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<document id="D6CE481AC623A9A3CFA392FAE4D4A880" ID-DOI="10.1649/0010-065X(2001)055[0378:CCNTBC]2.0.CO;2" ID-ISSN="1938-4394" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630155932255" checkinUser="tatiana" docAuthor="Jarrett, J. R. &amp; Scudder, G. G E." docDate="2001" docId="03A387CBFFDE5F2EEBF0FAEA5D0DFE93" docLanguage="en" docName="0010-065X_2001_055_0378_CCNTBC_2.0.CO_2.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (3)" docSource="http://dx.doi.org/10.1649/0010-065x(2001)055[0378:ccntbc]2.0.co;2" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Bembidion quadrimaculatum subsp. oppositum Say 1823" docType="treatment" docVersion="2" lastPageNumber="382" masterDocId="FF9AFFB3FFDD5F2AEA39FFB95E5AFFFB" masterDocTitle="Carabidae (Coleoptera) New To British Columbia, With One Species New To Canada" masterLastPageNumber="384" masterPageNumber="378" pageNumber="381" updateTime="1699167444876" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EAC39FA8E201AA9F9F342CBB6DBC983A">Carabidae (Coleoptera) New To British Columbia, With One Species New To Canada</mods:title>
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<mods:namePart id="97BD3C508A04ED5158199F8B6919EEEF">Jarrett, J. R.</mods:namePart>
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<mods:namePart id="092039490113B6FCFBF234941765CE44">Scudder, G. G E.</mods:namePart>
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<mods:title id="BA84BC34823CD536E46F1FF637D6A435">The Coleopterists Bulletin</mods:title>
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<mods:date id="AFFEB880DDBE5569F262AD43CAD112CE">2001</mods:date>
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<mods:number id="6E9F22233D51CA18C31A5BA596D9BB3F">2001-09-30</mods:number>
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<mods:identifier id="4CDA9AD2ABA7647F1AFBE941CD8EEADE" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03A387CBFFDE5F2EEBF0FAEA5D0DFE93" LSID="urn:lsid:plazi:treatment:03A387CBFFDE5F2EEBF0FAEA5D0DFE93" httpUri="http://treatment.plazi.org/id/03A387CBFFDE5F2EEBF0FAEA5D0DFE93" lastPageId="4" lastPageNumber="382" pageId="3" pageNumber="381">
<subSubSection id="C3106556FFDE5F29EBF0FAEA5D53FA68" pageId="3" pageNumber="381" type="nomenclature">
<paragraph id="8BB536DDFFDE5F29EBF0FAEA5DF8FA93" blockId="3.[457,930,1363,1384]" box="[457,930,1363,1384]" pageId="3" pageNumber="381">
<heading id="D0FD81B1FFDE5F29EBF0FAEA5DF8FA93" box="[457,930,1363,1384]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="381" reason="2">
<taxonomicName id="4C0A4D5EFFDE5F29EBF0FAEA5DF8FA93" ID-CoL="8LJHM" authority="Say" authorityName="Say" authorityYear="1823" box="[457,930,1363,1384]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="subSpecies" species="quadrimaculatum" subSpecies="oppositum">
<emphasis id="B97EEACFFFDE5F29EBF0FAEA5D29FA93" box="[457,883,1363,1384]" italics="true" pageId="3" pageNumber="381">Bembidion quadrimaculatum oppositum</emphasis>
Say
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB536DDFFDE5F29EB2FFAC75D53FA68" blockId="3.[278,1111,1406,1614]" box="[278,777,1406,1427]" pageId="3" pageNumber="381">
<materialsCitation id="3B623C80FFDE5F29EB2FFAC75D5CFA68" box="[278,774,1406,1427]" collectingDate="1923-04-06" collectionCode="UBC" collectorName="Mrs. W. W. H." pageId="3" pageNumber="381" specimenCount="1">
BC: Terrace,
<date id="FFB4101DFFDE5F29EB9FFAC75C57FA68" box="[422,525,1406,1427]" pageId="3" pageNumber="381" value="1923-04-06">
<collectingDate id="EFF0E9F5FFDE5F29EB9FFAC75C57FA68" box="[422,525,1406,1427]" pageId="3" pageNumber="381" value="1923-04-06">6.IV.1923</collectingDate>
</date>
(
<collectorName id="26FF530BFFDE5F29E824FAC75CF1FA68" box="[541,683,1406,1427]" pageId="3" pageNumber="381">Mrs. W.W.H.</collectorName>
) [
<collectionCode id="ED1BAE18FFDE5F29E8F8FAC75CA4FA68" box="[705,766,1406,1427]" country="Canada" httpUri="http://grbio.org/cool/m0m3-x1cz" name="University of British Columbia" pageId="3" pageNumber="381">UBC</collectionCode>
]
</materialsCitation>
.
</paragraph>
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<subSubSection id="C3106556FFDE5F2EEB09FA215D0DFE93" lastPageId="4" lastPageNumber="382" pageId="3" pageNumber="381" type="description">
<paragraph id="8BB536DDFFDE5F29EB09FA215A0CF9B6" blockId="3.[278,1111,1406,1614]" pageId="3" pageNumber="381">
Holarctic in distribution,
<taxonomicName id="4C0A4D5EFFDE5F29E807FA205DD6FA55" authorityName="Say" authorityYear="1823" box="[574,908,1433,1454]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="subSpecies" species="quadrimaculatum" subSpecies="oppositum">
<emphasis id="B97EEACFFFDE5F29E807FA205DD6FA55" box="[574,908,1433,1454]" italics="true" pageId="3" pageNumber="381">B. quadrimaculatum oppositum</emphasis>
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is known in North America from AB, CO, CT, DC, GA, IA, IL, IN, KS, MA, MB, MD, ME, MI, MN, MO, NB, NC, ND, NE, NF, NH, NJ, NS, NY, OH, ON, PA, PE, PQ, RI, SC, SD, SK, TN, TX, VA, VT, WI and WV (Bousquet and Larochelle 1993). The BC record seems to be the first west of the Rocky Mountains. Along with
<emphasis id="B97EEACFFFDE5F29EBAEF9A75C49F9C8" box="[407,531,1566,1587]" italics="true" pageId="3" pageNumber="381">
<taxonomicName id="4C0A4D5EFFDE5F29EBAEF9A75C54F9C8" baseAuthorityName="Herbst" baseAuthorityYear="1784" box="[407,526,1566,1587]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="species" species="lampros">B. lampros</taxonomicName>
,
</emphasis>
it is one of the least humidity loving species of
<taxonomicName id="4C0A4D5EFFDE5F29EE26F9A75C4BF9B6" authority="(Lindroth 1992)" baseAuthorityName="Lindroth" baseAuthorityYear="1992" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEACFFFDE5F29EE26F9A75F03F9B5" italics="true" pageId="3" pageNumber="381">Bembidion</emphasis>
(
<bibRefCitation id="EF9B4B2CFFDE5F29EB50F9815C52F9B6" author="Lindroth" box="[361,520,1592,1613]" pageId="3" pageNumber="381" refId="ref4191" refString="Lindroth, C. H. 1992. Ground Beetles (Carabidae) of Fennoscandia, Part 1. Smithsonian Institution Libraries and The National Science Foundation, Washington, D. C." type="book" year="1992">Lindroth 1992</bibRefCitation>
)
</taxonomicName>
. It has been frequently found on dry barren banks of
</paragraph>
<paragraph id="8BB536DDFFD95F2EEB2EFF515D0DFE93" blockId="4.[279,1112,232,360]" pageId="4" pageNumber="382">
fresh water, and also in open, preferably cultivated fields, quite independent of water (
<bibRefCitation id="EF9B4B2CFFD95F2EEB44FEBA5C40FEE3" author="Lindroth" box="[381,538,259,280]" pageId="4" pageNumber="382" refId="ref4191" refString="Lindroth, C. H. 1992. Ground Beetles (Carabidae) of Fennoscandia, Part 1. Smithsonian Institution Libraries and The National Science Foundation, Washington, D. C." type="book" year="1992">Lindroth 1992</bibRefCitation>
). It occurs on slightly moist, clay or clay­mixed sandy soil with sparse vegetation, allowing complete exposure to the sun (
<bibRefCitation id="EF9B4B2CFFD95F2EE9C3FEA75F15FEB6" author="Lindroth" pageId="4" pageNumber="382" refId="ref4148" refString="Lindroth, C. H. 1986. The Carabidae (Coleoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica, Vol. 15, Part 2. E. J. Brill / Scandinavian Science Press Ltd., Copenhagen." type="journal volume" year="1986">Lindroth 1986</bibRefCitation>
). This species breeds in spring, and hibernates as an adult (
<bibRefCitation id="EF9B4B2CFFD95F2EE9C2FE815F15FE93" author="Lindroth" pageId="4" pageNumber="382" refId="ref4191" refString="Lindroth, C. H. 1992. Ground Beetles (Carabidae) of Fennoscandia, Part 1. Smithsonian Institution Libraries and The National Science Foundation, Washington, D. C." type="book" year="1992">Lindroth 1992</bibRefCitation>
). Its wings are fully developed (
<bibRefCitation id="EF9B4B2CFFD95F2EE894FEEA5D11FE93" author="Lindroth" box="[685,843,339,360]" pageId="4" pageNumber="382" refId="ref4191" refString="Lindroth, C. H. 1992. Ground Beetles (Carabidae) of Fennoscandia, Part 1. Smithsonian Institution Libraries and The National Science Foundation, Washington, D. C." type="book" year="1992">Lindroth 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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