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<document id="0BC3D61FD1E38B54E49658046B12D9C3" ID-CLB-Dataset="6186" ID-DOI="10.11646/phytotaxa.28.1.2" ID-GBIF-Dataset="1d365dfe-b498-4540-aa83-c246f1c5a06e" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622648641615" checkinUser="felipe" docAuthor="Heatubun, Charlie D." docDate="2011" docId="03C4695899183A24FF76FBCCFF50F919" docLanguage="en" docName="Phytotaxa.28.6-26.pdf.imf" docOrigin="Phytotaxa 28" docSource="http://biotaxa.org/Phytotaxa/article/view/phytotaxa.28.1.2" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Areca riparia Heatubun 2011, sp. nov." docType="treatment" docVersion="7" lastPageNumber="21" masterDocId="FFFD112099163A2BFFE1FFDDFF8AFFFA" masterDocTitle="Seven New Species of Areca (Arecaceae)" masterLastPageNumber="26" masterPageNumber="6" pageNumber="20" updateTime="1698998193322" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="065DEEDAE32596850928A4391FB3F948" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0095C8BB1FD8CD8E57F1D06DC4479137">
<mods:title id="28FE7051BEBB813F5EF62B4F714888F1">Seven New Species of Areca (Arecaceae)</mods:title>
</mods:titleInfo>
<mods:name id="01D3A9F73DC2FA7CA952C45DF2672A21" type="personal">
<mods:role id="B7B5D1100E8CE4A79312CD9D764D2193">
<mods:roleTerm id="C730823670C7020625DC35B265F7AA0F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E8B9EAB41D95488441BDD2FFE5EF4C0">Heatubun, Charlie D.</mods:namePart>
<mods:affiliation id="B731B39907241E4218B057D1AD61D58B">Fakultas Kehutanan, Universitas Papua, Jl. Gunung Salju, Amban, Manokwari 98314, Papua Barat, Indonesia &amp; Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AB, United Kingdom. E-mail: charlie _ deheatboen @ yahoo. com</mods:affiliation>
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<mods:typeOfResource id="414BDDE209887535D437DC8198375615">text</mods:typeOfResource>
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<mods:title id="3F32ECC57E3826B5F5C51DDC06A2368F">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="0734A08F23505A0A5287E23424A85FC7">
<mods:date id="9393C8EB34D9F2DDD826A01E1EA7BA69">2011</mods:date>
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<mods:number id="7E65615AFF6B4333F80590E8BAA88280">2011-09-14</mods:number>
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<mods:number id="589D3994ADFA7E605139A05717B4DEBD">28</mods:number>
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<mods:location id="EB1AEE21EC368E4E78CC4DF38D205159">
<mods:url id="65AFD569B1907002215DD3666C6EBE54">http://biotaxa.org/Phytotaxa/article/view/phytotaxa.28.1.2</mods:url>
</mods:location>
<mods:classification id="EAAB8AEBA7EE91828F4BFF499F5601D7">journal article</mods:classification>
<mods:identifier id="E5ADC85D7AB68C71A6CFF9F2E5E3C52A" type="CLB-Dataset">6186</mods:identifier>
<mods:identifier id="A04D374344C77AB5D3D7698D66D4A7E8" type="DOI">10.11646/phytotaxa.28.1.2</mods:identifier>
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<treatment id="03C4695899183A24FF76FBCCFF50F919" ID-DOI="http://doi.org/10.5281/zenodo.4923805" ID-GBIF-Taxon="182645066" ID-Zenodo-Dep="4923805" LSID="urn:lsid:plazi:treatment:03C4695899183A24FF76FBCCFF50F919" httpUri="http://treatment.plazi.org/id/03C4695899183A24FF76FBCCFF50F919" lastPageId="15" lastPageNumber="21" pageId="14" pageNumber="20">
<subSubSection id="C3778BC599183A25FF76FBCCFDC1FBD7" box="[151,587,1040,1069]" pageId="14" pageNumber="20" type="nomenclature">
<paragraph id="8BD2D84E99183A25FF76FBCCFDC1FBD7" blockId="14.[151,1436,1040,1229]" box="[151,587,1040,1069]" pageId="14" pageNumber="20">
<heading id="D09A6F2299183A25FF76FBCCFDC1FBD7" bold="true" box="[151,587,1040,1069]" fontSize="12" level="1" pageId="14" pageNumber="20" reason="1">
<emphasis id="B919045C99183A25FF76FBCCFDC1FBD7" bold="true" box="[151,587,1040,1069]" pageId="14" pageNumber="20">
<taxonomicName id="4C6DA3CD99183A25FF76FBCCFE51FBD7" ID-CoL="5VRGJ" authority="Heatubun" authorityName="Heatubun" authorityYear="2011" box="[151,475,1040,1069]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="14" pageNumber="20" phylum="Tracheophyta" rank="species" species="riparia" status="sp. nov.">
<emphasis id="B919045C99183A25FF76FBCCFEC2FBD7" bold="true" box="[151,328,1041,1069]" italics="true" pageId="14" pageNumber="20">Areca riparia</emphasis>
Heatubun
</taxonomicName>
,
<emphasis id="B919045C99183A25FE09FBCCFDC1FBD7" bold="true" box="[488,587,1041,1069]" italics="true" pageId="14" pageNumber="20">
<taxonomicNameLabel id="A22AB92799183A25FE09FBCCFDC1FBD7" box="[488,587,1041,1069]" pageId="14" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599183A25FF76FBEBFEE1FB77" pageId="14" pageNumber="20" type="diagnosis">
<paragraph id="8BD2D84E99183A25FF76FBEBFEE1FB77" blockId="14.[151,1436,1040,1229]" pageId="14" pageNumber="20">
<emphasis id="B919045C99183A25FF76FBEBFEE1FB77" italics="true" pageId="14" pageNumber="20">
<taxonomicName id="4C6DA3CD99183A25FF76FBEBFE77FBB7" authority="Cambodianis" authorityName="Cambodianis" box="[151,509,1078,1101]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="14" pageNumber="20" phylum="Tracheophyta" rank="subSpecies" species="ceteris" subSpecies="speciebus">A ceteris speciebus Cambodianis</taxonomicName>
distinctissima; palma
<taxonomicName id="4C6DA3CD99183A25FD15FBEBFCFFFBB7" authorityName="Dransfield" authorityYear="1984" box="[756,885,1078,1101]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="14" pageNumber="20" phylum="Tracheophyta" rank="species" species="rheophytica">rheophytica</taxonomicName>
,
<taxonomicName id="4C6DA3CD99183A25FC63FBEBFB38FBB7" authority="Roxb. ex Buch." authorityName="Buch." box="[898,1202,1078,1101]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="14" pageNumber="20" phylum="Tracheophyta" rank="species" species="triandrae">A. triandrae Roxb. ex Buch.</taxonomicName>
-Ham. similis, foliolis angustissimis semper simplicem plicatis, inflorescentiis in 2 ordines ramosis, floribus femineis semper singulis in rachilla differt.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599183A24FF76FB4BFDFFFEB6" lastPageId="15" lastPageNumber="21" pageId="14" pageNumber="20" type="materials_examined">
<paragraph id="8BD2D84E99183A25FF76FB4BFE04FB37" blockId="14.[151,1436,1040,1229]" pageId="14" pageNumber="20">
<materialsCitation id="3B05D21399183A25FF76FB4BFE00FB37" ID-GBIF-Occurrence="3128532315" collectingDate="2008-09-27" collectionCode="K" country="Cambodia" latitude="11.583333" location="Tatai Chveng" longLatPrecision="1295" longitude="103.1" municipality="Type" pageId="14" pageNumber="20" specimenCount="1" stateProvince="Koh Kong" typeStatus="holotype">
<collectingMunicipality id="6BB6423499183A25FF76FB4BFF45FB57" box="[151,207,1174,1197]" pageId="14" pageNumber="20">Type</collectingMunicipality>
:––
<collectingCountry id="F37A98DE99183A25FF17FB4BFE00FB57" box="[246,394,1174,1197]" name="Cambodia" pageId="14" pageNumber="20">CAMBODIA</collectingCountry>
.
<collectingRegion id="49A916AC99183A25FE76FB4BFDFEFB57" box="[407,628,1174,1197]" country="Cambodia" name="Koh Kong" pageId="14" pageNumber="20">Koh Kong Province</collectingRegion>
:
<location id="8EB28E9599183A25FD61FB4BFC9AFB57" LSID="urn:lsid:plazi:treatment:03C4695899183A24FF76FBCCFF50F919:8EB28E9599183A25FD61FB4BFC9AFB57" box="[640,784,1174,1197]" country="Cambodia" latitude="11.583333" longLatPrecision="1295" longitude="103.1" municipality="Type" name="Tatai Chveng" pageId="14" pageNumber="20" stateProvince="Koh Kong">Tatai Chveng</location>
waterfall,
<geoCoordinate id="EE59BE8999183A25FC68FB4BFC64FB57" box="[905,1006,1174,1197]" degrees="11" direction="north" minutes="35" orientation="latitude" pageId="14" pageNumber="20" precision="925" value="11.583333">11º 35' N</geoCoordinate>
<geoCoordinate id="EE59BE8999183A25FC16FB4BFBE3FB57" box="[1015,1129,1174,1197]" degrees="103" direction="east" minutes="06" orientation="longitude" pageId="14" pageNumber="20" precision="925" value="103.1">103º 06' E</geoCoordinate>
,
<date id="FFD3FE8E99183A25FB99FB4BFAC4FB57" box="[1144,1358,1174,1197]" pageId="14" pageNumber="20" value="2008-09-27">
<collectingDate id="EF97076699183A25FB99FB4BFAC4FB57" box="[1144,1358,1174,1197]" pageId="14" pageNumber="20" value="2008-09-27">27 September 2008</collectingDate>
</date>
,
<emphasis id="B919045C99183A25FABAFB4BFF65FB37" italics="true" pageId="14" pageNumber="20">Evans 174</emphasis>
(
<typeStatus id="54D666EC99183A25FF1CFB6BFED6FB37" box="[253,348,1206,1229]" pageId="14" pageNumber="20" type="holotype">holotype</typeStatus>
<collectionCode id="ED7C408B99183A25FE83FB6BFEF6FB37" box="[354,380,1206,1229]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="14" pageNumber="20" type="Herbarium">K</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD2D84E99183A24FF76FB20FDFFFEB6" blockId="14.[151,1436,1277,2038]" lastBlockId="15.[151,1436,152,1763]" lastPageId="15" lastPageNumber="21" pageId="14" pageNumber="20">
Clustering, small and slender palm, snaking, rarely vertical. Stem to
<quantity id="4C9575AB99183A25FC31FB20FB9EFAED" box="[976,1044,1277,1303]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="20" unit="m" value="2.5">2.5 m</quantity>
tall,
<quantity id="4C9575AB99183A25FBB2FB20FB4EFAED" box="[1107,1220,1277,1303]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="14" pageNumber="20" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1−1.5 cm</quantity>
diam.; internodes
<quantity id="4C9575AB99183A25FF76FAF9FE82FAC4" box="[151,264,1316,1342]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="14" pageNumber="20" unit="cm" value="2.75" valueMax="3.0" valueMin="2.5">2.5−3 cm</quantity>
long, green near the crown, shiny, brown near the base, nodal scars conspicuous, white to whitish brown. Leaves about
<quantity id="4C9575AB99183A25FE74FA97FE48FA9E" box="[405,450,1354,1380]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="14" pageNumber="20" unit="in" value="7.0">7 in</quantity>
the crown, pinnate, to
<quantity id="4C9575AB99183A25FD30FA97FC90FA9E" box="[721,794,1354,1380]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="20" unit="cm" value="80.0">80 cm</quantity>
long (including petiole); sheath tubular, ca. 19 ×
<quantity id="4C9575AB99183A25FA83FA97FA16FA9E" box="[1378,1436,1354,1380]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="20" unit="cm" value="4.0">4 cm</quantity>
long, smooth, greenish cream, speckled with reddish scurf-like ramenta; crown shaft well defined,
<quantity id="4C9575AB99183A25FAC4FAACFA16FA71" box="[1317,1436,1393,1419]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="14" pageNumber="20" unit="cm" value="29.0" valueMax="30.0" valueMin="28.0">28−30 cm</quantity>
long and
<quantity id="4C9575AB99183A25FEE6FA45FEE8FA48" box="[263,354,1432,1458]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="14" pageNumber="20" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1−2 cm</quantity>
diam.; petiole short,
<quantity id="4C9575AB99183A25FD82FA45FD56FA48" box="[611,732,1432,1458]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="14" pageNumber="20" unit="cm" value="12.5" valueMax="13.0" valueMin="12.0">12−13 cm</quantity>
long, 5 ×
<quantity id="4C9575AB99183A25FCB8FA45FC14FA48" box="[857,926,1432,1458]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="20" unit="mm" value="5.0">5 mm</quantity>
at the base, channelled adaxially, rounded abaxially; rachis somewhat arching, very slender with adaxial longitudinal ridge, rounded abaxially, covered by rusty brown indumenta; blade with regularly arranged leaflets, about 25 leaflets on each side; leaflets somewhat arching, single fold, linear,
<quantity id="4C9575AB99183A25FD81F9D1FD52F9DC" box="[608,728,1548,1574]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="14" pageNumber="20" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20−30 cm</quantity>
long, acuminate, with pointed tips, papery, green adaxially and light green abaxially, slightly discolorous when dried; continuous white woolly ramenta present along the mid-ribs on abaxial surface. Inflorescence infrafoliar, erect, small and slender,
<quantity id="4C9575AB99183A25FBAFF984FB4DF989" box="[1102,1223,1625,1651]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="14" pageNumber="20" unit="cm" value="14.0" valueMax="15.0" valueMin="13.0">13−15 cm</quantity>
long and
<quantity id="4C9575AB99183A25FAA0F984FA16F989" box="[1345,1436,1625,1651]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="14" pageNumber="20" unit="cm" value="7.0" valueMax="8.0" valueMin="6.0">6−8 cm</quantity>
wide, protandrous, branching to 1 order and sometimes the basalmost rachilla branches to 2 order; peduncle
<quantity id="4C9575AB99183A25FF76F97BFE8CF93A" box="[151,262,1702,1728]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="14" pageNumber="20" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1−1.5 cm</quantity>
long; prophyll ca. 14 ×
<quantity id="4C9575AB99183A25FDC5F97BFDFDF93A" box="[548,631,1702,1728]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="20" unit="cm" value="4.5">4.5 cm</quantity>
, lanceolate, 2-keeled, papery, cream coloured, entirely enclosing the inflorescence, then splitting longitudinally and falling before staminate flower anthesis; rachis cream to greenish; rachillae ca. 16, divaricate, elongate, yellowish. Floral clusters uniseriate, only one complete triad including female flower occurring at the base of each rachilla, rarely several triads. Staminate flowers small, triangular, ca. 2 ×
<quantity id="4C9575AB99183A25FE8BF89CFE5BF8A1" box="[362,465,1857,1883]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="20" unit="mm" value="0.75">0.75 mm</quantity>
(young stage), asymmetric, cream to pale brown; calyx low, triangular cup-shaped, ca.
<quantity id="4C9575AB99183A25FF5EF8B5FEA2F878" box="[191,296,1896,1922]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="20" unit="mm" value="0.75">0.75 mm</quantity>
wide and
<quantity id="4C9575AB99183A25FE41F8B5FE73F878" box="[416,505,1896,1922]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="20" unit="mm" value="0.5">0.5 mm</quantity>
high (at young stage), 3-lobed; petals 3, strongly keeled, valvate, ca. 1.5 ×
<quantity id="4C9575AB99183A25FA89F8B5FF4FF852" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="20" unit="mm" value="0.75">0.75 mm</quantity>
(at young stage); stamens 3, about
<quantity id="4C9575AB99183A25FD95F853FD33F852" box="[628,697,1934,1960]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="20" unit="mm" value="1.0">1 mm</quantity>
long (at young stage), basifixed; anthers ca.
<quantity id="4C9575AB99183A25FB3BF853FABCF852" box="[1242,1334,1934,1960]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="20" unit="mm" value="0.8">0.8 mm</quantity>
long (at young stage), white to cream-coloured, longer than the filaments; filaments ca.
<quantity id="4C9575AB99183A25FB89F868FB4FF835" box="[1128,1221,1973,1999]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="20" unit="mm" value="0.2">0.2 mm</quantity>
(at young stage), orange to brown; pistillode lacking. Pistillate flowers larger than the staminate, triangular, ca.
<quantity id="4C9575AB99183A25FAE2F801FAD4F80C" box="[1283,1374,2012,2038]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="14" pageNumber="20" unit="mm" value="6.2">6.2 mm</quantity>
long and
<quantity id="4C9575AB99193A24FF29FF45FE81FF48" box="[200,267,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="21" unit="mm" value="4.0">4 mm</quantity>
wide (at young stage); sepals 3, strongly imbricate, ca. 4 ×
<quantity id="4C9575AB99193A24FC23FF45FB8FFF48" box="[962,1029,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="21" unit="mm" value="3.0">3 mm</quantity>
(at young stage), triangular; petals 3, imbricate, triangular, ca. 5 ×
<quantity id="4C9575AB99193A24FDD5FF63FDF6FF22" box="[564,636,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="21" unit="mm" value="3.0">3 mm</quantity>
at anthesis; gynoecium about
<quantity id="4C9575AB99193A24FBE8FF63FBDBFF22" box="[1033,1105,190,216]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="21" unit="mm" value="3.0">3 mm</quantity>
(at young stage), lacking staminodes. Fruits 2–2.8 × 1–1.5 ×
<quantity id="4C9575AB99193A24FDA3FF38FD3EFF05" box="[578,692,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="15" pageNumber="21" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 cm</quantity>
, elongate, fusiform, apical stigmatic remains, epicarp smooth, mesocarp fleshy, endocarp fibrous, bright crimson when ripe. Seeds ca. 1.6 ×
<quantity id="4C9575AB99193A24FBA4FED1FB10FEDC" box="[1093,1178,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="21" unit="cm" value="1.2">1.2 cm</quantity>
, obovoid, endosperm ruminate, embryo basal. (
<figureCitation id="1356C4CB99193A24FE20FEEFFDBBFEB6" box="[449,561,306,332]" captionStart="FIGURE 8" captionStartId="16.[151,255,1861,1885]" captionTargetBox="[179,1406,206,1830]" captionTargetId="figure-83@16.[172,1415,194,1840]" captionTargetPageId="16" captionText="FIGURE 8. Areca riparia. A. Stem, leaf and inflorescence. B. Detail of rachilla showing staminate and pistillate flowers in bud. C. Staminate flower in section. D. Pistillate flower. E, F. Fruit whole and in section. Scale bar: A = 4 cm; B = 7 mm; C = 1 mm; D = 3 mm; E, F = 1 cm. All from Evans 174. Drawn by Lucy T. Smith." figureDoi="http://doi.org/10.5281/zenodo.4894326" httpUri="https://zenodo.org/record/4894326/files/figure.png" pageId="15" pageNumber="21">Figures 8</figureCitation>
&amp;
<figureCitation id="1356C4CB99193A24FDB7FEEFFDEDFEB6" box="[598,615,306,332]" captionStart="FIGURE 9" captionStartId="17.[151,255,1921,1945]" captionTargetBox="[151,1436,186,1899]" captionTargetId="figure-74@17.[151,1436,186,1899]" captionTargetPageId="17" captionText="FIGURE 9. Areca riparia in its natural habitat in Cambodia. A. Close up view of the crown with leaves and leaf sheath, inflorescence and the ripe red fruits. B. Areca riparia surrounded by other riverside vegetation. C. The flowing rocky river bank is the habitat of Areca riparia. All photo by Tom D. Evans." figureDoi="http://doi.org/10.5281/zenodo.4894328" httpUri="https://zenodo.org/record/4894328/files/figure.png" pageId="15" pageNumber="21">9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599193A24FF27FE85FAFBFE1D" pageId="15" pageNumber="21" type="distribution">
<paragraph id="8BD2D84E99193A24FF27FE85FD17FE60" blockId="15.[151,1436,152,1763]" pageId="15" pageNumber="21">
<emphasis id="B919045C99193A24FF27FE85FEE3FE88" bold="true" box="[198,361,344,370]" pageId="15" pageNumber="21">Distribution:</emphasis>
—This palm is only known from one collection made by Dr. Tom Evans in Tatai Chveng waterfall in
<collectingRegion id="49A916AC99193A24FEC5FE5DFD9BFE60" box="[292,529,384,410]" country="Cambodia" name="Koh Kong" pageId="15" pageNumber="21">Koh Kong Province</collectingRegion>
,
<collectingCountry id="F37A98DE99193A24FDFCFE5DFD13FE60" box="[541,665,384,410]" name="Cambodia" pageId="15" pageNumber="21">Cambodia</collectingCountry>
.
</paragraph>
<paragraph id="8BD2D84E99193A24FF27FE7BFAFBFE1D" blockId="15.[151,1436,152,1763]" pageId="15" pageNumber="21">
<emphasis id="B919045C99193A24FF27FE7BFEBAFE3A" bold="true" box="[198,304,422,448]" pageId="15" pageNumber="21">Habitat:</emphasis>
––This species grows as a rheophyte in a wet, vegetated rocky river bank at elevation less than
<quantity id="4C9575AB99193A24FF76FE10FF59FE1D" box="[151,211,461,487]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="21" unit="m" value="20.0">20 m</quantity>
above sea level, downstream of the falls and outside the splash zone, but below the peak flood level.
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599193A24FF27FE2EFEA4FD78" pageId="15" pageNumber="21" type="description">
<paragraph id="8BD2D84E99193A24FF27FE2EFD1DFDF4" blockId="15.[151,1436,152,1763]" box="[198,663,499,526]" pageId="15" pageNumber="21">
<emphasis id="B919045C99193A24FF27FE2EFE52FDF7" bold="true" box="[198,472,499,525]" pageId="15" pageNumber="21">Local Name and uses:</emphasis>
—Not recorded.
</paragraph>
<paragraph id="8BD2D84E99193A24FF27FDC7FEA4FD78" blockId="15.[151,1436,152,1763]" pageId="15" pageNumber="21">
<emphasis id="B919045C99193A24FF27FDC7FE46FDCE" bold="true" box="[198,460,538,564]" pageId="15" pageNumber="21">Conservation Status:</emphasis>
—Data Deficient. Further population studies are needed to assess the conservation status of this palm, although the collector had stated that this palm is common on the river system near the
<typeStatus id="54D666EC99193A24FF76FDB5FF43FD78" box="[151,201,616,642]" pageId="15" pageNumber="21">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599193A24FF27FD53FBE5FD52" box="[198,1135,654,680]" pageId="15" pageNumber="21" type="etymology">
<paragraph id="8BD2D84E99193A24FF27FD53FBE5FD52" blockId="15.[151,1436,152,1763]" box="[198,1135,654,680]" pageId="15" pageNumber="21">
<emphasis id="B919045C99193A24FF27FD53FEDDFD52" bold="true" box="[198,343,654,680]" pageId="15" pageNumber="21">Etymology:</emphasis>
—The specific epithet reflects the habitat where this species grows.
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599193A24FF27FD68FAD5F995" pageId="15" pageNumber="21" type="discussion">
<paragraph id="8BD2D84E99193A24FF27FD68FD1BFA9A" blockId="15.[151,1436,152,1763]" pageId="15" pageNumber="21">
<emphasis id="B919045C99193A24FF27FD68FEDEFD35" bold="true" box="[198,340,693,719]" pageId="15" pageNumber="21">Discussion:</emphasis>
––
<taxonomicName id="4C6DA3CD99193A24FE90FD68FD98FD34" authorityName="Heatubun" authorityYear="2011" box="[369,530,693,718]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FE90FD68FD98FD34" box="[369,530,693,718]" italics="true" pageId="15" pageNumber="21">Areca riparia</emphasis>
</taxonomicName>
is the third species of
<taxonomicName id="4C6DA3CD99193A24FCC5FD68FCE3FD34" authorityName="Linnaeus (1753:" authorityYear="1189" box="[804,873,693,718]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="B919045C99193A24FCC5FD68FCE3FD34" box="[804,873,693,718]" italics="true" pageId="15" pageNumber="21">Areca</emphasis>
</taxonomicName>
that has been reported from
<collectingCountry id="F37A98DE99193A24FB22FD68FAB6FD35" box="[1219,1340,693,719]" name="Cambodia" pageId="15" pageNumber="21">Cambodia</collectingCountry>
and the fifth for Indo-China (
<bibRefCitation id="EFFCA5BF99193A24FE6FFD01FD54FD0C" author="Govaerts, R. &amp; Dransfield, J." box="[398,734,732,758]" pageId="15" pageNumber="21" refId="ref10692" refString="Govaerts, R. &amp; Dransfield, J. (2005) World checklist of palms. The Board of Trustees of the Royal Botanic Gardens, Kew, 223 pp." type="book" year="2005">Govaerts &amp; Dransfield 2005</bibRefCitation>
,
<bibRefCitation id="EFFCA5BF99193A24FD0BFD01FC27FD0C" author="Henderson, A." box="[746,941,732,758]" pageId="15" pageNumber="21" refId="ref10750" refString="Henderson, A. (2009) Field guide to the palms of southern Asia. Princeton University Press, Princeton, NJ, USA, 199 pp." type="book" year="2009">Henderson 2009</bibRefCitation>
,
<bibRefCitation id="EFFCA5BF99193A24FC58FD01FB4AFD0C" author="Henderson, A. &amp; Ban, N. K. &amp; Thanh, B. V." box="[953,1216,732,758]" pageId="15" pageNumber="21" pagination="34 - 40" refId="ref10779" refString="Henderson, A., Ban, N. K. &amp; Thanh, B. V. (2010) New species of Areca, Pinanga, and Licuala (Arecaceae) from Vietnam. Phytotaxa 8: 34 - 40." type="journal article" year="2010">
Henderson
<emphasis id="B919045C99193A24FBDEFD01FBF3FD0F" box="[1087,1145,732,757]" italics="true" pageId="15" pageNumber="21">et al.</emphasis>
2010
</bibRefCitation>
). This palm differs from other
<taxonomicName id="4C6DA3CD99193A24FEC9FCDFFE5CFCE6" box="[296,470,770,796]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B919045C99193A24FEC9FCDFFEFBFCE1" box="[296,369,770,795]" italics="true" pageId="15" pageNumber="21">Areca</emphasis>
species
</taxonomicName>
in the region by its growth-form being truly rheophytic (
<bibRefCitation id="EFFCA5BF99193A24FB59FCDFFA1DFCE6" author="Van Steenis, C. G. G. J." box="[1208,1431,770,796]" pageId="15" pageNumber="21" refId="ref11026" refString="Van Steenis, C. G. G. J. (1981) Rheophytes of the world. An account of flood-resistant flowering plants and ferns and the theory of autonomous evolution. Sijthoff and Noorhoff, Alphen aan den Rijn. 417 pp." type="book" year="1981">van Steenis 1981</bibRefCitation>
,
<bibRefCitation id="EFFCA5BF99193A24FF76FCF4FE10FCB9" author="Dransfield, J. &amp; Uhl, N. W. &amp; Asmussen, C. B. &amp; Baker, W. J. &amp; Harley, M. M. &amp; Lewis, C. E." box="[151,410,809,835]" pageId="15" pageNumber="21" refId="ref10598" refString="Dransfield, J., Uhl, N. W., Asmussen, C. B., Baker, W. J., Harley, M. M. &amp; Lewis, C. E. (2008) Genera Palmarum: The evolution and classification of palms. Royal Botanic Gardens, Kew, 732 pp." type="book" year="2008">
Dransfield
<emphasis id="B919045C99193A24FEFAFCF4FEDCFCB8" box="[283,342,809,834]" italics="true" pageId="15" pageNumber="21">et al.</emphasis>
2008
</bibRefCitation>
) in the flood zone of fast-flowing rocky rivers. This species has three stamens, similar to
<taxonomicName id="4C6DA3CD99193A24FF57FC8DFD6AFC90" authority="Henderson et al. (2010: 34)" authorityName="Henderson" authorityPageNumber="34" authorityYear="2010" box="[182,736,848,874]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="songthanhensis">
<emphasis id="B919045C99193A24FF57FC8DFE07FC93" box="[182,397,848,873]" italics="true" pageId="15" pageNumber="21">A. songthanhensis</emphasis>
<bibRefCitation id="EFFCA5BF99193A24FE74FC8DFD6AFC90" author="Henderson, A. &amp; Ban, N. K. &amp; Thanh, B. V." box="[405,736,848,874]" pageId="15" pageNumber="21" pagination="34 - 40" refId="ref10779" refString="Henderson, A., Ban, N. K. &amp; Thanh, B. V. (2010) New species of Areca, Pinanga, and Licuala (Arecaceae) from Vietnam. Phytotaxa 8: 34 - 40." type="journal article" year="2010">
Henderson
<emphasis id="B919045C99193A24FDFCFC8DFDD2FC93" box="[541,600,848,873]" italics="true" pageId="15" pageNumber="21">et al.</emphasis>
(2010: 34)
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6DA3CD99193A24FD0CFC8DFC4AFC90" authority="Roxb." authorityName="Roxb." box="[749,960,848,874]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="triandra">
<emphasis id="B919045C99193A24FD0CFC8DFCFBFC93" box="[749,881,848,873]" italics="true" pageId="15" pageNumber="21">A. triandra</emphasis>
Roxb.
</taxonomicName>
in
<bibRefCitation id="EFFCA5BF99193A24FC06FC8DFAE3FC90" author="Buchanan-Hamilton, F." box="[999,1385,848,874]" pageId="15" pageNumber="21" pagination="307 - 383" refId="ref10427" refString="Buchanan-Hamilton, F. (1826) Commentary on the Herbarium Amboinense. Memoirs of the Wernerian Natural History Society 5: 307 - 383." type="journal article" year="1826">Buchanan-Hamilton (1826: 310)</bibRefCitation>
and
<taxonomicName id="4C6DA3CD99193A24FF76FCAAFD9FFC6A" authority="Ridley (1907: 136)" authorityName="Ridley" authorityPageNumber="136" authorityYear="1907" box="[151,533,886,912]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="B919045C99193A24FF76FCAAFEAEFC6A" box="[151,292,887,912]" italics="true" pageId="15" pageNumber="21">A. montana</emphasis>
<bibRefCitation id="EFFCA5BF99193A24FECCFCABFD9FFC6A" author="Ridley, H." box="[301,533,886,912]" pageId="15" pageNumber="21" refId="ref10964" refString="Ridley, H. (1907) Materials for a Flora of the Malayan Peninsula. Part II. Methodist Publishing House, Singapore, 235 pp." type="book" year="1907">Ridley (1907: 136)</bibRefCitation>
</taxonomicName>
—this last species was synonymized by Andrew Henderson in his recent monograph of palms of Southern Asia (
<bibRefCitation id="EFFCA5BF99193A24FD87FC40FCA7FC4D" author="Henderson, A." box="[614,813,925,951]" pageId="15" pageNumber="21" refId="ref10750" refString="Henderson, A. (2009) Field guide to the palms of southern Asia. Princeton University Press, Princeton, NJ, USA, 199 pp." type="book" year="2009">Henderson 2009</bibRefCitation>
), a taxonomic decision questioned here. However,
<taxonomicName id="4C6DA3CD99193A24FA62FC40FF60FC27" authorityName="Heatubun" authorityYear="2011" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FA62FC40FF60FC27" italics="true" pageId="15" pageNumber="21">A. riparia</emphasis>
</taxonomicName>
can easily be distinguished from
<emphasis id="B919045C99193A24FD93FC19FC51FC27" box="[626,987,964,989]" italics="true" pageId="15" pageNumber="21">
<taxonomicName id="4C6DA3CD99193A24FD93FC19FCC3FC27" box="[626,841,964,989]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="songthanhensis">A. songthanhensis</taxonomicName>
,
<taxonomicName id="4C6DA3CD99193A24FCB4FC19FC51FC27" authorityName="Ridley" authorityYear="1907" box="[853,987,964,989]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="montana">A. montana</taxonomicName>
</emphasis>
and
<taxonomicName id="4C6DA3CD99193A24FBF5FC19FB1CFC27" box="[1044,1174,964,989]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="triandra">
<emphasis id="B919045C99193A24FBF5FC19FB1CFC27" box="[1044,1174,964,989]" italics="true" pageId="15" pageNumber="21">A. triandra</emphasis>
</taxonomicName>
by having narrow and flexible single-folded leaflets adapted to flowing flood water. Additional morphological characters found in
<taxonomicName id="4C6DA3CD99193A24FA62FC37FEC5FBD0" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="songthanhensis">
<emphasis id="B919045C99193A24FA62FC37FEC5FBD0" italics="true" pageId="15" pageNumber="21">A. songthanhensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DA3CD99193A24FE6AFBCCFD9FFBD0" authorityName="Ridley" authorityYear="1907" box="[395,533,1041,1066]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="montana">
<emphasis id="B919045C99193A24FE6AFBCCFD9FFBD0" box="[395,533,1041,1066]" italics="true" pageId="15" pageNumber="21">A. montana</emphasis>
</taxonomicName>
, such as solitary habit and pistillate flowers borne only on distal rachillae, further distinguish this new species, which is clustering and bears pistillate flowers on all rachillae. Differences from
<taxonomicName id="4C6DA3CD99193A24FE86FB82FE63FB82" box="[359,489,1119,1144]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="triandra">
<emphasis id="B919045C99193A24FE86FB82FE63FB82" box="[359,489,1119,1144]" italics="true" pageId="15" pageNumber="21">A. triandra</emphasis>
</taxonomicName>
are in dimension and inflorescence structure in general.
<taxonomicName id="4C6DA3CD99193A24FB64FB82FAAFFB82" authorityName="Heatubun" authorityYear="2011" box="[1157,1317,1119,1144]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FB64FB82FAAFFB82" box="[1157,1317,1119,1144]" italics="true" pageId="15" pageNumber="21">Areca riparia</emphasis>
</taxonomicName>
is a small and slender palm with a stem diameter to
<quantity id="4C9575AB99193A24FD70FB58FD69FB65" box="[657,739,1157,1183]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="21" unit="cm" value="1.5">1.5 cm</quantity>
and stem height to
<quantity id="4C9575AB99193A24FC33FB58FB90FB65" box="[978,1050,1157,1183]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="21" unit="m" value="2.5">2.5 m</quantity>
, inflorescences branched to two orders, rachillae relatively few with only one pistillate flower on each rachilla. In contrast,
<taxonomicName id="4C6DA3CD99193A24FB16FB71FAF5FB3F" box="[1271,1407,1196,1221]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="triandra">
<emphasis id="B919045C99193A24FB16FB71FAF5FB3F" box="[1271,1407,1196,1221]" italics="true" pageId="15" pageNumber="21">A. triandra</emphasis>
</taxonomicName>
is typically medium-sized with a stem diameter to
<quantity id="4C9575AB99193A24FD28FB0FFC88FB16" box="[713,770,1234,1260]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="21" unit="cm" value="7.0">7 cm</quantity>
or more and stem height to
<quantity id="4C9575AB99193A24FBA9FB0FFBFCFB16" box="[1096,1142,1234,1260]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="21" unit="m" value="6.0">6 m</quantity>
, inflorescences branched to three orders, numerous rachillae and 3 to 5 (sometimes up to 10) pistillate flowers on each rachilla. Also the inflorescence of
<taxonomicName id="4C6DA3CD99193A24FE85FAFDFE61FAC3" box="[356,491,1312,1337]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="triandra">
<emphasis id="B919045C99193A24FE85FAFDFE61FAC3" box="[356,491,1312,1337]" italics="true" pageId="15" pageNumber="21">A. triandra</emphasis>
</taxonomicName>
also has a powerful lemon scent as emphasized by
<bibRefCitation id="EFFCA5BF99193A24FB87FAFDFA1DFAC0" author="Lim, C. K. &amp; Whitmore, T." box="[1126,1431,1312,1338]" pageId="15" pageNumber="21" pagination="12 - 24" refId="ref10845" refString="Lim, C. K. &amp; Whitmore, T. (2001) The endangered Areca palms of Peninsular Malaysia. Folia Malaysiana 2: 12 - 24." type="journal article" year="2001">Lim &amp; Whitmore (2001)</bibRefCitation>
, which has not been recorded for
<taxonomicName id="4C6DA3CD99193A24FDF6FA9AFD00FA9A" authorityName="Heatubun" authorityYear="2011" box="[535,650,1351,1376]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FDF6FA9AFD00FA9A" box="[535,650,1351,1376]" italics="true" pageId="15" pageNumber="21">A. riparia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD2D84E99193A24FF27FAB0FAD5F995" blockId="15.[151,1436,152,1763]" pageId="15" pageNumber="21">
A rheophytic form of
<taxonomicName id="4C6DA3CD99193A24FE2AFAB0FDC7FA7C" box="[459,589,1389,1414]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="triandra">
<emphasis id="B919045C99193A24FE2AFAB0FDC7FA7C" box="[459,589,1389,1414]" italics="true" pageId="15" pageNumber="21">A. triandra</emphasis>
</taxonomicName>
has been reported recently from
<collectingCountry id="F37A98DE99193A24FC36FAB0FBB0FA7D" box="[983,1082,1389,1415]" name="Vietnam" pageId="15" pageNumber="21">Vietnam</collectingCountry>
(
<bibRefCitation id="EFFCA5BF99193A24FBABFAB0FA9AFA7D" author="Henderson, A." box="[1098,1296,1389,1415]" pageId="15" pageNumber="21" refId="ref10750" refString="Henderson, A. (2009) Field guide to the palms of southern Asia. Princeton University Press, Princeton, NJ, USA, 199 pp." type="book" year="2009">Henderson 2009</bibRefCitation>
,
<bibRefCitation id="EFFCA5BF99193A24FAFCFAB0FE91FA54" author="Henderson, A. &amp; Ban, N. K. &amp; Thanh, B. V." pageId="15" pageNumber="21" pagination="34 - 40" refId="ref10779" refString="Henderson, A., Ban, N. K. &amp; Thanh, B. V. (2010) New species of Areca, Pinanga, and Licuala (Arecaceae) from Vietnam. Phytotaxa 8: 34 - 40." type="journal article" year="2010">
Henderson
<emphasis id="B919045C99193A24FF76FA49FF59FA57" box="[151,211,1428,1453]" italics="true" pageId="15" pageNumber="21">et al.</emphasis>
2010
</bibRefCitation>
). One herbarium specimen collected by Andrew Henderson and his colleagues (
<emphasis id="B919045C99193A24FB37FA49FF58FA2E" italics="true" pageId="15" pageNumber="21">Henderson et al. 3623</emphasis>
) from a lowland area of
<collectingCountry id="F37A98DE99193A24FE13FA67FDDFFA2E" box="[498,597,1466,1492]" name="Vietnam" pageId="15" pageNumber="21">Vietnam</collectingCountry>
has been studied at Kew and is clearly not the same taxon as
<taxonomicName id="4C6DA3CD99193A24FAC0FA66FA1FFA2E" authorityName="Heatubun" authorityYear="2011" box="[1313,1429,1467,1492]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FAC0FA66FA1FFA2E" box="[1313,1429,1467,1492]" italics="true" pageId="15" pageNumber="21">A. riparia</emphasis>
</taxonomicName>
. In this specimen, the typical features of
<taxonomicName id="4C6DA3CD99193A24FD8CFA3CFD7AFA00" box="[621,752,1505,1530]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="triandra">
<emphasis id="B919045C99193A24FD8CFA3CFD7AFA00" box="[621,752,1505,1530]" italics="true" pageId="15" pageNumber="21">A. triandra</emphasis>
</taxonomicName>
can be observed, such as the relatively broad leaflets with multiple folds (at least two-folded with prominent nerves in the leaflet, in contrast to the single-folded, smooth leaflets without prominent nerves in
<taxonomicName id="4C6DA3CD99193A24FD55F9F2FCA1F9B2" authorityName="Heatubun" authorityYear="2011" box="[692,811,1583,1608]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FD55F9F2FCA1F9B2" box="[692,811,1583,1608]" italics="true" pageId="15" pageNumber="21">A. riparia</emphasis>
</taxonomicName>
), the relatively large inflorescence (about 2.5 times longer and wider than
<taxonomicName id="4C6DA3CD99193A24FE7FF988FD98F994" authorityName="Heatubun" authorityYear="2011" box="[414,530,1621,1646]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FE7FF988FD98F994" box="[414,530,1621,1646]" italics="true" pageId="15" pageNumber="21">A. riparia</emphasis>
</taxonomicName>
) that is branched to three orders, as opposed to two orders in
<taxonomicName id="4C6DA3CD99193A24FB05F988FAD2F994" authorityName="Heatubun" authorityYear="2011" box="[1252,1368,1621,1646]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FB05F988FAD2F994" box="[1252,1368,1621,1646]" italics="true" pageId="15" pageNumber="21">A. riparia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599193A24FF27F9A1FF50F919" pageId="15" pageNumber="21" type="reference_group">
<paragraph id="8BD2D84E99193A24FF27F9A1FF50F919" blockId="15.[151,1436,152,1763]" pageId="15" pageNumber="21">
<treatmentCitationGroup id="AB7DFF6099193A24FF27F9A1FF50F919" pageId="15" pageNumber="21">
<taxonomicName id="4C6DA3CD99193A24FF27F9A1FEEFF96F" ID-CoL="5VRGJ" authorityName="Heatubun" authorityYear="2011" box="[198,357,1660,1685]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B919045C99193A24FF27F9A1FEEFF96F" box="[198,357,1660,1685]" italics="true" pageId="15" pageNumber="21">Areca riparia</emphasis>
</taxonomicName>
is the second rheophytic species in the genus, the other species is
<treatmentCitation id="0ACCFE5F99193A24FB8CF9A1FE99F946" author="Dransfield, J." page=")" pageId="15" pageNumber="21" year="1984">
<taxonomicName id="4C6DA3CD99193A24FB8CF9A1FE99F946" ID-CoL="67RGK" authority="Dransfield (1984: 18)" authorityName="Dransfield" authorityPageNumber="18" authorityYear="1984" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="15" pageNumber="21" phylum="Tracheophyta" rank="species" species="rheophytica">
<emphasis id="B919045C99193A24FB8CF9A1FA92F96F" box="[1133,1304,1660,1685]" italics="true" pageId="15" pageNumber="21">A. rheophytica</emphasis>
<bibRefCitation id="EFFCA5BF99193A24FAC1F9A1FE99F946" author="Dransfield, J." pageId="15" pageNumber="21" pagination="1 - 22" refId="ref10479" refString="Dransfield, J. (1984) The genus Areca (Palmae: Arecoideae) in Borneo. Kew Bulletin 39: 1 - 22." type="journal article" year="1984">Dransfield (1984: 18)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
from Borneo. Besides being rheophytes, these two species are different from each other in many ways.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>
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