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<document id="D6B6738555B1779AFE52309843BCFF65" ID-DOI="10.1093/zoolinnean/zlae113" ID-ISSN="0024-4082" ID-Zenodo-Dep="14542355" ID-ZooBank="9ECD785-00BF-494C-9888-0B0D073F302C" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1732849427912" checkinUser="plazi" docAuthor="Sun, Chenhui, Rasnitsyn, Alexandr P., Wedmann, Sonja, Zhuang, Jialiang, Shih, Chungkun, Ren, Dong &amp; Gao" docDate="2024" docId="03C487AD1F0C2360FC70DCF0FAE7F900" docLanguage="en" docName="zlae113.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (1)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlae113" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Enspeletoma Sun &amp; Rasnitsyn &amp; Wedmann &amp; Zhuang &amp; Shih &amp; Ren &amp; Gao 2024, gen. nov." docType="treatment" docVersion="6" lastPageNumber="3" masterDocId="FFFDFFD51F0E2362FFF2DF42FF5AFF8B" masterDocTitle="Revisions * new taxa * and venation transformations of the sawfly family Blasticotomidae sensu lato (Hymenoptera: ºenthredinoidea) highlight the evolution of the basal Hymenoptera" masterLastPageNumber="19" masterPageNumber="1" pageNumber="3" updateTime="1734832297368" updateUser="ExternalLinkService">
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<mods:title id="4CDA0413D19148704F5C68125FAF27B7">Revisions * new taxa * and venation transformations of the sawfly family Blasticotomidae sensu lato (Hymenoptera: ºenthredinoidea) highlight the evolution of the basal Hymenoptera</mods:title>
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<mods:namePart id="68E03BDBBC781C0438FD3A389F9E76A9">Sun, Chenhui</mods:namePart>
<mods:affiliation id="E0AD29C518D44AF0893317957E9BEA20">College of Life Sciences * Capital Normal University * Beijing 100048 * China</mods:affiliation>
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<mods:namePart id="614EB99B2F2055F35F72709E3D0732BC">Rasnitsyn, Alexandr P.</mods:namePart>
<mods:affiliation id="B31D391F8E382C851FA25FFFF9780FF7">Paleontological Institute * Russian Academy of Sciences * 123 * 117647 Moscow * Russia &amp; Department of Palaeontology * Natural History Museum * Cromwell Road * London SW 7 5 BD * UK</mods:affiliation>
<mods:nameIdentifier id="4CD6A9D68C4DDE5980984E1B99E96FD1" type="email">alex.rasnitsyn@gmail.com&amp;tpgao@cnu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="C19BA5C47F5B8DCFDD0E6E46B3A8AC38">Wedmann, Sonja</mods:namePart>
<mods:affiliation id="651E3FAC0B3EB5275922B6A8EFDA48E1">Senckenberg Forschungsstation Grube Messel * Senckenberg Forschungsinstitut und Naturmuseum Frankfurt am Main * 64409 Messel * Germany</mods:affiliation>
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<mods:namePart id="0F724E7108D8A106DFCA26731E63DD5F">Zhuang, Jialiang</mods:namePart>
<mods:affiliation id="A864565D68839345D4FE11DA84D7DC20">College of Life Sciences * Capital Normal University * Beijing 100048 * China</mods:affiliation>
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<mods:namePart id="AC130CE53BB8F7CA6700F6026910DA41">Shih, Chungkun</mods:namePart>
<mods:affiliation id="8C100622F026DA2513731DBD080DC707">College of Life Sciences * Capital Normal University * Beijing 100048 * China &amp; Department of Paleobiology * National Museum of Natural History * Smithsonian Institution * Washington * DC 20013 - 7012 * USA</mods:affiliation>
<mods:nameIdentifier id="53884FD19AE8264B67D4C31B8766174B" type="email">alex.rasnitsyn@gmail.com&amp;tpgao@cnu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="B21C9ACA5E61F6ECC18931E508B585A9">Ren, Dong</mods:namePart>
<mods:affiliation id="924C5B5DF499FD55D1F2C5DE8C60D859">College of Life Sciences * Capital Normal University * Beijing 100048 * China</mods:affiliation>
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<mods:date id="FF00757663A44FA0CCE1F340666700FD">2024</mods:date>
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<treatment id="03C487AD1F0C2360FC70DCF0FAE7F900" ID-DOI="http://doi.org/10.5281/zenodo.14522742" ID-Zenodo-Dep="14522742" LSID="urn:lsid:plazi:treatment:03C487AD1F0C2360FC70DCF0FAE7F900" httpUri="http://treatment.plazi.org/id/03C487AD1F0C2360FC70DCF0FAE7F900" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection id="C37765301F0C2360FC70DCF0FA23FC47" box="[898,1401,946,973]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BD236BB1F0C2360FC70DCF0FA23FC47" blockId="2.[898,1401,946,973]" box="[898,1401,946,973]" pageId="2" pageNumber="3">
<heading id="D09A81D71F0C2360FC70DCF0FA23FC47" bold="true" box="[898,1401,946,973]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<emphasis id="B919EAA91F0C2360FC70DCF0FA23FC47" bold="true" box="[898,1401,946,973]" pageId="2" pageNumber="3">
<taxonomicName id="4C6D4D381F0C2360FC70DCF0FA56FC47" authority="Rasnitsyn &amp; Wedmann" authorityName="Sun &amp; Rasnitsyn &amp; Wedmann &amp; Zhuang &amp; Shih &amp; Ren &amp; Gao" authorityYear="2024" box="[898,1292,946,973]" class="Insecta" family="Xyelotomidae" genus="Enspeletoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B919EAA91F0C2360FC70DCF0FB48FC46" bold="true" box="[898,1042,946,973]" italics="true" pageId="2" pageNumber="3">Enspeletoma</emphasis>
Rasnitsyn &amp; Wedmann
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<taxonomicNameLabel id="A22A57D21F0C2360FAE3DCF0FA23FC47" box="[1297,1401,946,972]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C37765301F0C2360FCCBDCB1FBF9FBE3" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="8BD236BB1F0C2360FCCBDCB1FBF9FBE3" blockId="2.[825,1474,1010,1128]" pageId="2" pageNumber="3">
<emphasis id="B919EAA91F0C2360FCCBDCB1FCECFB81" box="[825,950,1011,1034]" italics="true" pageId="2" pageNumber="3">
<typeStatus id="54D688191F0C2360FCCBDCB1FC33FB81" box="[825,873,1011,1034]" pageId="2" pageNumber="3">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C6D4D381F0C2360FC4DDCB0FA98FB80" authority="Rasnitsyn &amp; Wedmann" authorityName="Sun &amp; Rasnitsyn &amp; Wedmann &amp; Zhuang &amp; Shih &amp; Ren &amp; Gao" authorityYear="2024" box="[959,1474,1010,1035]" class="Insecta" family="Xyelotomidae" genus="Enspeletoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="oligocaenica" status="gen. et sp. nov.">
<emphasis id="B919EAA91F0C2360FC4DDCB0FB96FB81" bold="true" box="[959,1228,1010,1034]" italics="true" pageId="2" pageNumber="3">Enspeletoma oligocaenica</emphasis>
Rasnitsyn &amp; Wedmann
</taxonomicName>
<emphasis id="B919EAA91F0C2360FCC9DB50FC83FBA1" bold="true" box="[827,985,1042,1066]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="A22A57D21F0C2360FCC9DB50FC83FBA1" box="[827,985,1042,1066]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
http://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="FFCB0C6E1F0C2360FAC1DB73FBF9FBE3" pageId="2" pageNumber="3">C92A60A4- F156-4903-BCCE-07644EE4140C</uuid>
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<subSubSection id="C37765301F0C2360FCCBDBC8FA77FB6A" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BD236BB1F0C2360FCCBDBC8FA77FB6A" blockId="2.[825,1476,1162,1249]" pageId="2" pageNumber="3">
<emphasis id="B919EAA91F0C2360FCCBDBC8FCF2FB29" box="[825,936,1162,1186]" italics="true" pageId="2" pageNumber="3">Etymology:</emphasis>
The first part of the genus name is derived from the
<typeStatus id="54D688191F0C2360FCC9DBE8FC33FB49" box="[827,873,1194,1218]" pageId="2" pageNumber="3">type</typeStatus>
locality Enspel; the second part* -toma* refers to the
<typeStatus id="54D688191F0C2360FA64DBE8FA9EFB49" box="[1430,1476,1194,1218]" pageId="2" pageNumber="3">type</typeStatus>
genus
<taxonomicName id="4C6D4D381F0C2360FC88DB8BFB76FB6A" authority="Klug" authorityName="Klug" authorityYear="1834" box="[890,1068,1225,1249]" class="Insecta" family="Blasticotomidae" genus="Blasticotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B919EAA91F0C2360FC88DB8BFCA2FB6A" box="[890,1016,1225,1249]" italics="true" pageId="2" pageNumber="3">Blasticotoma</emphasis>
Klug
</taxonomicName>
* 1834. Gender feminine.
</paragraph>
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<subSubSection id="C37765301F0C2360FCCBDA40FA46FA91" box="[825,1308,1282,1306]" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="8BD236BB1F0C2360FCCBDA40FA46FA91" blockId="2.[825,1308,1282,1306]" box="[825,1308,1282,1306]" pageId="2" pageNumber="3">
<emphasis id="B919EAA91F0C2360FCCBDA40FC82FA91" box="[825,984,1282,1306]" italics="true" pageId="2" pageNumber="3">Included species:</emphasis>
Type species only (
<figureCitation id="13562A3E1F0C2360FB51DA40FA56FA91" box="[1187,1292,1282,1306]" captionStart="Figure 2" captionStartId="4.[129,194,1859,1883]" captionTargetBox="[132,1469,152,1828]" captionTargetId="figure-9@4.[129,1473,149,1831]" captionTargetPageId="4" captionText="Figure 2. Cenozoic extinct Blasticotominae.AC* Enspeletoma oligocaenica. Holotype* NHMMZ PE 1995/9108-LS* sex unknown* Upper Oligocene of Enspel* Germany.A* dorsal view.B* C* line drawings of forewing* reconstructed from both preserved wings* with veins labelled (B) and with cells labelled (C). D* E* Paremphytus ostentus. Holotype No. 2042* Museum of Comparative Zoology* Harvard University* Cambridge* MA* USA; female* uppermost Eocene* Colorado* USA. D* dorsal view.E* line drawing (Zhelochovtzev and Rasnitsyn 1972). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.14542359" httpUri="https://zenodo.org/record/14542359/files/figure.png" pageId="2" pageNumber="3">Fig. 2AC</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37765301F0C2360FCCBDA7DFB5FFA3E" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BD236BB1F0C2360FCCBDA7DFB5FFA3E" blockId="2.[825,1476,1343,1461]" pageId="2" pageNumber="3">
<emphasis id="B919EAA91F0C2360FCCBDA7DFCC5FADD" box="[825,927,1343,1366]" italics="true" pageId="2" pageNumber="3">Diagnosis:</emphasis>
Pterostigma almost as long and wide as cell 1mcu; cell 3rm with veins 2r-rs and 2m-cu opposite each other; vein 2r-m about as long as vein 2r-rs; cell 2mcu relatively short* about as long as cell 1r+2rm.
</paragraph>
</subSubSection>
<subSubSection id="C37765301F0C2360FCCBDA95FAE7F900" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BD236BB1F0C2360FCCBDA95FAE7F900" blockId="2.[825,1475,1495,1676]" pageId="2" pageNumber="3">
<emphasis id="B919EAA91F0C2360FCCBDA95FCCEFA64" box="[825,916,1495,1519]" italics="true" pageId="2" pageNumber="3">Remarks:</emphasis>
Although known from a fragmentary fossil (
<figureCitation id="13562A3E1F0C2360FAB5DA95FAD4FA64" box="[1351,1422,1495,1519]" captionStart="Figure 2" captionStartId="4.[129,194,1859,1883]" captionTargetBox="[132,1469,152,1828]" captionTargetId="figure-9@4.[129,1473,149,1831]" captionTargetPageId="4" captionText="Figure 2. Cenozoic extinct Blasticotominae.AC* Enspeletoma oligocaenica. Holotype* NHMMZ PE 1995/9108-LS* sex unknown* Upper Oligocene of Enspel* Germany.A* dorsal view.B* C* line drawings of forewing* reconstructed from both preserved wings* with veins labelled (B) and with cells labelled (C). D* E* Paremphytus ostentus. Holotype No. 2042* Museum of Comparative Zoology* Harvard University* Cambridge* MA* USA; female* uppermost Eocene* Colorado* USA. D* dorsal view.E* line drawing (Zhelochovtzev and Rasnitsyn 1972). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.14542359" httpUri="https://zenodo.org/record/14542359/files/figure.png" pageId="2" pageNumber="3">Fig. 2A</figureCitation>
)* the new genus demonstrates diagnostic characters of the subfamily
<taxonomicName id="4C6D4D381F0C2360FCCBD957FC82F9A6" authorityName="Thomson" authorityYear="1871" box="[825,984,1557,1581]" class="Insecta" family="Argidae" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Blasticotominae">Blasticotominae</taxonomicName>
as defined above and in the key below. It differs clearly from other genera and particularly from the only other Cenozoic fossil*
<taxonomicName id="4C6D4D381F0C2360FC2FD916FBB9F9E7" authority="Brues" authorityName="Brues" authorityYear="1908" box="[989,1251,1620,1644]" class="Insecta" family="Blasticotomidae" genus="Paremphytus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ostentus">
<emphasis id="B919EAA91F0C2360FC2FD916FBF2F9E7" box="[989,1192,1620,1644]" italics="true" pageId="2" pageNumber="3">Paremphytus ostentus</emphasis>
Brues
</taxonomicName>
* 1908 (
<figureCitation id="13562A3E1F0C2360FAC7D916FA24F9E7" box="[1333,1406,1620,1644]" captionStart="Figure 2" captionStartId="4.[129,194,1859,1883]" captionTargetBox="[132,1469,152,1828]" captionTargetId="figure-9@4.[129,1473,149,1831]" captionTargetPageId="4" captionText="Figure 2. Cenozoic extinct Blasticotominae.AC* Enspeletoma oligocaenica. Holotype* NHMMZ PE 1995/9108-LS* sex unknown* Upper Oligocene of Enspel* Germany.A* dorsal view.B* C* line drawings of forewing* reconstructed from both preserved wings* with veins labelled (B) and with cells labelled (C). D* E* Paremphytus ostentus. Holotype No. 2042* Museum of Comparative Zoology* Harvard University* Cambridge* MA* USA; female* uppermost Eocene* Colorado* USA. D* dorsal view.E* line drawing (Zhelochovtzev and Rasnitsyn 1972). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.14542359" httpUri="https://zenodo.org/record/14542359/files/figure.png" pageId="2" pageNumber="3">Fig. 2D</figureCitation>
* E)* in having a huge pterostigma and a very short cell 2mcu (
<figureCitation id="13562A3E1F0C2360FABFD931FACBF900" box="[1357,1425,1651,1675]" captionStart="Figure 2" captionStartId="4.[129,194,1859,1883]" captionTargetBox="[132,1469,152,1828]" captionTargetId="figure-9@4.[129,1473,149,1831]" captionTargetPageId="4" captionText="Figure 2. Cenozoic extinct Blasticotominae.AC* Enspeletoma oligocaenica. Holotype* NHMMZ PE 1995/9108-LS* sex unknown* Upper Oligocene of Enspel* Germany.A* dorsal view.B* C* line drawings of forewing* reconstructed from both preserved wings* with veins labelled (B) and with cells labelled (C). D* E* Paremphytus ostentus. Holotype No. 2042* Museum of Comparative Zoology* Harvard University* Cambridge* MA* USA; female* uppermost Eocene* Colorado* USA. D* dorsal view.E* line drawing (Zhelochovtzev and Rasnitsyn 1972). Scale bars:1 mm." figureDoi="http://doi.org/10.5281/zenodo.14542359" httpUri="https://zenodo.org/record/14542359/files/figure.png" pageId="2" pageNumber="3">Fig. 2B</figureCitation>
* C).
</paragraph>
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