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<document id="2AD7813C4ADC43FFEA9E616F474620E2" ID-CLB-Dataset="93239" ID-DOI="10.11646/zootaxa.5162.4.1" ID-GBIF-Dataset="7d343017-483e-43be-9439-d06e748fc54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810290" ID-ZooBank="DD2279FA-E8F1-4951-A5CA-91082E875580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657270124299" checkinUser="plazi" docAuthor="Fortey, Richard A., Wernette, Shelly J. &amp; Hughes, Nigel C." docDate="2022" docId="03CB87846E4F1D12FF3AC4D64885F134" docLanguage="en" docName="zootaxa.5162.4.1.pdf" docOrigin="Zootaxa 5162 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Niobe Angelin 1851" docType="treatment" docVersion="8" lastPageNumber="327" masterDocId="FFF2FFFC6E551D08FFADC4414A1BF54C" masterDocTitle="Revision of F. R. C. Reeds Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China" masterLastPageNumber="356" masterPageNumber="301" pageNumber="327" updateTime="1732822491633" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F4AFE225327BAB2E3FDD9A4CD9631594">Revision of F. R. C. Reeds Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:affiliation id="CAFF9475A2AE56D7D86ACB41A34FCB1E">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. &amp; Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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<treatment id="03CB87846E4F1D12FF3AC4D64885F134" ID-DOI="http://doi.org/10.5281/zenodo.6816301" ID-GBIF-Taxon="197214784" ID-Zenodo-Dep="6816301" LSID="urn:lsid:plazi:treatment:03CB87846E4F1D12FF3AC4D64885F134" httpUri="http://treatment.plazi.org/id/03CB87846E4F1D12FF3AC4D64885F134" lastPageNumber="327" pageId="26" pageNumber="327" scope_family="Asaphidae" scope_subFamily="Niobinae">
<subSubSection id="C37865196E4F1D12FF3AC4D64B03F5FD" box="[151,280,151,178]" pageId="26" pageNumber="327" type="nomenclature">
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<taxonomicName id="4C624D116E4F1D12FF3AC4D64AC4F5FE" ID-CoL="7PMH9" authorityName="Angelin" authorityYear="1851" box="[151,223,151,178]" class="Trilobita" family="Asaphidae" genus="Niobe" higherTaxonomySource="CoL" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Platyhelminthes" rank="genus">
<emphasis id="B916EA806E4F1D12FF3AC4D64AC4F5FE" bold="true" box="[151,223,151,178]" italics="true" pageId="26" pageNumber="327">Niobe</emphasis>
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? sp.
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37865196E4F1D12FF3AC4FA4978F454" pageId="26" pageNumber="327" type="description">
<paragraph id="8BDD36926E4F1D12FF3AC4FA4B39F599" blockId="26.[151,290,151,213]" box="[151,290,187,213]" pageId="26" pageNumber="327">
<figureCitation id="13592A176E4F1D12FF3AC4FA4B1AF599" box="[151,257,187,213]" captionStart="FIGURE 8" captionStartId="22.[151,250,1587,1611]" captionTargetBox="[273,1314,181,1563]" captionTargetId="figure-29@22.[273,1314,181,1563]" captionTargetPageId="22" captionText="FIGURE 8. (16) Birmanites yunnanensis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1. Librigena (original of Reed 1917, pl. 7, fig. 3), GSI 11892. 2. Pygidium (original of Reed 1917, pl. 7, fig. 6), GSI 11895. 3. Pygidium (original of Reed 1917, pl. 7, fig. 8), GSI 11897. 4. Librigena (original of Reed 1917, pl. 7, fig. 5), GSI 11894. 5. Articulated specimen (original of Reed 1917, pl. 6, fig. 12), GSI 11887. 6. Cephalon (original of Reed 1917, pl. 6, fig. 14), GSI 11889. (7, 10) Remopleuriidae gen. et sp. indet. (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 7. Cranidium, (original of Reed 1917, pl. 6, fig. 9), GSI 11884. 10. Librigena (original of Reed 1917, pl. 6, fig. 10), GSI 11885. (8) Dolerobasilicus? sp. (Reed), Pygidium (original of Reed 1915, pl. 7, fig. 3), GSI 11535, Lower Naungkangyi Beds, Nawa, northern Shan State, Myanmar. (9) Nileus sp., Hypostome (original of Reed 1917, pl. 8, fig. 5), GSI 11904, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (11, 12) Niobe sp. (Reed). 11a, b. Pygidium (original of Reed 1915, pl. 5, fig. 14), GSI 11516, Hwe Mawng Beds (uppermost Katian), Nati, northern part of Shan State, Myanmar, original and retrodeformation. 12. Pygidium (original of Reed 1917, pl. 6, fig. 11), GSI 11886, Shihtien Formation (Darriwilian), Shidian, Baoshan Prefecture, western Yunnan, China. Scale bars = 5 mm for 16, 8, 9; = 2 mm for 7, 1012)" figureDoi="http://doi.org/10.5281/zenodo.6810312" httpUri="https://zenodo.org/record/6810312/files/figure.png" pageId="26" pageNumber="327">Figs 8.11</figureCitation>
,
<figureCitation id="13592A176E4F1D12FEABC4FA4B39F599" box="[262,290,187,213]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (16,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,1115) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 16, 810, 1215; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="26" pageNumber="327">12</figureCitation>
</paragraph>
<paragraph id="8BDD36926E4F1D12FF3AC4BE4978F454" blockId="26.[151,867,255,280]" box="[151,867,255,280]" pageId="26" pageNumber="327">
1915
<taxonomicName id="4C624D116E4F1D12FF5AC541482BF454" authority="Pompeckj" authorityName="Pompeckj" box="[247,560,256,280]" class="Trilobita" family="Asaphidae" genus="Asaphus" isUncertain="true" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="B916EA806E4F1D12FF5AC5414B56F454" box="[247,333,256,280]" italics="true" pageId="26" pageNumber="327">Asaphus</emphasis>
cf.
<emphasis id="B916EA806E4F1D12FED9C5414BDAF45B" box="[372,449,256,279]" italics="true" pageId="26" pageNumber="327">ornatus</emphasis>
Pompeckj
</taxonomicName>
; Reed, p. 2930, pl. 5, fig. 14.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E4F1D12FF3AC5044BC3F4E4" pageId="26" pageNumber="327" type="materials_examined">
<paragraph id="8BDD36926E4F1D12FF3AC5044BC3F4E4" blockId="26.[151,1437,325,1144]" pageId="26" pageNumber="327">
<emphasis id="B916EA806E4F1D12FF3AC5044B1DF413" bold="true" box="[151,262,325,351]" pageId="26" pageNumber="327">Material.</emphasis>
Pygidium from the Hwe Mawng Beds (uppermost Katian) at Nati,
<figureCitation id="13592A176E4F1D12FBAEC5074E7CF42C" box="[1027,1127,326,352]" captionStart="FIGURE 8" captionStartId="22.[151,250,1587,1611]" captionTargetBox="[273,1314,181,1563]" captionTargetId="figure-29@22.[273,1314,181,1563]" captionTargetPageId="22" captionText="FIGURE 8. (16) Birmanites yunnanensis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1. Librigena (original of Reed 1917, pl. 7, fig. 3), GSI 11892. 2. Pygidium (original of Reed 1917, pl. 7, fig. 6), GSI 11895. 3. Pygidium (original of Reed 1917, pl. 7, fig. 8), GSI 11897. 4. Librigena (original of Reed 1917, pl. 7, fig. 5), GSI 11894. 5. Articulated specimen (original of Reed 1917, pl. 6, fig. 12), GSI 11887. 6. Cephalon (original of Reed 1917, pl. 6, fig. 14), GSI 11889. (7, 10) Remopleuriidae gen. et sp. indet. (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 7. Cranidium, (original of Reed 1917, pl. 6, fig. 9), GSI 11884. 10. Librigena (original of Reed 1917, pl. 6, fig. 10), GSI 11885. (8) Dolerobasilicus? sp. (Reed), Pygidium (original of Reed 1915, pl. 7, fig. 3), GSI 11535, Lower Naungkangyi Beds, Nawa, northern Shan State, Myanmar. (9) Nileus sp., Hypostome (original of Reed 1917, pl. 8, fig. 5), GSI 11904, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (11, 12) Niobe sp. (Reed). 11a, b. Pygidium (original of Reed 1915, pl. 5, fig. 14), GSI 11516, Hwe Mawng Beds (uppermost Katian), Nati, northern part of Shan State, Myanmar, original and retrodeformation. 12. Pygidium (original of Reed 1917, pl. 6, fig. 11), GSI 11886, Shihtien Formation (Darriwilian), Shidian, Baoshan Prefecture, western Yunnan, China. Scale bars = 5 mm for 16, 8, 9; = 2 mm for 7, 1012)" figureDoi="http://doi.org/10.5281/zenodo.6810312" httpUri="https://zenodo.org/record/6810312/files/figure.png" pageId="26" pageNumber="327">Fig. 8.11</figureCitation>
(
<bibRefCitation id="EFF34B636E4F1D12FBDAC5074EE2F42D" author="Reed, F. R. C." box="[1143,1273,326,353]" pageId="26" pageNumber="327" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
pl. 5, fig. 14),
<collectionCode id="ED73AE576E4F1D12FF3AC52B4ADFF4C8" box="[151,196,362,388]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="26" pageNumber="327">GSI</collectionCode>
11516.
<bibRefCitation id="EFF34B636E4F1D12FE8DC52B4BBEF4C9" author="Reed, F. R. C." box="[288,421,362,389]" pageId="26" pageNumber="327" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915</bibRefCitation>
, p. 30) also mentions an asaphid thorax from a much larger individual, unfigured, but we did not locate this specimen.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E4F1D12FF6AC5F04885F134" pageId="26" pageNumber="327" type="discussion">
<paragraph id="8BDD36926E4F1D12FF6AC5F04885F134" blockId="26.[151,1437,325,1144]" pageId="26" pageNumber="327">
<emphasis id="B916EA806E4F1D12FF6AC5F04B57F487" bold="true" box="[199,332,433,459]" pageId="26" pageNumber="327">Discussion.</emphasis>
A cast from the counterpart of the specimen figured by
<bibRefCitation id="EFF34B636E4F1D12FC6FC5F34E55F481" author="Reed, F. R. C." box="[962,1102,434,461]" pageId="26" pageNumber="327" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915)</bibRefCitation>
shows some lateral extensive deformation, and a simple retrodeformation of this specimen is shown in
<figureCitation id="13592A176E4F1D12FC78C5974E51F4BC" box="[981,1098,470,496]" captionStart="FIGURE 8" captionStartId="22.[151,250,1587,1611]" captionTargetBox="[273,1314,181,1563]" captionTargetId="figure-29@22.[273,1314,181,1563]" captionTargetPageId="22" captionText="FIGURE 8. (16) Birmanites yunnanensis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1. Librigena (original of Reed 1917, pl. 7, fig. 3), GSI 11892. 2. Pygidium (original of Reed 1917, pl. 7, fig. 6), GSI 11895. 3. Pygidium (original of Reed 1917, pl. 7, fig. 8), GSI 11897. 4. Librigena (original of Reed 1917, pl. 7, fig. 5), GSI 11894. 5. Articulated specimen (original of Reed 1917, pl. 6, fig. 12), GSI 11887. 6. Cephalon (original of Reed 1917, pl. 6, fig. 14), GSI 11889. (7, 10) Remopleuriidae gen. et sp. indet. (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 7. Cranidium, (original of Reed 1917, pl. 6, fig. 9), GSI 11884. 10. Librigena (original of Reed 1917, pl. 6, fig. 10), GSI 11885. (8) Dolerobasilicus? sp. (Reed), Pygidium (original of Reed 1915, pl. 7, fig. 3), GSI 11535, Lower Naungkangyi Beds, Nawa, northern Shan State, Myanmar. (9) Nileus sp., Hypostome (original of Reed 1917, pl. 8, fig. 5), GSI 11904, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (11, 12) Niobe sp. (Reed). 11a, b. Pygidium (original of Reed 1915, pl. 5, fig. 14), GSI 11516, Hwe Mawng Beds (uppermost Katian), Nati, northern part of Shan State, Myanmar, original and retrodeformation. 12. Pygidium (original of Reed 1917, pl. 6, fig. 11), GSI 11886, Shihtien Formation (Darriwilian), Shidian, Baoshan Prefecture, western Yunnan, China. Scale bars = 5 mm for 16, 8, 9; = 2 mm for 7, 1012)" figureDoi="http://doi.org/10.5281/zenodo.6810312" httpUri="https://zenodo.org/record/6810312/files/figure.png" pageId="26" pageNumber="327">Fig. 8.11b</figureCitation>
, suggesting that the pygidium was originally slightly less than twice as wide as long. It is not otherwise badly preserved. Four progressively shorter and relatively well-defined pleural ribs fade under halfway across the pleural fields, and interpleural furrows are not developed. Four (faint fifth) axial rings on the axis, of which only the first two pass across the mid part of the axis, posterior 40% of the axis slightly convex and unfurrowed. The termination of the ribs usually marks the inner edge of the doublure in asaphids, which would indicate a straight line converging on the posterior part of the axis, around which it would be deeply embayed. The pygidial border is hardly developed. The structure of the pygidium is more like that of a niobinid than any other asaphid group. In the type species of the genus the pygidium has a well-developed border and lobe-like pleural rib terminations, but this is not true of all species that have been attributed to
<taxonomicName id="4C624D116E4F1D12FE85C75B4B70F678" authorityName="Angelin" authorityYear="1851" box="[296,363,794,820]" class="Trilobita" family="Asaphidae" genus="Niobe" higherTaxonomySource="CoL" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Platyhelminthes" rank="genus">
<emphasis id="B916EA806E4F1D12FE85C75B4B70F678" box="[296,363,794,820]" italics="true" pageId="26" pageNumber="327">Niobe</emphasis>
</taxonomicName>
. No post-Tremadoc species are currently recognised in
<collectingCountry id="F37576026E4F1D12FC4EC75B4E3DF678" box="[995,1062,794,820]" name="China" pageId="26" pageNumber="327">China</collectingCountry>
(
<bibRefCitation id="EFF34B636E4F1D12FB98C75B4F06F678" author="Zhou, Z. - Y. &amp; Zhen, Y. - Y." box="[1077,1309,794,820]" pageId="26" pageNumber="327" refId="ref49762" refString="Zhou, Z. - Y. &amp; Zhen, Y. - Y. (Eds.) (2008) Trilobite Record of China. Science Press, Beijing, 20 pp." type="book" year="2008">Zhou &amp; Zhen, 2008</bibRefCitation>
). However,
<specimenCount id="9D64FD1B6E4F1D12FF3AC77F4B26F614" box="[151,317,830,856]" pageId="26" pageNumber="327" type="generic">two specimens</specimenCount>
of thorax + pygidium figured by
<bibRefCitation id="EFF34B636E4F1D12FD1AC77F497BF615" author="Lu, Y. - H. &amp; Chang, W. - T. &amp; Chu, C. - L. &amp; Chien, Y. - Y. &amp; Hsiang, L. - W." box="[695,864,830,857]" pageId="26" pageNumber="327" refId="ref45722" refString="Lu, Y. - H., Chang, W. - T., Chu, C. - L., Chien, Y. - Y. &amp; Hsiang, L. - W. (1965) Chinese fossils of all groups, Trilobita, 2 volumes. Science Press, Beijing, 766 pp. [in Chinese]." type="book" year="1965">
Lu
<emphasis id="B916EA806E4F1D12FD70C77E490DF614" box="[733,790,830,856]" italics="true" pageId="26" pageNumber="327">et al.</emphasis>
(1965
</bibRefCitation>
, pl. 94, figs 15, 16) under the name
<taxonomicName id="4C624D116E4F1D12FB55C77F4B55F630" authority="(Reed)" baseAuthorityName="Reed" class="Trilobita" family="Asaphidae" genus="Basiliella" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B916EA806E4F1D12FB55C77F4AE2F630" italics="true" pageId="26" pageNumber="327">Basiliella yunnanensis</emphasis>
(Reed)
</taxonomicName>
are much more like the
<taxonomicName id="4C624D116E4F1D12FDCBC72348B2F630" authorityName="Angelin" authorityYear="1851" box="[614,681,866,892]" class="Trilobita" family="Asaphidae" genus="Niobe" higherTaxonomySource="CoL" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Platyhelminthes" rank="genus">
<emphasis id="B916EA806E4F1D12FDCBC72348B2F630" box="[614,681,866,892]" italics="true" pageId="26" pageNumber="327">Niobe</emphasis>
</taxonomicName>
considered here in their short pygidial pleural furrows, and unlike the type material of
<taxonomicName id="4C624D116E4F1D12FEDAC7C6481AF6EC" box="[375,513,903,928]" class="Trilobita" family="Asaphidae" genus="Basiliella" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B916EA806E4F1D12FEDAC7C6481AF6EC" box="[375,513,903,928]" italics="true" pageId="26" pageNumber="327">yunnanensis</emphasis>
</taxonomicName>
considered above. Outside
<collectingCountry id="F37576026E4F1D12FC9BC7C74967F6EC" box="[822,892,902,928]" name="China" pageId="26" pageNumber="327">China</collectingCountry>
, the closest match is probably with the pygidium of the
<typeStatus id="54D988306E4F1D12FF4CC7EB4B58F688" box="[225,323,938,964]" pageId="26" pageNumber="327" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C624D116E4F1D12FEC6C7EB4884F689" authority="Fortey, 1975" authorityName="Fortey" authorityYear="1975" box="[363,671,938,965]" class="Trilobita" family="Asaphidae" genus="Niobe" higherTaxonomySource="CoL" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Platyhelminthes" rank="species" species="occulta">
<emphasis id="B916EA806E4F1D12FEC6C7EB4813F688" box="[363,520,938,964]" italics="true" pageId="26" pageNumber="327">Niobe occulta</emphasis>
<bibRefCitation id="EFF34B636E4F1D12FDBDC7EB4884F689" author="Fortey, R. A." box="[528,671,938,965]" pageId="26" pageNumber="327" pagination="1 - 207" refId="ref43877" refString="Fortey, R. A. (1975) The Ordovician trilobites of Spitsbergen. II. Asaphidae, Nileidae, Raphiophoridae and Telephinidae of the Valhallfonna Formation. Norsk Polarinstitutt Skrifter, 162, 1 - 207." type="journal article" year="1975">Fortey, 1975</bibRefCitation>
</taxonomicName>
from the late Floian of Spitsbergen (
<bibRefCitation id="EFF34B636E4F1D12FBEEC7EB4ECCF689" author="Fortey, R. A." box="[1091,1239,938,965]" pageId="26" pageNumber="327" pagination="1 - 207" refId="ref43877" refString="Fortey, R. A. (1975) The Ordovician trilobites of Spitsbergen. II. Asaphidae, Nileidae, Raphiophoridae and Telephinidae of the Valhallfonna Formation. Norsk Polarinstitutt Skrifter, 162, 1 - 207." type="journal article" year="1975">Fortey, 1975</bibRefCitation>
, pl. 6, fig.1) with slightly longer pleural furrows and more axial rings, but without cephalic sclerites it is not possible to make further comparison. In view of the stratigraphical difference from other niobines it is identified with question accordingly. A small pygidium from the Shihtien Formation at Pupiao,
<collectingRegion id="49A6F8706E4F1D12FCA8C0574945F17C" box="[773,862,1046,1072]" country="China" name="Yunnan" pageId="26" pageNumber="327">Yunnan</collectingRegion>
, described by
<bibRefCitation id="EFF34B636E4F1D12FC56C0574E9BF17C" author="Reed, F. R. C." box="[1019,1152,1046,1072]" pageId="26" pageNumber="327" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed (1917</bibRefCitation>
, pl. 6, fig. 11) as
<taxonomicName id="4C624D116E4F1D12FA90C0574B30F118" authorityName="Wahlenberg" authorityYear="1818" class="Trilobita" family="Asaphidae" genus="Asaphus" isUncertain="true" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="expansus">
<emphasis id="B916EA806E4F1D12FA90C0574F87F17C" box="[1341,1436,1046,1072]" italics="true" pageId="26" pageNumber="327">Asaphus</emphasis>
aff.
<emphasis id="B916EA806E4F1D12FF69C07A4B30F118" box="[196,299,1083,1108]" italics="true" pageId="26" pageNumber="327">expansus</emphasis>
</taxonomicName>
is more similar to that of
<taxonomicName id="4C624D116E4F1D12FDFCC07B48A7F118" box="[593,700,1082,1108]" class="Trilobita" family="Asaphidae" genus="Niobe" higherTaxonomySource="CoL" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Platyhelminthes" rank="species" species="undetermined">
<emphasis id="B916EA806E4F1D12FDFCC07B488FF118" box="[593,660,1082,1108]" italics="true" pageId="26" pageNumber="327">Niobe</emphasis>
sp.
</taxonomicName>
than other asaphid pygidia (see
<figureCitation id="13592A176E4F1D12FB84C07B4E8BF118" box="[1065,1168,1082,1108]" captionStart="FIGURE 8" captionStartId="22.[151,250,1587,1611]" captionTargetBox="[273,1314,181,1563]" captionTargetId="figure-29@22.[273,1314,181,1563]" captionTargetPageId="22" captionText="FIGURE 8. (16) Birmanites yunnanensis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1. Librigena (original of Reed 1917, pl. 7, fig. 3), GSI 11892. 2. Pygidium (original of Reed 1917, pl. 7, fig. 6), GSI 11895. 3. Pygidium (original of Reed 1917, pl. 7, fig. 8), GSI 11897. 4. Librigena (original of Reed 1917, pl. 7, fig. 5), GSI 11894. 5. Articulated specimen (original of Reed 1917, pl. 6, fig. 12), GSI 11887. 6. Cephalon (original of Reed 1917, pl. 6, fig. 14), GSI 11889. (7, 10) Remopleuriidae gen. et sp. indet. (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 7. Cranidium, (original of Reed 1917, pl. 6, fig. 9), GSI 11884. 10. Librigena (original of Reed 1917, pl. 6, fig. 10), GSI 11885. (8) Dolerobasilicus? sp. (Reed), Pygidium (original of Reed 1915, pl. 7, fig. 3), GSI 11535, Lower Naungkangyi Beds, Nawa, northern Shan State, Myanmar. (9) Nileus sp., Hypostome (original of Reed 1917, pl. 8, fig. 5), GSI 11904, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (11, 12) Niobe sp. (Reed). 11a, b. Pygidium (original of Reed 1915, pl. 5, fig. 14), GSI 11516, Hwe Mawng Beds (uppermost Katian), Nati, northern part of Shan State, Myanmar, original and retrodeformation. 12. Pygidium (original of Reed 1917, pl. 6, fig. 11), GSI 11886, Shihtien Formation (Darriwilian), Shidian, Baoshan Prefecture, western Yunnan, China. Scale bars = 5 mm for 16, 8, 9; = 2 mm for 7, 1012)" figureDoi="http://doi.org/10.5281/zenodo.6810312" httpUri="https://zenodo.org/record/6810312/files/figure.png" pageId="26" pageNumber="327">Fig. 8.12</figureCitation>
) although with one less pleural furrow. It is recorded here as
<taxonomicName id="4C624D116E4F1D12FD98C01F4885F134" box="[565,670,1118,1144]" class="Trilobita" family="Asaphidae" genus="Niobe" higherTaxonomySource="CoL" kingdom="Animalia" order="Asaphida" pageId="26" pageNumber="327" phylum="Platyhelminthes" rank="species" species="undetermined">
<emphasis id="B916EA806E4F1D12FD98C01F4863F134" box="[565,632,1118,1144]" italics="true" pageId="26" pageNumber="327">Niobe</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>