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<document id="5116F120154B321582C1BAD31116DDB4" ID-DOI="10.11646/zootaxa.3335.1.1" ID-GBIF-Dataset="ad866625-8c24-4739-b287-e70ac2f3ac00" ID-ISSN="1175-5326" ID-Zenodo-Dep="281297" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460486437628" checkinUser="plazi" docAuthor="Castilho, Raphael C., De, Gilberto J. &amp; Halliday, Bruce" docDate="2012" docId="03CB87F0FF2FFFECFF2A5562FC9A1D5C" docLanguage="en" docName="zt03335p028.pdf" docOrigin="Zootaxa 3335" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pennarhodeus brevipennatus Karg" docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="FFF2FF88FF2BFFEAFFBD530DFFD71824" masterDocTitle="Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="5" updateTime="1698309252394" updateUser="plazi">
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<mods:title id="DE50348276BCD59559EC3D5FD38FEF0D">Revision of the genera Interrhodeus, Pennarhodeus and Poropodalius (Acari: Rhodacaridae)</mods:title>
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<mods:namePart id="54683E0BD803ED76960D327F1E8C5676">Castilho, Raphael C.</mods:namePart>
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<mods:namePart id="AD2068FBD13D6C3095FF6D717ED2B322">De, Gilberto J.</mods:namePart>
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<mods:namePart id="7121B5A7592CC952BAC111626CD000C7">Halliday, Bruce</mods:namePart>
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<mods:title id="6A74A2B18229D15D332F18A5006C499D">Zootaxa</mods:title>
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<mods:part id="6EFDEA151D3004EAB215E787F19EEEC8">
<mods:date id="EAFD1C3076EDA2C75691000042457A7C">2012</mods:date>
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<mods:number id="5988CCC0B3592539000E88B689865C15">3335</mods:number>
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<treatment id="03CB87F0FF2FFFECFF2A5562FC9A1D5C" ID-DOI="http://doi.org/10.5281/zenodo.6171793" ID-GBIF-Taxon="119518468" ID-Zenodo-Dep="6171793" LSID="urn:lsid:plazi:treatment:03CB87F0FF2FFFECFF2A5562FC9A1D5C" httpUri="http://treatment.plazi.org/id/03CB87F0FF2FFFECFF2A5562FC9A1D5C" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="C378656DFF2FFFEEFF2A5562FD961EAD" box="[151,577,1647,1673]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BDD36E6FF2FFFEEFF2A5562FD961EAD" blockId="4.[151,577,1647,1673]" box="[151,577,1647,1673]" pageId="4" pageNumber="5">
<heading id="D095818AFF2FFFEEFF2A5562FD961EAD" bold="true" box="[151,577,1647,1673]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="4C624D65FF2FFFEEFF2A5562FD961EAD" ID-CoL="76JN4" authority="Karg" authorityName="Karg" box="[151,577,1647,1673]" class="Arachnida" family="Rhodacaridae" genus="Pennarhodeus" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brevipennatus">
<emphasis id="B916EAF4FF2FFFEEFF2A5562FD961EAD" bold="true" box="[151,577,1647,1673]" pageId="4" pageNumber="5">
<emphasis id="B916EAF4FF2FFFEEFF2A5562FE2E1EAD" bold="true" box="[151,505,1647,1673]" italics="true" pageId="4" pageNumber="5">Pennarhodeus brevipennatus</emphasis>
Karg
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C378656DFF2FFFEEFF2A55BAFAE01F12" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8BDD36E6FF2FFFEEFF2A55BAFDAC1EE8" blockId="4.[151,635,1718,1741]" box="[151,635,1718,1741]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB7211C8FF2FFFEEFF2A55BAFDAC1EE8" box="[151,635,1718,1741]" pageId="4" pageNumber="5">
<taxonomicName id="4C624D65FF2FFFEEFF2A55BAFDA01EE8" ID-CoL="76JN4" authority="Karg, 2000b: 211" authorityName="Karg" authorityPageNumber="211" authorityYear="2000" box="[151,631,1718,1741]" class="Arachnida" family="Rhodacaridae" genus="Pennarhodeus" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="brevipennatus">
<emphasis id="B916EAF4FF2FFFEEFF2A55BAFE691EE9" box="[151,446,1719,1741]" italics="true" pageId="4" pageNumber="5">Pennarhodeus brevipennatus</emphasis>
<treatmentCitation id="0AC310F7FF2FFFEEFE7955BAFDA01EE8" author="Karg" box="[452,631,1718,1740]" page="211" pageId="4" pageNumber="5" year="2000">
<bibRefCitation id="EFF34B17FF2FFFEEFE7955BAFD911EE8" author="Karg" box="[452,582,1718,1740]" pageId="4" pageNumber="5" pagination="207 - 213" refId="ref22818" refString="Karg, W. (2000 b) Neue Raubmilbenarten der Pionierartengruppe Rhodacaridae Oudemans (Acarina, Parasitiformes). Abhandlungen und Berichte des Naturkundemuseums Gorlitz, 72, 207 - 213." type="journal article" year="2000" yearSuffix="b">Karg, 2000b</bibRefCitation>
: 211
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDD36E6FF2FFFEEFF2A55F4FAE01F12" blockId="4.[151,1437,1785,2026]" pageId="4" pageNumber="5">
<emphasis id="B916EAF4FF2FFFEEFF2A55F4FE4F1F36" bold="true" box="[151,408,1785,1810]" pageId="4" pageNumber="5">Specimens examined.</emphasis>
<typeStatus id="54D98844FF2FFFEEFE1C55F7FDDE1F36" box="[417,521,1786,1810]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
male: La Selva,
<collectingCountry id="F3757676FF2FFFEEFD7355F7FC9C1F36" box="[718,843,1786,1810]" name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
(10°26
<geoCoordinate id="EE565021FF2FFFEEFC1A55F7FBC61F36" box="[935,1041,1785,1811]" direction="north" orientation="latitude" pageId="4" pageNumber="5" precision="925" value="84.01667">N 84°01</geoCoordinate>
W), elevation
<quantity id="4C9A9B03FF2FFFEEFB0A55F7FAFB1F36" box="[1207,1324,1786,1810]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="4" pageNumber="5" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50150 m</quantity>
, Primario (indicated as soil sample in the original description),
<date id="FFDC1026FF2FFFEEFD575413FCA11F12" box="[746,886,1822,1846]" pageId="4" pageNumber="5" value="1993-08-03">3-VIII-1993</date>
, Coll. B., SMNG 93/38525 (in SMNG).
</paragraph>
</subSubSection>
<subSubSection id="C378656DFF2FFFECFF7A544FFB011C10" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BDD36E6FF2FFFEFFF7A544FFE7118D3" blockId="4.[151,1437,1785,2026]" lastBlockId="5.[151,1437,151,644]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="B916EAF4FF2FFFEEFF7A544FFE0E1F7F" bold="true" box="[199,473,1858,1883]" pageId="4" pageNumber="5">Diagnosis (adult male).</emphasis>
Arthrodial process of chelicera with a short coronet-like fringe; epistome with anteromedial projection about 1.2 times as long as antero-lateral projections, and with margin serrate laterad to antero-lateral projections; corniculi rather straight; dorsal idiosomal setae pilose, except
<emphasis id="B916EAF4FF2FFFEEFBB75486FBF41F87" box="[1034,1059,1931,1955]" italics="true" pageId="4" pageNumber="5">r3</emphasis>
,
<emphasis id="B916EAF4FF2FFFEEFB8C5486FB981F86" box="[1073,1103,1931,1954]" italics="true" pageId="4" pageNumber="5">Z5</emphasis>
and
<emphasis id="B916EAF4FF2FFFEEFB385486FB761F86" box="[1157,1185,1931,1954]" italics="true" pageId="4" pageNumber="5">S5</emphasis>
, feather-like; podonotal and opisthonotal shields imbricate; with two crescent-shaped scleronoduli between setae
<emphasis id="B916EAF4FF2FFFEEFB0254A2FB021FE2" box="[1215,1237,1967,1990]" italics="true" pageId="4" pageNumber="5">j5</emphasis>
and
<emphasis id="B916EAF4FF2FFFEEFAAC54A3FAF01FE2" box="[1297,1319,1966,1990]" italics="true" pageId="4" pageNumber="5">j6</emphasis>
; posterior margin of opisthonotal shield slightly concave between setae
<emphasis id="B916EAF4FF2FFFEEFCFB54DEFCB31FCE" box="[838,868,2003,2026]" italics="true" pageId="4" pageNumber="5">Z5</emphasis>
; seta
<emphasis id="B916EAF4FF2FFFEEFC1854DEFC121FCE" box="[933,965,2003,2026]" italics="true" pageId="4" pageNumber="5">R5</emphasis>
absent; sternogenital shield lightly reticulate; unsclerotised integument between sternogenital and ventrianal shields with a subtriangular accessory platelet; peritreme extending anteriorly to level of posterior margin of coxa II; with two spur-like ventral setae on each of femur, genu and tibia II.
</paragraph>
<paragraph id="8BDD36E6FF2EFFEFFF7A520EFDB21940" blockId="5.[151,1437,151,644]" pageId="5" pageNumber="6">
<emphasis id="B916EAF4FF2EFFEFFF7A520EFE991938" bold="true" box="[199,334,259,284]" pageId="5" pageNumber="6">Adult male</emphasis>
(
<figureCitation id="13592A63FF2EFFEFFEE35209FE751938" box="[350,418,260,284]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
AG) (
<typeStatus id="54D98844FF2EFFEFFE415209FDB31938" box="[508,612,260,284]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
). Dorsal idiosomal setae (except
<emphasis id="B916EAF4FF2EFFEFFC635209FC201938" box="[990,1015,260,284]" italics="true" pageId="5" pageNumber="6">r3</emphasis>
,
<emphasis id="B916EAF4FF2EFFEFFBBA5208FBF21938" box="[1031,1061,261,284]" italics="true" pageId="5" pageNumber="6">Z5</emphasis>
and
<emphasis id="B916EAF4FF2EFFEFFBE35208FBAD1938" box="[1118,1146,261,284]" italics="true" pageId="5" pageNumber="6">S5</emphasis>
), para-anal and post-anal setae pilose; setae
<emphasis id="B916EAF4FF2EFFEFFED15225FE521964" box="[364,389,296,320]" italics="true" pageId="5" pageNumber="6">r3</emphasis>
,
<emphasis id="B916EAF4FF2EFFEFFE295225FE65191B" box="[404,434,296,319]" italics="true" pageId="5" pageNumber="6">Z5</emphasis>
,
<emphasis id="B916EAF4FF2EFFEFFE7F5225FE09191B" box="[450,478,296,319]" italics="true" pageId="5" pageNumber="6">S5</emphasis>
and
<emphasis id="B916EAF4FF2EFFEFFDA55225FDE8191B" box="[536,575,296,319]" italics="true" pageId="5" pageNumber="6">Jv5</emphasis>
feather-like; hypostomal, subcapitular, ventral idiosomal (except
<emphasis id="B916EAF4FF2EFFEFFA915225FA86191B" box="[1324,1361,296,319]" italics="true" pageId="5" pageNumber="6">Jv5</emphasis>
, paraanal and post-anal) and leg setae smooth.
</paragraph>
<paragraph id="8BDD36E6FF2EFFEFFF7A5262FAB71AA0" blockId="5.[151,1437,151,644]" pageId="5" pageNumber="6">
<emphasis id="B916EAF4FF2EFFEFFF7A5262FE8819AC" bold="true" box="[199,351,367,392]" pageId="5" pageNumber="6">Gnathosoma</emphasis>
. Fixed cheliceral digit 36 long; movable digit 37 long; teeth and pilus dentilis not visible because of position of chelicera. Spermatodactyl 77 long, abruptly bent ventrally at an acute angle, distal portion narrower than proximal portion (
<figureCitation id="13592A63FF2EFFEFFE2352BAFE3619EB" box="[414,481,439,463]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
A). Arthrodial process of chelicera with a short coronet-like fringe. Epistome with a smooth antero-medial projection slightly wider at the base and provided with three apical denticles, flanked on each side by shorter, smooth projection with four or five apical denticles (
<figureCitation id="13592A63FF2EFFEFFC7352F2FBC61A33" box="[974,1041,511,535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
B); margin of epistome with two or three large teeth between antero-medial and antero-lateral projections and serrate laterad to antero-lateral projections. Deutosternal denticles in eight rows, each with seven to 13 denticles; anterior row “V shaped, other rows straight and transverse (
<figureCitation id="13592A63FF2EFFEFFE1F5161FE331AA0" box="[418,484,620,644]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
C). Corniculi rather straight. Measurements of setae:
<emphasis id="B916EAF4FF2EFFEFFB845161FB901AA0" box="[1081,1095,620,644]" italics="true" pageId="5" pageNumber="6">h</emphasis>
1 14,
<emphasis id="B916EAF4FF2EFFEFFB3B5161FB431AA0" box="[1158,1172,620,644]" italics="true" pageId="5" pageNumber="6">h</emphasis>
2 10,
<emphasis id="B916EAF4FF2EFFEFFB6E5161FB361AA0" box="[1235,1249,620,644]" italics="true" pageId="5" pageNumber="6">h</emphasis>
3 11,
<emphasis id="B916EAF4FF2EFFEFFAA25160FAE11AA0" box="[1311,1334,621,644]" italics="true" pageId="5" pageNumber="6">sc</emphasis>
12.
</paragraph>
<caption id="DF1D666EFF2EFFEFFF2A545DFDE51F80" httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="5" pageNumber="6" targetBox="[394,1161,705,1809]" targetPageId="5">
<paragraph id="8BDD36E6FF2EFFEFFF2A545DFDE51F80" blockId="5.[151,1436,1872,1956]" pageId="5" pageNumber="6">
<emphasis id="B916EAF4FF2EFFEFFF2A545DFEDA1F42" bold="true" box="[151,269,1872,1895]" pageId="5" pageNumber="6">FIGURE 3</emphasis>
.
<taxonomicName id="4C624D65FF2EFFEFFEA6545CFD541F43" authority="Karg." authorityName="Karg." box="[283,643,1873,1895]" class="Arachnida" family="Rhodacaridae" genus="Pennarhodeus" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brevipennatus">
<emphasis id="B916EAF4FF2EFFEFFEA6545CFD951F43" box="[283,578,1873,1895]" italics="true" pageId="5" pageNumber="6">Pennarhodeus brevipennatus</emphasis>
Karg.
</taxonomicName>
Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility.
</paragraph>
</caption>
<paragraph id="8BDD36E6FF2DFFECFF7A5395FE2819D0" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="7">
<emphasis id="B916EAF4FF2DFFECFF7A5395FE501894" bold="true" box="[199,391,151,176]" pageId="6" pageNumber="7">Dorsal idiosoma</emphasis>
(
<figureCitation id="13592A63FF2DFFECFE2B539AFE0F188B" box="[406,472,151,175]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
D). Podonotal shield imbricate, except for a punctate band along posterior margin; 160 long and 173 wide at widest point; with 22 pairs of setae (seta
<emphasis id="B916EAF4FF2DFFECFCDA53B1FCA818F0" box="[871,895,188,212]" italics="true" pageId="6" pageNumber="7">r1</emphasis>
absent); with two crescent-shaped scleronoduli between setae
<emphasis id="B916EAF4FF2DFFECFE8453EDFE9818D3" box="[313,335,224,247]" italics="true" pageId="6" pageNumber="7">j5</emphasis>
and
<emphasis id="B916EAF4FF2DFFECFE3953EDFE4D18DC" box="[388,410,224,248]" italics="true" pageId="6" pageNumber="7">j6</emphasis>
and four pairs of distinguishable lyrifissures. Unsclerotised integument laterad to podonotal shield with a pair of setae (
<emphasis id="B916EAF4FF2DFFECFE6E5209FE3C1938" box="[467,491,260,284]" italics="true" pageId="6" pageNumber="7">r4</emphasis>
). Opisthonotal shield imbricate, except for a punctate band along anterior margin; posterior margin slightly concave between setae
<emphasis id="B916EAF4FF2DFFECFD015225FD0D191B" box="[700,730,296,319]" italics="true" pageId="6" pageNumber="7">Z5</emphasis>
; 125 long and 141 wide at widest point; with 18 pairs of setae and three pairs of distinguishable lyrifissures. Seta
<emphasis id="B916EAF4FF2DFFECFD6E5241FD241940" box="[723,755,332,356]" italics="true" pageId="6" pageNumber="7">R1</emphasis>
on ventrianal shield; seta
<emphasis id="B916EAF4FF2DFFECFBA55240FBE01940" box="[1048,1079,333,356]" italics="true" pageId="6" pageNumber="7">R5</emphasis>
absent. Measurements of setae:
<emphasis id="B916EAF4FF2DFFECFF2A527DFF7A19AC" box="[151,173,368,392]" italics="true" pageId="6" pageNumber="7">j1</emphasis>
9,
<emphasis id="B916EAF4FF2DFFECFF6C527DFF3019AC" box="[209,231,368,392]" italics="true" pageId="6" pageNumber="7">j2</emphasis>
8,
<emphasis id="B916EAF4FF2DFFECFEB1527DFEF619AC" box="[268,289,368,392]" italics="true" pageId="6" pageNumber="7">j3</emphasis>
8,
<emphasis id="B916EAF4FF2DFFECFEFB527DFE8B19AC" box="[326,348,368,392]" italics="true" pageId="6" pageNumber="7">j4</emphasis>
10,
<emphasis id="B916EAF4FF2DFFECFE32527DFE7219A3" box="[399,421,368,391]" italics="true" pageId="6" pageNumber="7">j5</emphasis>
10,
<emphasis id="B916EAF4FF2DFFECFE6A527DFE3A19AC" box="[471,493,368,392]" italics="true" pageId="6" pageNumber="7">j6</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFDA2527DFDE019AC" box="[543,567,368,392]" italics="true" pageId="6" pageNumber="7">z1</emphasis>
8,
<emphasis id="B916EAF4FF2DFFECFDE1527DFDA219AC" box="[604,629,368,392]" italics="true" pageId="6" pageNumber="7">z2</emphasis>
not seen (broken),
<emphasis id="B916EAF4FF2DFFECFCEF527DFCBD19AC" box="[850,874,368,392]" italics="true" pageId="6" pageNumber="7">z3</emphasis>
10,
<emphasis id="B916EAF4FF2DFFECFC20527DFC6119AC" box="[925,950,368,392]" italics="true" pageId="6" pageNumber="7">z4</emphasis>
10,
<emphasis id="B916EAF4FF2DFFECFC54527DFBD519A3" box="[1001,1026,368,391]" italics="true" pageId="6" pageNumber="7">z5</emphasis>
not seen (broken),
<emphasis id="B916EAF4FF2DFFECFB63527DFB2019AC" box="[1246,1271,368,392]" italics="true" pageId="6" pageNumber="7">z6</emphasis>
not seen (broken),
<geoCoordinate id="EE565021FF2DFFECFF6A5299FEC41988" box="[215,275,404,428]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-1.1833334">
<emphasis id="B916EAF4FF2DFFECFF6A5299FF271988" box="[215,240,404,428]" italics="true" pageId="6" pageNumber="7">s1</emphasis>
11
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFE9D5299FE871988" box="[288,336,404,428]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-2.15">
<emphasis id="B916EAF4FF2DFFECFE9D5299FEEE1988" box="[288,313,404,428]" italics="true" pageId="6" pageNumber="7">s2</emphasis>
9
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFEE15299FE5B1988" box="[348,396,404,428]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-3.15">
<emphasis id="B916EAF4FF2DFFECFEE15299FEA21988" box="[348,373,404,428]" italics="true" pageId="6" pageNumber="7">s3</emphasis>
9
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFE255299FE021988" box="[408,469,404,428]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-4.1833334">
<emphasis id="B916EAF4FF2DFFECFE255299FE671988" box="[408,432,404,428]" italics="true" pageId="6" pageNumber="7">s4</emphasis>
11
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFE5C5298FDC81988" box="[481,543,404,428]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-5.1666665">
<emphasis id="B916EAF4FF2DFFECFE5C5298FE2E1988" box="[481,505,405,428]" italics="true" pageId="6" pageNumber="7">s5</emphasis>
10
</geoCoordinate>
,
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<emphasis id="B916EAF4FF2DFFECFD965299FD931988" box="[555,580,404,428]" italics="true" pageId="6" pageNumber="7">s6</emphasis>
10
</geoCoordinate>
,
<emphasis id="B916EAF4FF2DFFECFDC85299FD591988" box="[629,654,404,428]" italics="true" pageId="6" pageNumber="7">r2</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFD035299FD001988" box="[702,727,404,428]" italics="true" pageId="6" pageNumber="7">r3</emphasis>
15,
<emphasis id="B916EAF4FF2DFFECFCB55299FCF61988" box="[776,801,404,428]" italics="true" pageId="6" pageNumber="7">r4</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFCEC5298FCBD1988" box="[849,874,405,428]" italics="true" pageId="6" pageNumber="7">r5</emphasis>
not seen (broken),
<emphasis id="B916EAF4FF2DFFECFBF95299FB8B1988" box="[1092,1116,404,428]" italics="true" pageId="6" pageNumber="7">r6</emphasis>
14,
<emphasis id="B916EAF4FF2DFFECFB335299FB7F1988" box="[1166,1192,404,428]" italics="true" pageId="6" pageNumber="7">J1</emphasis>
not seen (broken),
<emphasis id="B916EAF4FF2DFFECFA3C5299FA4C1988" box="[1409,1435,404,428]" italics="true" pageId="6" pageNumber="7">J2</emphasis>
not seen (broken),
<emphasis id="B916EAF4FF2DFFECFEDA52B5FE5619F4" box="[359,385,440,464]" italics="true" pageId="6" pageNumber="7">J3</emphasis>
not seen (broken),
<emphasis id="B916EAF4FF2DFFECFDE452B5FDA319F4" box="[601,628,440,464]" italics="true" pageId="6" pageNumber="7">J4</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFD1E52B5FD6A19EB" box="[675,701,440,463]" italics="true" pageId="6" pageNumber="7">J5</emphasis>
12,
<emphasis id="B916EAF4FF2DFFECFD5052B5FCDC19F4" box="[749,779,440,464]" italics="true" pageId="6" pageNumber="7">Z1</emphasis>
12,
<emphasis id="B916EAF4FF2DFFECFC8152B5FC8E19F4" box="[828,857,440,464]" italics="true" pageId="6" pageNumber="7">Z2</emphasis>
13,
<emphasis id="B916EAF4FF2DFFECFC3452B5FC7019F4" box="[905,935,440,464]" italics="true" pageId="6" pageNumber="7">Z3</emphasis>
13,
<emphasis id="B916EAF4FF2DFFECFC6A52B5FC2219F4" box="[983,1013,440,464]" italics="true" pageId="6" pageNumber="7">Z4</emphasis>
13,
<emphasis id="B916EAF4FF2DFFECFB9852B5FB9419EB" box="[1061,1091,440,463]" italics="true" pageId="6" pageNumber="7">Z5</emphasis>
14,
<geoCoordinate id="EE565021FF2DFFECFBCE52B5FB6419EB" box="[1139,1203,439,464]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-1.2">
<emphasis id="B916EAF4FF2DFFECFBCE52B5FB5819F4" box="[1139,1167,440,464]" italics="true" pageId="6" pageNumber="7">S1</emphasis>
12
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFB0252B5FB2819EB" box="[1215,1279,439,464]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-2.1833334">
<emphasis id="B916EAF4FF2DFFECFB0252B5FB0C19F4" box="[1215,1243,440,464]" italics="true" pageId="6" pageNumber="7">S2</emphasis>
11
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFAB752B5FA9C19EB" box="[1290,1355,439,464]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-3.2">
<emphasis id="B916EAF4FF2DFFECFAB752B5FAF119F4" box="[1290,1318,440,464]" italics="true" pageId="6" pageNumber="7">S3</emphasis>
12
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFAEB52B5FA4019EB" box="[1366,1431,439,464]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-4.2">
<emphasis id="B916EAF4FF2DFFECFAEB52B5FAA519F4" box="[1366,1394,440,464]" italics="true" pageId="6" pageNumber="7">S4</emphasis>
12
</geoCoordinate>
,
<geoCoordinate id="EE565021FF2DFFECFF2A52D0FF0F19D0" box="[151,216,476,500]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-5.2166667">
<emphasis id="B916EAF4FF2DFFECFF2A52D0FF6419D0" box="[151,179,477,500]" italics="true" pageId="6" pageNumber="7">S5</emphasis>
13
</geoCoordinate>
,
<emphasis id="B916EAF4FF2DFFECFF5952D1FED419D0" box="[228,259,476,500]" italics="true" pageId="6" pageNumber="7">R1</emphasis>
8,
<emphasis id="B916EAF4FF2DFFECFE9B52D1FE9219D0" box="[294,325,476,500]" italics="true" pageId="6" pageNumber="7">R2</emphasis>
9,
<emphasis id="B916EAF4FF2DFFECFED552D1FE5019D0" box="[360,391,476,500]" italics="true" pageId="6" pageNumber="7">R3</emphasis>
10,
<emphasis id="B916EAF4FF2DFFECFE0552D1FE0F19D0" box="[440,472,476,500]" italics="true" pageId="6" pageNumber="7">R4</emphasis>
11.
</paragraph>
<paragraph id="8BDD36E6FF2DFFECFF7A510DFB481BEC" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="7">
<emphasis id="B916EAF4FF2DFFECFF7A510DFE441A3C" bold="true" box="[199,403,511,536]" pageId="6" pageNumber="7">Ventral idiosoma</emphasis>
(
<figureCitation id="13592A63FF2DFFECFE1E52F2FE311A33" box="[419,486,511,535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
E). Base of tritosternum 15 long and 12 wide proximally (
<figureCitation id="13592A63FF2DFFECFBCB52F2FB6E1A33" box="[1142,1209,511,535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
F); laciniae 46 long, separated for about 90% of their total length, pilose. Sternogenital shield lightly reticulate, with indistinct anterior margin and punctate band along straight posterior margin; region anterior to the first pair of lyrifissures (
<date id="FFDC1026FF2DFFECFA9C5144FA961A7B" box="[1313,1345,583,608]" pageId="6" pageNumber="7">
<emphasis id="B916EAF4FF2DFFECFA9C5144FAE21A44" box="[1313,1333,585,608]" italics="true" pageId="6" pageNumber="7">iv</emphasis>
1
</date>
) lightly sclerotised and punctate; approximately 132 long, including the lightly sclerotised and punctate region, and 65 wide at widest point; with five pairs of setae and three pairs of lyrifissures; distance between
<emphasis id="B916EAF4FF2DFFECFB72519CFB351A8C" box="[1231,1250,657,680]" italics="true" pageId="6" pageNumber="7">st</emphasis>
5-
<emphasis id="B916EAF4FF2DFFECFB44519CFADB1A8C" box="[1273,1292,657,680]" italics="true" pageId="6" pageNumber="7">st</emphasis>
5 28; genital opening on anterior margin of shield. Elongate remnant of endopodal shield adjacent to coxa IV. Unsclerotised integument between sternogenital and ventrianal shields with a subtriangular accessory platelet and punctate near anterior margin of ventrianal shield. Ventrianal shield imbricate; 95 long and 170 wide at widest point and extending to dorso-lateral region of idiosoma, but not fused to dorsal shield; with nine pairs of setae (
<emphasis id="B916EAF4FF2DFFECFB0D502DFB181B1C" box="[1200,1231,800,824]" italics="true" pageId="6" pageNumber="7">R1</emphasis>
,
<emphasis id="B916EAF4FF2DFFECFB61502DFAE31B13" box="[1244,1332,800,824]" italics="true" pageId="6" pageNumber="7">Jv1-Jv5</emphasis>
and
<emphasis id="B916EAF4FF2DFFECFAD4502DFF161B78" italics="true" pageId="6" pageNumber="7">Zv1- Zv3</emphasis>
) in addition to circum-anal setae, a distinguishable pore and two distinguishable lyrifissures; post-anal seta about as long as para-anal setae. Peritreme extending anteriorly to level of posterior margin of coxa II. Peritrematal shield extending posteriorly beyond mid-level of coxa IV. Measurements of setae:
<emphasis id="B916EAF4FF2DFFECFB885081FB811B80" box="[1077,1110,908,932]" italics="true" pageId="6" pageNumber="7">st1</emphasis>
13,
<emphasis id="B916EAF4FF2DFFECFB3A5081FB701B80" box="[1159,1191,908,932]" italics="true" pageId="6" pageNumber="7">st2</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFB655081FB2E1B80" box="[1240,1273,908,932]" italics="true" pageId="6" pageNumber="7">st3</emphasis>
13,
<emphasis id="B916EAF4FF2DFFECFA975081FA9C1B80" box="[1322,1355,908,932]" italics="true" pageId="6" pageNumber="7">st4</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFAC65080FA4B1B80" box="[1403,1436,909,932]" italics="true" pageId="6" pageNumber="7">st5</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFF7D50BDFF311BEC" box="[192,230,944,968]" italics="true" pageId="6" pageNumber="7">Jv1</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFEAB50BDFEEA1BEC" box="[278,317,944,968]" italics="true" pageId="6" pageNumber="7">Jv2</emphasis>
12,
<emphasis id="B916EAF4FF2DFFECFED350BDFE421BEC" box="[366,405,944,968]" italics="true" pageId="6" pageNumber="7">Jv3</emphasis>
12,
<emphasis id="B916EAF4FF2DFFECFE7B50BDFE3B1BEC" box="[454,492,944,968]" italics="true" pageId="6" pageNumber="7">Jv4</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFDA050BDFD931BE3" box="[541,580,944,967]" italics="true" pageId="6" pageNumber="7">Jv5</emphasis>
12,
<emphasis id="B916EAF4FF2DFFECFDC850BDFD481BEC" box="[629,671,944,968]" italics="true" pageId="6" pageNumber="7">Zv1</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFD7250BDFD2E1BEC" box="[719,761,944,968]" italics="true" pageId="6" pageNumber="7">Zv2</emphasis>
11,
<emphasis id="B916EAF4FF2DFFECFC9450BDFC841BEC" box="[809,851,944,968]" italics="true" pageId="6" pageNumber="7">Zv3</emphasis>
9, para-anal 13, post-anal 13.
</paragraph>
<paragraph id="8BDD36E6FF2DFFECFF7A50D9FB011C10" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="7">
<emphasis id="B916EAF4FF2DFFECFF7A50D9FED11BC8" bold="true" box="[199,262,980,1004]" pageId="6" pageNumber="7">Legs.</emphasis>
Lengths: I: 240; II: 204; III: 177; IV: 230. Genu IV with ten setae and tibia IV with nine setae. With two spur-like ventral setae on each of femur, genu and tibia II (
<figureCitation id="13592A63FF2DFFECFC9450F5FCBC1C34" box="[809,875,1016,1040]" captionStart="FIGURE 3" captionStartId="5.[151,250,1872,1895]" captionTargetBox="[394,1161,705,1809]" captionTargetId="figure@5.[385,1168,697,1811]" captionTargetPageId="5" captionText="FIGURE 3. Pennarhodeus brevipennatus Karg. Male. A. Lateral (antiaxial) view of spermatodactyl; B. Epistome; C. Hypostome; D. Dorsal idiosoma; E. Ventral idiosoma; F. Tritosternum; G. Anterolateral view of femur, genu and tibia of leg II. Lyrifissures enlarged for improved visibility." httpUri="https://zenodo.org/record/281300/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
G). Pretarsus I, a single, sessile, curved claw; pretarsi IIIV, an elongate ambulacral stalk, a pair of strong claws and three rounded pulvillar lobes.
</paragraph>
</subSubSection>
<subSubSection id="C378656DFF2DFFECFF7A574DFC9A1D5C" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BDD36E6FF2DFFECFF7A574DFC9A1D5C" blockId="6.[151,1437,151,1400]" pageId="6" pageNumber="7">
<emphasis id="B916EAF4FF2DFFECFF7A574DFEEC1C7C" bold="true" box="[199,315,1088,1112]" pageId="6" pageNumber="7">Remarks.</emphasis>
This species is known only from the adult male
<typeStatus id="54D98844FF2DFFECFCD3574DFC031C7C" box="[878,980,1088,1112]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
. The following characteristics appear in the original description and illustrations of the species, but do not agree with our observations of the
<typeStatus id="54D98844FF2DFFECFB455769FA891C58" box="[1272,1374,1124,1148]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
: margin of epistome smooth between antero-medial and antero-lateral projections; podonotal shield with 22 pairs of setae and four crescent-shaped scleronoduli between setae
<emphasis id="B916EAF4FF2DFFECFC9757A0FC971CE0" box="[810,832,1197,1220]" italics="true" pageId="6" pageNumber="7">j5</emphasis>
and
<emphasis id="B916EAF4FF2DFFECFCCA57A1FC5B1CE0" box="[887,908,1196,1220]" italics="true" pageId="6" pageNumber="7">j6</emphasis>
; unsclerotised integument laterad to podonotal shield without setae; opisthonotal shield with 17 pairs of setae; unsclerotised integument laterad to opisthonotal shield with a pair of setae.
<bibRefCitation id="EFF34B17FF2DFFECFE7557F8FDB71D28" author="Karg" box="[456,608,1268,1293]" pageId="6" pageNumber="7" pagination="207 - 213" refId="ref22818" refString="Karg, W. (2000 b) Neue Raubmilbenarten der Pionierartengruppe Rhodacaridae Oudemans (Acarina, Parasitiformes). Abhandlungen und Berichte des Naturkundemuseums Gorlitz, 72, 207 - 213." type="journal article" year="2000" yearSuffix="b">Karg (2000b)</bibRefCitation>
provided no information about the hypostome, tritosternum, ventral idiosoma, lyrifissures and leg setal counts. The only measurements provided in the original description referred to the length of the idiosoma (290), width of idiosoma (180), length of seta
<emphasis id="B916EAF4FF2DFFECFC235630FC6B1D70" box="[926,956,1341,1364]" italics="true" pageId="6" pageNumber="7">Z5</emphasis>
(13), range of other dorsal idiosomal setae (810), length of legs I (220), II (190), III (170) and IV (220).
</paragraph>
</subSubSection>
</treatment>
</document>