treatments-xml/data/03/CB/87/03CB8782994F89171F99FB43D193C1EB.xml
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<document id="9E1B9F7A01483D9D5BA847506436C2E3" ID-CLB-Dataset="111179" ID-DOI="10.11646/zootaxa.5169.5.4" ID-GBIF-Dataset="7567e591-3c05-4f51-bb7d-00c04500d9f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6952689" ID-ZooBank="D91260D4-D451-4BFB-AF7B-43C1BE9515F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659441016244" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="03CB8782994F89171F99FB43D193C1EB" docLanguage="en" docName="zootaxa.5169.5.4.pdf" docOrigin="Zootaxa 5169 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carcinocephalus cuccodoroi Shavrin 2022, sp.n." docType="treatment" docVersion="7" lastPageNumber="460" masterDocId="FFF2FFFA994D89141F0EFF87D014C476" masterDocTitle="New species and records of Omaliini McLeay, 1825 from Eastern Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="471" masterPageNumber="457" pageNumber="459" updateTime="1699363382017" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="83DAD79A3436B47926629F55C22CFA5E">New species and records of Omaliini McLeay, 1825 from Eastern Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<mods:namePart id="E692561BE456BDCD60A160820CBDE7D1">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="04ECE03939C4BF694E9EFBCEECE51068">2022</mods:date>
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<treatment id="03CB8782994F89171F99FB43D193C1EB" ID-DOI="http://doi.org/10.5281/zenodo.6952629" ID-GBIF-Taxon="198497309" ID-Zenodo-Dep="6952629" LSID="urn:lsid:plazi:treatment:03CB8782994F89171F99FB43D193C1EB" httpUri="http://treatment.plazi.org/id/03CB8782994F89171F99FB43D193C1EB" lastPageId="3" lastPageNumber="460" pageId="2" pageNumber="459">
<subSubSection id="C378651F994F89161F99FB43D220C0A9" box="[151,564,1220,1247]" pageId="2" pageNumber="459" type="nomenclature">
<paragraph id="8BDD3694994F89161F99FB43D220C0A9" blockId="2.[151,564,1220,1283]" box="[151,564,1220,1247]" pageId="2" pageNumber="459">
<heading id="D09581F8994F89161F99FB43D220C0A9" bold="true" box="[151,564,1220,1247]" fontSize="11" level="1" pageId="2" pageNumber="459" reason="1">
<emphasis id="B916EA86994F89161F99FB43D220C0A9" bold="true" box="[151,564,1220,1247]" pageId="2" pageNumber="459">
<taxonomicName id="4C624D17994F89161F99FB43D1E6C0A9" ID-CoL="9GDK8" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,498,1220,1247]" class="Insecta" family="Staphylinidae" genus="Carcinocephalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="cuccodoroi" status="sp. nov.">
<emphasis id="B916EA86994F89161F99FB43D1E6C0A9" bold="true" box="[151,498,1220,1247]" italics="true" pageId="2" pageNumber="459">Carcinocephalus cuccodoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557FD994F89161EF7FB42D220C0A9" box="[505,564,1221,1247]" pageId="2" pageNumber="459" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C378651F994F89161F99FB6FD134C175" box="[151,288,1256,1283]" pageId="2" pageNumber="459" type="description">
<paragraph id="8BDD3694994F89161F99FB6FD134C175" blockId="2.[151,564,1220,1283]" box="[151,288,1256,1283]" pageId="2" pageNumber="459">
(
<figureCitation id="13592A11994F89161F91FB6FD10CC175" box="[159,280,1256,1283]" captionStart="FIGURES 915" captionStartId="4.[151,264,1892,1916]" captionTargetBox="[160,1421,499,1856]" captionTargetId="figure-160@4.[151,1436,489,1869]" captionTargetPageId="4" captionText="FIGURES 915. Carcinocephalus cuccodoroi (Figs. 912) and Paraphloeostiba cooteri (Figs. 1315): 9aedeagus, parameral view, 10aedeagus, lateral view, 11male abdominal tergite VIII, 12male abdominal sternite VIII, 13female abdominal tergite VIII, 14female abdominal sternite VIII, 15female accessory sclerite. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6952695" httpUri="https://zenodo.org/record/6952695/files/figure.png" pageId="2" pageNumber="459">Figs. 912</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C378651F994F89161F99FAB6D39FC1E5" pageId="2" pageNumber="459" type="materials_examined">
<paragraph id="8BDD3694994F89161F99FAB6D39FC1E5" blockId="2.[151,1437,1328,2039]" pageId="2" pageNumber="459">
<emphasis id="B916EA86994F89161F99FAB6D1DDC13C" bold="true" box="[151,457,1328,1355]" pageId="2" pageNumber="459">Type material examined.</emphasis>
<materialsCitation id="3B0A3CC9994F89161EDBFAB7D39FC1E5" ID-GBIF-Occurrence="3874686314" collectingDate="1998-11-24" collectionCode="MHNG" collectorName="G. Cuccodoro" country="India" determinerName="Shavrin A. V." elevation="0" latitude="25.132223" location="Mt. Barail" longLatPrecision="21" longitude="93.04972" municipality="Dima Hasao" pageId="2" pageNumber="459" specimenCount="1" specimenCount-male="1" stateProvince="Assam" typeStatus="holotype">
<typeStatus id="54D98836994F89161EDBFAB7D229C13C" box="[469,573,1328,1354]" pageId="2" pageNumber="459" type="holotype">Holotype</typeStatus>
<specimenCount id="9D64FD1D994F89161D47FAB6D24AC13D" box="[585,606,1329,1355]" pageId="2" pageNumber="459" type="male"></specimenCount>
(dissected):
<collectingCountry id="F3757604994F89161DF2FAB6D340C13D" box="[764,852,1329,1355]" name="India" pageId="2" pageNumber="459">INDIA</collectingCountry>
:
<collectingRegion id="49A6F876994F89161C6CFAB6D3A7C13D" box="[866,947,1329,1355]" country="India" name="Assam" pageId="2" pageNumber="459">Assam</collectingRegion>
:
<collectingMunicipality id="6BB9ACEE994F89161CCCFAB7D447C13D" box="[962,1107,1328,1355]" pageId="2" pageNumber="459">Dima Hasao</collectingMunicipality>
|
<location id="8EBD604F994F89161B7EFAB7D4CBC13C" LSID="urn:lsid:plazi:treatment:03CB8782994F89171F99FB43D193C1EB:8EBD604F994F89161B7EFAB7D4CBC13C" box="[1136,1247,1328,1354]" country="India" latitude="25.132223" longLatPrecision="21" longitude="93.04972" municipality="Dima Hasao" name="Mt. Barail" pageId="2" pageNumber="459" stateProvince="Assam">Mt.Barail</location>
,
<location id="8EBD604F994F89161BE1FAB6D59EC13C" LSID="urn:lsid:plazi:treatment:03CB8782994F89171F99FB43D193C1EB:8EBD604F994F89161BE1FAB6D59EC13C" box="[1263,1418,1328,1355]" country="India" latitude="25.132223" longLatPrecision="21" longitude="93.04972" municipality="Dima Hasao" name="Notun Leikul" pageId="2" pageNumber="459" stateProvince="Assam">Notun Leikul</location>
|
<geoCoordinate id="EE565053994F89161F99FAD3D13EC119" box="[151,298,1364,1391]" degrees="25" direction="north" minutes="07" orientation="latitude" pageId="2" pageNumber="459" precision="15" seconds="56" value="25.132223">25°07`56``N</geoCoordinate>
;
<geoCoordinate id="EE565053994F89161E38FAD3D1D7C119" box="[310,451,1364,1391]" degrees="93" direction="east" minutes="02" orientation="longitude" pageId="2" pageNumber="459" precision="15" seconds="59" value="93.04972">93°02`59``E</geoCoordinate>
|
<date id="FFDC1054994F89161ED6FAD2D24CC119" box="[472,600,1365,1391]" pageId="2" pageNumber="459" value="1998-11-24">
<collectingDate id="EF98E9BC994F89161ED6FAD2D24CC119" box="[472,600,1365,1391]" pageId="2" pageNumber="459" value="1998-11-24">24.11.1998</collectingDate>
</date>
, 850 m |
<collectorName id="26975342994F89161DCCFAD2D374C118" box="[706,864,1364,1391]" pageId="2" pageNumber="459">G. Cuccodoro</collectorName>
#4a &lt;printed&gt;, HOLOTYPE |
<originalDetermination id="A87C8E68994F89161BDEFAD3D144C1E5" pageId="2" pageNumber="459">
<taxonomicName id="4C624D17994F89161BDEFAD3D59AC118" authorityName="Bernhauer" authorityYear="1903" box="[1232,1422,1364,1390]" class="Insecta" family="Staphylinidae" genus="Carcinocephalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA86994F89161BDEFAD3D59AC118" box="[1232,1422,1364,1390]" italics="true" pageId="2" pageNumber="459">Carcinocephalus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C624D17994F89161F99FAFFD100C1E4" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,276,1400,1426]" class="Insecta" family="Staphylinidae" genus="Carcinocephalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="cuccodoroi" status="sp. nov.">
<emphasis id="B916EA86994F89161F99FAFFD100C1E4" box="[151,276,1400,1426]" italics="true" pageId="2" pageNumber="459">cuccodoroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557FD994F89161E15FAFED144C1E5" box="[283,336,1401,1427]" pageId="2" pageNumber="459" rank="species">sp.n.</taxonomicNameLabel>
</originalDetermination>
|
<determinerName id="7E01DA50994F89161E6AFAFFD1E7C1E5" box="[356,499,1400,1427]" pageId="2" pageNumber="459">Shavrin A.V.</determinerName>
des. 2022 &lt;red, printed&gt; (
<collectionCode id="ED73AE51994F89161C2DFAFED36BC1E5" box="[803,895,1401,1427]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="459" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C378651F994F89171FC9FA1BD4BDC09F" lastPageId="3" lastPageNumber="460" pageId="2" pageNumber="459" type="description">
<paragraph id="8BDD3694994F89161FC9FA1BD4F5C1AD" blockId="2.[151,1437,1328,2039]" pageId="2" pageNumber="459">
<emphasis id="B916EA86994F89161FC9FA1BD145C1C0" bold="true" box="[199,337,1436,1462]" pageId="2" pageNumber="459">Description</emphasis>
. Measurements: HW: 0.36; HL: 0.22; AL: 0.53; OL: 0.12; PL: 0.27; PW: 0.42; ESL: 0.51; EW: 0.51; AW: 0.47; MTbL: 0.22; MTrL: 0.13 (MTrL 14: 0.05; MTrL 5: 0.08); AedL: 0.27; TL: 1.82.
</paragraph>
<paragraph id="8BDD3694994F89161FC9FA63D269C3FD" blockId="2.[151,1437,1328,2039]" pageId="2" pageNumber="459">Body somewhat elongate, slightly convex. Head reddish-brown, with slightly paler frontal portion; pronotum, elytra and abdomen yellow-brown, with paler laterobasal portions of pronotum, latero-apical portions of elytra and paratergites; antennomeres 511 brown; mouthparts, antennomeres 14 and legs yellow. Head with several very fine punctures in middle; neck without visible punctation; pronotum with sparse and fine punctation, sparser and finer in apical and basal portions; punctation of elytra denser and deeper than that on pronotum, finer along suture and sparser in apical portion, punctures on each elytron arranged to vague and tangled five rows; abdomen with indistinct, very fine and sparse punctation. Body glossy, with strong transverse microsculpture in middle of vertex and diagonal meshes between infraorbital ridges and anteocellar foveae; neck with indistinct isodiametric microreticulation; pronotum, scutellum and elytra without microsculpture; abdomen with moderately strong and dense isodiametric meshes. Head with long and moderately dense setation in frontal portion, with several semierect setae between apical margins of eyes and median portion; lateral portions of pronotum, elytra and paratergites of abdomen with sparse and short, erect setae.</paragraph>
<paragraph id="8BDD3694994F89171FC9F813D402C64B" blockId="2.[151,1437,1328,2039]" lastBlockId="3.[151,1437,151,1437]" lastPageId="3" lastPageNumber="460" pageId="2" pageNumber="459">Head 1.6 times as broad as long, with slightly elevated medioapical and infraorbital portions, with moderately deep, suboval impressions between supra-antennal prominences and middle part of vertex; anteocellar foveae deep and long, diagonally stretching toward level of anterior third of eyes; middle part between level of apical third of eyes with narrow transverse impression; anterolateral portions of head between antennal insertion and anterior margin of eye slightly concaved. Eyes large and convex; postocular ridge small, subacute; surface between postocular ridge and posterior margin of eye as long as diameter of one nearest ommatidium. Ocelli large, convex, located at level of postocular ridges; distance between ocelli about one and a half times less than distance between ocellus and posterior margin of eye; neck constriction without visible occipital furrow. Last segment of maxillary palpomeres slightly broader than preceding palpomere, from middle gradually narrowed toward subacute apex; preapical segment small, transverse, slightly broader than long. Antenna short, reaching apical margin of pronotum when reclined, with transverse antennomeres 410; antennomeres with long setation, distinctly denser on 510; basal antennomere more than twice as long as broad, antennomere 2 suboval, distinctly narrower than basal antennomere, 3 very narrow, slightly shorter than 2, 4 very small, slightly broader than 3, 5 distinctly broader and slightly longer than 4, 6 longer and broader than 5, 710 longer and broader than 6, apical antennomere about as long as two preapical antennomeres, from middle gradually narrowed toward rounded apex.</paragraph>
<paragraph id="8BDD3694994E89171FC9FDC0D35DC687" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="460">Pronotum slightly convex, 1.5 times as broad as long, slightly broader than head, from widest anterior third strongly narrowed posteriad; anterior angles widely rounded and slightly protruded anteriad; anterior margin rounded, narrower than posterior margin, with shallow subtriangular emargination in middle; posterior angles subacute, distinctly protruded posteriad; lateral edges with smooth irregular crenulation; lateral portions without mediolateral pits, with widely impressed laterobasal portions.</paragraph>
<paragraph id="8BDD3694994E89171FC9FD7CD3E2C74F" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="460">Elytra slightly convex, as long as broad, 1.8 times as long as pronotum, lateral sides bordered, slightly explanate in middle; posterior margins somewhat straight. Hind wings fully developed.</paragraph>
<paragraph id="8BDD3694994E89171FC9FCC4D3CEC72B" blockId="3.[151,1437,151,1437]" box="[199,986,835,861]" pageId="3" pageNumber="460">Legs moderately short; metatarsomeres 1.6 times as long as metatibia.</paragraph>
<paragraph id="8BDD3694994E89171FC9FCE0D150C7BF" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="460">Abdomen convex, slightly narrower than elytra, with two oval tomentose spots (wing-folding patches) in middle of abdominal tergite IV and two smaller spots in middle of tergite V; apical margin of tergite VII with very narrow palisade fringe.</paragraph>
<paragraph id="8BDD3694994E89171FC9FC54D3DBC00B" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="460">
Male. Apical margin of abdominal tergite VIII slightly concave (
<figureCitation id="13592A11994E89171C91FC54D3E5C79B" box="[927,1009,979,1005]" captionStart="FIGURES 915" captionStartId="4.[151,264,1892,1916]" captionTargetBox="[160,1421,499,1856]" captionTargetId="figure-160@4.[151,1436,489,1869]" captionTargetPageId="4" captionText="FIGURES 915. Carcinocephalus cuccodoroi (Figs. 912) and Paraphloeostiba cooteri (Figs. 1315): 9aedeagus, parameral view, 10aedeagus, lateral view, 11male abdominal tergite VIII, 12male abdominal sternite VIII, 13female abdominal tergite VIII, 14female abdominal sternite VIII, 15female accessory sclerite. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6952695" httpUri="https://zenodo.org/record/6952695/files/figure.png" pageId="3" pageNumber="460">Fig. 11</figureCitation>
). Apical margin of abdominal sternite VIII widely rounded (
<figureCitation id="13592A11994E89171E83FC70D1F5C067" box="[397,481,1015,1041]" captionStart="FIGURES 915" captionStartId="4.[151,264,1892,1916]" captionTargetBox="[160,1421,499,1856]" captionTargetId="figure-160@4.[151,1436,489,1869]" captionTargetPageId="4" captionText="FIGURES 915. Carcinocephalus cuccodoroi (Figs. 912) and Paraphloeostiba cooteri (Figs. 1315): 9aedeagus, parameral view, 10aedeagus, lateral view, 11male abdominal tergite VIII, 12male abdominal sternite VIII, 13female abdominal tergite VIII, 14female abdominal sternite VIII, 15female accessory sclerite. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6952695" httpUri="https://zenodo.org/record/6952695/files/figure.png" pageId="3" pageNumber="460">Fig. 12</figureCitation>
). Aedeagus narrow, from widest basal part gradually narrowed toward truncate apex; parameres very wide and long, but slightly not exceeding apex of median lobe, with two apical and two longer preapical setae; internal sac with paired sclerotized structures in apical, median and basal parts and moderately long flagellum between them (
<figureCitation id="13592A11994E89171EBBFBE4D1EDC00B" box="[437,505,1123,1149]" captionStart="FIGURES 915" captionStartId="4.[151,264,1892,1916]" captionTargetBox="[160,1421,499,1856]" captionTargetId="figure-160@4.[151,1436,489,1869]" captionTargetPageId="4" captionText="FIGURES 915. Carcinocephalus cuccodoroi (Figs. 912) and Paraphloeostiba cooteri (Figs. 1315): 9aedeagus, parameral view, 10aedeagus, lateral view, 11male abdominal tergite VIII, 12male abdominal sternite VIII, 13female abdominal tergite VIII, 14female abdominal sternite VIII, 15female accessory sclerite. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6952695" httpUri="https://zenodo.org/record/6952695/files/figure.png" pageId="3" pageNumber="460">Fig. 9</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="13592A11994E89171C77FBE4D3DFC00B" box="[889,971,1123,1149]" captionStart="FIGURES 915" captionStartId="4.[151,264,1892,1916]" captionTargetBox="[160,1421,499,1856]" captionTargetId="figure-160@4.[151,1436,489,1869]" captionTargetPageId="4" captionText="FIGURES 915. Carcinocephalus cuccodoroi (Figs. 912) and Paraphloeostiba cooteri (Figs. 1315): 9aedeagus, parameral view, 10aedeagus, lateral view, 11male abdominal tergite VIII, 12male abdominal sternite VIII, 13female abdominal tergite VIII, 14female abdominal sternite VIII, 15female accessory sclerite. Scale bar: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6952695" httpUri="https://zenodo.org/record/6952695/files/figure.png" pageId="3" pageNumber="460">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="8BDD3694994E89171FC9FB00D185C0D7" blockId="3.[151,1437,151,1437]" box="[199,401,1159,1185]" pageId="3" pageNumber="460">Female unknown.</paragraph>
<paragraph id="8BDD3694994E89171FC9FB2DD4BDC09F" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="460">
<emphasis id="B916EA86994E89171FC9FB2DD1A2C0B3" bold="true" box="[199,438,1194,1221]" pageId="3" pageNumber="460">Comparative notes.</emphasis>
<taxonomicName id="4C624D17994E89171ECCFB2CD31EC0B3" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[450,778,1195,1221]" class="Insecta" family="Staphylinidae" genus="Carcinocephalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="460" phylum="Arthropoda" rank="species" species="cuccodoroi" status="sp. nov.">
<emphasis id="B916EA86994E89171ECCFB2CD31EC0B3" box="[450,778,1195,1221]" italics="true" pageId="3" pageNumber="460">Carcinocephalus cuccodoroi</emphasis>
</taxonomicName>
<emphasis id="B916EA86994E89171C18FB2CD359C0B3" bold="true" box="[790,845,1195,1221]" pageId="3" pageNumber="460">
<taxonomicNameLabel id="A22557FD994E89171C18FB2CD359C0B3" box="[790,845,1195,1221]" pageId="3" pageNumber="460" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from all congeners (
<bibRefCitation id="EFF34B65994E89171B91FB2CD52AC0B3" author="Shavrin, A. V." box="[1183,1342,1195,1221]" pageId="3" pageNumber="460" pagination="81 - 87" refId="ref8302" refString="Shavrin, A. V. (2018) A new species of the genus Carcinocephalus Bernhauer (Coleoptera: Staphylinidae: Omaliinae: Omaliini) from Taiwan. Fragmenta Faunistica, 61 (2), 81 - 87. https: // doi. org / 10.3161 / 00159301 FF 2018.61.2081" type="journal article" year="2018">Shavrin 2018</bibRefCitation>
) by the narrower and flattened body, and different internal and external morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C378651F994E89171FC9FB75D22CC123" pageId="3" pageNumber="460" type="distribution">
<paragraph id="8BDD3694994E89171FC9FB75D4FBC17B" blockId="3.[151,1437,151,1437]" box="[199,1263,1266,1293]" pageId="3" pageNumber="460">
<emphasis id="B916EA86994E89171FC9FB75D175C17A" bold="true" box="[199,353,1266,1292]" pageId="3" pageNumber="460">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="54D98836994E89171CA8FB74D3C2C17B" box="[934,982,1267,1293]" pageId="3" pageNumber="460">type</typeStatus>
locality in
<collectingRegion id="49A6F876994E89171B5AFB74D4B7C17B" box="[1108,1187,1267,1293]" country="India" name="Assam" pageId="3" pageNumber="460">Assam</collectingRegion>
,
<collectingCountry id="F3757604994E89171BA0FB74D4FFC17B" box="[1198,1259,1267,1293]" name="India" pageId="3" pageNumber="460">India</collectingCountry>
.
</paragraph>
<paragraph id="8BDD3694994E89171FC9FA91D22CC123" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="460">
<emphasis id="B916EA86994E89171FC9FA91D15FC146" bold="true" box="[199,331,1302,1328]" pageId="3" pageNumber="460">Bionomics.</emphasis>
The
<typeStatus id="54D98836994E89171E8FFA90D1F7C147" box="[385,483,1303,1329]" pageId="3" pageNumber="460" type="holotype">holotype</typeStatus>
was collected at elevation
<quantity id="4C9A9B71994E89171C02FA91D346C147" box="[780,850,1302,1329]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="460" unit="m" value="850.0">850 m</quantity>
a.s.l. in gallery forest by sifting of rotten fruits in dry stream (G. Cuccodoro, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="C378651F994E89171FC9FAD8D193C1EB" pageId="3" pageNumber="460" type="etymology">
<paragraph id="8BDD3694994E89171FC9FAD8D193C1EB" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="460">
<emphasis id="B916EA86994E89171FC9FAD8D15AC10F" bold="true" box="[199,334,1375,1401]" pageId="3" pageNumber="460">Etymology.</emphasis>
Patronymic, the species is named to honour my colleague, Giulio Cuccodoro (
<collectingRegion id="49A6F876994E89171BA9FAD8D516C10F" box="[1191,1282,1375,1401]" country="Switzerland" name="Geneve" pageId="3" pageNumber="460">Genève</collectingRegion>
), the collector of the
<typeStatus id="54D98836994E89171FD1FA04D11BC1EB" box="[223,271,1411,1437]" pageId="3" pageNumber="460">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
</treatment>
</document>