122 lines
13 KiB
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122 lines
13 KiB
XML
<document id="0D3A1A433D9F19D82C2A40DF21858168" ID-DOI="10.11646/zootaxa.3731.4.4" ID-GBIF-Dataset="26facfec-e615-4814-88c5-c966fcbc9efc" ID-ISSN="1175-5326" ID-Zenodo-Dep="248259" ID-ZooBank="2066C0D1-2ED6-41EB-AF3C-3DAB58EA1718" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460694621800" checkinUser="plazi" docAuthor="Krikken, Jan & Li, Chun-Lin" docDate="2013" docId="03ABBA06FFBFFFDFA181FC65FCFFF95B" docLanguage="en" docName="zt03731p519.pdf" docOrigin="Zootaxa 3731 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolbochromus (Metabolbochromus) catenatus Lansberge 1886" docType="treatment" docVersion="9" lastPageNumber="515" masterDocId="FF92C27EFFABFFCBA116FFB7FFCBFFE1" masterDocTitle="Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae)" masterLastPageNumber="519" masterPageNumber="495" pageNumber="515" updateTime="1698353700889" updateUser="plazi">
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<mods:mods id="544307337A5441814E349DA471C5CA0D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="30BC297F8D27CB904058716996E327BB">
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<mods:title id="D8D20C87AA726A2EBC50FF60D86A6395">Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae)</mods:title>
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<mods:namePart id="DDD8E5365ABA72AAE50B3D5017A1B777">Krikken, Jan</mods:namePart>
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<mods:roleTerm id="60F5AAB0B2135C1B9AB1CE2BA89FC2BF">Author</mods:roleTerm>
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<mods:namePart id="4EE446C3E1FA60A8FF3329E9E0F43241">Li, Chun-Lin</mods:namePart>
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<mods:title id="21878FFEB6C92F776202F1FCC09FFE6D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9397F1D627C6C810F9445056748B15E6">
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<mods:date id="C93C6085132F32892EDECAA8ED1F5245">2013</mods:date>
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<mods:detail id="1EE4E4A8257EC5EE1DC15B9FC38EC20C" type="volume">
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<mods:number id="0BEE5276CBE6B321850E59E291CE47CF">3731</mods:number>
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<mods:detail id="A2F0BA80CB3041B35DAAF2FD4711D80E" type="issue">
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<mods:number id="FB744DEB9F5A81AF4EFE55DD1E4278FC">4</mods:number>
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<mods:classification id="D4FCA36C30EFAE522E7F773CDC7022B0">journal article</mods:classification>
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<mods:identifier id="24808711FFD19039CE370F096F1BCB75" type="DOI">10.11646/zootaxa.3731.4.4</mods:identifier>
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<mods:identifier id="3ACA93CDF492D83F38D5434A62274986" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B68FE6D5F13FD7FF0A05E63A0EC6171D" type="Zenodo-Dep">248259</mods:identifier>
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<treatment id="03ABBA06FFBFFFDFA181FC65FCFFF95B" ID-DOI="http://doi.org/10.5281/zenodo.5680771" ID-GBIF-Taxon="119581752" ID-Zenodo-Dep="5680771" LSID="urn:lsid:plazi:treatment:03ABBA06FFBFFFDFA181FC65FCFFF95B" httpUri="http://treatment.plazi.org/id/03ABBA06FFBFFFDFA181FC65FCFFF95B" lastPageNumber="515" pageId="20" pageNumber="515">
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<subSubSection id="C318589BFFBFFFDFA181FC65FCDEFA5C" pageId="20" pageNumber="515" type="nomenclature">
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<paragraph id="8BBD0B10FFBFFFDFA181FC65FC64FC0D" blockId="20.[151,943,978,1038]" box="[151,943,978,1004]" pageId="20" pageNumber="515">
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<heading id="D0F5BC7CFFBFFFDFA181FC65FC64FC0D" bold="true" box="[151,943,978,1004]" fontSize="11" level="1" pageId="20" pageNumber="515" reason="1">
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<taxonomicName id="4C027093FFBFFFDFA181FC65FC64FC0D" authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[151,943,978,1004]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="catenatus" subGenus="Metabolbochromus">
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<emphasis id="B976D702FFBFFFDFA181FC65FC64FC0D" bold="true" box="[151,943,978,1004]" pageId="20" pageNumber="515">
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<emphasis id="B976D702FFBFFFDFA181FC65FE82FC0D" bold="true" box="[151,329,978,1004]" italics="true" pageId="20" pageNumber="515">Bolbochromus</emphasis>
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(
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<emphasis id="B976D702FFBFFFDFA04FFC65FD88FC0D" bold="true" box="[345,579,978,1004]" italics="true" pageId="20" pageNumber="515">Metabolbochromus</emphasis>
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)
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<emphasis id="B976D702FFBFFFDFA342FC65FD0CFC0D" bold="true" box="[596,711,978,1004]" italics="true" pageId="20" pageNumber="515">catenatus</emphasis>
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(Lansberge, 1886)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BBD0B10FFBFFFDFA181FC42FE93FBEF" blockId="20.[151,943,978,1038]" box="[151,344,1013,1038]" pageId="20" pageNumber="515">
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(
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<figureCitation id="13391795FFBFFFDFA189FC42FEECFBEF" box="[159,295,1013,1038]" captionStart="FIGURES 9 – 16" captionStartId="9.[151,264,1972,1994]" captionTargetBox="[167,1418,202,1950]" captionTargetId="figure@9.[151,1436,193,1951]" captionTargetPageId="9" captionText="FIGURES 9 – 16. Habitus of male Bolbochromus (oblique and lateral views). 9 – 10, B. sinensis; 11 – 12, B. pumilus; 13 – 14, B. catenatus; 15 – 16, B. niger. Scale lines 1 mm." httpUri="https://zenodo.org/record/248261/files/figure.png" pageId="20" pageNumber="515">Figs. 13–14</figureCitation>
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, 39)
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</paragraph>
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<paragraph id="8BBD0B10FFBFFFDFA181FB8AFCDEFA5C" blockId="20.[151,1436,1085,1722]" pageId="20" pageNumber="515">
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<emphasis id="B976D702FFBFFFDFA181FB8AFE8BFBB7" bold="true" box="[151,320,1085,1110]" pageId="20" pageNumber="515">Identification.</emphasis>
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Basic colour predominantly black or dark-brown, usually with anterior and posterior yellow-orange marking on bottom of lateral declivity of pronotum (they may be contiguous) and one yellow-orange marking near scutellum on elytral base (at least over interstriae 2–3)—colour pattern, however, very variable, at least in parts of its apparently extensive range (note that
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<taxonomicName id="4C027093FFBFFFDFA372FB1DFD3BFB20" box="[612,752,1194,1217]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="catenatus">
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<emphasis id="B976D702FFBFFFDFA372FB1DFD3BFB20" box="[612,752,1194,1217]" italics="true" pageId="20" pageNumber="515">B. catenatus</emphasis>
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</taxonomicName>
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is on Java and Bali replaced by the two other species in the subgenus). Apicomedian tubercle of clypeus and postfrontal tubercle usually distinct, but short, clypeogenal tubercle on either side lacking (and in this character distinct from
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<taxonomicName id="4C027093FFBFFFDFA2BFFB45FBF5FAE8" box="[937,1086,1265,1289]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="sulcicollis">
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<emphasis id="B976D702FFBFFFDFA2BFFB45FBF5FAE8" box="[937,1086,1265,1289]" italics="true" pageId="20" pageNumber="515">B. sulcicollis</emphasis>
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</taxonomicName>
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). Median sulcus of pronotum distinct, adjacent surface usually evenly convex. Lateral declivity of pronotum with abundant punctation. The more common general colour pattern (black-brown with well delimited yellow-orange markings on pronotum and elytra) also occurs, with slight variations, in other groups of species (mainly continental and
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<collectingCountry id="F3154B80FFBFFFDFA54BFAEAFB17FA97" box="[1117,1244,1373,1398]" name="Philippines" pageId="20" pageNumber="515">Philippines</collectingCountry>
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)—these are here allocated to the subgenus
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<taxonomicName id="4C027093FFBFFFDFA0DCFA36FDA7FA78" box="[458,620,1409,1433]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="genus">
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<emphasis id="B976D702FFBFFFDFA0DCFA36FDA7FA78" box="[458,620,1409,1433]" italics="true" pageId="20" pageNumber="515">Bolbochromus</emphasis>
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</taxonomicName>
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(different aedeagi, different cephalic protrusions, see two of the new species described above). Body length usually
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<quantity id="4CFAA6F5FFBFFFDFA3B2FA11FCDAFA5C" box="[676,785,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="20" pageNumber="515" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C318589BFFBFFFDFA1D1FA7EFE4BF977" pageId="20" pageNumber="515" type="discussion">
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<paragraph id="8BBD0B10FFBFFFDFA1D1FA7EFE4BF977" blockId="20.[151,1436,1085,1722]" pageId="20" pageNumber="515">
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<emphasis id="B976D702FFBFFFDFA1D1FA7EFE8FFA03" bold="true" box="[199,324,1481,1506]" pageId="20" pageNumber="515">Comment.</emphasis>
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Reported from Sumatra and Borneo, whence longer, most variable series were seen (many collected in flight interception traps by A.J. Davis, in the Danum area, Sabah). The name
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<taxonomicName id="4C027093FFBFFFDFA582FA59FAEBF9E4" box="[1172,1312,1518,1541]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="catenatus">
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<emphasis id="B976D702FFBFFFDFA582FA59FAEBF9E4" box="[1172,1312,1518,1541]" italics="true" pageId="20" pageNumber="515">B. catenatus</emphasis>
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</taxonomicName>
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may stand for a complex of species and/or subspecies, but an examination of more informative material (at least from the
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<collectingCountry id="F3154B80FFBFFFDFA181F982FEDCF9AF" box="[151,279,1589,1614]" name="Philippines" pageId="20" pageNumber="515">Philippines</collectingCountry>
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) is needed to draw conclusions. This species, as here tentatively conceived, is the only
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<taxonomicName id="4C027093FFBFFFDFA5ECF982FA57F9AC" box="[1274,1436,1589,1613]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="genus">
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<emphasis id="B976D702FFBFFFDFA5ECF982FA57F9AC" box="[1274,1436,1589,1613]" italics="true" pageId="20" pageNumber="515">Bolbochromus</emphasis>
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</taxonomicName>
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common on Borneo; occurring up to approximately
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<quantity id="4CFAA6F5FFBFFFDFA3FBF9EEFC8FF990" box="[749,836,1625,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="515" unit="m" value="1500.0">1500 m</quantity>
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on Mt Kinabalu (note doubtful records of
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<taxonomicName id="4C027093FFBFFFDFA424F9EDFA57F990" box="[1330,1436,1625,1649]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="515" phylum="Arthropoda" rank="species" species="walshi">
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<emphasis id="B976D702FFBFFFDFA424F9EDFA88F990" box="[1330,1347,1626,1649]" italics="true" pageId="20" pageNumber="515">B</emphasis>
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.
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<emphasis id="B976D702FFBFFFDFA445F9EEFA57F990" box="[1363,1436,1625,1649]" italics="true" pageId="20" pageNumber="515">walshi</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="F3154B80FFBFFFDFA1C2F9CAFEBFF977" box="[212,372,1661,1686]" name="Malaysia" pageId="20" pageNumber="515">North Borneo</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C318589BFFBFFFDFA1D1F916FCFFF95B" box="[199,820,1697,1722]" pageId="20" pageNumber="515" type="distribution">
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<paragraph id="8BBD0B10FFBFFFDFA1D1F916FCFFF95B" blockId="20.[151,1436,1085,1722]" box="[199,820,1697,1722]" pageId="20" pageNumber="515">
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<emphasis id="B976D702FFBFFFDFA1D1F916FEABF95B" bold="true" box="[199,352,1697,1722]" pageId="20" pageNumber="515">Distribution.</emphasis>
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<collectingCountry id="F3154B80FFBFFFDFA071F916FE13F95B" box="[359,472,1697,1722]" name="Indonesia" pageId="20" pageNumber="515">Indonesia</collectingCountry>
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: Sumatra, Borneo,
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<collectingCountry id="F3154B80FFBFFFDFA3B8F916FCFBF95B" box="[686,816,1697,1722]" name="Philippines" pageId="20" pageNumber="515">Philippines</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |