rostral organ (white arrowhead). 5,Teeth on the prearticular. B Semi-interpretative line drawing of the specimen" figureDoi="http://doi.org/10.5281/zenodo.13126231" httpUri="https://zenodo.org/record/13126231/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
and
<figureCitation id="13762A37F34AFF8612348FE6BE26FD3D" box="[286,299,683,707]" captionStart="Fig" captionStartId="9.[168,199,1857,1877]" captionTargetBox="[152,1438,237,1889]" captionTargetId="figure-11@9.[284,1304,253,1844]" captionTargetPageId="9" captionText="Fig. 5 Skull of Libys callolepis sp. nov. on the part (holotype, NMBE 5034073). A Photos and B outline of the skull" figureDoi="http://doi.org/10.5281/zenodo.11998937" httpUri="https://zenodo.org/record/11998937/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
). Te skull roof is divided into a parietonasal and postparietal shields, free from one to the other and separated by the intracranial joint, which appears to be straight. Te postparietal shield appears to be smaller than the parietonasal shield, this last being about 1.45 longer. Although slightly lower, this ratio is however close to the ratios of Jurassic and Cretaceous coelacanths that have typically a parietonasal shield circa 1.5 to 2 times longer than the postparietal shield (e.g.,
<taxonomicName id="4C4D4D31F34AFF8612EF8EE6BD7CFC3D" authorityName="Munster" authorityYear="1842" box="[453,625,939,963]" class="Coelacanthi" family="Mawsoniidae" genus="Libys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coelacanthiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="polypterus">
<emphasis id="B939EAA0F34AFF8612EF8EE6BD7CFC3D" box="[453,625,939,963]" italics="true" pageId="5" pageNumber="6">Libys polypterus</emphasis>
</taxonomicName>
has a ratio of about 1.7). On the Teysachaux specimen, the dermal bones of the skull appear to be smooth and unornamented.
</paragraph>
</subSubSection>
</treatment>
</document>