treatments-xml/data/03/C4/69/03C4695899163A2FFF73F8DBFD34FDF4.xml

<document id="FB32A7D6AE6D3D90DF184DDFAA76E867" ID-CLB-Dataset="6186" ID-DOI="10.11646/phytotaxa.28.1.2" ID-GBIF-Dataset="1d365dfe-b498-4540-aa83-c246f1c5a06e" ID-ISSN="1179-3163" ID-Zenodo-Dep="4894308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622648641615" checkinUser="felipe" docAuthor="Heatubun, Charlie D." docDate="2011" docId="03C4695899163A2FFF73F8DBFD34FDF4" docLanguage="en" docName="Phytotaxa.28.6-26.pdf.imf" docOrigin="Phytotaxa 28" docSource="http://biotaxa.org/Phytotaxa/article/view/phytotaxa.28.1.2" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Areca bakeri Heatubun 2011, sp. nov." docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="FFFD112099163A2BFFE1FFDDFF8AFFFA" masterDocTitle="Seven New Species of Areca (Arecaceae)" masterLastPageNumber="26" masterPageNumber="6" pageNumber="6" updateTime="1698998193322" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A3DF6FDA376EEA3014DBCCDC569705FC">Seven New Species of Areca (Arecaceae)</mods:title>
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<mods:namePart id="1A17B2760662BB2DB57283B965ECF61E">Heatubun, Charlie D.</mods:namePart>
<mods:affiliation id="491E8A3EB4FA9FACF4BBE01170BB449A">Fakultas Kehutanan, Universitas Papua, Jl. Gunung Salju, Amban, Manokwari 98314, Papua Barat, Indonesia &amp; Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AB, United Kingdom. E-mail: charlie _ deheatboen @ yahoo. com</mods:affiliation>
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<treatment id="03C4695899163A2FFF73F8DBFD34FDF4" ID-DOI="http://doi.org/10.5281/zenodo.4923795" ID-GBIF-Taxon="182645067" ID-Zenodo-Dep="4923795" LSID="urn:lsid:plazi:treatment:03C4695899163A2FFF73F8DBFD34FDF4" httpUri="http://treatment.plazi.org/id/03C4695899163A2FFF73F8DBFD34FDF4" lastPageId="4" lastPageNumber="10" pageId="0" pageNumber="6">
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<heading id="D09A6F2299163A2BFF73F8DBFDB5F8D9" bold="true" box="[146,575,1798,1827]" fontSize="12" level="1" pageId="0" pageNumber="6" reason="1">
<emphasis id="B919045C99163A2BFF73F8DBFDB5F8D9" bold="true" box="[146,575,1798,1827]" pageId="0" pageNumber="6">
<taxonomicName id="4C6DA3CD99163A2BFF73F8DBFE45F8D9" ID-CoL="5VRFG" authority="Heatubun" authorityName="Heatubun" authorityYear="2011" box="[146,463,1798,1827]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="0" pageNumber="6" phylum="Tracheophyta" rank="species" species="bakeri" status="sp. nov.">
<emphasis id="B919045C99163A2BFF73F8DBFEB6F8D8" bold="true" box="[146,316,1798,1826]" italics="true" pageId="0" pageNumber="6">Areca bakeri</emphasis>
Heatubun
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, 
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<emphasis id="B919045C99163A2BFE3DF8DBFDBDF8D8" bold="true" box="[476,567,1798,1826]" italics="true" pageId="0" pageNumber="6">sp. nov</emphasis>
.
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<taxonomicName id="4C6DA3CD99163A2BFF73F8F6FD8AF8B8" authority="Borneensibus" authorityName="Borneensibus" box="[146,512,1835,1858]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="0" pageNumber="6" phylum="Tracheophyta" rank="subSpecies" species="ceteris" subSpecies="speciebus">A ceteris speciebus Borneensibus</taxonomicName>
habitu acaule–caespitoso, petiolis longis 
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, inflorescentia erecta rigida, floribus staminatis distichisis, floribus foemineis singulariter dispositis distincta.
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<materialsCitation id="3B05D21399163A2BFF73F8B6FD38F858" ID-GBIF-Occurrence="3128532301" collectingDate="1996-04-22" collectionCode="K, KEP, SING, SAR" country="Malaysia" county="Baker" location="Type" municipality="Semengoh Forest" pageId="0" pageNumber="6" specimenCount="1" typeStatus="holotype">
<location id="8EB28E9599163A2BFF73F8B6FF4DF878" LSID="urn:lsid:plazi:treatment:03C4695899163A2FFF73F8DBFD34FDF4:8EB28E9599163A2BFF73F8B6FF4DF878" box="[146,199,1899,1922]" country="Malaysia" county="Baker" municipality="Semengoh Forest" name="Type" pageId="0" pageNumber="6">Type</location>
:––CULTIVATED. 
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, 
<collectingCountry id="F37A98DE99163A2BFE16F8B6FDDDF878" box="[503,599,1899,1922]" name="Malaysia" pageId="0" pageNumber="6">Sarawak</collectingCountry>
: 
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at the 
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<collectingMunicipality id="6BB6423499163A2BFCC0F8B6FC54F878" box="[801,990,1899,1922]" pageId="0" pageNumber="6">Semengoh Forest</collectingMunicipality>
Arboretum
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, 
<date id="FFD3FE8E99163A2BFB8DF8B6FA83F878" box="[1132,1289,1899,1922]" pageId="0" pageNumber="6" value="1996-04-22">
<collectingDate id="EF97076699163A2BFB8DF8B6FA83F878" box="[1132,1289,1899,1922]" pageId="0" pageNumber="6" value="1996-04-22">22 April 1996</collectingDate>
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, 
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et al. 716
</emphasis>
(
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<collectionCode id="ED7C408B99163A2BFEBCF856FEFDF858" box="[349,375,1931,1954]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="0" pageNumber="6" type="Herbarium">K</collectionCode>
!, isotypes 
<collectionCode id="ED7C408B99163A2BFE04F856FD9DF858" box="[485,535,1931,1954]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="0" pageNumber="6" type="Herbarium">KEP</collectionCode>
, 
<collectionCode id="ED7C408B99163A2BFDC0F856FDD5F858" box="[545,607,1931,1954]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="0" pageNumber="6" type="Herbarium">SING</collectionCode>
, 
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!)
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.
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<paragraph id="8BD2D84E99173A2AFF76FF45FDBBFB82" blockId="1.[151,1436,152,1926]" pageId="1" pageNumber="7">
Undergrowth palmlet densely clustering with ca. 20 stems in a clump to 
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in height. Stem subterranean to very short (to 
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), ca. 
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diam.; internodes very close (
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long), not conspicuous, covered by marcescent leaf sheaths. Leaves 
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crown, litter trapping, pinnate, 
<quantity id="4C9575AB99173A2AFC3DFF38FBFBFF05" box="[988,1137,229,255]" metricMagnitude="0" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="1" pageNumber="7" unit="cm" value="170.0" valueMax="200.0" valueMin="140.0">140−200 cm</quantity>
long (excluding petiole); sheath tubular, 
<quantity id="4C9575AB99173A2AFEADFED1FE49FEDC" box="[332,451,268,294]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.1" metricValueMin="2.0" pageId="1" pageNumber="7" unit="cm" value="25.5" valueMax="31.0" valueMin="20.0">20−31 cm</quantity>
long and 
<quantity id="4C9575AB99173A2AFDDBFED1FD1EFEDC" box="[570,660,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="7" unit="cm" value="3.5" valueMax="4.0" valueMin="3.0">3−4 cm</quantity>
wide, smooth, not fibrous, yellowish green; crown shaft not well defined, up to 
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long and up to 
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diam.; petiole 
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long, channelled adaxially, rounded abaxially; rachis ascending but not arching, with adaxial longitudinal ridge, rounded abaxially; blade with slightly irregularly arranged leaflets, 7−8 leaflets on each side; basal leaflets ca. 52 × 
<quantity id="4C9575AB99173A2AFB41FE5DFAA9FE60" box="[1184,1315,384,410]" metricMagnitude="-1" metricUnit="m" metricValue="1.02" metricValueMax="1.4" metricValueMin="0.64" pageId="1" pageNumber="7" unit="cm" value="10.2" valueMax="14.0" valueMin="6.4">6.4−14 cm</quantity>
, with 7−9 folds, sigmoid, the middle leaflets 57−59 × 
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, with 5−9 folds, slightly sigmoid and the terminal leaflets ca. 42 × 
<quantity id="4C9575AB99173A2AFE82FE10FE3BFE1D" box="[355,433,461,487]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="7" unit="cm" value="11.0">11 cm</quantity>
, with up to 11 folds, slightly sigmoid, tips pointed except for the terminal leaflets slightly oblique-lobed, papery, discolorous, darker adaxially than abaxially. Inflorescence infrafoliar, erect, bursting out among marcescent sheaths, 
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long and 
<quantity id="4C9575AB99173A2AFC8AFDC7FB8BFDCE" box="[875,1025,538,564]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="12.5" metricValueMin="4.5" pageId="1" pageNumber="7" unit="cm" value="8.5" valueMax="12.5" valueMin="4.5">4.5−12.5 cm</quantity>
wide, protandrous, branching to 1 order; prophyll elongated, up to 
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long, ca. 
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wide, two-keeled, leathery, cream, light green near the apex; peduncle 
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long, pale yellowish green; rachis yellowish green; rachis bracts caducous; rachillae 11−13, 
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long and 
<quantity id="4C9575AB99173A2AFE3EFD53FDE4FD52" box="[479,622,654,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.6" metricValueMin="3.5" pageId="1" pageNumber="7" unit="mm" value="4.55" valueMax="5.6" valueMin="3.5">3.5−5.6 mm</quantity>
wide, very stiff, stout and straight, slightly swollen and flattened near the base, pale green, elongate. Floral clusters distichous on rachillae, only one complete triad including female flower occurring at the base of each rachilla. Staminate flowers small, sessile, triangular, ca. 6.2 × 
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, asymmetric; sepals 3, low; petals 3, triangular, small, striate; stamens 6, small, anthers shorter than the filaments; filaments slender, elongate; pistillode ca. 2.5 × 1.0 mm, pointed. Pistillate flowers larger than the staminate, triangular, borne on the enlarged basal portion of rachillae, only one flower on each rachillae, buds varying greatly in size depending on stage of development, ca. 
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long and 
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wide in late anthesis; sepals 3, strongly imbricate, ca. 7 × 
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, triangular, asymmetrical, striate; petals 3, imbricate, triangular, ca. 12.5 × 
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, striate; gynoecium ca. 
<quantity id="4C9575AB99173A2AFD97FC19FD69FC24" box="[630,739,964,990]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="7" unit="mm" value="10.5">10.5 mm</quantity>
long and 
<quantity id="4C9575AB99173A2AFC81FC19FC2CFC24" box="[864,934,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="7" unit="mm" value="4.0">4 mm</quantity>
wide at the base; stigma ca. 
<quantity id="4C9575AB99173A2AFAEEFC19FADFFC24" box="[1295,1365,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="7" unit="mm" value="8.0">8 mm</quantity>
long, pointed with 3 lobes, split 
<quantity id="4C9575AB99173A2AFE34FC37FD90FBFE" box="[469,538,1002,1028]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="7" unit="mm" value="8.0">8 mm</quantity>
to the base; style ca. 
<quantity id="4C9575AB99173A2AFCFCFC37FCF2FBFE" box="[797,888,1002,1028]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="7" unit="mm" value="1.5">1.5 mm</quantity>
long; staminodes ca. 6, irregularly dentiform, 0.5−1 × 
<quantity id="4C9575AB99173A2AFF1FFBCCFE2EFBD1" box="[254,420,1041,1067]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="1" pageNumber="7" unit="mm" value="0.375" valueMax="0.5" valueMin="0.25">0.25−0.5 mm</quantity>
. Fruits elongate, sickle-shaped, 6.0−6.5 × 
<quantity id="4C9575AB99173A2AFC46FBCCFBB5FBD1" box="[935,1087,1041,1067]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="1" pageNumber="7" unit="cm" value="1.375" valueMax="1.45" valueMin="1.3">1.3−1.45 cm</quantity>
(young fruits), beak 
<quantity id="4C9575AB99173A2AFAA1FBCCFF43FBA8" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="7" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4−1.5 mm</quantity>
; epicarp smooth, shiny, dark green (young), mature fruits not known. Seed very young; endosperm sparsely ruminate. (
<figureCitation id="1356C4CB99173A2AFE9CFB83FE67FB82" box="[381,493,1118,1144]" captionStart="FIGURE 1" captionStartId="2.[151,257,1891,1915]" captionTargetBox="[159,1420,193,1854]" captionTargetId="figure-109@2.[155,1432,181,1871]" captionTargetPageId="2" captionText="FIGURE 1. Areca bakeri. A. Apical portion of leaf. B. Middle portion of leaf. C. Basal portion of leaf and petiole attached to crown shaft. D. Inflorescence attached to stem just below the crown shaft and showing acaulescent habit. E, F. Staminate flower whole and in section. G. Pistillate flower. Scale bar: A, B = 8 cm; C, D = 3 cm; E, F = 2.2 mm; G = 7 mm. A–D from Baker et al. 716; E–G from Mokhtar &amp; Othman S 44726. Drawn by Lucy T. Smith." figureDoi="http://doi.org/10.5281/zenodo.4894310" httpUri="https://zenodo.org/record/4894310/files/figure.png" pageId="1" pageNumber="7">Figures 1</figureCitation>
&amp; 
<figureCitation id="1356C4CB99173A2AFDF3FB83FDA9FB82" box="[530,547,1118,1144]" captionStart="FIGURE 2" captionStartId="3.[151,258,1916,1940]" captionTargetBox="[151,1436,182,1895]" captionTargetId="figure-85@3.[151,1436,182,1895]" captionTargetPageId="3" captionText="FIGURE 2. Areca bakeri. A. Acaulescent habit with densely suckering clump; the palm was mis-identified as A. jugahpunya, as shown on the label. B. Infructescence bursting out from the marcescent leaf sheaths. C. Prophyll, inflorescence, infructescence and seedling. All photos were taken from the specimen cultivated at Semengoh Forest Arboretum near Kuching, Sarawak, Malaysia. Photos: A (Charlie D. Heatubun), B–C (John Dransfield)." figureDoi="http://doi.org/10.5281/zenodo.4894312" httpUri="https://zenodo.org/record/4894312/files/figure.png" pageId="1" pageNumber="7">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599173A2AFF27FB55FE77F966" pageId="1" pageNumber="7" type="distribution">
<paragraph id="8BD2D84E99173A2AFF27FB55FB48FB33" blockId="1.[151,1436,152,1926]" pageId="1" pageNumber="7">
<emphasis id="B919045C99173A2AFF27FB55FEE3FB58" bold="true" box="[198,361,1160,1186]" pageId="1" pageNumber="7">Distribution:</emphasis>
—Only known from very limited collections from Miri (4 
<superScript id="7C18750699173A2AFBAEFB58FBD1FB69" attach="none" box="[1103,1115,1157,1171]" fontSize="6" pageId="1" pageNumber="7">th</superScript>
) Division of 
<collectingCountry id="F37A98DE99173A2AFAF0FB55FAFCFB58" box="[1297,1398,1160,1186]" name="Malaysia" pageId="1" pageNumber="7">Sarawak</collectingCountry>
in northern Borneo and from a plant cultivated in Semengoh Forest Arboretum near Kuching.
</paragraph>
<paragraph id="8BD2D84E99173A2AFF27FB08FE8EFAEC" blockId="1.[151,1436,152,1926]" pageId="1" pageNumber="7">
<emphasis id="B919045C99173A2AFF27FB08FEBAFB15" bold="true" box="[198,304,1237,1263]" pageId="1" pageNumber="7">Habitat:</emphasis>
––This species grows in primary mixed dipterocarp forest in river valleys at about 
<quantity id="4C9575AB99173A2AFAEEFB08FAC7FB15" box="[1295,1357,1237,1263]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="7" unit="m" value="42.0">42 m</quantity>
above sea level.
</paragraph>
<paragraph id="8BD2D84E99173A2AFF27FAFFFDA6FAC6" blockId="1.[151,1436,152,1926]" box="[198,556,1314,1340]" pageId="1" pageNumber="7">
<emphasis id="B919045C99173A2AFF27FAFFFEE7FAC6" bold="true" box="[198,365,1314,1340]" pageId="1" pageNumber="7">Local names:</emphasis>
—Not recorded.
</paragraph>
<paragraph id="8BD2D84E99173A2AFF27FA94FB65FA99" blockId="1.[151,1436,152,1926]" box="[198,1263,1353,1379]" pageId="1" pageNumber="7">
<emphasis id="B919045C99173A2AFF27FA94FE82FA99" bold="true" box="[198,264,1353,1379]" pageId="1" pageNumber="7">Uses:</emphasis>
—Traditional uses are not known. However, the palm has potential as an ornamental.
</paragraph>
<paragraph id="8BD2D84E99173A2AFF27FAADFE77F966" blockId="1.[151,1436,152,1926]" pageId="1" pageNumber="7">
<emphasis id="B919045C99173A2AFF27FAADFE46FA70" bold="true" box="[198,460,1392,1418]" pageId="1" pageNumber="7">Conservation status:</emphasis>
—Critically Endangered (CR B2ab). This palm meets the criteria for the threat category “Critically Endangered” (
<bibRefCitation id="EFFCA5BF99173A2AFDA3FA4AFD5FFA4B" author="IUCN" box="[578,725,1431,1457]" pageId="1" pageNumber="7" refId="ref10822" refString="IUCN (2001) IUCN red list categories and criteria. Version 3.1. IUCN, Gland &amp; Cambridge, 418 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
) because it is known only from one locality at 
<taxonomicName id="4C6DA3CD99173A2AFAFCFA4AFE94FA20" authority="Anap, Tatau, Miri" authorityName="Anap, Tatau, Miri" genus="Ulu" pageId="1" pageNumber="7" rank="genus">Ulu Anap, Tatau, Miri</taxonomicName>
, 
<collectingCountry id="F37A98DE99173A2AFECAFA1DFE1AFA20" box="[299,400,1472,1498]" name="Malaysia" pageId="1" pageNumber="7">Sarawak</collectingCountry>
and its area of occupancy is estimated to be less than 
<quantity id="4C9575AB99173A2AFBF5FA1DFBD5FA20" box="[1044,1119,1472,1498]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="7" unit="km" value="10.0">10 km</quantity>
<superScript id="7C18750699173A2AFBBEFA60FBEDFA31" attach="left" box="[1119,1127,1469,1483]" fontSize="6" pageId="1" pageNumber="7">2</superScript>
. In addition, rain forest is highly threatened in general in 
<collectingCountry id="F37A98DE99173A2AFDF1FA3AFDF0F9FB" box="[528,634,1511,1537]" name="Malaysia" pageId="1" pageNumber="7">Sarawak</collectingCountry>
, for example due to logging activities and oil palm plantation, and thus a decline in extent of occurrence, area of occupancy and quality of habitat is inferred. One clump of this palm has been successfully established and fruits in Semengoh Forest Arboretum as part of the 
<emphasis id="B919045C99173A2AFAAAF9E9FA16F9B7" box="[1355,1436,1588,1613]" italics="true" pageId="1" pageNumber="7">ex situ</emphasis>
conservation program run by Biodiversity Research Centre, 
<collectingCountry id="F37A98DE99173A2AFCB6F986FC36F98F" box="[855,956,1627,1653]" name="Malaysia" pageId="1" pageNumber="7">Sarawak</collectingCountry>
Forestry Corporation (previously known as 
<collectingCountry id="F37A98DE99173A2AFF57F95FFE91F966" box="[182,283,1666,1692]" name="Malaysia" pageId="1" pageNumber="7">Sarawak</collectingCountry>
Forestry Institute).
</paragraph>
</subSubSection>
<subSubSection id="C3778BC599173A2AFF27F975FEA8F87C" pageId="1" pageNumber="7" type="etymology">
<paragraph id="8BD2D84E99173A2AFF27F975FC31F913" blockId="1.[151,1436,152,1926]" pageId="1" pageNumber="7">
<emphasis id="B919045C99173A2AFF27F975FEDDF938" bold="true" box="[198,343,1704,1730]" pageId="1" pageNumber="7">Etymology:</emphasis>
—This new 
<taxonomicName id="4C6DA3CD99173A2AFE0FF974FDB9F938" authorityName="Linnaeus (1753:" authorityYear="1189" box="[494,563,1705,1730]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="1" pageNumber="7" phylum="Tracheophyta" rank="genus">
<emphasis id="B919045C99173A2AFE0FF974FDB9F938" box="[494,563,1705,1730]" italics="true" pageId="1" pageNumber="7">Areca</emphasis>
</taxonomicName>
is named after Dr. William J. Baker, the Head of Palm Research at the Royal Botanic Gardens, Kew and the collector of the 
<typeStatus id="54D666EC99173A2AFCEAF912FCB7F913" box="[779,829,1743,1769]" pageId="1" pageNumber="7">type</typeStatus>
specimen.
</paragraph>
<paragraph id="8BD2D84E99173A2AFF27F925FEA8F87C" blockId="1.[151,1436,152,1926]" pageId="1" pageNumber="7">
<emphasis id="B919045C99173A2AFF27F925FDE0F8E8" bold="true" box="[198,618,1784,1810]" pageId="1" pageNumber="7">Additional specimens examined:</emphasis>
–– 
<collectingCountry id="F37A98DE99173A2AFD69F924FCA9F8E9" box="[648,803,1785,1811]" name="Malaysia" pageId="1" pageNumber="7">MALAYSIA</collectingCountry>
. Borneo, 
<collectingRegion id="49A916AC99173A2AFC43F924FB81F8E9" box="[930,1035,1785,1811]" country="Malaysia" name="Sarawak" pageId="1" pageNumber="7">Sarawak</collectingRegion>
: Miri (4 
<superScript id="7C18750699173A2AFB98F928FB0FF8F9" attach="none" box="[1145,1157,1781,1795]" fontSize="6" pageId="1" pageNumber="7">th</superScript>
) 
<taxonomicName id="4C6DA3CD99173A2AFB7FF924FAD7F8E9" authority=", Tatau" authorityName="Tatau" box="[1182,1373,1785,1811]" genus="Division" pageId="1" pageNumber="7" rank="genus">Division, Tatau</taxonomicName>
, 
<taxonomicName id="4C6DA3CD99173A2AFA91F924FF56F8C3" authority="Anap" authorityName="Anap" genus="Ulu" pageId="1" pageNumber="7" rank="genus">Ulu Anap</taxonomicName>
, 
<date id="FFD3FE8E99173A2AFF0BF8C2FE1EF8C3" box="[234,404,1823,1849]" pageId="1" pageNumber="7" value="1982-06-22">22 June 1982</date>
, 
<emphasis id="B919045C99173A2AFE43F8FDFC8CF8C3" box="[418,774,1824,1849]" italics="true" pageId="1" pageNumber="7">
Mokhtar &amp; Othman 
<accessionNumber id="943E45AD99173A2AFD43F8FDFC8CF8C3" box="[674,774,1824,1849]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S44726" pageId="1" pageNumber="7" type="EnaNcbi">S 44726</accessionNumber>
</emphasis>
(K!, L!, KEP, SAN, SAR!). CULTIVATED. Borneo, 
<collectingRegion id="49A916AC99173A2AFF76F89BFE8AF89A" box="[151,256,1862,1888]" country="Malaysia" name="Sarawak" pageId="1" pageNumber="7">Sarawak</collectingRegion>
: Semengoh Forest Arboretum, 
<geoCoordinate id="EE59BE8999173A2AFD8CF89BFC84F89A" box="[621,782,1861,1888]" degrees="01" direction="north" minutes="24.059" orientation="latitude" pageId="1" pageNumber="7" precision="1" value="1.4009833">01º 24.059ʹ N</geoCoordinate>
<geoCoordinate id="EE59BE8999173A2AFCF4F89BFC49F89A" box="[789,963,1861,1888]" degrees="110" direction="east" minutes="19.423" orientation="longitude" pageId="1" pageNumber="7" precision="1" value="110.323715">110º 19.423ʹ E</geoCoordinate>
, 
<date id="FFD3FE8E99173A2AFC2EF89BFB0EF89A" box="[975,1156,1862,1888]" pageId="1" pageNumber="7" value="2008-03-31">31 March 2008</date>
, 
<emphasis id="B919045C99173A2AFB71F89BFA16F8A5" box="[1168,1436,1862,1887]" italics="true" pageId="1" pageNumber="7">Heatubun &amp; Kuda 903</emphasis>
(K!, SAR!).
</paragraph>
</subSubSection>
<caption id="DF1288C699143A29FF76F8BEFB36F821" ID-DOI="http://doi.org/10.5281/zenodo.4894310" ID-Zenodo-Dep="4894310" httpUri="https://zenodo.org/record/4894310/files/figure.png" pageId="2" pageNumber="8" startId="2.[151,257,1891,1915]" targetBox="[159,1420,193,1854]" targetPageId="2">
<paragraph id="8BD2D84E99143A29FF76F8BEFB36F821" blockId="2.[151,1436,1891,2011]" pageId="2" pageNumber="8">
<emphasis id="B919045C99143A29FF76F8BEFE9CF881" bold="true" box="[151,278,1891,1915]" pageId="2" pageNumber="8">FIGURE 1</emphasis>
. 
<taxonomicName id="4C6DA3CD99143A29FEC7F8B9FE3FF881" authorityName="Heatubun" authorityYear="2011" box="[294,437,1892,1915]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="8" phylum="Tracheophyta" rank="species" species="bakeri">
<emphasis id="B919045C99143A29FEC7F8B9FE3FF881" box="[294,437,1892,1915]" italics="true" pageId="2" pageNumber="8">Areca bakeri</emphasis>
</taxonomicName>
. A. Apical portion of leaf. B. Middle portion of leaf. C. Basal portion of leaf and petiole attached to crown shaft. D. Inflorescence attached to stem just below the crown shaft and showing acaulescent habit. E, F. Staminate flower whole and in section. G. Pistillate flower. Scale bar: A, B = 8 cm; C, D = 3 cm; E, F = 2.2 mm; G = 7 mm. A–D from 
<emphasis id="B919045C99143A29FEA2F819FE67F821" box="[323,493,1988,2011]" italics="true" pageId="2" pageNumber="8">Baker et al. 716</emphasis>
; E–G from 
<emphasis id="B919045C99143A29FD8CF819FC14F821" box="[621,926,1988,2011]" italics="true" pageId="2" pageNumber="8">
Mokhtar &amp; Othman 
<accessionNumber id="943E45AD99143A29FCA6F819FC14F821" box="[839,926,1988,2011]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S44726" pageId="2" pageNumber="8" type="EnaNcbi">S 44726</accessionNumber>
</emphasis>
. Drawn by Lucy T. Smith.
</paragraph>
</caption>
<caption id="DF1288C699153A28FF76F8A1FB7AF80E" ID-DOI="http://doi.org/10.5281/zenodo.4894312" ID-Zenodo-Dep="4894312" httpUri="https://zenodo.org/record/4894312/files/figure.png" pageId="3" pageNumber="9" startId="3.[151,258,1916,1940]" targetBox="[151,1436,182,1895]" targetPageId="3">
<paragraph id="8BD2D84E99153A28FF76F8A1FB7AF80E" blockId="3.[151,1436,1916,2036]" pageId="3" pageNumber="9">
<emphasis id="B919045C99153A28FF76F8A1FE95F86E" bold="true" box="[151,287,1916,1940]" pageId="3" pageNumber="9">FIGURE 2.</emphasis>
<taxonomicName id="4C6DA3CD99153A28FEC6F8A0FE32F86E" authorityName="Heatubun" authorityYear="2011" box="[295,440,1917,1940]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="3" pageNumber="9" phylum="Tracheophyta" rank="species" species="bakeri">
<emphasis id="B919045C99153A28FEC6F8A0FE32F86E" box="[295,440,1917,1940]" italics="true" pageId="3" pageNumber="9">Areca bakeri</emphasis>
</taxonomicName>
. A. Acaulescent habit with densely suckering clump; the palm was mis-identified as 
<taxonomicName id="4C6DA3CD99153A28FA65F8A0FE91F84E" authorityName="Dransfield" authorityYear="1984" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="3" pageNumber="9" phylum="Tracheophyta" rank="species" species="jugahpunya">
<emphasis id="B919045C99153A28FA65F8A0FE91F84E" italics="true" pageId="3" pageNumber="9">A. jugahpunya</emphasis>
</taxonomicName>
, as shown on the label. B. Infructescence bursting out from the marcescent leaf sheaths. C. Prophyll, inflorescence, infructescence and seedling. All photos were taken from the specimen cultivated at Semengoh Forest Arboretum near Kuching, Sarawak, Malaysia. Photos: A (Charlie D. Heatubun), B–C (John Dransfield).
</paragraph>
</caption>
<subSubSection id="C3778BC599123A2FFF27FF4AFD34FDF4" pageId="4" pageNumber="10" type="discussion">
<paragraph id="8BD2D84E99123A2FFF27FF4AFD34FDF4" blockId="4.[151,1436,151,526]" pageId="4" pageNumber="10">
<emphasis id="B919045C99123A2FFF27FF4AFEDEFF4B" bold="true" box="[198,340,151,177]" pageId="4" pageNumber="10">Discussion:</emphasis>
–– 
<taxonomicName id="4C6DA3CD99123A2FFE90FF45FD82FF4B" authorityName="Heatubun" authorityYear="2011" box="[369,520,152,177]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="bakeri">
<emphasis id="B919045C99123A2FFE90FF45FD82FF4B" box="[369,520,152,177]" italics="true" pageId="4" pageNumber="10">Areca bakeri</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C6DA3CD99123A2FFD7FFF45FBC6FF48" authority="Dransfield (1984: 13)" authorityName="Dransfield" authorityPageNumber="13" authorityYear="1984" box="[670,1100,152,178]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="jugahpunya">
<emphasis id="B919045C99123A2FFD7FFF45FCC3FF4B" box="[670,841,152,177]" italics="true" pageId="4" pageNumber="10">A. jugahpunya</emphasis>
<bibRefCitation id="EFFCA5BF99123A2FFCAEFF45FBC6FF48" author="Dransfield, J." box="[847,1100,152,178]" pageId="4" pageNumber="10" pagination="1 - 22" refId="ref10479" refString="Dransfield, J. (1984) The genus Areca (Palmae: Arecoideae) in Borneo. Kew Bulletin 39: 1 - 22." type="journal article" year="1984">Dransfield (1984: 13)</bibRefCitation>
</taxonomicName>
in its acaulescent, clustering habit and broad leaflets, but can immediately be distinguished by the densely clustering habit with ca. 20 stems in the clump, very long and slender petioles, leaflets more papery and the tips of terminal leaflets slightly obliquely lobed. The inflorescence of 
<taxonomicName id="4C6DA3CD99123A2FFD0FFED1FCE9FEDF" authorityName="Heatubun" authorityYear="2011" box="[750,867,268,293]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="bakeri">
<emphasis id="B919045C99123A2FFD0FFED1FCE9FEDF" box="[750,867,268,293]" italics="true" pageId="4" pageNumber="10">A. bakeri</emphasis>
</taxonomicName>
is more similar to 
<taxonomicName id="4C6DA3CD99123A2FFBBDFED1FA9FFEDF" authorityName="Heatubun" authorityYear="2011" box="[1116,1301,268,293]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="dransfieldii">
<emphasis id="B919045C99123A2FFBBDFED1FA9FFEDF" box="[1116,1301,268,293]" italics="true" pageId="4" pageNumber="10">A. dransfieldii</emphasis>
</taxonomicName>
than to 
<taxonomicName id="4C6DA3CD99123A2FFA63FED1FEAFFEB1" authorityName="Dransfield" authorityYear="1984" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="jugahpunya">
<emphasis id="B919045C99123A2FFA63FED1FEAFFEB1" italics="true" pageId="4" pageNumber="10">A. jugahpunya</emphasis>
</taxonomicName>
, especially with its swollen-flattened rachillae and only one pistillate flower on each rachilla. However, 
<taxonomicName id="4C6DA3CD99123A2FFEF0FE84FEF7FE88" authorityName="Heatubun" authorityYear="2011" box="[273,381,345,370]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="bakeri">
<emphasis id="B919045C99123A2FFEF0FE84FEF7FE88" box="[273,381,345,370]" italics="true" pageId="4" pageNumber="10">A. bakeri</emphasis>
</taxonomicName>
is easily distinguished from 
<taxonomicName id="4C6DA3CD99123A2FFD35FE84FCF6FE88" authorityName="Heatubun" authorityYear="2011" box="[724,892,345,370]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="dransfieldii">
<emphasis id="B919045C99123A2FFD35FE84FCF6FE88" box="[724,892,345,370]" italics="true" pageId="4" pageNumber="10">A. dransfieldii</emphasis>
</taxonomicName>
by its acaulescent litter-trapping habit, leaves with very long petioles (
<quantity id="4C9575AB99123A2FFE31FE5DFDE5FE60" box="[464,623,384,410]" metricMagnitude="0" metricUnit="m" metricValue="1.025" metricValueMax="1.05" metricValueMin="1.0" pageId="4" pageNumber="10" unit="cm" value="102.5" valueMax="105.0" valueMin="100.0">100–105 cm</quantity>
) and broad leaflets, and infrafoliar inflorescences borne among marcescent leaf sheaths with somewhat long peduncles. In contrast, 
<taxonomicName id="4C6DA3CD99123A2FFC35FE7AFB08FE3A" authorityName="Heatubun" authorityYear="2011" box="[980,1154,423,448]" class="Liliopsida" family="Arecaceae" genus="Areca" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="10" phylum="Tracheophyta" rank="species" species="dransfieldii">
<emphasis id="B919045C99123A2FFC35FE7AFB08FE3A" box="[980,1154,423,448]" italics="true" pageId="4" pageNumber="10">A. dransfieldii</emphasis>
</taxonomicName>
has conspicuous stems with aerial branching, leaves with long petioles (
<quantity id="4C9575AB99123A2FFCE0FE10FC0BFE1D" box="[769,897,461,487]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" metricValueMax="8.1" metricValueMin="2.5" pageId="4" pageNumber="10" unit="cm" value="53.0" valueMax="81.0" valueMin="25.0">25–81 cm</quantity>
) and numerous finely-regular leaflets, and infrafoliar inflorescences with short peduncles.
</paragraph>
</subSubSection>
</treatment>
</document>