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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3857.2.8" ID-GBIF-Dataset="09b8963f-3aa7-4914-9242-09d5968cfe35" ID-ISSN="1175-5326" ID-Zenodo-Dep="229717" ID-ZooBank="6D902CD0-7A28-485F-A91E-E23EE74C8907" checkinTime="1460899676357" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel Ángel" docDate="2014" docId="03FA87D00000FF9CFF7349550ADFF883" docLanguage="en" docName="zt03857p300.pdf" docOrigin="Zootaxa 3857 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.10:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="10" docTitle="Macrodactylus Dejean" docType="key" docVersion="8" lastPageNumber="299" masterDocId="FFC3FFA80007FF9BFFE44A110F43FFAE" masterDocTitle="New species and records of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Bolivia" masterLastPageNumber="300" masterPageNumber="292" pageNumber="299" updateTime="1665413707506" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species and records of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Bolivia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arce-Pérez, Roberto</mods:namePart>
<mods:affiliation>Red Sistemática y Biodiversidad, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, Mexico.</mods:affiliation>
<mods:nameIdentifier type="email">roberto.arce@inecol.mx</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Morón, Miguel Ángel</mods:namePart>
<mods:affiliation>Red Sistemática y Biodiversidad, Instituto de Ecología, A. C. Apdo. Postal 63, Xalapa, Veracruz 91000, Mexico.</mods:affiliation>
<mods:nameIdentifier type="email">miguel.moron@inecol.mx</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-08-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3857</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>292</mods:start>
<mods:end>300</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3857.2.8</mods:identifier>
<mods:identifier type="GBIF-Dataset">09b8963f-3aa7-4914-9242-09d5968cfe35</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">229717</mods:identifier>
<mods:identifier type="ZooBank">6D902CD0-7A28-485F-A91E-E23EE74C8907</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6123041" ID-GBIF-Taxon="119587634" ID-Zenodo-Dep="6123041" LSID="urn:lsid:plazi:treatment:03FA87D00000FF9CFF7349550ADFF883" httpUri="http://treatment.plazi.org/id/03FA87D00000FF9CFF7349550ADFF883" lastPageNumber="299" pageId="7" pageNumber="299">
<subSubSection box="[151,755,836,862]" pageId="7" pageNumber="299" type="nomenclature">
<paragraph blockId="7.[151,755,836,862]" box="[151,755,836,862]" pageId="7" pageNumber="299">
<heading bold="true" box="[151,755,836,862]" fontSize="11" level="1" pageId="7" pageNumber="299" reason="1">
<emphasis bold="true" box="[151,755,836,862]" pageId="7" pageNumber="299">
Key to the species of
<taxonomicName authorityName="Dejean" box="[412,591,836,862]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[412,591,836,862]" italics="true" pageId="7" pageNumber="299">Macrodactylus</emphasis>
</taxonomicName>
from
<collectingCountry box="[667,755,836,862]" name="Bolivia" pageId="7" pageNumber="299">Bolivia</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="299" type="key">
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" box="[151,1435,905,927]" pageId="7" pageNumber="299">
<keyLead box="[151,1435,905,927]" pageId="7" pageNumber="299">
<number box="[151,163,905,927]" pageId="7" pageNumber="299" value="1.0">1</number>
Body length greater than
<quantity box="[468,527,905,927]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="299" unit="mm" value="9.0">9 mm</quantity>
..........................................................................
<number box="[1423,1435,905,927]" pageId="7" pageNumber="299" value="2.0">2</number>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" box="[151,1435,935,957]" pageId="7" pageNumber="299">
<keyLead box="[151,1435,935,957]" pageId="7" pageNumber="299">
- Body length less than
<quantity box="[438,493,935,957]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="299" unit="mm" value="9.0">9 mm</quantity>
.............................................................................
<number box="[1423,1435,935,956]" pageId="7" pageNumber="299" value="7.0">7</number>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
<number box="[151,163,964,986]" pageId="7" pageNumber="299" value="2.0">2</number>
Antennal club black, length equal to length of antennal stalk. Vestiture does not completely obscure dorsal surface. Length
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="299" unit="mm" value="14.0">14 mm</quantity>
(
<figureCitation box="[275,367,993,1015]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1620,1643]" captionTargetBox="[337,1249,407,1598]" captionTargetId="figure@1.[337,1250,407,1598]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Macrodactylus megaphyllus: 1) dorsal view, 2) lateral view. Macrodactylus yunganus: 3) dorsal view, 4) lateral view. Macrodactylus nobilis: 5) dorsal view, 6) lateral view." httpUri="https://zenodo.org/record/229718/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[329,367,993,1015]" pageId="7" pageNumber="299" value="1.5" valueMax="2.0" valueMin="1.0">1–2</number>
</figureCitation>
), male genitalia (
<figureCitation box="[533,625,993,1015]" captionStart="FIGURES 7 – 17" captionStartId="2.[151,264,1584,1607]" captionTargetBox="[372,1211,631,1558]" captionTargetId="figure@2.[366,1220,625,1563]" captionTargetPageId="2" captionText="FIGURES 7 – 17. Diagnostic structures of Macrodactylus megaphyllus: 7) parameres, distal view, 8) parameres, lateral view, 9) basal piece, dorsal view. Macrodactylus yunganus: 10) parameres, distal view, 11) parameres, lateral view, 12) basal piece, dorsal view, 13) female genital plates, ventral view. Macrodactylus nobilis: 14) parameres, distal view, 15) parameres, lateral view, 16) basal piece, dorsal view, 17) female genital plates, ventral view. Scales in mm." httpUri="https://zenodo.org/record/229719/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[587,625,993,1015]" pageId="7" pageNumber="299" value="8.0" valueMax="9.0" valueMin="7.0">7–9</number>
</figureCitation>
)................................... ..
<taxonomicName authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" box="[1073,1435,993,1015]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="megaphyllus">
<emphasis box="[1073,1226,993,1015]" italics="true" pageId="7" pageNumber="299">M. megaphyllus</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
- Antennal club yellow or black, length ¾ length of antennal stalk. Dorsally glabrous or with vestiture that does not completely obscure dorsal surface..................................................................................
<number box="[1423,1435,1052,1074]" pageId="7" pageNumber="299" value="3.0">3</number>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
<number box="[151,163,1081,1103]" pageId="7" pageNumber="299" value="3.0">3</number>
Dorsally aeneus-green, glabrous. Elytra striated, greenish yellow. Clypeus, antennae, legs, and pygidium yellowish red. Length
<quantity box="[225,292,1111,1133]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="299" unit="mm" value="12.0">12 mm</quantity>
.............................................................................
<taxonomicName authority="Blanchard" authorityName="Blanchard" box="[1224,1436,1111,1133]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis box="[1224,1331,1112,1133]" italics="true" pageId="7" pageNumber="299">M. cupreus</emphasis>
Blanchard
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" box="[151,1435,1140,1162]" pageId="7" pageNumber="299">
<keyLead box="[151,1435,1140,1162]" pageId="7" pageNumber="299">
- Dorsally yellowish red or intense green, glabrous or vestiture in wide rows........................................
<number box="[1423,1435,1140,1162]" pageId="7" pageNumber="299" value="4.0">4</number>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" box="[151,1435,1169,1191]" pageId="7" pageNumber="299">
<keyLead box="[151,1435,1169,1191]" pageId="7" pageNumber="299">
<specimenCount box="[151,284,1169,1191]" pageId="7" pageNumber="299" type="male">
<number box="[151,163,1169,1191]" pageId="7" pageNumber="299" value="4.0">4</number>
Males
</specimenCount>
dorsally glabrous or only with vestiture on scutellum....................................................
<number box="[1423,1435,1169,1191]" pageId="7" pageNumber="299" value="5.0">5</number>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" box="[151,1435,1199,1221]" pageId="7" pageNumber="299">
<keyLead box="[151,1435,1199,1221]" pageId="7" pageNumber="299">
- Males dorsally with vestiture in wide rows.................................................................
<number box="[1423,1435,1199,1221]" pageId="7" pageNumber="299" value="6.0">6</number>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
<specimenCount box="[151,284,1228,1250]" pageId="7" pageNumber="299" type="male">
<number box="[151,163,1228,1250]" pageId="7" pageNumber="299" value="5.0">5</number>
Males
</specimenCount>
with pronotum intense red; with sparse, yellow setae on lateral margins; scutellum with yellowish-gray setae. Elytra intense dark green, with yellowish-green and red shine; with long and short setae intermixed. Female with elytra reddish yellow; with setae on disc; pygidium prominent, red. Length
<quantity box="[782,884,1287,1309]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="7" pageNumber="299" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
..............................
<taxonomicName authority="Moser" authorityName="Moser" box="[1246,1435,1287,1309]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="disparilis">
<emphasis box="[1246,1369,1287,1309]" italics="true" pageId="7" pageNumber="299">M. disparilis</emphasis>
Moser
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
- Males dorsally glabrous; pronotum black; elytra yellowish red, with inner suture dark, dark shadows on disc (
<figureCitation box="[1333,1424,1316,1338]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1620,1643]" captionTargetBox="[337,1249,407,1598]" captionTargetId="figure@1.[337,1250,407,1598]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Macrodactylus megaphyllus: 1) dorsal view, 2) lateral view. Macrodactylus yunganus: 3) dorsal view, 4) lateral view. Macrodactylus nobilis: 5) dorsal view, 6) lateral view." httpUri="https://zenodo.org/record/229718/files/figure.png" pageId="7" pageNumber="299">
Figs
<number box="[1385,1424,1316,1338]" pageId="7" pageNumber="299" value="3.5" valueMax="4.0" valueMin="3.0">3–4</number>
</figureCitation>
). Female with elytra darkened, basal third yellowish red; pygidium subtriangular, short, weakly convex. Length
<quantity box="[1289,1378,1345,1367]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="299" unit="mm" value="12.5">12.5 mm</quantity>
. Male and female genitalia (
<figureCitation box="[432,547,1375,1397]" captionStart="FIGURES 7 – 17" captionStartId="2.[151,264,1584,1607]" captionTargetBox="[372,1211,631,1558]" captionTargetId="figure@2.[366,1220,625,1563]" captionTargetPageId="2" captionText="FIGURES 7 – 17. Diagnostic structures of Macrodactylus megaphyllus: 7) parameres, distal view, 8) parameres, lateral view, 9) basal piece, dorsal view. Macrodactylus yunganus: 10) parameres, distal view, 11) parameres, lateral view, 12) basal piece, dorsal view, 13) female genital plates, ventral view. Macrodactylus nobilis: 14) parameres, distal view, 15) parameres, lateral view, 16) basal piece, dorsal view, 17) female genital plates, ventral view. Scales in mm." httpUri="https://zenodo.org/record/229719/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[486,547,1375,1397]" pageId="7" pageNumber="299" value="11.5" valueMax="13.0" valueMin="10.0">10–13</number>
</figureCitation>
).............................................
<taxonomicName authority="Arce-Perez &amp; Moron" authorityName="Arce-Perez &amp; Moron" box="[1102,1435,1375,1397]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="yunganus">
<emphasis box="[1102,1226,1376,1397]" italics="true" pageId="7" pageNumber="299">M. yunganus</emphasis>
Arce-Pérez &amp; Morón
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
<number box="[151,163,1404,1426]" pageId="7" pageNumber="299" value="6.0">6</number>
Head, pronotum, and inner half of elytra nearly black; with dense, white vestiture. Pronotum with three wide rows of setae, elytra with wide band along inner half. Length
<quantity box="[677,783,1433,1455]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="7" pageNumber="299" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
. Male and female genitalia (
<figureCitation box="[1054,1172,1433,1455]" captionStart="FIGURES 7 – 17" captionStartId="2.[151,264,1584,1607]" captionTargetBox="[372,1211,631,1558]" captionTargetId="figure@2.[366,1220,625,1563]" captionTargetPageId="2" captionText="FIGURES 7 – 17. Diagnostic structures of Macrodactylus megaphyllus: 7) parameres, distal view, 8) parameres, lateral view, 9) basal piece, dorsal view. Macrodactylus yunganus: 10) parameres, distal view, 11) parameres, lateral view, 12) basal piece, dorsal view, 13) female genital plates, ventral view. Macrodactylus nobilis: 14) parameres, distal view, 15) parameres, lateral view, 16) basal piece, dorsal view, 17) female genital plates, ventral view. Scales in mm." httpUri="https://zenodo.org/record/229719/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[1108,1172,1433,1455]" pageId="7" pageNumber="299" value="15.5" valueMax="17.0" valueMin="14.0">14–17</number>
</figureCitation>
).........
<taxonomicName authority="Frey" authorityName="Frey" box="[1288,1435,1433,1455]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="nobilis">
<emphasis box="[1288,1385,1433,1455]" italics="true" pageId="7" pageNumber="299">M. nobilis</emphasis>
Frey
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
- Head and pronotum dark green; elytra yellowish red with green shine; clypeus, legs, and pygidium yellowish red. Males without wide rows of setae on pronotum. Length
<quantity box="[655,746,1492,1514]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="7" pageNumber="299" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
...........................................
<taxonomicName authority="Frey" authorityName="Frey" box="[1276,1435,1492,1514]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="zischkai">
<emphasis box="[1276,1385,1492,1514]" italics="true" pageId="7" pageNumber="299">M. zischkai</emphasis>
Frey
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
<number box="[151,163,1522,1543]" pageId="7" pageNumber="299" value="7.0">7</number>
Dorsal surface completely covered with yellow, short setae that obscure surface (
<figureCitation box="[994,1111,1521,1543]" captionStart="FIGURES 18 – 21" captionStartId="5.[151,264,1878,1901]" captionTargetBox="[345,1229,631,1844]" captionTargetId="figure@5.[343,1244,625,1857]" captionTargetPageId="5" captionText="FIGURES 18 – 21. Macrodactylus bolivianus: 18) dorsal view, 20) lateral view. Macrodactylus gracilis: 19) dorsal view, 21) lateral view." httpUri="https://zenodo.org/record/229720/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[1048,1111,1521,1543]" pageId="7" pageNumber="299" value="20.5" valueMax="21.0" valueMin="20.0">20–21</number>
</figureCitation>
). Male and female genitalia (
<figureCitation captionStart="FIGURES 22 – 29" captionStartId="6.[151,264,975,998]" captionTargetBox="[342,1235,197,950]" captionTargetId="figure@6.[339,1248,193,954]" captionTargetPageId="6" captionText="FIGURES 22 – 29. Diagnostic structures of Macrodactylus bolivianus: 22) parameres, distal view, 23) parameres, lateral view, 24) basal piece, dorsal view, 25) female genital plates, ventral view. Macrodactylus gracilis: 26) parameres, distal view, 27) parameres, lateral view, 28) basal piece, dorsal view, 29) female genital plates, ventral view. Scales in mm." httpUri="https://zenodo.org/record/229721/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[225,286,1551,1573]" pageId="7" pageNumber="299" value="27.5" valueMax="29.0" valueMin="26.0">26–29</number>
</figureCitation>
)................................................................................
<taxonomicName authority="Moser" authorityName="Moser" box="[1263,1435,1551,1573]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis box="[1263,1368,1551,1573]" italics="true" pageId="7" pageNumber="299">M. gracilis</emphasis>
Moser
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" box="[151,1436,1580,1602]" pageId="7" pageNumber="299">
<keyLead box="[151,1436,1580,1602]" pageId="7" pageNumber="299">
- Dorsal surface with sparse setae that exposes surface; setae arranged in wide rows, narrow rows, or dispersed.............
<number box="[1424,1436,1580,1602]" pageId="7" pageNumber="299" value="8.0">8</number>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
<number box="[151,163,1609,1631]" pageId="7" pageNumber="299" value="8.0">8</number>
Pronotum and scutellum dark red, with dense yellow setae that obscure surface. Elytra reddish yellow; with two yellow, wide rows. Pygidium with abundant setae. Length
<quantity box="[658,713,1639,1661]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="299" unit="mm" value="9.0">9 mm</quantity>
........................................
<taxonomicName authority="Moser" authorityName="Moser" box="[1209,1435,1639,1661]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="signatipennis">
<emphasis box="[1209,1369,1640,1661]" italics="true" pageId="7" pageNumber="299">M. signatipennis</emphasis>
Moser
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
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- Pronotum and scutellum reddish yellow or dark red with sparse setae, surface visible. Elytral setae arranged in narrow rows. Pygidium with sparse setae.............................................................................
<number box="[1423,1435,1697,1719]" pageId="7" pageNumber="299" value="9.0">9</number>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="7" pageNumber="299">
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
<keyLead pageId="7" pageNumber="299">
<number box="[151,163,1727,1749]" pageId="7" pageNumber="299" value="9.0">9</number>
Pronotum dark red; elytra yellowish red; dorsally with short, white setae, and surface visible. Length
<quantity box="[1232,1289,1727,1749]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="299" unit="mm" value="8.0">8 mm</quantity>
(
<figureCitation box="[1305,1425,1727,1749]" captionStart="FIGURES 18 – 21" captionStartId="5.[151,264,1878,1901]" captionTargetBox="[345,1229,631,1844]" captionTargetId="figure@5.[343,1244,625,1857]" captionTargetPageId="5" captionText="FIGURES 18 – 21. Macrodactylus bolivianus: 18) dorsal view, 20) lateral view. Macrodactylus gracilis: 19) dorsal view, 21) lateral view." httpUri="https://zenodo.org/record/229720/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[1361,1425,1727,1749]" pageId="7" pageNumber="299" value="18.5" valueMax="19.0" valueMin="18.0">18–19</number>
</figureCitation>
). Male and female genitalia (
<figureCitation box="[487,606,1756,1778]" captionStart="FIGURES 22 – 29" captionStartId="6.[151,264,975,998]" captionTargetBox="[342,1235,197,950]" captionTargetId="figure@6.[339,1248,193,954]" captionTargetPageId="6" captionText="FIGURES 22 – 29. Diagnostic structures of Macrodactylus bolivianus: 22) parameres, distal view, 23) parameres, lateral view, 24) basal piece, dorsal view, 25) female genital plates, ventral view. Macrodactylus gracilis: 26) parameres, distal view, 27) parameres, lateral view, 28) basal piece, dorsal view, 29) female genital plates, ventral view. Scales in mm." httpUri="https://zenodo.org/record/229721/files/figure.png" pageId="7" pageNumber="299">
Figs.
<number box="[542,606,1756,1778]" pageId="7" pageNumber="299" value="23.5" valueMax="25.0" valueMin="22.0">22–25</number>
</figureCitation>
)...................................................
<taxonomicName authority="Moser" authorityName="Moser" box="[1231,1429,1756,1778]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis box="[1231,1363,1756,1778]" italics="true" pageId="7" pageNumber="299">M. bolivianus</emphasis>
Moser
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="7.[151,1436,905,1837]" pageId="7" pageNumber="299">
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- Dorsally bright reddish yellow; pronotum thickly dotted and with white setae towards lateral margins; elytra black with metallic luster, transversely rugose, with few rows of setae. Length
<quantity box="[819,878,1815,1837]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="299" unit="mm" value="9.0">9 mm</quantity>
................................
<taxonomicName authority="Kirsch" authorityName="Kirsch" box="[1267,1436,1815,1837]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="aeneus">
<emphasis box="[1267,1366,1816,1837]" italics="true" pageId="7" pageNumber="299">M. aeneus</emphasis>
Kirsch
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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