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<document id="5920F691404D182D71C8B087AF8136D0" ID-CLB-Dataset="54645" ID-DOI="10.11646/zootaxa.5278.1.4" ID-GBIF-Dataset="743d5014-f7d0-425d-8103-182ee59d4377" ID-ISSN="1175-5326" ID-Zenodo-Dep="7895044" ID-ZooBank="0DE3A459-D9E2-4DB5-9C61-483A0763EFAB" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683183250860" checkinUser="plazi" docAuthor="Ng, Shing-Lai, Ho, Hsuan-Ching, Joung, Shoou-Jeng &amp; Liu, Kwang-Ming" docDate="2023" docId="039D2D44A23AFFBEFF2345AFFDE1FA1E" docLanguage="en" docName="zootaxa.5278.1.4.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Okamejei picta Ng &amp; Ho &amp; Joung &amp; Liu 2023, sp. nov." docType="treatment" docUuid="5CD9D2D9-582C-4A97-9BA3-E5AEE6AE2297" docUuidSource="ZooBank" docVersion="11" lastPageNumber="117" masterDocId="FFA4553CA23BFFB0FFB44009FFC6FFF2" masterDocTitle="Okamejei picta sp. nov., a new rajid skate from the South China Sea (Rajiformes Rajidae)" masterLastPageNumber="118" masterPageNumber="103" pageNumber="104" updateTime="1701998452478" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E9D3B2D79803C1457FBAD2C184761B53">Okamejei picta sp. nov., a new rajid skate from the South China Sea (Rajiformes Rajidae)</mods:title>
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<figureCitation id="130F80D7A23AFFB1FF2345C2FEC4FA17" box="[151,258,1483,1509]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStartId-0="5.[152,255,1456,1480]" captionStartId-1="6.[152,255,1608,1632]" captionStartId-2="7.[152,255,1048,1072]" captionStartId-3="7.[152,255,1614,1638]" captionStartId-4="8.[152,255,1009,1033]" captionStartId-5="8.[152,255,1797,1821]" captionStartId-6="9.[152,255,1803,1827]" captionStartId-7="10.[152,255,1040,1064]" captionStartId-8="10.[152,255,1871,1895]" captionTargetBox-0="[151,1436,181,1432]" captionTargetBox-1="[151,1436,181,1584]" captionTargetBox-2="[280,1308,181,1024]" captionTargetBox-3="[280,1308,1205,1591]" captionTargetBox-4="[640,947,381,973]" captionTargetBox-5="[290,1297,1093,1773]" captionTargetBox-6="[347,1241,849,1777]" captionTargetBox-7="[303,1284,181,1017]" captionTargetBox-8="[312,1275,1125,1848]" captionTargetId-0="figure-17@5.[151,1436,181,1433]" captionTargetId-1="figure-31@6.[151,1436,181,1584]" captionTargetId-2="figure-17@7.[280,1308,181,1024]" captionTargetId-3="figure-89@7.[280,1308,1205,1591]" captionTargetId-4="figure-125@8.[640,947,381,985]" captionTargetId-5="figure-167@8.[290,1297,1093,1773]" captionTargetId-6="figure-348@9.[344,1244,849,1780]" captionTargetId-7="figure-31@10.[303,1284,181,1017]" captionTargetId-8="figure-72@10.[312,1275,1125,1848]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="10" captionText-0="FIGURE 1. Fresh coloration of Okamejei picta sp. nov., NMMB-P36013, 429 mm TL, adult male holotype: top, dorsal surface; bottom, ventral surface." captionText-1="FIGURE 2. Fresh coloration of Okamejei picta sp. nov., ASIZP0081116, 405 mm TL, adult male paratype: top, dorsal surface; bottom, ventral surface." captionText-2="FIGURE 3. Details of dorsal (A, B) and ventral (C, D) views of snout area of Okamejei picta sp. nov.: A, dorsal view of adult male holotype (NMMB-P36013, 429 mm TL); B, dorsal view of adult male paratype (ASIZP0081116, 405 mm TL); C, ventral view of adult male holotype (NMMB-P36013, 429 mm TL); D, ventral view of adult male paratype (ASIZP0081116, 405 mm TL)." captionText-3="FIGURE 4. Details of thorns on midtail (A, B) and the dorsal-caudal area (C, D) of Okamejei picta sp. nov.: A, adult male holotype (NMMB-P36013, 429 mm TL); B, adult male paratype (ASIZP0081116, 405 mm TL); C, adult male holotype (NMMB- P36013, 429 mm TL); D, adult male paratype (ASIZP0081116, 405 mm TL)." captionText-4="FIGURE 5. Okamejei picta sp. nov., adult male holotype, NMMB-P36013, 429 mm TL, left clasper spread opened. bs, boss; cf, cleft; sk, spike; tb, terminal bridge; ps, pseudorhipidion; rh, rhipidion; sh, shield; st, sentinel." captionText-5="FIGURE 6. Okamejei pictasp.nov., adult male holotype,NMMB-P36013,429 mmTL,X-radiograph of pelvic girdleandclasper basal cartilages (ventral view). ax, axial cartilage; β, beta cartilage; b12, intermediate segments 1 and 2; bpv, basipterygium; ip, iliac process; lp, lateral process; lpp, lateral prepelvic process of obturator foramina; pvp, pelvic propterygium." captionText-6="FIGURE 7. Terminal skeleton of left clasper of (A) Okamejei picta sp. nov., adult male holotype, NMMB-P36013, 429 mm TL; and (B) Okamejei hollandi, EBFS adult male, uncatalogued specimen, 432 mm TL. ax, axial cartilage; at12, accessory terminal cartilage 1 and 2; dm, dorsal marginal cartilage; dt13, dorsal terminal cartilage 13; vm, ventral marginal cartilage; vt, ventral terminal cartilage." captionText-7="FIGURE 8. Ventral terminal cartilage from left clasper of Okamejei picta sp. nov., adult male paratype, ASIZP0081116, 405 mm TL. (A) Dorsal view; (B) ventral view; (C) proximal tip viewed from left side." captionText-8="FIGURE 9. Okamejei picta sp. nov., adult male holotype, NMMB-P36013, 429 mm TL, X-radiograph of neurocranium, snout and pectoral area. ac, antorbital cartilage; af, anterior fontanelle; cr, neurocranium; ms, mesopterygium; mt, metapterygium; nc, nasal capsule; pf, posterior fontanelle; poc, preorbital canal foramen, postp, postorbital process; pr, propterygium; prep, preorbital process; rb, rostral base; rc, rostral cartilage." figureDoi-0="http://doi.org/10.5281/zenodo.7895046" figureDoi-1="http://doi.org/10.5281/zenodo.7895048" figureDoi-2="http://doi.org/10.5281/zenodo.7895052" figureDoi-3="http://doi.org/10.5281/zenodo.7895054" figureDoi-4="http://doi.org/10.5281/zenodo.7895056" figureDoi-5="http://doi.org/10.5281/zenodo.7895058" figureDoi-6="http://doi.org/10.5281/zenodo.7895060" figureDoi-7="http://doi.org/10.5281/zenodo.7895062" figureDoi-8="http://doi.org/10.5281/zenodo.7895064" httpUri-0="https://zenodo.org/record/7895046/files/figure.png" httpUri-1="https://zenodo.org/record/7895048/files/figure.png" httpUri-2="https://zenodo.org/record/7895052/files/figure.png" httpUri-3="https://zenodo.org/record/7895054/files/figure.png" httpUri-4="https://zenodo.org/record/7895056/files/figure.png" httpUri-5="https://zenodo.org/record/7895058/files/figure.png" httpUri-6="https://zenodo.org/record/7895060/files/figure.png" httpUri-7="https://zenodo.org/record/7895062/files/figure.png" httpUri-8="https://zenodo.org/record/7895064/files/figure.png" pageId="1" pageNumber="104">Figs. 19</figureCitation>
,
<tableCitation id="C6B6A9E9A23AFFB1FEB945C2FE40FA17" box="[269,390,1483,1509]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="2.[152,245,623,649]" captionStartId-1="12.[152,245,944,968]" captionTargetBox-0="[162,2021,699,1525]" captionTargetPageId-0="2" captionTargetPageId-1="12" captionText-0="TABLE 1. Morphometric data of O. picta sp. nov., and O. hollandi. Mean values are given in parentheses. Data of the holotypes of O. hollandi and O. mengae were obtained from Ishihara (1986) and Jeong et al. (2007), respectively. *: data transformed to %TL. HT=Holotype; PT=Paratype; NT=Non-types" captionText-1="TABLE 2. Comparisons of selected characters of O. picta sp. nov. and the similar species. Data of the holotypes of O. hollandi and O. mengae were obtained from Ishihara (1986) and Jeong et al. (2007), respectively." httpUri-0="http://table.plazi.org/id/DF4BCCDAA239FFB2FF2C4266F9E5FD5F" httpUri-1="http://table.plazi.org/id/DF4BCCDAA237FFBCFF2C43B9FB6FFC1E" pageId="1" pageNumber="104" tableUuid-0="DF4BCCDAA239FFB2FF2C4266F9E5FD5F" tableUuid-1="DF4BCCDAA237FFBCFF2C43B9FB6FFC1E">Tables 12</tableCitation>
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TL, adult male, ca.
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, depth ca.
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, bottom trawl,
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<emphasis id="B9404040A23AFFB1FF234676FECDF96B" bold="true" box="[151,267,1663,1689]" pageId="1" pageNumber="104">
<typeStatus id="548F22F0A23AFFB1FF234676FEC0F96B" box="[151,262,1663,1689]" pageId="1" pageNumber="104" type="paratype">Paratype</typeStatus>
.
</emphasis>
ASIZP0081116, adult male,
<quantity id="4CCC31B7A23AFFB1FDE54677FD68F96B" box="[593,686,1662,1689]" metricMagnitude="-1" metricUnit="m" metricValue="4.05" pageId="1" pageNumber="104" unit="mm" value="405.0">405 mm</quantity>
TL (snout slightly damaged), collected together with the
<typeStatus id="548F22F0A23AFFB1FA864676FA5EF96B" box="[1330,1432,1663,1689]" pageId="1" pageNumber="104" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C32ECFD9A23AFFB1FF7346ABFC35F84B" pageId="1" pageNumber="104" type="diagnosis">
<paragraph id="8B8B9C52A23AFFB1FF7346ABFC35F84B" blockId="1.[151,1437,1591,1977]" pageId="1" pageNumber="104">
<emphasis id="B9404040A23AFFB1FF7346ABFE84F94E" bold="true" box="[199,322,1698,1724]" pageId="1" pageNumber="104">Diagnosis.</emphasis>
A medium-sized species of
<taxonomicName id="4C34E7D1A23AFFB1FDCB46AAFD2CF94F" authorityName="Ishiyama" authorityYear="1958" box="[639,746,1699,1725]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="104" phylum="Chordata" rank="genus">
<emphasis id="B9404040A23AFFB1FDCB46AAFD2CF94F" box="[639,746,1699,1725]" italics="true" pageId="1" pageNumber="104">Okamejei</emphasis>
</taxonomicName>
(~
<quantity id="4CCC31B7A23AFFB1FCB346AAFCA1F94F" box="[775,871,1699,1725]" metricMagnitude="-1" metricUnit="m" metricValue="4.29" pageId="1" pageNumber="104" unit="mm" value="429.0">429 mm</quantity>
TL) with yellowish brown dorsal surface, densely mottled with small black circular to irregular spots; pale blotches present but indistinct; absence of prominent pair of pectoral markings and posterior ocellus; ventral surface entirely whitish; ventral sensory pores dark edged; and the following combination of characters: disc width 57.663.2% TL, disc length 75.478.1% disc width; tail long, length 116118% distance from snout tip to rear of cloaca; tail very slender, height 80.084.4% width at its midlength; snout narrow and pointed, preorbital snout length 22.832.1% disc length; first dorsal-fin base length 20.925.0% distance from first dorsal-fin origin to tail tip; tip of clasper without component funnel; nuchal thorn small; total pectoral radials 7376; trunk centra 2731; total centra 123126.
</paragraph>
</subSubSection>
<caption id="DF4BCCDAA239FFB2FF2C4266F9E5FD5F" ID-Table-UUID="DF4BCCDAA239FFB2FF2C4266F9E5FD5F" httpUri="http://table.plazi.org/id/DF4BCCDAA239FFB2FF2C4266F9E5FD5F" pageId="2" pageNumber="105" startId="2.[152,245,623,649]" targetBox="[162,2021,699,1525]" targetIsTable="true" targetPageId="2">
<paragraph id="8B8B9C52A239FFB2FF2C4266F9E5FD5F" blockId="2.[151,2037,623,685]" pageId="2" pageNumber="105">
<emphasis id="B9404040A239FFB2FF2C4266FED6FD7B" bold="true" box="[152,272,623,649]" pageId="2" pageNumber="105">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="4C34E7D1A239FFB2FDBF4266FDA7FD7B" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[523,609,623,649]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A239FFB2FDBF4266FDA7FD7B" box="[523,609,623,649]" italics="true" pageId="2" pageNumber="105">O. picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A239FFB2FDDD4266FD79FD7B" bold="true" box="[617,703,623,649]" pageId="2" pageNumber="105">
<taxonomicNameLabel id="A273FD3BA239FFB2FDDD4266FD79FD7B" box="[617,703,623,649]" pageId="2" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C34E7D1A239FFB2FD4F4266FCB0FD7B" box="[763,886,623,649]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A239FFB2FD4F4266FCB0FD7B" box="[763,886,623,649]" italics="true" pageId="2" pageNumber="105">O. hollandi</emphasis>
</taxonomicName>
. Mean values are given in parentheses. Data of the holotypes of
<taxonomicName id="4C34E7D1A239FFB2F9864266F96BFD7B" box="[1586,1709,623,649]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A239FFB2F9864266F96BFD7B" box="[1586,1709,623,649]" italics="true" pageId="2" pageNumber="105">O. hollandi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34E7D1A239FFB2F9564266F890FD7B" authorityName="Jeong, Nakabo &amp; Wu" authorityYear="2007" box="[1762,1878,623,649]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A239FFB2F9564266F890FD7B" box="[1762,1878,623,649]" italics="true" pageId="2" pageNumber="105">O. mengae</emphasis>
</taxonomicName>
were obtained from Ishihara (1986) and
<bibRefCitation id="EFA5E1A3A239FFB2FE1D429AFDB2FD5F" author="Jeong, C. - H. &amp; Nakabo, T. &amp; Wu, H. - L." box="[425,628,659,685]" pageId="2" pageNumber="105" pagination="57 - 65" refId="ref7491" refString="Jeong, C. - H., Nakabo, T. &amp; Wu, H. - L. (2007) A new species of skate (Chondrichthyes: Rajidae), Okamejei mengae from the South China Sea. Korean Journal of Ichthyology, 19 (1), 57 - 65." type="journal article" year="2007">
Jeong
<emphasis id="B9404040A239FFB2FE5A429AFDE2FD5F" box="[494,548,659,685]" italics="true" pageId="2" pageNumber="105">et al.</emphasis>
(2007)
</bibRefCitation>
, respectively. *: data transformed to %TL. HT=Holotype; PT=Paratype; NT=Non-types
</paragraph>
</caption>
<paragraph id="8B8B9C52A239FFB2F997429AF9E1FD5F" blockId="2.[151,2037,623,685]" box="[1571,1575,659,685]" pageId="2" pageNumber="105">.</paragraph>
<paragraph id="8B8B9C52A239FFB2FE5642B2F804FA27" pageId="2" pageNumber="105">
<table id="F9346EF2A239004FFF1642B2F823FA07" box="[162,2021,699,1525]" gridcols="12" gridrows="12" pageId="2" pageNumber="105">
<tr id="35049E10A239004FFF1642B2F823FD26" box="[162,2021,699,724]" gridrow="0" pageId="2" pageNumber="105" rowspan-0="1" rowspan-3="1" rowspan-6="1" rowspan-9="1">
<th id="76D5F76CA239004FFE7742B2FD8DFD26" box="[451,587,699,724]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="2" pageNumber="105">
<taxonomicName id="4C34E7D1A239FFB2FE5642B2FDF2FD26" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[482,564,699,724]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="picta">O. picta</taxonomicName>
</th>
<th id="76D5F76CA239004FFD6242B2FC50FD26" box="[726,918,699,724]" gridcol="4" gridrow="0" pageId="2" pageNumber="105">
<taxonomicName id="4C34E7D1A239FFB2FD6242B2FC8AFD26" box="[726,844,699,724]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="hollandi">O. hollandi</taxonomicName>
</th>
<th id="76D5F76CA239004FFC7842B2FBFBFD26" box="[972,1085,699,724]" gridcol="5" gridrow="0" pageId="2" pageNumber="105">
<taxonomicName id="4C34E7D1A239FFB2FC7842B2FBFBFD26" authorityName="Jeong, Nakabo &amp; Wu" authorityYear="2007" box="[972,1085,699,724]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="mengae">O. mengae</taxonomicName>
</th>
<th id="76D5F76CA239004FFACB42B2F9C3FD26" box="[1407,1541,699,724]" colspan="2" colspanRight="1" gridcol="7" gridrow="0" pageId="2" pageNumber="105">
<taxonomicName id="4C34E7D1A239FFB2FA2042B2FA20FD26" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[1428,1510,699,724]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="picta">O. picta</taxonomicName>
</th>
<th id="76D5F76CA239004FF93642B2F8FBFD26" box="[1666,1853,699,724]" gridcol="10" gridrow="0" pageId="2" pageNumber="105">
<taxonomicName id="4C34E7D1A239FFB2F93642B2F93EFD26" box="[1666,1784,699,724]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="hollandi">O. hollandi</taxonomicName>
</th>
<th id="76D5F76CA239004FF8C042B2F823FD26" box="[1908,2021,699,724]" gridcol="11" gridrow="0" pageId="2" pageNumber="105">
<taxonomicName id="4C34E7D1A239FFB2F8C042B2F823FD26" authorityName="Jeong, Nakabo &amp; Wu" authorityYear="2007" box="[1908,2021,699,724]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="105" phylum="Chordata" rank="species" species="mengae">O. mengae</taxonomicName>
</th>
</tr>
<tr id="35049E10A239004FFF164303F823FCD1" box="[162,2021,778,803]" gridrow="1" pageId="2" pageNumber="105" rowspan-0="1" rowspan-6="1">
<td id="76D5F76CA239004FFE774303FE29FCD1" box="[451,495,778,803]" gridcol="1" gridrow="1" pageId="2" pageNumber="105">HT</td>
<td id="76D5F76CA239004FFDAB4303FD8DFCD1" box="[543,587,778,803]" gridcol="2" gridrow="1" pageId="2" pageNumber="105">PT</td>
<td id="76D5F76CA239004FFD3A4303FD7CFCD1" box="[654,698,778,803]" gridcol="3" gridrow="1" pageId="2" pageNumber="105">HT</td>
<td id="76D5F76CA239004FFD624303FC50FCD1" box="[726,918,778,803]" gridcol="4" gridrow="1" pageId="2" pageNumber="105">NT (n=8)</td>
<td id="76D5F76CA239004FFC784303FBFBFCD1" box="[972,1085,778,803]" gridcol="5" gridrow="1" pageId="2" pageNumber="105">HT</td>
<td id="76D5F76CA239004FFACB4303FA6DFCD1" box="[1407,1451,778,803]" gridcol="7" gridrow="1" pageId="2" pageNumber="105">HT</td>
<td id="76D5F76CA239004FFA6D4303F9C3FCD1" box="[1497,1541,778,803]" gridcol="8" gridrow="1" pageId="2" pageNumber="105">PT</td>
<td id="76D5F76CA239004FF98F4303F9A0FCD1" box="[1595,1638,778,803]" gridcol="9" gridrow="1" pageId="2" pageNumber="105">HT</td>
<td id="76D5F76CA239004FF9364303F8FBFCD1" box="[1666,1853,778,803]" gridcol="10" gridrow="1" pageId="2" pageNumber="105">NT (n=8)</td>
<td id="76D5F76CA239004FF8C04303F823FCD1" box="[1908,2021,778,803]" gridcol="11" gridrow="1" pageId="2" pageNumber="105">HT</td>
</tr>
<tr id="35049E10A239004FFF16433BF823FCB9" box="[162,2021,818,843]" gridrow="2" pageId="2" pageNumber="105" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA239004FFF16433BFE5BFCB9" box="[162,413,818,843]" gridcol="0" gridrow="2" pageId="2" pageNumber="105">Total length (mm)</th>
<td id="76D5F76CA239004FFE77433BFE29FCB9" box="[451,495,818,843]" gridcol="1" gridrow="2" pageId="2" pageNumber="105">429</td>
<td id="76D5F76CA239004FFDAB433BFD8DFCB9" box="[543,587,818,843]" gridcol="2" gridrow="2" pageId="2" pageNumber="105">405</td>
<td id="76D5F76CA239004FFD3A433BFD7CFCB9" box="[654,698,818,843]" gridcol="3" gridrow="2" pageId="2" pageNumber="105">412</td>
<td id="76D5F76CA239004FFD62433BFC50FCB9" box="[726,918,818,843]" gridcol="4" gridrow="2" pageId="2" pageNumber="105">350480</td>
<td id="76D5F76CA239004FFC78433BFBFBFCB9" box="[972,1085,818,843]" gridcol="5" gridrow="2" pageId="2" pageNumber="105">295</td>
<td id="76D5F76CA239004FF936433BF8FBFCB9" box="[1666,1853,818,843]" gridcol="10" gridrow="2" pageId="2" pageNumber="105">% TL</td>
</tr>
<tr id="35049E10A239004FFF164373F823FC41" box="[162,2021,890,947]" gridrow="3" pageId="2" pageNumber="105" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
<td id="76D5F76CA239004FFD624373FC50FC41" box="[726,918,890,947]" gridcol="4" gridrow="3" pageId="2" pageNumber="105">% TL</td>
<td id="76D5F76CA239004FFBC54373FA86FC41" box="[1137,1344,890,947]" gridcol="6" gridrow="3" pageId="2" pageNumber="105">Nasal curtain (total width)</td>
<td id="76D5F76CA239004FFACB4373FA6DFC41" box="[1407,1451,890,947]" gridcol="7" gridrow="3" pageId="2" pageNumber="105">8.8</td>
<td id="76D5F76CA239004FFA6D4373F9C3FC41" box="[1497,1541,890,947]" gridcol="8" gridrow="3" pageId="2" pageNumber="105">7.8</td>
<td id="76D5F76CA239004FF98F4373F9A0FC41" box="[1595,1638,890,947]" gridcol="9" gridrow="3" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF9364373F8FBFC41" box="[1666,1853,890,947]" gridcol="10" gridrow="3" pageId="2" pageNumber="105">8.19.1 (8.5)</td>
<td id="76D5F76CA239004FF8C04373F823FC41" box="[1908,2021,890,947]" gridcol="11" gridrow="3" pageId="2" pageNumber="105">-</td>
</tr>
<tr id="35049E10A239004FFF1643CAF823FC0E" box="[162,2021,963,1020]" gridrow="4" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF1643CAFE5BFC0E" box="[162,413,963,1020]" gridcol="0" gridrow="4" pageId="2" pageNumber="105">Disc width</th>
<td id="76D5F76CA239004FFE7743CAFE29FC0E" box="[451,495,963,1020]" gridcol="1" gridrow="4" pageId="2" pageNumber="105">57.6</td>
<td id="76D5F76CA239004FFDAB43CAFD8DFC0E" box="[543,587,963,1020]" gridcol="2" gridrow="4" pageId="2" pageNumber="105">63.2</td>
<td id="76D5F76CA239004FFD3A43CAFD7CFC0E" box="[654,698,963,1020]" gridcol="3" gridrow="4" pageId="2" pageNumber="105">63.6</td>
<td id="76D5F76CA239004FFD6243CAFC50FC0E" box="[726,918,963,1020]" gridcol="4" gridrow="4" pageId="2" pageNumber="105">62.768.5 (65.0)</td>
<td id="76D5F76CA239004FFC7843CAFBFBFC0E" box="[972,1085,963,1020]" gridcol="5" gridrow="4" pageId="2" pageNumber="105">63.6</td>
<td id="76D5F76CA239004FFBC543CAFA86FC0E" box="[1137,1344,963,1020]" gridcol="6" gridrow="4" pageId="2" pageNumber="105">Nasal curtain (min. width)</td>
<td id="76D5F76CA239004FFACB43CAFA6DFC0E" box="[1407,1451,963,1020]" gridcol="7" gridrow="4" pageId="2" pageNumber="105">5.0</td>
<td id="76D5F76CA239004FFA6D43CAF9C3FC0E" box="[1497,1541,963,1020]" gridcol="8" gridrow="4" pageId="2" pageNumber="105">3.9</td>
<td id="76D5F76CA239004FF98F43CAF9A0FC0E" box="[1595,1638,963,1020]" gridcol="9" gridrow="4" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF93643CAF8FBFC0E" box="[1666,1853,963,1020]" gridcol="10" gridrow="4" pageId="2" pageNumber="105">4.25.6 (5.2)</td>
<td id="76D5F76CA239004FF8C043CAF823FC0E" box="[1908,2021,963,1020]" gridcol="11" gridrow="4" pageId="2" pageNumber="105">4.6</td>
</tr>
<tr id="35049E10A239004FFF164402F823FBB6" box="[162,2021,1035,1092]" gridrow="5" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF164402FE5BFBB6" box="[162,413,1035,1092]" gridcol="0" gridrow="5" pageId="2" pageNumber="105">Disc length</th>
<td id="76D5F76CA239004FFE774402FE29FBB6" box="[451,495,1035,1092]" gridcol="1" gridrow="5" pageId="2" pageNumber="105">45.0</td>
<td id="76D5F76CA239004FFDAB4402FD8DFBB6" box="[543,587,1035,1092]" gridcol="2" gridrow="5" pageId="2" pageNumber="105">47.7</td>
<td id="76D5F76CA239004FFD3A4402FD7CFBB6" box="[654,698,1035,1092]" gridcol="3" gridrow="5" pageId="2" pageNumber="105">52.7</td>
<td id="76D5F76CA239004FFD624402FC50FBB6" box="[726,918,1035,1092]" gridcol="4" gridrow="5" pageId="2" pageNumber="105">51.655.8 (53.2)</td>
<td id="76D5F76CA239004FFC784402FBFBFBB6" box="[972,1085,1035,1092]" gridcol="5" gridrow="5" pageId="2" pageNumber="105">55.7</td>
<td id="76D5F76CA239004FFBC54402FA86FBB6" box="[1137,1344,1035,1092]" gridcol="6" gridrow="5" pageId="2" pageNumber="105">Nasal curtain (lobe width)</td>
<td id="76D5F76CA239004FFACB4402FA6DFBB6" box="[1407,1451,1035,1092]" gridcol="7" gridrow="5" pageId="2" pageNumber="105">2.2</td>
<td id="76D5F76CA239004FFA6D4402F9C3FBB6" box="[1497,1541,1035,1092]" gridcol="8" gridrow="5" pageId="2" pageNumber="105">2.6</td>
<td id="76D5F76CA239004FF98F4402F9A0FBB6" box="[1595,1638,1035,1092]" gridcol="9" gridrow="5" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF9364402F8FBFBB6" box="[1666,1853,1035,1092]" gridcol="10" gridrow="5" pageId="2" pageNumber="105">1.62.1 (1.8)</td>
<td id="76D5F76CA239004FF8C04402F823FBB6" box="[1908,2021,1035,1092]" gridcol="11" gridrow="5" pageId="2" pageNumber="105">1.6</td>
</tr>
<tr id="35049E10A239004FFF16445AF823FB7E" box="[162,2021,1107,1164]" gridrow="6" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF16445AFE5BFB7E" box="[162,413,1107,1164]" gridcol="0" gridrow="6" pageId="2" pageNumber="105">Snout length to maximum width</th>
<td id="76D5F76CA239004FFE77445AFE29FB7E" box="[451,495,1107,1164]" gridcol="1" gridrow="6" pageId="2" pageNumber="105">33.0</td>
<td id="76D5F76CA239004FFDAB445AFD8DFB7E" box="[543,587,1107,1164]" gridcol="2" gridrow="6" pageId="2" pageNumber="105">28.0</td>
<td id="76D5F76CA239004FFD3A445AFD7CFB7E" box="[654,698,1107,1164]" gridcol="3" gridrow="6" pageId="2" pageNumber="105">32.0</td>
<td id="76D5F76CA239004FFD62445AFC50FB7E" box="[726,918,1107,1164]" gridcol="4" gridrow="6" pageId="2" pageNumber="105">29.832.1 (30.9)</td>
<td id="76D5F76CA239004FFC78445AFBFBFB7E" box="[972,1085,1107,1164]" gridcol="5" gridrow="6" pageId="2" pageNumber="105">32.9</td>
<td id="76D5F76CA239004FFBC5445AFA86FB7E" box="[1137,1344,1107,1164]" gridcol="6" gridrow="6" pageId="2" pageNumber="105">Width of first gill opening</td>
<td id="76D5F76CA239004FFACB445AFA6DFB7E" box="[1407,1451,1107,1164]" gridcol="7" gridrow="6" pageId="2" pageNumber="105">0.8</td>
<td id="76D5F76CA239004FFA6D445AF9C3FB7E" box="[1497,1541,1107,1164]" gridcol="8" gridrow="6" pageId="2" pageNumber="105">1.2</td>
<td id="76D5F76CA239004FF98F445AF9A0FB7E" box="[1595,1638,1107,1164]" gridcol="9" gridrow="6" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF936445AF8FBFB7E" box="[1666,1853,1107,1164]" gridcol="10" gridrow="6" pageId="2" pageNumber="105">1.12.0 (1.7)</td>
<td id="76D5F76CA239004FF8C0445AF823FB7E" box="[1908,2021,1107,1164]" gridcol="11" gridrow="6" pageId="2" pageNumber="105">1.6</td>
</tr>
<tr id="35049E10A239004FFF164492F823FB26" box="[162,2021,1179,1236]" gridrow="7" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF164492FE5BFB26" box="[162,413,1179,1236]" gridcol="0" gridrow="7" pageId="2" pageNumber="105">Snout length (preorbital)</th>
<td id="76D5F76CA239004FFE774492FE29FB26" box="[451,495,1179,1236]" gridcol="1" gridrow="7" pageId="2" pageNumber="105">14.5</td>
<td id="76D5F76CA239004FFDAB4492FD8DFB26" box="[543,587,1179,1236]" gridcol="2" gridrow="7" pageId="2" pageNumber="105">10.9</td>
<td id="76D5F76CA239004FFD3A4492FD7CFB26" box="[654,698,1179,1236]" gridcol="3" gridrow="7" pageId="2" pageNumber="105">14.1</td>
<td id="76D5F76CA239004FFD624492FC50FB26" box="[726,918,1179,1236]" gridcol="4" gridrow="7" pageId="2" pageNumber="105">12.815.3 (13.9)</td>
<td id="76D5F76CA239004FFC784492FBFBFB26" box="[972,1085,1179,1236]" gridcol="5" gridrow="7" pageId="2" pageNumber="105">17.6</td>
<td id="76D5F76CA239004FFBC54492FA86FB26" box="[1137,1344,1179,1236]" gridcol="6" gridrow="7" pageId="2" pageNumber="105">Width of fifth gill opening</td>
<td id="76D5F76CA239004FFACB4492FA6DFB26" box="[1407,1451,1179,1236]" gridcol="7" gridrow="7" pageId="2" pageNumber="105">0.8</td>
<td id="76D5F76CA239004FFA6D4492F9C3FB26" box="[1497,1541,1179,1236]" gridcol="8" gridrow="7" pageId="2" pageNumber="105">0.8</td>
<td id="76D5F76CA239004FF98F4492F9A0FB26" box="[1595,1638,1179,1236]" gridcol="9" gridrow="7" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF9364492F8FBFB26" box="[1666,1853,1179,1236]" gridcol="10" gridrow="7" pageId="2" pageNumber="105">1.01.5 (1.1)</td>
<td id="76D5F76CA239004FF8C04492F823FB26" box="[1908,2021,1179,1236]" gridcol="11" gridrow="7" pageId="2" pageNumber="105">0.7</td>
</tr>
<tr id="35049E10A239004FFF1644EDF823FAEF" box="[162,2021,1252,1309]" gridrow="8" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF1644EDFE5BFAEF" box="[162,413,1252,1309]" gridcol="0" gridrow="8" pageId="2" pageNumber="105">Snout to spiracle</th>
<td id="76D5F76CA239004FFE7744EDFE29FAEF" box="[451,495,1252,1309]" gridcol="1" gridrow="8" pageId="2" pageNumber="105">19.1</td>
<td id="76D5F76CA239004FFDAB44EDFD8DFAEF" box="[543,587,1252,1309]" gridcol="2" gridrow="8" pageId="2" pageNumber="105">13.7</td>
<td id="76D5F76CA239004FFD3A44EDFD7CFAEF" box="[654,698,1252,1309]" gridcol="3" gridrow="8" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FFD6244EDFC50FAEF" box="[726,918,1252,1309]" gridcol="4" gridrow="8" pageId="2" pageNumber="105">18.120.5 (19.0)</td>
<td id="76D5F76CA239004FFC7844EDFBFBFAEF" box="[972,1085,1252,1309]" gridcol="5" gridrow="8" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FFBC544EDFA86FAEF" box="[1137,1344,1252,1309]" gridcol="6" gridrow="8" pageId="2" pageNumber="105">Distance between first gill openings</td>
<td id="76D5F76CA239004FFACB44EDFA6DFAEF" box="[1407,1451,1252,1309]" gridcol="7" gridrow="8" pageId="2" pageNumber="105">12.1</td>
<td id="76D5F76CA239004FFA6D44EDF9C3FAEF" box="[1497,1541,1252,1309]" gridcol="8" gridrow="8" pageId="2" pageNumber="105">13.7</td>
<td id="76D5F76CA239004FF98F44EDF9A0FAEF" box="[1595,1638,1252,1309]" gridcol="9" gridrow="8" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF93644EDF8FBFAEF" box="[1666,1853,1252,1309]" gridcol="10" gridrow="8" pageId="2" pageNumber="105">11.513.0 (12.3)</td>
<td id="76D5F76CA239004FF8C044EDF823FAEF" box="[1908,2021,1252,1309]" gridcol="11" gridrow="8" pageId="2" pageNumber="105">14.1</td>
</tr>
<tr id="35049E10A239004FFF164525F823FA97" box="[162,2021,1324,1381]" gridrow="9" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF164525FE5BFA97" box="[162,413,1324,1381]" gridcol="0" gridrow="9" pageId="2" pageNumber="105">Head (dorsal length)</th>
<td id="76D5F76CA239004FFE774525FE29FA97" box="[451,495,1324,1381]" gridcol="1" gridrow="9" pageId="2" pageNumber="105">24.5</td>
<td id="76D5F76CA239004FFDAB4525FD8DFA97" box="[543,587,1324,1381]" gridcol="2" gridrow="9" pageId="2" pageNumber="105">24.8</td>
<td id="76D5F76CA239004FFD3A4525FD7CFA97" box="[654,698,1324,1381]" gridcol="3" gridrow="9" pageId="2" pageNumber="105">20.9</td>
<td id="76D5F76CA239004FFD624525FC50FA97" box="[726,918,1324,1381]" gridcol="4" gridrow="9" pageId="2" pageNumber="105">19.321.5 (20.2)</td>
<td id="76D5F76CA239004FFC784525FBFBFA97" box="[972,1085,1324,1381]" gridcol="5" gridrow="9" pageId="2" pageNumber="105">24.7</td>
<td id="76D5F76CA239004FFBC54525FA86FA97" box="[1137,1344,1324,1381]" gridcol="6" gridrow="9" pageId="2" pageNumber="105">Distance between fifth gill openings</td>
<td id="76D5F76CA239004FFACB4525FA6DFA97" box="[1407,1451,1324,1381]" gridcol="7" gridrow="9" pageId="2" pageNumber="105">6.6</td>
<td id="76D5F76CA239004FFA6D4525F9C3FA97" box="[1497,1541,1324,1381]" gridcol="8" gridrow="9" pageId="2" pageNumber="105">7.2</td>
<td id="76D5F76CA239004FF98F4525F9A0FA97" box="[1595,1638,1324,1381]" gridcol="9" gridrow="9" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF9364525F8FBFA97" box="[1666,1853,1324,1381]" gridcol="10" gridrow="9" pageId="2" pageNumber="105">6.37.9 (7.0)</td>
<td id="76D5F76CA239004FF8C04525F823FA97" box="[1908,2021,1324,1381]" gridcol="11" gridrow="9" pageId="2" pageNumber="105">0.7</td>
</tr>
<tr id="35049E10A239004FFF16457DF823FA5F" box="[162,2021,1396,1453]" gridrow="10" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF16457DFE5BFA5F" box="[162,413,1396,1453]" gridcol="0" gridrow="10" pageId="2" pageNumber="105">Orbit diameter</th>
<td id="76D5F76CA239004FFE77457DFE29FA5F" box="[451,495,1396,1453]" gridcol="1" gridrow="10" pageId="2" pageNumber="105">4.1</td>
<td id="76D5F76CA239004FFDAB457DFD8DFA5F" box="[543,587,1396,1453]" gridcol="2" gridrow="10" pageId="2" pageNumber="105">4.5</td>
<td id="76D5F76CA239004FFD3A457DFD7CFA5F" box="[654,698,1396,1453]" gridcol="3" gridrow="10" pageId="2" pageNumber="105">3.7</td>
<td id="76D5F76CA239004FFD62457DFC50FA5F" box="[726,918,1396,1453]" gridcol="4" gridrow="10" pageId="2" pageNumber="105">3.74.2 (4.0)</td>
<td id="76D5F76CA239004FFC78457DFBFBFA5F" box="[972,1085,1396,1453]" gridcol="5" gridrow="10" pageId="2" pageNumber="105">4.3</td>
<td id="76D5F76CA239004FFBC5457DFA86FA5F" box="[1137,1344,1396,1453]" gridcol="6" gridrow="10" pageId="2" pageNumber="105">Clasper (post cloacal length)</td>
<td id="76D5F76CA239004FFACB457DFA6DFA5F" box="[1407,1451,1396,1453]" gridcol="7" gridrow="10" pageId="2" pageNumber="105">23.5</td>
<td id="76D5F76CA239004FFA6D457DF9C3FA5F" box="[1497,1541,1396,1453]" gridcol="8" gridrow="10" pageId="2" pageNumber="105">26.3</td>
<td id="76D5F76CA239004FF98F457DF9A0FA5F" box="[1595,1638,1396,1453]" gridcol="9" gridrow="10" pageId="2" pageNumber="105">-</td>
<td id="76D5F76CA239004FF936457DF8FBFA5F" box="[1666,1853,1396,1453]" gridcol="10" gridrow="10" pageId="2" pageNumber="105">21.923.5 (22.7)</td>
<td id="76D5F76CA239004FF8C0457DF823FA5F" box="[1908,2021,1396,1453]" gridcol="11" gridrow="10" pageId="2" pageNumber="105">-</td>
</tr>
<tr id="35049E10A239004FFF1645B5F823FA07" box="[162,2021,1468,1525]" gridrow="11" pageId="2" pageNumber="105">
<th id="76D5F76CA239004FFF1645B5FE5BFA07" box="[162,413,1468,1525]" gridcol="0" gridrow="11" pageId="2" pageNumber="105">Orbit + spiracle length</th>
<td id="76D5F76CA239004FFE7745B5FE29FA07" box="[451,495,1468,1525]" gridcol="1" gridrow="11" pageId="2" pageNumber="105">5.6</td>
<td id="76D5F76CA239004FFDAB45B5FD8DFA07" box="[543,587,1468,1525]" gridcol="2" gridrow="11" pageId="2" pageNumber="105">5.6</td>
<td id="76D5F76CA239004FFD3A45B5FD7CFA07" box="[654,698,1468,1525]" gridcol="3" gridrow="11" pageId="2" pageNumber="105">5.2</td>
<td id="76D5F76CA239004FFD6245B5FC50FA07" box="[726,918,1468,1525]" gridcol="4" gridrow="11" pageId="2" pageNumber="105">5.05.9 (5.5)</td>
<td id="76D5F76CA239004FFC7845B5FBFBFA07" box="[972,1085,1468,1525]" gridcol="5" gridrow="11" pageId="2" pageNumber="105">5.6</td>
<td id="76D5F76CA239004FFBC545B5FA86FA07" box="[1137,1344,1468,1525]" gridcol="6" gridrow="11" pageId="2" pageNumber="105">Length of anterior pelvic lobe</td>
<td id="76D5F76CA239004FFACB45B5FA6DFA07" box="[1407,1451,1468,1525]" gridcol="7" gridrow="11" pageId="2" pageNumber="105">13.0</td>
<td id="76D5F76CA239004FFA6D45B5F9C3FA07" box="[1497,1541,1468,1525]" gridcol="8" gridrow="11" pageId="2" pageNumber="105">12.8</td>
<td id="76D5F76CA239004FF98F45B5F9A0FA07" box="[1595,1638,1468,1525]" gridcol="9" gridrow="11" pageId="2" pageNumber="105">11.9</td>
<td id="76D5F76CA239004FF93645B5F8FBFA07" box="[1666,1853,1468,1525]" gridcol="10" gridrow="11" pageId="2" pageNumber="105">10.312.1 (11.4)</td>
<td id="76D5F76CA239004FF8C045B5F823FA07" box="[1908,2021,1468,1525]" gridcol="11" gridrow="11" pageId="2" pageNumber="105">14.2</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8B9C52A239FFB2F975460CF82CF9EC" blockId="2.[1729,2026,1541,1566]" box="[1729,2026,1541,1566]" pageId="2" pageNumber="105">
<emphasis id="B9404040A239FFB2F975460CF82CF9EC" box="[1729,2026,1541,1566]" italics="true" pageId="2" pageNumber="105">
<tableNote id="76D29DDCA239FFB2F975460CF82CF9EC" box="[1729,2026,1541,1566]" pageId="2" pageNumber="105" targetBox="[162,1986,778,1525]" targetPageId="2">...Continued on the next page</tableNote>
</emphasis>
</paragraph>
<caption id="DF4BCCDAA238FFB3FF234250FE43FD80" box="[151,389,601,626]" isContinuationCaption="true" pageId="3" pageNumber="106" startId="3.[151,235,601,626]" targetBox="[151,2009,643,1546]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="8B8B9C52A238FFB3FF234250FE43FD80" blockId="3.[151,389,601,626]" box="[151,389,601,626]" pageId="3" pageNumber="106">
<emphasis id="B9404040A238FFB3FF234250FEC3FD80" bold="true" box="[151,261,601,626]" pageId="3" pageNumber="106">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B8B9C52A238FFB3FE63428AF862FA18" pageId="3" pageNumber="106">
<table id="F9346EF2A238004FFF23428AF81FF9F8" box="[151,2009,643,1546]" gridcols="12" gridrows="14" pageId="3" pageNumber="106">
<tr id="35049E10A238004FFF23428AF81FFD6E" box="[151,2009,643,668]" gridrow="0" pageId="3" pageNumber="106" rowspan-0="1" rowspan-3="1" rowspan-6="1" rowspan-9="1">
<th id="76D5F76CA238004FFE0C428AFD86FD6E" box="[440,576,643,668]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="3" pageNumber="106">
<taxonomicName id="4C34E7D1A238FFB3FE63428AFDEFFD6E" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[471,553,643,668]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="106" phylum="Chordata" rank="species" species="picta">O. picta</taxonomicName>
</th>
<th id="76D5F76CA238004FFD7F428AFC4CFD6E" box="[715,906,643,668]" gridcol="4" gridrow="0" pageId="3" pageNumber="106">
<taxonomicName id="4C34E7D1A238FFB3FD7F428AFC87FD6E" box="[715,833,643,668]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="106" phylum="Chordata" rank="species" species="hollandi">O. hollandi</taxonomicName>
</th>
<th id="76D5F76CA238004FFC75428AFBF4FD6E" box="[961,1074,643,668]" gridcol="5" gridrow="0" pageId="3" pageNumber="106">
<taxonomicName id="4C34E7D1A238FFB3FC75428AFBF4FD6E" authorityName="Jeong, Nakabo &amp; Wu" authorityYear="2007" box="[961,1074,643,668]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="106" phylum="Chordata" rank="species" species="mengae">O. mengae</taxonomicName>
</th>
<th id="76D5F76CA238004FFAC0428AFA3CFD6E" box="[1396,1530,643,668]" colspan="2" colspanRight="1" gridcol="7" gridrow="0" pageId="3" pageNumber="106">
<taxonomicName id="4C34E7D1A238FFB3FA3D428AFA1DFD6E" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[1417,1499,643,668]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="106" phylum="Chordata" rank="species" species="picta">O. picta</taxonomicName>
</th>
<th id="76D5F76CA238004FF9C2428AF8F4FD6E" box="[1654,1842,643,668]" gridcol="10" gridrow="0" pageId="3" pageNumber="106">
<taxonomicName id="4C34E7D1A238FFB3F9C2428AF92AFD6E" box="[1654,1772,643,668]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="106" phylum="Chordata" rank="species" species="hollandi">O. hollandi</taxonomicName>
</th>
<th id="76D5F76CA238004FF8DC428AF81FFD6E" box="[1896,2009,643,668]" gridcol="11" gridrow="0" pageId="3" pageNumber="106">
<taxonomicName id="4C34E7D1A238FFB3F8DC428AF81FFD6E" box="[1896,2009,643,668]" pageId="3" pageNumber="106">
<emphasis id="B9404040A238FFB3F8DC428AF81FFD6E" bold="true" box="[1896,2009,643,668]" italics="true" pageId="3" pageNumber="106">O. mengae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35049E10A238004FFF2342A2F81FFD36" box="[151,2009,683,708]" gridrow="1" pageId="3" pageNumber="106" rowspan-0="1" rowspan-6="1">
<td id="76D5F76CA238004FFE0C42A2FE22FD36" box="[440,484,683,708]" gridcol="1" gridrow="1" pageId="3" pageNumber="106">HT</td>
<td id="76D5F76CA238004FFDA042A2FD86FD36" box="[532,576,683,708]" gridcol="2" gridrow="1" pageId="3" pageNumber="106">PT</td>
<td id="76D5F76CA238004FFD3642A2FD68FD36" box="[642,686,683,708]" gridcol="3" gridrow="1" pageId="3" pageNumber="106">HT</td>
<td id="76D5F76CA238004FFD7F42A2FC4CFD36" box="[715,906,683,708]" gridcol="4" gridrow="1" pageId="3" pageNumber="106">NT (n=8)</td>
<td id="76D5F76CA238004FFC7542A2FBF4FD36" box="[961,1074,683,708]" gridcol="5" gridrow="1" pageId="3" pageNumber="106">HT</td>
<td id="76D5F76CA238004FFAC042A2FA66FD36" box="[1396,1440,683,708]" gridcol="7" gridrow="1" pageId="3" pageNumber="106">HT</td>
<td id="76D5F76CA238004FFA7A42A2FA3CFD36" box="[1486,1530,683,708]" gridcol="8" gridrow="1" pageId="3" pageNumber="106">PT</td>
<td id="76D5F76CA238004FF98442A2F99AFD36" box="[1584,1628,683,708]" gridcol="9" gridrow="1" pageId="3" pageNumber="106">HT</td>
<td id="76D5F76CA238004FF9C242A2F8F4FD36" box="[1654,1842,683,708]" gridcol="10" gridrow="1" pageId="3" pageNumber="106">NT (n=8)</td>
<td id="76D5F76CA238004FF8DC42A2F81FFD36" box="[1896,2009,683,708]" gridcol="11" gridrow="1" pageId="3" pageNumber="106">HT</td>
</tr>
<tr id="35049E10A238004FFF2342DDF81FFCFF" box="[151,2009,724,781]" gridrow="2" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF2342DDFE56FCFF" box="[151,400,724,781]" gridcol="0" gridrow="2" pageId="3" pageNumber="106">Spiracle length</th>
<td id="76D5F76CA238004FFE0C42DDFE22FCFF" box="[440,484,724,781]" gridcol="1" gridrow="2" pageId="3" pageNumber="106">2.4</td>
<td id="76D5F76CA238004FFDA042DDFD86FCFF" box="[532,576,724,781]" gridcol="2" gridrow="2" pageId="3" pageNumber="106">2.7</td>
<td id="76D5F76CA238004FFD3642DDFD68FCFF" box="[642,686,724,781]" gridcol="3" gridrow="2" pageId="3" pageNumber="106">5.2</td>
<td id="76D5F76CA238004FFD7F42DDFC4CFCFF" box="[715,906,724,781]" gridcol="4" gridrow="2" pageId="3" pageNumber="106">2.23.2 (2.8)</td>
<td id="76D5F76CA238004FFC7542DDFBF4FCFF" box="[961,1074,724,781]" gridcol="5" gridrow="2" pageId="3" pageNumber="106">3.1</td>
<td id="76D5F76CA238004FFBFA42DDFA8AFCFF" box="[1102,1356,724,781]" gridcol="6" gridrow="2" pageId="3" pageNumber="106">Length of posterior pelvic lobe</td>
<td id="76D5F76CA238004FFAC042DDFA66FCFF" box="[1396,1440,724,781]" gridcol="7" gridrow="2" pageId="3" pageNumber="106">17.6</td>
<td id="76D5F76CA238004FFA7A42DDFA3CFCFF" box="[1486,1530,724,781]" gridcol="8" gridrow="2" pageId="3" pageNumber="106">18.5</td>
<td id="76D5F76CA238004FF98442DDF99AFCFF" box="[1584,1628,724,781]" gridcol="9" gridrow="2" pageId="3" pageNumber="106">15.0</td>
<td id="76D5F76CA238004FF9C242DDF8F4FCFF" box="[1654,1842,724,781]" gridcol="10" gridrow="2" pageId="3" pageNumber="106">14.517.4 (15.8)</td>
<td id="76D5F76CA238004FF8DC42DDF81FFCFF" box="[1896,2009,724,781]" gridcol="11" gridrow="2" pageId="3" pageNumber="106">16.8</td>
</tr>
<tr id="35049E10A238004FFF234315F81FFCC7" box="[151,2009,796,821]" gridrow="3" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF234315FE56FCC7" box="[151,400,796,821]" gridcol="0" gridrow="3" pageId="3" pageNumber="106">Distance between orbits</th>
<td id="76D5F76CA238004FFE0C4315FE22FCC7" box="[440,484,796,821]" gridcol="1" gridrow="3" pageId="3" pageNumber="106">3.3</td>
<td id="76D5F76CA238004FFDA04315FD86FCC7" box="[532,576,796,821]" gridcol="2" gridrow="3" pageId="3" pageNumber="106">3.5</td>
<td id="76D5F76CA238004FFD364315FD68FCC7" box="[642,686,796,821]" gridcol="3" gridrow="3" pageId="3" pageNumber="106">4.0</td>
<td id="76D5F76CA238004FFD7F4315FC4CFCC7" box="[715,906,796,821]" gridcol="4" gridrow="3" pageId="3" pageNumber="106">3.44.5 (3.9)</td>
<td id="76D5F76CA238004FFC754315FBF4FCC7" box="[961,1074,796,821]" gridcol="5" gridrow="3" pageId="3" pageNumber="106">4.0</td>
<td id="76D5F76CA238004FFBFA4315FA8AFCC7" box="[1102,1356,796,821]" gridcol="6" gridrow="3" pageId="3" pageNumber="106">Pelvic base width</td>
<td id="76D5F76CA238004FFAC04315FA66FCC7" box="[1396,1440,796,821]" gridcol="7" gridrow="3" pageId="3" pageNumber="106">12.7</td>
<td id="76D5F76CA238004FFA7A4315FA3CFCC7" box="[1486,1530,796,821]" gridcol="8" gridrow="3" pageId="3" pageNumber="106">12.9</td>
<td id="76D5F76CA238004FF9844315F99AFCC7" box="[1584,1628,796,821]" gridcol="9" gridrow="3" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C24315F8F4FCC7" box="[1654,1842,796,821]" gridcol="10" gridrow="3" pageId="3" pageNumber="106">9.914.4 (53.2)</td>
<td id="76D5F76CA238004FF8DC4315F81FFCC7" box="[1896,2009,796,821]" gridcol="11" gridrow="3" pageId="3" pageNumber="106">-</td>
</tr>
<tr id="35049E10A238004FFF23434CF81FFC8C" box="[151,2009,837,894]" gridrow="4" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF23434CFE56FC8C" box="[151,400,837,894]" gridcol="0" gridrow="4" pageId="3" pageNumber="106">Distance between spiracles</th>
<td id="76D5F76CA238004FFE0C434CFE22FC8C" box="[440,484,837,894]" gridcol="1" gridrow="4" pageId="3" pageNumber="106">5.5</td>
<td id="76D5F76CA238004FFDA0434CFD86FC8C" box="[532,576,837,894]" gridcol="2" gridrow="4" pageId="3" pageNumber="106">5.5</td>
<td id="76D5F76CA238004FFD36434CFD68FC8C" box="[642,686,837,894]" gridcol="3" gridrow="4" pageId="3" pageNumber="106">9.3</td>
<td id="76D5F76CA238004FFD7F434CFC4CFC8C" box="[715,906,837,894]" gridcol="4" gridrow="4" pageId="3" pageNumber="106">5.66.1 (5.8)</td>
<td id="76D5F76CA238004FFC75434CFBF4FC8C" box="[961,1074,837,894]" gridcol="5" gridrow="4" pageId="3" pageNumber="106">6.4</td>
<td id="76D5F76CA238004FFBFA434CFA8AFC8C" box="[1102,1356,837,894]" gridcol="6" gridrow="4" pageId="3" pageNumber="106">Tail at axil of pelvic fin (width)</td>
<td id="76D5F76CA238004FFAC0434CFA66FC8C" box="[1396,1440,837,894]" gridcol="7" gridrow="4" pageId="3" pageNumber="106">2.5</td>
<td id="76D5F76CA238004FFA7A434CFA3CFC8C" box="[1486,1530,837,894]" gridcol="8" gridrow="4" pageId="3" pageNumber="106">3.4</td>
<td id="76D5F76CA238004FF984434CF99AFC8C" box="[1584,1628,837,894]" gridcol="9" gridrow="4" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C2434CF8F4FC8C" box="[1654,1842,837,894]" gridcol="10" gridrow="4" pageId="3" pageNumber="106">2.84.1 (3.4)</td>
<td id="76D5F76CA238004FF8DC434CF81FFC8C" box="[1896,2009,837,894]" gridcol="11" gridrow="4" pageId="3" pageNumber="106">5.4</td>
</tr>
<tr id="35049E10A238004FFF234386F81FFC3A" box="[151,2009,911,968]" gridrow="5" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF234386FE56FC3A" box="[151,400,911,968]" gridcol="0" gridrow="5" pageId="3" pageNumber="106">Snout to cloaca</th>
<td id="76D5F76CA238004FFE0C4386FE22FC3A" box="[440,484,911,968]" gridcol="1" gridrow="5" pageId="3" pageNumber="106">45.6</td>
<td id="76D5F76CA238004FFDA04386FD86FC3A" box="[532,576,911,968]" gridcol="2" gridrow="5" pageId="3" pageNumber="106">47.5</td>
<td id="76D5F76CA238004FFD364386FD68FC3A" box="[642,686,911,968]" gridcol="3" gridrow="5" pageId="3" pageNumber="106">51.5</td>
<td id="76D5F76CA238004FFD7F4386FC4CFC3A" box="[715,906,911,968]" gridcol="4" gridrow="5" pageId="3" pageNumber="106">46.951.5 (48.6)</td>
<td id="76D5F76CA238004FFC754386FBF4FC3A" box="[961,1074,911,968]" gridcol="5" gridrow="5" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FFBFA4386FA8AFC3A" box="[1102,1356,911,968]" gridcol="6" gridrow="5" pageId="3" pageNumber="106">Tail at axil of pelvic fin (height)</td>
<td id="76D5F76CA238004FFAC04386FA66FC3A" box="[1396,1440,911,968]" gridcol="7" gridrow="5" pageId="3" pageNumber="106">2.0</td>
<td id="76D5F76CA238004FFA7A4386FA3CFC3A" box="[1486,1530,911,968]" gridcol="8" gridrow="5" pageId="3" pageNumber="106">2.0</td>
<td id="76D5F76CA238004FF9844386F99AFC3A" box="[1584,1628,911,968]" gridcol="9" gridrow="5" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C24386F8F4FC3A" box="[1654,1842,911,968]" gridcol="10" gridrow="5" pageId="3" pageNumber="106">1.41.8 (1.6)</td>
<td id="76D5F76CA238004FF8DC4386F81FFC3A" box="[1896,2009,911,968]" gridcol="11" gridrow="5" pageId="3" pageNumber="106">2.3</td>
</tr>
<tr id="35049E10A238004FFF2343DEF81FFBE2" box="[151,2009,983,1040]" gridrow="6" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF2343DEFE56FBE2" box="[151,400,983,1040]" gridcol="0" gridrow="6" pageId="3" pageNumber="106">Cloaca to first dorsal-fin origin</th>
<td id="76D5F76CA238004FFE0C43DEFE22FBE2" box="[440,484,983,1040]" gridcol="1" gridrow="6" pageId="3" pageNumber="106">32.3</td>
<td id="76D5F76CA238004FFDA043DEFD86FBE2" box="[532,576,983,1040]" gridcol="2" gridrow="6" pageId="3" pageNumber="106">32.1</td>
<td id="76D5F76CA238004FFD3643DEFD68FBE2" box="[642,686,983,1040]" gridcol="3" gridrow="6" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FFD7F43DEFC4CFBE2" box="[715,906,983,1040]" gridcol="4" gridrow="6" pageId="3" pageNumber="106">25.129.2 (27.2)</td>
<td id="76D5F76CA238004FFC7543DEFBF4FBE2" box="[961,1074,983,1040]" gridcol="5" gridrow="6" pageId="3" pageNumber="106">30.3</td>
<td id="76D5F76CA238004FFBFA43DEFA8AFBE2" box="[1102,1356,983,1040]" gridcol="6" gridrow="6" pageId="3" pageNumber="106">Tail at midlength (width)</td>
<td id="76D5F76CA238004FFAC043DEFA66FBE2" box="[1396,1440,983,1040]" gridcol="7" gridrow="6" pageId="3" pageNumber="106">1.8</td>
<td id="76D5F76CA238004FFA7A43DEFA3CFBE2" box="[1486,1530,983,1040]" gridcol="8" gridrow="6" pageId="3" pageNumber="106">2.0</td>
<td id="76D5F76CA238004FF98443DEF99AFBE2" box="[1584,1628,983,1040]" gridcol="9" gridrow="6" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C243DEF8F4FBE2" box="[1654,1842,983,1040]" gridcol="10" gridrow="6" pageId="3" pageNumber="106">1.42.0 (1.6)</td>
<td id="76D5F76CA238004FF8DC43DEF81FFBE2" box="[1896,2009,983,1040]" gridcol="11" gridrow="6" pageId="3" pageNumber="106">2.1</td>
</tr>
<tr id="35049E10A238004FFF234429F81FFBAB" box="[151,2009,1056,1113]" gridrow="7" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF234429FE56FBAB" box="[151,400,1056,1113]" gridcol="0" gridrow="7" pageId="3" pageNumber="106">Cloaca to second dorsal- fin origin</th>
<td id="76D5F76CA238004FFE0C4429FE22FBAB" box="[440,484,1056,1113]" gridcol="1" gridrow="7" pageId="3" pageNumber="106">41.1</td>
<td id="76D5F76CA238004FFDA04429FD86FBAB" box="[532,576,1056,1113]" gridcol="2" gridrow="7" pageId="3" pageNumber="106">41.9</td>
<td id="76D5F76CA238004FFD364429FD68FBAB" box="[642,686,1056,1113]" gridcol="3" gridrow="7" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FFD7F4429FC4CFBAB" box="[715,906,1056,1113]" gridcol="4" gridrow="7" pageId="3" pageNumber="106">34.139.6 (36.8)</td>
<td id="76D5F76CA238004FFC754429FBF4FBAB" box="[961,1074,1056,1113]" gridcol="5" gridrow="7" pageId="3" pageNumber="106">38.1</td>
<td id="76D5F76CA238004FFBFA4429FA8AFBAB" box="[1102,1356,1056,1113]" gridcol="6" gridrow="7" pageId="3" pageNumber="106">Tail at midlength (height)</td>
<td id="76D5F76CA238004FFAC04429FA66FBAB" box="[1396,1440,1056,1113]" gridcol="7" gridrow="7" pageId="3" pageNumber="106">1.4</td>
<td id="76D5F76CA238004FFA7A4429FA3CFBAB" box="[1486,1530,1056,1113]" gridcol="8" gridrow="7" pageId="3" pageNumber="106">1.7</td>
<td id="76D5F76CA238004FF9844429F99AFBAB" box="[1584,1628,1056,1113]" gridcol="9" gridrow="7" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C24429F8F4FBAB" box="[1654,1842,1056,1113]" gridcol="10" gridrow="7" pageId="3" pageNumber="106">1.11.3 (1.1)</td>
<td id="76D5F76CA238004FF8DC4429F81FFBAB" box="[1896,2009,1056,1113]" gridcol="11" gridrow="7" pageId="3" pageNumber="106">1.2</td>
</tr>
<tr id="35049E10A238004FFF234461F81FFB53" box="[151,2009,1128,1185]" gridrow="8" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF234461FE56FB53" box="[151,400,1128,1185]" gridcol="0" gridrow="8" pageId="3" pageNumber="106">Cloaca to caudal origin</th>
<td id="76D5F76CA238004FFE0C4461FE22FB53" box="[440,484,1128,1185]" gridcol="1" gridrow="8" pageId="3" pageNumber="106">46.2</td>
<td id="76D5F76CA238004FFDA04461FD86FB53" box="[532,576,1128,1185]" gridcol="2" gridrow="8" pageId="3" pageNumber="106">48.9</td>
<td id="76D5F76CA238004FFD364461FD68FB53" box="[642,686,1128,1185]" gridcol="3" gridrow="8" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FFD7F4461FC4CFB53" box="[715,906,1128,1185]" gridcol="4" gridrow="8" pageId="3" pageNumber="106">40.947.3 (43.8)</td>
<td id="76D5F76CA238004FFC754461FBF4FB53" box="[961,1074,1128,1185]" gridcol="5" gridrow="8" pageId="3" pageNumber="106">43.7</td>
<td id="76D5F76CA238004FFBFA4461FA8AFB53" box="[1102,1356,1128,1185]" gridcol="6" gridrow="8" pageId="3" pageNumber="106">Tail at first dorsal-fin origin (width)</td>
<td id="76D5F76CA238004FFAC04461FA66FB53" box="[1396,1440,1128,1185]" gridcol="7" gridrow="8" pageId="3" pageNumber="106">1.2</td>
<td id="76D5F76CA238004FFA7A4461FA3CFB53" box="[1486,1530,1128,1185]" gridcol="8" gridrow="8" pageId="3" pageNumber="106">1.1</td>
<td id="76D5F76CA238004FF9844461F99AFB53" box="[1584,1628,1128,1185]" gridcol="9" gridrow="8" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C24461F8F4FB53" box="[1654,1842,1128,1185]" gridcol="10" gridrow="8" pageId="3" pageNumber="106">1.01.4 (1.2)</td>
<td id="76D5F76CA238004FF8DC4461F81FFB53" box="[1896,2009,1128,1185]" gridcol="11" gridrow="8" pageId="3" pageNumber="106">2.0</td>
</tr>
<tr id="35049E10A238004FFF2344B9F81FFAFB" box="[151,2009,1200,1289]" gridrow="9" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF2344B9FE56FAFB" box="[151,400,1200,1289]" gridcol="0" gridrow="9" pageId="3" pageNumber="106">Distance from cloaca to caudal-fin tip (tail length)</th>
<td id="76D5F76CA238004FFE0C44B9FE22FAFB" box="[440,484,1200,1289]" gridcol="1" gridrow="9" pageId="3" pageNumber="106">53.6</td>
<td id="76D5F76CA238004FFDA044B9FD86FAFB" box="[532,576,1200,1289]" gridcol="2" gridrow="9" pageId="3" pageNumber="106">55.1</td>
<td id="76D5F76CA238004FFD3644B9FD68FAFB" box="[642,686,1200,1289]" gridcol="3" gridrow="9" pageId="3" pageNumber="106">47.8</td>
<td id="76D5F76CA238004FFD7F44B9FC4CFAFB" box="[715,906,1200,1289]" gridcol="4" gridrow="9" pageId="3" pageNumber="106">48.453.4 (51.0)</td>
<td id="76D5F76CA238004FFC7544B9FBF4FAFB" box="[961,1074,1200,1289]" gridcol="5" gridrow="9" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FFBFA44B9FA8AFAFB" box="[1102,1356,1200,1289]" gridcol="6" gridrow="9" pageId="3" pageNumber="106">Tail at first dorsal-fin origin (height)</td>
<td id="76D5F76CA238004FFAC044B9FA66FAFB" box="[1396,1440,1200,1289]" gridcol="7" gridrow="9" pageId="3" pageNumber="106">1.2</td>
<td id="76D5F76CA238004FFA7A44B9FA3CFAFB" box="[1486,1530,1200,1289]" gridcol="8" gridrow="9" pageId="3" pageNumber="106">1.0</td>
<td id="76D5F76CA238004FF98444B9F99AFAFB" box="[1584,1628,1200,1289]" gridcol="9" gridrow="9" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C244B9F8F4FAFB" box="[1654,1842,1200,1289]" gridcol="10" gridrow="9" pageId="3" pageNumber="106">0.91.1 (1.0)</td>
<td id="76D5F76CA238004FF8DC44B9F81FFAFB" box="[1896,2009,1200,1289]" gridcol="11" gridrow="9" pageId="3" pageNumber="106">1.2</td>
</tr>
<tr id="35049E10A238004FFF234511F81FFAA3" box="[151,2009,1304,1361]" gridrow="10" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF234511FE56FAA3" box="[151,400,1304,1361]" gridcol="0" gridrow="10" pageId="3" pageNumber="106">Pre-upper jaw length</th>
<td id="76D5F76CA238004FFE0C4511FE22FAA3" box="[440,484,1304,1361]" gridcol="1" gridrow="10" pageId="3" pageNumber="106">14.7</td>
<td id="76D5F76CA238004FFDA04511FD86FAA3" box="[532,576,1304,1361]" gridcol="2" gridrow="10" pageId="3" pageNumber="106">11.9</td>
<td id="76D5F76CA238004FFD364511FD68FAA3" box="[642,686,1304,1361]" gridcol="3" gridrow="10" pageId="3" pageNumber="106">14.3</td>
<td id="76D5F76CA238004FFD7F4511FC4CFAA3" box="[715,906,1304,1361]" gridcol="4" gridrow="10" pageId="3" pageNumber="106">14.116.9 (15.4)</td>
<td id="76D5F76CA238004FFC754511FBF4FAA3" box="[961,1074,1304,1361]" gridcol="5" gridrow="10" pageId="3" pageNumber="106">18.0</td>
<td id="76D5F76CA238004FFBFA4511FA8AFAA3" box="[1102,1356,1304,1361]" gridcol="6" gridrow="10" pageId="3" pageNumber="106">First dorsal-fin base length</td>
<td id="76D5F76CA238004FFAC04511FA66FAA3" box="[1396,1440,1304,1361]" gridcol="7" gridrow="10" pageId="3" pageNumber="106">4.4</td>
<td id="76D5F76CA238004FFA7A4511FA3CFAA3" box="[1486,1530,1304,1361]" gridcol="8" gridrow="10" pageId="3" pageNumber="106">5.2</td>
<td id="76D5F76CA238004FF9844511F99AFAA3" box="[1584,1628,1304,1361]" gridcol="9" gridrow="10" pageId="3" pageNumber="106">4.3</td>
<td id="76D5F76CA238004FF9C24511F8F4FAA3" box="[1654,1842,1304,1361]" gridcol="10" gridrow="10" pageId="3" pageNumber="106">3.55.1 (4.7)</td>
<td id="76D5F76CA238004FF8DC4511F81FFAA3" box="[1896,2009,1304,1361]" gridcol="11" gridrow="10" pageId="3" pageNumber="106">4.7</td>
</tr>
<tr id="35049E10A238004FFF234568F81FFA88" box="[151,2009,1377,1402]" gridrow="11" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF234568FE56FA88" box="[151,400,1377,1402]" gridcol="0" gridrow="11" pageId="3" pageNumber="106">Pre-nasal length</th>
<td id="76D5F76CA238004FFE0C4568FE22FA88" box="[440,484,1377,1402]" gridcol="1" gridrow="11" pageId="3" pageNumber="106">11.7</td>
<td id="76D5F76CA238004FFDA04568FD86FA88" box="[532,576,1377,1402]" gridcol="2" gridrow="11" pageId="3" pageNumber="106">9.1</td>
<td id="76D5F76CA238004FFD364568FD68FA88" box="[642,686,1377,1402]" gridcol="3" gridrow="11" pageId="3" pageNumber="106">11.7</td>
<td id="76D5F76CA238004FFD7F4568FC4CFA88" box="[715,906,1377,1402]" gridcol="4" gridrow="11" pageId="3" pageNumber="106">10.513.0 (11.7)</td>
<td id="76D5F76CA238004FFC754568FBF4FA88" box="[961,1074,1377,1402]" gridcol="5" gridrow="11" pageId="3" pageNumber="106">14.9</td>
<td id="76D5F76CA238004FFBFA4568FA8AFA88" box="[1102,1356,1377,1402]" gridcol="6" gridrow="11" pageId="3" pageNumber="106">First dorsal-fin height</td>
<td id="76D5F76CA238004FFAC04568FA66FA88" box="[1396,1440,1377,1402]" gridcol="7" gridrow="11" pageId="3" pageNumber="106">2.3</td>
<td id="76D5F76CA238004FFA7A4568FA3CFA88" box="[1486,1530,1377,1402]" gridcol="8" gridrow="11" pageId="3" pageNumber="106">2.8</td>
<td id="76D5F76CA238004FF9844568F99AFA88" box="[1584,1628,1377,1402]" gridcol="9" gridrow="11" pageId="3" pageNumber="106">1.9</td>
<td id="76D5F76CA238004FF9C24568F8F4FA88" box="[1654,1842,1377,1402]" gridcol="10" gridrow="11" pageId="3" pageNumber="106">2.33.7 (2.7)</td>
<td id="76D5F76CA238004FF8DC4568F81FFA88" box="[1896,2009,1377,1402]" gridcol="11" gridrow="11" pageId="3" pageNumber="106">2.9</td>
</tr>
<tr id="35049E10A238004FFF234580F81FFA30" box="[151,2009,1417,1474]" gridrow="12" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF234580FE56FA30" box="[151,400,1417,1474]" gridcol="0" gridrow="12" pageId="3" pageNumber="106">Ventral head length (to fifth gill)</th>
<td id="76D5F76CA238004FFE0C4580FE22FA30" box="[440,484,1417,1474]" gridcol="1" gridrow="12" pageId="3" pageNumber="106">28.1</td>
<td id="76D5F76CA238004FFDA04580FD86FA30" box="[532,576,1417,1474]" gridcol="2" gridrow="12" pageId="3" pageNumber="106">28.3</td>
<td id="76D5F76CA238004FFD364580FD68FA30" box="[642,686,1417,1474]" gridcol="3" gridrow="12" pageId="3" pageNumber="106">27.6</td>
<td id="76D5F76CA238004FFD7F4580FC4CFA30" box="[715,906,1417,1474]" gridcol="4" gridrow="12" pageId="3" pageNumber="106">27.229.8 (28.2)</td>
<td id="76D5F76CA238004FFC754580FBF4FA30" box="[961,1074,1417,1474]" gridcol="5" gridrow="12" pageId="3" pageNumber="106">30.6</td>
<td id="76D5F76CA238004FFBFA4580FA8AFA30" box="[1102,1356,1417,1474]" gridcol="6" gridrow="12" pageId="3" pageNumber="106">First dorsal-fin origin to caudal-fin tip</td>
<td id="76D5F76CA238004FFAC04580FA66FA30" box="[1396,1440,1417,1474]" gridcol="7" gridrow="12" pageId="3" pageNumber="106">20.9</td>
<td id="76D5F76CA238004FFA7A4580FA3CFA30" box="[1486,1530,1417,1474]" gridcol="8" gridrow="12" pageId="3" pageNumber="106">20.9</td>
<td id="76D5F76CA238004FF9844580F99AFA30" box="[1584,1628,1417,1474]" gridcol="9" gridrow="12" pageId="3" pageNumber="106">22.2</td>
<td id="76D5F76CA238004FF9C24580F8F4FA30" box="[1654,1842,1417,1474]" gridcol="10" gridrow="12" pageId="3" pageNumber="106">22.225.7 (23.5)</td>
<td id="76D5F76CA238004FF8DC4580F81FFA30" box="[1896,2009,1417,1474]" gridcol="11" gridrow="12" pageId="3" pageNumber="106">18.1</td>
</tr>
<tr id="35049E10A238004FFF2345D8F81FF9F8" box="[151,2009,1489,1546]" gridrow="13" pageId="3" pageNumber="106">
<th id="76D5F76CA238004FFF2345D8FE56F9F8" box="[151,400,1489,1546]" gridcol="0" gridrow="13" pageId="3" pageNumber="106">Mouth width</th>
<td id="76D5F76CA238004FFE0C45D8FE22F9F8" box="[440,484,1489,1546]" gridcol="1" gridrow="13" pageId="3" pageNumber="106">7.4</td>
<td id="76D5F76CA238004FFDA045D8FD86F9F8" box="[532,576,1489,1546]" gridcol="2" gridrow="13" pageId="3" pageNumber="106">7.4</td>
<td id="76D5F76CA238004FFD3645D8FD68F9F8" box="[642,686,1489,1546]" gridcol="3" gridrow="13" pageId="3" pageNumber="106">8.3</td>
<td id="76D5F76CA238004FFD7F45D8FC4CF9F8" box="[715,906,1489,1546]" gridcol="4" gridrow="13" pageId="3" pageNumber="106">7.08.0 (7.6)</td>
<td id="76D5F76CA238004FFC7545D8FBF4F9F8" box="[961,1074,1489,1546]" gridcol="5" gridrow="13" pageId="3" pageNumber="106">7.8</td>
<td id="76D5F76CA238004FFBFA45D8FA8AF9F8" box="[1102,1356,1489,1546]" gridcol="6" gridrow="13" pageId="3" pageNumber="106">Second dorsal-fin origin to caudal-fin tip</td>
<td id="76D5F76CA238004FFAC045D8FA66F9F8" box="[1396,1440,1489,1546]" gridcol="7" gridrow="13" pageId="3" pageNumber="106">12.2</td>
<td id="76D5F76CA238004FFA7A45D8FA3CF9F8" box="[1486,1530,1489,1546]" gridcol="8" gridrow="13" pageId="3" pageNumber="106">11.4</td>
<td id="76D5F76CA238004FF98445D8F99AF9F8" box="[1584,1628,1489,1546]" gridcol="9" gridrow="13" pageId="3" pageNumber="106">-</td>
<td id="76D5F76CA238004FF9C245D8F8F4F9F8" box="[1654,1842,1489,1546]" gridcol="10" gridrow="13" pageId="3" pageNumber="106">13.214.5 (14.0)</td>
<td id="76D5F76CA238004FF8DC45D8F81FF9F8" box="[1896,2009,1489,1546]" gridcol="11" gridrow="13" pageId="3" pageNumber="106">-</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8B9C52A238FFB3F9014613F819F9C1" blockId="3.[1717,2015,1562,1587]" box="[1717,2015,1562,1587]" pageId="3" pageNumber="106">
<emphasis id="B9404040A238FFB3F9014613F819F9C1" box="[1717,2015,1562,1587]" italics="true" pageId="3" pageNumber="106">
<tableNote id="76D29DDCA238FFB3F9014613F819F9C1" box="[1717,2015,1562,1587]" pageId="3" pageNumber="106" targetBox="[151,2009,643,1546]" targetPageId="3">...Continued on the next page</tableNote>
</emphasis>
</paragraph>
<caption id="DF4BCCDAA23FFFB4FF2342A4FE43FD34" box="[151,389,685,710]" isContinuationCaption="true" pageId="4" pageNumber="107" startId="4.[151,235,685,710]" targetBox="[151,2009,726,1509]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="8B8B9C52A23FFFB4FF2342A4FE43FD34" blockId="4.[151,389,685,710]" box="[151,389,685,710]" pageId="4" pageNumber="107">
<emphasis id="B9404040A23FFFB4FF2342A4FEC3FD34" bold="true" box="[151,261,685,710]" pageId="4" pageNumber="107">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B8B9C52A23FFFB4FE6342DFFBC0FA17" pageId="4" pageNumber="107">
<table id="F9346EF2A23F004FFF2342DFF81FFA17" box="[151,2009,726,1509]" gridcols="12" gridrows="19" pageId="4" pageNumber="107">
<tr id="35049E10A23F004FFF2342DFF81FFD1D" box="[151,2009,726,751]" gridrow="0" pageId="4" pageNumber="107" rowspan-0="1" rowspan-3="1" rowspan-6="1" rowspan-9="1">
<th id="76D5F76CA23F004FFE0042DFFD82FD1D" box="[436,580,726,751]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="4" pageNumber="107">
<taxonomicName id="4C34E7D1A23FFFB4FE6342DFFDEFFD1D" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[471,553,726,751]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="107" phylum="Chordata" rank="species" species="picta">O. picta</taxonomicName>
</th>
<th id="76D5F76CA23F004FFD7F42DFFCB1FD1D" box="[715,887,726,751]" gridcol="4" gridrow="0" pageId="4" pageNumber="107">
<taxonomicName id="4C34E7D1A23FFFB4FD7F42DFFC87FD1D" box="[715,833,726,751]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="107" phylum="Chordata" rank="species" species="hollandi">O. hollandi</taxonomicName>
</th>
<th id="76D5F76CA23F004FFC7542DFFBF4FD1D" box="[961,1074,726,751]" gridcol="5" gridrow="0" pageId="4" pageNumber="107">
<taxonomicName id="4C34E7D1A23FFFB4FC7542DFFBF4FD1D" authorityName="Jeong, Nakabo &amp; Wu" authorityYear="2007" box="[961,1074,726,751]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="107" phylum="Chordata" rank="species" species="mengae">O. mengae</taxonomicName>
</th>
<th id="76D5F76CA23F004FFACD42DFFA32FD1D" box="[1401,1524,726,751]" colspan="2" colspanRight="1" gridcol="7" gridrow="0" pageId="4" pageNumber="107">
<taxonomicName id="4C34E7D1A23FFFB4FA3D42DFFA1DFD1D" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[1417,1499,726,751]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="107" phylum="Chordata" rank="species" species="picta">O. picta</taxonomicName>
</th>
<th id="76D5F76CA23F004FF9C242DFF8E6FD1D" box="[1654,1824,726,751]" gridcol="10" gridrow="0" pageId="4" pageNumber="107">
<taxonomicName id="4C34E7D1A23FFFB4F9C242DFF92AFD1D" box="[1654,1772,726,751]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="107" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A23FFFB4F9C242DFF92AFD1D" bold="true" box="[1654,1772,726,751]" italics="true" pageId="4" pageNumber="107">O. hollandi</emphasis>
</taxonomicName>
</th>
<th id="76D5F76CA23F004FF8DC42DFF81FFD1D" box="[1896,2009,726,751]" gridcol="11" gridrow="0" pageId="4" pageNumber="107">
<taxonomicName id="4C34E7D1A23FFFB4F8DC42DFF81FFD1D" box="[1896,2009,726,751]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="107" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A23FFFB4F8DC42DFF81FFD1D" bold="true" box="[1896,2009,726,751]" italics="true" pageId="4" pageNumber="107">O. mengae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35049E10A23F004FFF2342F6F81FFCEA" box="[151,2009,767,792]" gridrow="1" pageId="4" pageNumber="107" rowspan-0="1" rowspan-6="1">
<td id="76D5F76CA23F004FFE0042F6FE2EFCEA" box="[436,488,767,792]" gridcol="1" gridrow="1" pageId="4" pageNumber="107">HT</td>
<td id="76D5F76CA23F004FFDA442F6FD82FCEA" box="[528,580,767,792]" gridcol="2" gridrow="1" pageId="4" pageNumber="107">PT</td>
<td id="76D5F76CA23F004FFD3C42F6FD6CFCEA" box="[648,682,767,792]" gridcol="3" gridrow="1" pageId="4" pageNumber="107">HT</td>
<td id="76D5F76CA23F004FFD7F42F6FCB1FCEA" box="[715,887,767,792]" gridcol="4" gridrow="1" pageId="4" pageNumber="107">NT (n=8)</td>
<td id="76D5F76CA23F004FFC7542F6FBF4FCEA" box="[961,1074,767,792]" gridcol="5" gridrow="1" pageId="4" pageNumber="107">HT</td>
<td id="76D5F76CA23F004FFACD42F6FA5DFCEA" box="[1401,1435,767,792]" gridcol="7" gridrow="1" pageId="4" pageNumber="107">HT</td>
<td id="76D5F76CA23F004FFA6042F6FA32FCEA" box="[1492,1524,767,792]" gridcol="8" gridrow="1" pageId="4" pageNumber="107">PT</td>
<td id="76D5F76CA23F004FF98142F6F991FCEA" box="[1589,1623,767,792]" gridcol="9" gridrow="1" pageId="4" pageNumber="107">HT</td>
<td id="76D5F76CA23F004FF9C242F6F8E6FCEA" box="[1654,1824,767,792]" gridcol="10" gridrow="1" pageId="4" pageNumber="107">NT (n=8)</td>
<td id="76D5F76CA23F004FF8DC42F6F81FFCEA" box="[1896,2009,767,792]" gridcol="11" gridrow="1" pageId="4" pageNumber="107">HT</td>
</tr>
<tr id="35049E10A23F004FFF234321F81FFC93" box="[151,2009,808,865]" gridrow="2" pageId="4" pageNumber="107">
<th id="76D5F76CA23F004FFF234321FE52FC93" box="[151,404,808,865]" gridcol="0" gridrow="2" pageId="4" pageNumber="107">Distance between nostrils</th>
<td id="76D5F76CA23F004FFE004321FE2EFC93" box="[436,488,808,865]" gridcol="1" gridrow="2" pageId="4" pageNumber="107">6.6</td>
<td id="76D5F76CA23F004FFDA44321FD82FC93" box="[528,580,808,865]" gridcol="2" gridrow="2" pageId="4" pageNumber="107">6.7</td>
<td id="76D5F76CA23F004FFD3C4321FD6CFC93" box="[648,682,808,865]" gridcol="3" gridrow="2" pageId="4" pageNumber="107">7.3</td>
<td id="76D5F76CA23F004FFD7F4321FCB1FC93" box="[715,887,808,865]" gridcol="4" gridrow="2" pageId="4" pageNumber="107">6.87.5 (7.0)</td>
<td id="76D5F76CA23F004FFC754321FBF4FC93" box="[961,1074,808,865]" gridcol="5" gridrow="2" pageId="4" pageNumber="107">7.6</td>
<td id="76D5F76CA23F004FFBC04321FAE0FC93" box="[1140,1318,808,865]" gridcol="6" gridrow="2" pageId="4" pageNumber="107">Interdorsal space</td>
<td id="76D5F76CA23F004FFACD4321FA5DFC93" box="[1401,1435,808,865]" gridcol="7" gridrow="2" pageId="4" pageNumber="107">4.3</td>
<td id="76D5F76CA23F004FFA604321FA32FC93" box="[1492,1524,808,865]" gridcol="8" gridrow="2" pageId="4" pageNumber="107">4.4</td>
<td id="76D5F76CA23F004FF9814321F991FC93" box="[1589,1623,808,865]" gridcol="9" gridrow="2" pageId="4" pageNumber="107">5.2</td>
<td id="76D5F76CA23F004FF9C24321F8E6FC93" box="[1654,1824,808,865]" gridcol="10" gridrow="2" pageId="4" pageNumber="107">3.56.6 (4.8)</td>
<td id="76D5F76CA23F004FF8DC4321F81FFC93" box="[1896,2009,808,865]" gridcol="11" gridrow="2" pageId="4" pageNumber="107">2.9</td>
</tr>
<tr id="35049E10A23F004FFF234379F81FFC7B" box="[151,2009,880,905]" gridrow="3" pageId="4" pageNumber="107">
<th id="76D5F76CA23F004FFF234379FE52FC7B" box="[151,404,880,905]" gridcol="0" gridrow="3" pageId="4" pageNumber="107">Nasal curtain length</th>
<td id="76D5F76CA23F004FFE004379FE2EFC7B" box="[436,488,880,905]" gridcol="1" gridrow="3" pageId="4" pageNumber="107">3.8</td>
<td id="76D5F76CA23F004FFDA44379FD82FC7B" box="[528,580,880,905]" gridcol="2" gridrow="3" pageId="4" pageNumber="107">4.4</td>
<td id="76D5F76CA23F004FFD3C4379FD6CFC7B" box="[648,682,880,905]" gridcol="3" gridrow="3" pageId="4" pageNumber="107">4.1</td>
<td id="76D5F76CA23F004FFD7F4379FCB1FC7B" box="[715,887,880,905]" gridcol="4" gridrow="3" pageId="4" pageNumber="107">4.65.8 (5.3)</td>
<td id="76D5F76CA23F004FFC754379FBF4FC7B" box="[961,1074,880,905]" gridcol="5" gridrow="3" pageId="4" pageNumber="107">5.1</td>
<td id="76D5F76CA23F004FFBC04379FAE0FC7B" box="[1140,1318,880,905]" gridcol="6" gridrow="3" pageId="4" pageNumber="107">Caudal-fin length</td>
<td id="76D5F76CA23F004FFACD4379FA5DFC7B" box="[1401,1435,880,905]" gridcol="7" gridrow="3" pageId="4" pageNumber="107">5.7</td>
<td id="76D5F76CA23F004FFA604379FA32FC7B" box="[1492,1524,880,905]" gridcol="8" gridrow="3" pageId="4" pageNumber="107">5.3</td>
<td id="76D5F76CA23F004FF9814379F991FC7B" box="[1589,1623,880,905]" gridcol="9" gridrow="3" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FF9C24379F8E6FC7B" box="[1654,1824,880,905]" gridcol="10" gridrow="3" pageId="4" pageNumber="107">5.88.0 (7.1)</td>
<td id="76D5F76CA23F004FF8DC4379F81FFC7B" box="[1896,2009,880,905]" gridcol="11" gridrow="3" pageId="4" pageNumber="107">4.4</td>
</tr>
<tr id="35049E10A23F004FFF234391F81FFC43" box="[151,2009,920,945]" gridrow="4" pageId="4" pageNumber="107">
<th id="76D5F76CA23F004FFF234391F81FFC43" box="[151,2009,920,945]" colspan="12" colspanRight="11" gridcol="0" gridrow="4" pageId="4" pageNumber="107">
<emphasis id="B9404040A23FFFB4FF234391FF22FC43" bold="true" box="[151,228,920,945]" pageId="4" pageNumber="107">Counts</emphasis>
</th>
</tr>
<tr id="35049E10A23F004FFF2343C8F81FFC28" box="[151,2009,961,986]" gridrow="5" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF2343C8FE52FC28" box="[151,404,961,986]" gridcol="0" gridrow="5" pageId="4" pageNumber="107">Tooth rows on upper jaw</th>
<td id="76D5F76CA23F004FFE0043C8FE2EFC28" box="[436,488,961,986]" gridcol="1" gridrow="5" pageId="4" pageNumber="107">40</td>
<td id="76D5F76CA23F004FFDA443C8FD82FC28" box="[528,580,961,986]" gridcol="2" gridrow="5" pageId="4" pageNumber="107">41</td>
<td id="76D5F76CA23F004FFD3C43C8FD6CFC28" box="[648,682,961,986]" gridcol="3" gridrow="5" pageId="4" pageNumber="107">43</td>
<td id="76D5F76CA23F004FFD7F43C8FCB1FC28" box="[715,887,961,986]" gridcol="4" gridrow="5" pageId="4" pageNumber="107">4450</td>
<td id="76D5F76CA23F004FFC7543C8FBF4FC28" box="[961,1074,961,986]" gridcol="5" gridrow="5" pageId="4" pageNumber="107">42</td>
</tr>
<tr id="35049E10A23F004FFF2343E0F81FFBF0" box="[151,2009,1001,1026]" gridrow="6" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF2343E0FE52FBF0" box="[151,404,1001,1026]" gridcol="0" gridrow="6" pageId="4" pageNumber="107">Tooth rows on lower jaw</th>
<td id="76D5F76CA23F004FFE0043E0FE2EFBF0" box="[436,488,1001,1026]" gridcol="1" gridrow="6" pageId="4" pageNumber="107">40</td>
<td id="76D5F76CA23F004FFDA443E0FD82FBF0" box="[528,580,1001,1026]" gridcol="2" gridrow="6" pageId="4" pageNumber="107">40</td>
<td id="76D5F76CA23F004FFD3C43E0FD6CFBF0" box="[648,682,1001,1026]" gridcol="3" gridrow="6" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F43E0FCB1FBF0" box="[715,887,1001,1026]" gridcol="4" gridrow="6" pageId="4" pageNumber="107">4247</td>
<td id="76D5F76CA23F004FFC7543E0FBF4FBF0" box="[961,1074,1001,1026]" gridcol="5" gridrow="6" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF234418F81FFBD8" box="[151,2009,1041,1066]" gridrow="7" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF234418FE52FBD8" box="[151,404,1041,1066]" gridcol="0" gridrow="7" pageId="4" pageNumber="107">Trunk centra</th>
<td id="76D5F76CA23F004FFE004418FE2EFBD8" box="[436,488,1041,1066]" gridcol="1" gridrow="7" pageId="4" pageNumber="107">31</td>
<td id="76D5F76CA23F004FFDA44418FD82FBD8" box="[528,580,1041,1066]" gridcol="2" gridrow="7" pageId="4" pageNumber="107">27</td>
<td id="76D5F76CA23F004FFD3C4418FD6CFBD8" box="[648,682,1041,1066]" gridcol="3" gridrow="7" pageId="4" pageNumber="107">27</td>
<td id="76D5F76CA23F004FFD7F4418FCB1FBD8" box="[715,887,1041,1066]" gridcol="4" gridrow="7" pageId="4" pageNumber="107">2830</td>
<td id="76D5F76CA23F004FFC754418FBF4FBD8" box="[961,1074,1041,1066]" gridcol="5" gridrow="7" pageId="4" pageNumber="107">23</td>
</tr>
<tr id="35049E10A23F004FFF234433F81FFBA1" box="[151,2009,1082,1107]" gridrow="8" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF234433FE52FBA1" box="[151,404,1082,1107]" gridcol="0" gridrow="8" pageId="4" pageNumber="107">Monospondylous centra</th>
<td id="76D5F76CA23F004FFE004433FE2EFBA1" box="[436,488,1082,1107]" gridcol="1" gridrow="8" pageId="4" pageNumber="107">29</td>
<td id="76D5F76CA23F004FFDA44433FD82FBA1" box="[528,580,1082,1107]" gridcol="2" gridrow="8" pageId="4" pageNumber="107">24</td>
<td id="76D5F76CA23F004FFD3C4433FD6CFBA1" box="[648,682,1082,1107]" gridcol="3" gridrow="8" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F4433FCB1FBA1" box="[715,887,1082,1107]" gridcol="4" gridrow="8" pageId="4" pageNumber="107">2426</td>
<td id="76D5F76CA23F004FFC754433FBF4FBA1" box="[961,1074,1082,1107]" gridcol="5" gridrow="8" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF23446BF81FFB89" box="[151,2009,1122,1147]" gridrow="9" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF23446BFE52FB89" box="[151,404,1122,1147]" gridcol="0" gridrow="9" pageId="4" pageNumber="107">Predorsal caudal centra</th>
<td id="76D5F76CA23F004FFE00446BFE2EFB89" box="[436,488,1122,1147]" gridcol="1" gridrow="9" pageId="4" pageNumber="107">44</td>
<td id="76D5F76CA23F004FFDA4446BFD82FB89" box="[528,580,1122,1147]" gridcol="2" gridrow="9" pageId="4" pageNumber="107">43</td>
<td id="76D5F76CA23F004FFD3C446BFD6CFB89" box="[648,682,1122,1147]" gridcol="3" gridrow="9" pageId="4" pageNumber="107">42</td>
<td id="76D5F76CA23F004FFD7F446BFCB1FB89" box="[715,887,1122,1147]" gridcol="4" gridrow="9" pageId="4" pageNumber="107">3547</td>
<td id="76D5F76CA23F004FFC75446BFBF4FB89" box="[961,1074,1122,1147]" gridcol="5" gridrow="9" pageId="4" pageNumber="107">50</td>
</tr>
<tr id="35049E10A23F004FFF234483F81FFB51" box="[151,2009,1162,1187]" gridrow="10" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF234483FE52FB51" box="[151,404,1162,1187]" gridcol="0" gridrow="10" pageId="4" pageNumber="107">Predorsal centra</th>
<td id="76D5F76CA23F004FFE004483FE2EFB51" box="[436,488,1162,1187]" gridcol="1" gridrow="10" pageId="4" pageNumber="107">75</td>
<td id="76D5F76CA23F004FFDA44483FD82FB51" box="[528,580,1162,1187]" gridcol="2" gridrow="10" pageId="4" pageNumber="107">70</td>
<td id="76D5F76CA23F004FFD3C4483FD6CFB51" box="[648,682,1162,1187]" gridcol="3" gridrow="10" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F4483FCB1FB51" box="[715,887,1162,1187]" gridcol="4" gridrow="10" pageId="4" pageNumber="107">6376</td>
<td id="76D5F76CA23F004FFC754483FBF4FB51" box="[961,1074,1162,1187]" gridcol="5" gridrow="10" pageId="4" pageNumber="107">73</td>
</tr>
<tr id="35049E10A23F004FFF2344BBF81FFB39" box="[151,2009,1202,1227]" gridrow="11" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF2344BBFE52FB39" box="[151,404,1202,1227]" gridcol="0" gridrow="11" pageId="4" pageNumber="107">Inter-dorsal origin centra</th>
<td id="76D5F76CA23F004FFE0044BBFE2EFB39" box="[436,488,1202,1227]" gridcol="1" gridrow="11" pageId="4" pageNumber="107">16</td>
<td id="76D5F76CA23F004FFDA444BBFD82FB39" box="[528,580,1202,1227]" gridcol="2" gridrow="11" pageId="4" pageNumber="107">16</td>
<td id="76D5F76CA23F004FFD3C44BBFD6CFB39" box="[648,682,1202,1227]" gridcol="3" gridrow="11" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F44BBFCB1FB39" box="[715,887,1202,1227]" gridcol="4" gridrow="11" pageId="4" pageNumber="107">1721</td>
<td id="76D5F76CA23F004FFC7544BBFBF4FB39" box="[961,1074,1202,1227]" gridcol="5" gridrow="11" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF2344D3F81FFB01" box="[151,2009,1242,1267]" gridrow="12" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF2344D3FE52FB01" box="[151,404,1242,1267]" gridcol="0" gridrow="12" pageId="4" pageNumber="107">Diplospondylous centra</th>
<td id="76D5F76CA23F004FFE0044D3FE2EFB01" box="[436,488,1242,1267]" gridcol="1" gridrow="12" pageId="4" pageNumber="107">95</td>
<td id="76D5F76CA23F004FFDA444D3FD82FB01" box="[528,580,1242,1267]" gridcol="2" gridrow="12" pageId="4" pageNumber="107">96</td>
<td id="76D5F76CA23F004FFD3C44D3FD6CFB01" box="[648,682,1242,1267]" gridcol="3" gridrow="12" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F44D3FCB1FB01" box="[715,887,1242,1267]" gridcol="4" gridrow="12" pageId="4" pageNumber="107">6683</td>
<td id="76D5F76CA23F004FFC7544D3FBF4FB01" box="[961,1074,1242,1267]" gridcol="5" gridrow="12" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF23450AF81FFAEE" box="[151,2009,1283,1308]" gridrow="13" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF23450AFE52FAEE" box="[151,404,1283,1308]" gridcol="0" gridrow="13" pageId="4" pageNumber="107">Total centra</th>
<td id="76D5F76CA23F004FFE00450AFE2EFAEE" box="[436,488,1283,1308]" gridcol="1" gridrow="13" pageId="4" pageNumber="107">126</td>
<td id="76D5F76CA23F004FFDA4450AFD82FAEE" box="[528,580,1283,1308]" gridcol="2" gridrow="13" pageId="4" pageNumber="107">123</td>
<td id="76D5F76CA23F004FFD3C450AFD6CFAEE" box="[648,682,1283,1308]" gridcol="3" gridrow="13" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F450AFCB1FAEE" box="[715,887,1283,1308]" gridcol="4" gridrow="13" pageId="4" pageNumber="107">95112</td>
<td id="76D5F76CA23F004FFC75450AFBF4FAEE" box="[961,1074,1283,1308]" gridcol="5" gridrow="13" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF234522F81FFAB6" box="[151,2009,1323,1348]" gridrow="14" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF234522FE52FAB6" box="[151,404,1323,1348]" gridcol="0" gridrow="14" pageId="4" pageNumber="107">Pelvic radials</th>
<td id="76D5F76CA23F004FFE004522FE2EFAB6" box="[436,488,1323,1348]" gridcol="1" gridrow="14" pageId="4" pageNumber="107">1+19</td>
<td id="76D5F76CA23F004FFDA44522FD82FAB6" box="[528,580,1323,1348]" gridcol="2" gridrow="14" pageId="4" pageNumber="107">1+20</td>
<td id="76D5F76CA23F004FFD3C4522FD6CFAB6" box="[648,682,1323,1348]" gridcol="3" gridrow="14" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F4522FCB1FAB6" box="[715,887,1323,1348]" gridcol="4" gridrow="14" pageId="4" pageNumber="107">1+1822</td>
<td id="76D5F76CA23F004FFC754522FBF4FAB6" box="[961,1074,1323,1348]" gridcol="5" gridrow="14" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF23455AF81FFA9E" box="[151,2009,1363,1388]" gridrow="15" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF23455AFE52FA9E" box="[151,404,1363,1388]" gridcol="0" gridrow="15" pageId="4" pageNumber="107">Propterygials</th>
<td id="76D5F76CA23F004FFE00455AFE2EFA9E" box="[436,488,1363,1388]" gridcol="1" gridrow="15" pageId="4" pageNumber="107">29</td>
<td id="76D5F76CA23F004FFDA4455AFD82FA9E" box="[528,580,1363,1388]" gridcol="2" gridrow="15" pageId="4" pageNumber="107">28</td>
<td id="76D5F76CA23F004FFD3C455AFD6CFA9E" box="[648,682,1363,1388]" gridcol="3" gridrow="15" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F455AFCB1FA9E" box="[715,887,1363,1388]" gridcol="4" gridrow="15" pageId="4" pageNumber="107">2932</td>
<td id="76D5F76CA23F004FFC75455AFBF4FA9E" box="[961,1074,1363,1388]" gridcol="5" gridrow="15" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF234572F81FFA66" box="[151,2009,1403,1428]" gridrow="16" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF234572FE52FA66" box="[151,404,1403,1428]" gridcol="0" gridrow="16" pageId="4" pageNumber="107">Mesopterygials</th>
<td id="76D5F76CA23F004FFE004572FE2EFA66" box="[436,488,1403,1428]" gridcol="1" gridrow="16" pageId="4" pageNumber="107">10</td>
<td id="76D5F76CA23F004FFDA44572FD82FA66" box="[528,580,1403,1428]" gridcol="2" gridrow="16" pageId="4" pageNumber="107">8</td>
<td id="76D5F76CA23F004FFD3C4572FD6CFA66" box="[648,682,1403,1428]" gridcol="3" gridrow="16" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F4572FCB1FA66" box="[715,887,1403,1428]" gridcol="4" gridrow="16" pageId="4" pageNumber="107">911</td>
<td id="76D5F76CA23F004FFC754572FBF4FA66" box="[961,1074,1403,1428]" gridcol="5" gridrow="16" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF2345ADF81FFA4F" box="[151,2009,1444,1469]" gridrow="17" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF2345ADFE52FA4F" box="[151,404,1444,1469]" gridcol="0" gridrow="17" pageId="4" pageNumber="107">Metapterygials</th>
<td id="76D5F76CA23F004FFE0045ADFE2EFA4F" box="[436,488,1444,1469]" gridcol="1" gridrow="17" pageId="4" pageNumber="107">37</td>
<td id="76D5F76CA23F004FFDA445ADFD82FA4F" box="[528,580,1444,1469]" gridcol="2" gridrow="17" pageId="4" pageNumber="107">37</td>
<td id="76D5F76CA23F004FFD3C45ADFD6CFA4F" box="[648,682,1444,1469]" gridcol="3" gridrow="17" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F45ADFCB1FA4F" box="[715,887,1444,1469]" gridcol="4" gridrow="17" pageId="4" pageNumber="107">4145</td>
<td id="76D5F76CA23F004FFC7545ADFBF4FA4F" box="[961,1074,1444,1469]" gridcol="5" gridrow="17" pageId="4" pageNumber="107">-</td>
</tr>
<tr id="35049E10A23F004FFF2345C5F81FFA17" box="[151,2009,1484,1509]" gridrow="18" pageId="4" pageNumber="107" rowspan-10="1" rowspan-11="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76D5F76CA23F004FFF2345C5FE52FA17" box="[151,404,1484,1509]" gridcol="0" gridrow="18" pageId="4" pageNumber="107">Total basal radials</th>
<td id="76D5F76CA23F004FFE0045C5FE2EFA17" box="[436,488,1484,1509]" gridcol="1" gridrow="18" pageId="4" pageNumber="107">76</td>
<td id="76D5F76CA23F004FFDA445C5FD82FA17" box="[528,580,1484,1509]" gridcol="2" gridrow="18" pageId="4" pageNumber="107">73</td>
<td id="76D5F76CA23F004FFD3C45C5FD6CFA17" box="[648,682,1484,1509]" gridcol="3" gridrow="18" pageId="4" pageNumber="107">-</td>
<td id="76D5F76CA23F004FFD7F45C5FCB1FA17" box="[715,887,1484,1509]" gridcol="4" gridrow="18" pageId="4" pageNumber="107">8286</td>
<td id="76D5F76CA23F004FFC7545C5FBF4FA17" box="[961,1074,1484,1509]" gridcol="5" gridrow="18" pageId="4" pageNumber="107">96</td>
</tr>
</table>
</paragraph>
<caption id="DF4BCCDAA23EFFB5FF2C45B9FE19FA1E" ID-DOI="http://doi.org/10.5281/zenodo.7895046" ID-Zenodo-Dep="7895046" httpUri="https://zenodo.org/record/7895046/files/figure.png" pageId="5" pageNumber="108" startId="5.[152,255,1456,1480]" targetBox="[151,1436,181,1432]" targetPageId="5" targetType="figure">
<paragraph id="8B8B9C52A23EFFB5FF2C45B9FE19FA1E" blockId="5.[151,1436,1456,1516]" pageId="5" pageNumber="108">
<emphasis id="B9404040A23EFFB5FF2C45B9FEDAFA3A" bold="true" box="[152,284,1456,1480]" pageId="5" pageNumber="108">FIGURE 1.</emphasis>
Fresh coloration of
<taxonomicName id="4C34E7D1A23EFFB5FE4C45B9FD50FA3A" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[504,662,1456,1480]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="108" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A23EFFB5FE4C45B9FD50FA3A" box="[504,662,1456,1480]" italics="true" pageId="5" pageNumber="108">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A23EFFB5FD1445B8FD32FA3B" bold="true" box="[672,756,1457,1481]" pageId="5" pageNumber="108">
<taxonomicNameLabel id="A273FD3BA23EFFB5FD1445B8FD32FA3B" box="[672,756,1457,1481]" pageId="5" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, NMMB-P36013, 429 mm TL, adult male holotype: top, dorsal surface; bottom, ventral surface.
</paragraph>
</caption>
<subSubSection id="C32ECFD9A23EFFBCFF734638FA9BF807" lastPageId="12" lastPageNumber="115" pageId="5" pageNumber="108" type="description">
<paragraph id="8B8B9C52A23EFFB5FF734638FDF9F928" blockId="5.[151,1437,1585,2041]" pageId="5" pageNumber="108">
<emphasis id="B9404040A23EFFB5FF734638FE9FF9B9" bold="true" box="[199,345,1585,1611]" pageId="5" pageNumber="108">Description.</emphasis>
The following data are provided for
<typeStatus id="548F22F0A23EFFB5FD4A4638FCA5F9B9" box="[766,867,1585,1611]" pageId="5" pageNumber="108" type="holotype">holotype</typeStatus>
, followed by the value of
<typeStatus id="548F22F0A23EFFB5FB39463BFB2BF9BE" box="[1165,1261,1586,1612]" pageId="5" pageNumber="108" type="paratype">paratype</typeStatus>
in parentheses. Disc quadrangular, length 78.1% (75.4%) width; angle in front of spiracles 70° (64°); axis of greatest width 60.4% (48.4%) of disc length; anterior margin double concave, moderately concave anteriorly to the snout tip, convex besides eyes, considerably concave besides and posterior to spiracles; apex very narrowly rounded; posterior margin slightly convex; free rear tip rounded.
</paragraph>
<paragraph id="8B8B9C52A23EFFB5FF7346EDFCFAF80A" blockId="5.[151,1437,1585,2041]" pageId="5" pageNumber="108">
Details of dorsal and ventral view of head are shown in
<figureCitation id="130F80D7A23EFFB5FC8046EDFCB0F90C" box="[820,886,1764,1790]" captionStart="FIGURE 3" captionStartId="7.[152,255,1048,1072]" captionTargetBox="[280,1308,181,1024]" captionTargetId="figure-17@7.[280,1308,181,1024]" captionTargetPageId="7" captionText="FIGURE 3. Details of dorsal (A, B) and ventral (C, D) views of snout area of Okamejei picta sp. nov.: A, dorsal view of adult male holotype (NMMB-P36013, 429 mm TL); B, dorsal view of adult male paratype (ASIZP0081116, 405 mm TL); C, ventral view of adult male holotype (NMMB-P36013, 429 mm TL); D, ventral view of adult male paratype (ASIZP0081116, 405 mm TL)." figureDoi="http://doi.org/10.5281/zenodo.7895052" httpUri="https://zenodo.org/record/7895052/files/figure.png" pageId="5" pageNumber="108">Fig. 3</figureCitation>
. Head rather short, preorbital snout length 59.0% (43.9%) head length, distance between orbits 13.4% (13.9%) head length; pre-upper jaw length 52.2% (42.1%) ventral head length. Snout tip produced and very narrowly pointed; fleshy process absent at tip. Distance between orbits 81.2% (76.9%) orbit diameter. Spiracle medium-sized, length 58.9% (60.0%) orbit diameter; its opening in distorted teardrop-shape. Nostril suboval; anterior nasal flap somewhat expanded, lobelike, partly covered beneath nasal curtain; posterior inner margin fully covered by nasal curtain; posterior lobes forming nasal curtain, produced posterolaterally, posterior margin with prominent fringes; distance between nostrils 54.5% (48.5%) distance between first gill slits, 101% (93.0%) distance between fifth gill slits.
</paragraph>
<caption id="DF4BCCDAA23DFFB6FF2C4641FE4FF976" ID-DOI="http://doi.org/10.5281/zenodo.7895048" ID-Zenodo-Dep="7895048" httpUri="https://zenodo.org/record/7895048/files/figure.png" pageId="6" pageNumber="109" startId="6.[152,255,1608,1632]" targetBox="[151,1436,181,1584]" targetPageId="6" targetType="figure">
<paragraph id="8B8B9C52A23DFFB6FF2C4641FE4FF976" blockId="6.[151,1437,1608,1668]" pageId="6" pageNumber="109">
<emphasis id="B9404040A23DFFB6FF2C4641FEDEF992" bold="true" box="[152,280,1608,1632]" pageId="6" pageNumber="109">FIGURE 2.</emphasis>
Fresh coloration of
<taxonomicName id="4C34E7D1A23DFFB6FE574641FDBAF992" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[483,636,1608,1632]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="109" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A23DFFB6FE574641FDBAF992" box="[483,636,1608,1632]" italics="true" pageId="6" pageNumber="109">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A23DFFB6FD364641FD17F992" bold="true" box="[642,721,1608,1632]" pageId="6" pageNumber="109">
<taxonomicNameLabel id="A273FD3BA23DFFB6FD364641FD17F992" box="[642,721,1608,1632]" pageId="6" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ASIZP0081116, 405 mm TL, adult male paratype: top, dorsal surface; bottom, ventral surface.
</paragraph>
</caption>
<paragraph id="8B8B9C52A23DFFB6FF7346DEFE1EF8AE" blockId="6.[151,1437,1750,2028]" pageId="6" pageNumber="109">Upper jaw prominently arched, continuous at symphysis; lower jaw in wide inverted V-shape; part of lateral teeth covered by lobe of nasal curtain. Teeth unicuspid, with subcircular bases near symphysis; arranged in longitudinal rows; main cusps slender, long, sharp, directed posteriorly in upper jaw, directed anteriorly in lower jaw, becoming shorter and oblique laterally.</paragraph>
<paragraph id="8B8B9C52A23DFFB6FF73476FFEFFF81E" blockId="6.[151,1437,1750,2028]" pageId="6" pageNumber="109">Pelvic fins bilobed; anterior lobe short and slender, narrowly rounded distally, lateral margin entire, inner margin considerably incised; posterior lobe greatly elongate, length 17.6% (18.5%) TL, lateral margins straight, free rear tip rounded; inner margin rather shallow, connected to anterolateral margin of clasper; anterior lobe 73.9% (69.0%) posterior lobe.</paragraph>
<caption id="DF4BCCDAA23CFFB7FF2C4411FF00FB6E" ID-DOI="http://doi.org/10.5281/zenodo.7895052" ID-Zenodo-Dep="7895052" httpUri="https://zenodo.org/record/7895052/files/figure.png" pageId="7" pageNumber="110" startId="7.[152,255,1048,1072]" targetBox="[280,1308,181,1024]" targetPageId="7" targetType="figure">
<paragraph id="8B8B9C52A23CFFB7FF2C4411FF00FB6E" blockId="7.[151,1437,1048,1180]" pageId="7" pageNumber="110">
<emphasis id="B9404040A23CFFB7FF2C4411FEDEFBC2" bold="true" box="[152,280,1048,1072]" pageId="7" pageNumber="110">FIGURE 3.</emphasis>
Details of dorsal (A, B) and ventral (C, D) views of snout area of
<taxonomicName id="4C34E7D1A23CFFB7FC034411FB97FBC2" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[951,1105,1048,1072]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="110" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A23CFFB7FC034411FB97FBC2" box="[951,1105,1048,1072]" italics="true" pageId="7" pageNumber="110">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A23CFFB7FBE34411FB61FBC2" bold="true" box="[1111,1191,1048,1072]" pageId="7" pageNumber="110">
<taxonomicNameLabel id="A273FD3BA23CFFB7FBE34411FB61FBC2" box="[1111,1191,1048,1072]" pageId="7" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, dorsal view of adult male holotype (NMMB-P36013, 429 mm TL); B, dorsal view of adult male paratype (ASIZP0081116, 405 mm TL); C, ventral view of adult male holotype (NMMB-P36013, 429 mm TL); D, ventral view of adult male paratype (ASIZP0081116, 405 mm TL).
</paragraph>
</caption>
<caption id="DF4BCCDAA23CFFB7FF2C4647FC61F95C" ID-DOI="http://doi.org/10.5281/zenodo.7895054" ID-Zenodo-Dep="7895054" httpUri="https://zenodo.org/record/7895054/files/figure.png" pageId="7" pageNumber="110" startId="7.[152,255,1614,1638]" targetBox="[280,1308,1205,1591]" targetPageId="7" targetType="figure">
<paragraph id="8B8B9C52A23CFFB7FF2C4647FC61F95C" blockId="7.[151,1436,1614,1711]" pageId="7" pageNumber="110">
<emphasis id="B9404040A23CFFB7FF2C4647FEDCF994" bold="true" box="[152,282,1614,1638]" pageId="7" pageNumber="110">FIGURE 4.</emphasis>
Details of thorns on midtail (A, B) and the dorsal-caudal area (C, D) of
<taxonomicName id="4C34E7D1A23CFFB7FBB94647FB6FF994" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[1037,1193,1614,1638]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="110" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A23CFFB7FBB94647FB6FF994" box="[1037,1193,1614,1638]" italics="true" pageId="7" pageNumber="110">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A23CFFB7FB044647FAC4F994" bold="true" box="[1200,1282,1614,1638]" pageId="7" pageNumber="110">
<taxonomicNameLabel id="A273FD3BA23CFFB7FB044647FAC4F994" box="[1200,1282,1614,1638]" pageId="7" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, adult male holotype (NMMB-P36013, 429 mm TL); B, adult male paratype (ASIZP0081116, 405 mm TL); C, adult male holotype (NMMBP36013, 429 mm TL); D, adult male paratype (ASIZP0081116, 405 mm TL).
</paragraph>
</caption>
<paragraph id="8B8B9C52A23CFFB7FF734709FC54F83C" blockId="7.[151,1437,1792,2035]" pageId="7" pageNumber="110">Tail long and thin, moderately depressed; relatively narrow at base, slightly tapering gradually to tail tip, not expanded at midlength; width at first dorsal-fin 65.7% (54.7%) width at midlength of tail, 47.6% (32.6%) width at axil of pelvic fin; length from postcloaca 116118% distance from snout tip to precloaca; anterior cross-section suboval, flat on dorsal and ventral surface of the whole tail; height 81.3% (58.7%) width at axil of pelvic fin, 79.8% (84.4%) width at midlength, 98.0% (93.3%) width at first dorsal-fin origin; lateral tail fold poorly developed, its origin slightly anterior to first dorsal-fin origin, extending to tail tip.</paragraph>
<paragraph id="8B8B9C52A23CFFB8FF7347D1FAC8FE97" blockId="7.[151,1437,1792,2035]" lastBlockId="8.[151,1437,151,357]" lastPageId="8" lastPageNumber="111" pageId="7" pageNumber="110">
Dorsal fins small, subequal in size and shape (
<figureCitation id="130F80D7A23CFFB7FD7D47D1FC84F800" box="[713,834,2008,2034]" captionStart="FIGURE 4" captionStartId="7.[152,255,1614,1638]" captionTargetBox="[280,1308,1205,1591]" captionTargetId="figure-89@7.[280,1308,1205,1591]" captionTargetPageId="7" captionText="FIGURE 4. Details of thorns on midtail (A, B) and the dorsal-caudal area (C, D) of Okamejei picta sp. nov.: A, adult male holotype (NMMB-P36013, 429 mm TL); B, adult male paratype (ASIZP0081116, 405 mm TL); C, adult male holotype (NMMB- P36013, 429 mm TL); D, adult male paratype (ASIZP0081116, 405 mm TL)." figureDoi="http://doi.org/10.5281/zenodo.7895054" httpUri="https://zenodo.org/record/7895054/files/figure.png" pageId="7" pageNumber="110">Fig. 4C, D</figureCitation>
); first dorsal-fin height 51.6% (52.7%) in base length; second dorsal-fin height 64.9% (67.2%) in base length; fins subtriangular, anterior margins oblique, posterior margins rounded and slightly convex; interdorsal space moderately wide, 97.9% (83.5%) in first dorsal-fin base length; distance from first dorsal-fin origin to tail tip 38.9% (38.0%) tail length; caudal-fin length 27.4% (25.4%) distance from first dorsal-fin origin to tail tip; first dorsal-fin base length 76.3% (98.1%) caudal-fin length. Caudal-fin epichordal lobe well developed, base long and low; its height greater posteriorly; tip pointed, its posterodorsal margin somewhat convex; well separated from second dorsal fin; hypochordal lobe poorly developed.
</paragraph>
<caption id="DF4BCCDAA233FFB8FF2C43F8FBA5FBDF" ID-DOI="http://doi.org/10.5281/zenodo.7895056" ID-Zenodo-Dep="7895056" httpUri="https://zenodo.org/record/7895056/files/figure.png" pageId="8" pageNumber="111" startId="8.[152,255,1009,1033]" targetBox="[640,947,381,973]" targetPageId="8" targetType="figure">
<paragraph id="8B8B9C52A233FFB8FF2C43F8FBA5FBDF" blockId="8.[151,1436,1009,1069]" pageId="8" pageNumber="111">
<emphasis id="B9404040A233FFB8FF2C43F8FEDFFBFB" bold="true" box="[152,281,1009,1033]" pageId="8" pageNumber="111">FIGURE 5.</emphasis>
<taxonomicName id="4C34E7D1A233FFB8FEAB43F8FE7CFBFB" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[287,442,1009,1033]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="111" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A233FFB8FEAB43F8FE7CFBFB" box="[287,442,1009,1033]" italics="true" pageId="8" pageNumber="111">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A233FFB8FE7443F8FDD7FBFB" bold="true" box="[448,529,1009,1033]" pageId="8" pageNumber="111">
<taxonomicNameLabel id="A273FD3BA233FFB8FE7443F8FDD7FBFB" box="[448,529,1009,1033]" pageId="8" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male holotype, NMMB-P36013, 429 mm TL, left clasper spread opened. bs, boss; cf, cleft; sk, spike; tb, terminal bridge; ps, pseudorhipidion; rh, rhipidion; sh, shield; st, sentinel.
</paragraph>
</caption>
<caption id="DF4BCCDAA233FFB8FF2C470CFB3DF894" ID-DOI="http://doi.org/10.5281/zenodo.7895058" ID-Zenodo-Dep="7895058" httpUri="https://zenodo.org/record/7895058/files/figure.png" pageId="8" pageNumber="111" startId="8.[152,255,1797,1821]" targetBox="[290,1297,1093,1773]" targetPageId="8" targetType="figure">
<paragraph id="8B8B9C52A233FFB8FF2C470CFB3DF894" blockId="8.[151,1436,1797,1894]" pageId="8" pageNumber="111">
<emphasis id="B9404040A233FFB8FF2C470CFED3F8EF" bold="true" box="[152,277,1797,1821]" pageId="8" pageNumber="111">FIGURE 6.</emphasis>
<taxonomicName id="4C34E7D1A233FFB8FEAD470FFE77F8EF" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[281,433,1798,1822]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="111" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A233FFB8FEAD470FFE77F8EF" box="[281,433,1798,1822]" italics="true" pageId="8" pageNumber="111">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A233FFB8FE00470FFDC4F8EC" bold="true" box="[436,514,1798,1822]" pageId="8" pageNumber="111">
<taxonomicNameLabel id="A273FD3BA233FFB8FE00470FFDC4F8EC" box="[436,514,1798,1822]" pageId="8" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male holotype, NMMB-P36013, 429 mm TL, X-radiograph of pelvic girdle and clasper basal cartilages (ventral view). ax, axial cartilage; β, beta cartilage; b12, intermediate segments 1 and 2; bpv, basipterygium; ip, iliac process; lp, lateral process; lpp, lateral prepelvic process of obturator foramina; pvp, pelvic propterygium.
</paragraph>
</caption>
<paragraph id="8B8B9C52A233FFB9FF7347B1FA8CFD93" blockId="8.[151,1436,1976,2038]" lastBlockId="9.[151,1437,151,825]" lastPageId="9" lastPageNumber="112" pageId="8" pageNumber="111">
Orbital thorns small, smaller than alar and malar thorns, 2 (4) on preorbit, 5 (5) on orbit, directed posteriorly; one rudimentary nuchal thorn present; no thorns on scapular regions. Malar thorn patch with 20 (20) enlarged, strongly curved thorns in ca. 3 rows; main patch small, curved slightly, located at disc margin near eye and spiracle; thorns merging with bunches of smaller thornlets laterally, thorns near disc edge smaller; thornlets at anterior disc larger, decreasing in size posteriorly. Alar thorn patch with 34 (33) thorns, size subequal to tail thorns; embedded and directed posteromedially, in about 23 rows; patch narrow, length slightly longer than malar patch. Tail thorns variously developed, in three rows (
<figureCitation id="130F80D7A232FFB9FD99412EFD6EFEB3" box="[557,680,295,321]" captionStart="FIGURE 4" captionStartId="7.[152,255,1614,1638]" captionTargetBox="[280,1308,1205,1591]" captionTargetId="figure-89@7.[280,1308,1205,1591]" captionTargetPageId="7" captionText="FIGURE 4. Details of thorns on midtail (A, B) and the dorsal-caudal area (C, D) of Okamejei picta sp. nov.: A, adult male holotype (NMMB-P36013, 429 mm TL); B, adult male paratype (ASIZP0081116, 405 mm TL); C, adult male holotype (NMMB- P36013, 429 mm TL); D, adult male paratype (ASIZP0081116, 405 mm TL)." figureDoi="http://doi.org/10.5281/zenodo.7895054" httpUri="https://zenodo.org/record/7895054/files/figure.png" pageId="9" pageNumber="112">Fig. 4A, B</figureCitation>
); row not staggered; 26 (21) predorsal thorns, weakly curved, with very small bases, size subequal to orbital thorns; interdorsal thorns small, 3 (1); median row poorly developed; thorns in median row weak, 11 (5) thorns, emerging near pelvic-fin rear tip, extending along length of predorsal tail in a single row; dorsolateral rows of similar size; with 15 (16) similar thorns on each side, almost continuous, extending from near pelvic-fin rear tip to first dorsal-fin origin (to first dorsal-fin base end in the
<typeStatus id="548F22F0A232FFB9FB6041BEFAFFFE23" box="[1236,1337,439,465]" pageId="9" pageNumber="112" type="paratype">paratype</typeStatus>
), located above mid-lateral tail, closer to thorns of median row. Denticles poorly developed, disc and tail mostly naked; confined to a moderately developed patch on snout tip dorsally; long, widely spaced, thorn-like denticles along anterior margin of disc from snout tip to an area anterior to alar thorn patch; confined to a small patch at snout tip ventrally; tail and posterior disc entirely naked; denticles short, prickle-like, sparsely distributed in patches.
</paragraph>
<paragraph id="8B8B9C52A232FFB9FF734262FD70FCCB" blockId="9.[151,1437,151,825]" pageId="9" pageNumber="112">
Clasper well developed, stout, depressed, length 23.5% (26.3%) TL; tip pointed. Clasper external features included boss (bs), cleft (cf), spike (sk), terminal bridge (tb), pseudorhipidion (ps), rhipidion (rh), shield (sh) and sentinel (st) (
<figureCitation id="130F80D7A232FFB9FE9B42BAFEB0FD3F" box="[303,374,690,717]" captionStart="FIGURE 5" captionStartId="8.[152,255,1009,1033]" captionTargetBox="[640,947,381,973]" captionTargetId="figure-125@8.[640,947,381,985]" captionTargetPageId="8" captionText="FIGURE 5. Okamejei picta sp. nov., adult male holotype, NMMB-P36013, 429 mm TL, left clasper spread opened. bs, boss; cf, cleft; sk, spike; tb, terminal bridge; ps, pseudorhipidion; rh, rhipidion; sh, shield; st, sentinel." figureDoi="http://doi.org/10.5281/zenodo.7895056" httpUri="https://zenodo.org/record/7895056/files/figure.png" pageId="9" pageNumber="112">Fig. 5</figureCitation>
); bs rather expanded, situated between lower part of cf and sk; cf narrow, partitioned by tb; ps rod-like, situated between upper cleft and rh; rh bilobed, its distal tip barely extend to proximal end of cf; st moderately developed, positioned among rh, tb, bs and sk; sk sharp, rather straight, not hook-like; sh long, with sharp outer edge; component funnel not evident.
</paragraph>
<caption id="DF4BCCDAA232FFB9FF2C4702FE78F87D" ID-DOI="http://doi.org/10.5281/zenodo.7895060" ID-Zenodo-Dep="7895060" httpUri="https://zenodo.org/record/7895060/files/figure.png" pageId="9" pageNumber="112" startId="9.[152,255,1803,1827]" targetBox="[347,1241,849,1777]" targetPageId="9" targetType="figure">
<paragraph id="8B8B9C52A232FFB9FF2C4702FE78F87D" blockId="9.[151,1436,1803,1935]" pageId="9" pageNumber="112">
<emphasis id="B9404040A232FFB9FF2C4702FEDFF8D1" bold="true" box="[152,281,1803,1827]" pageId="9" pageNumber="112">FIGURE 7.</emphasis>
Terminal skeleton of left clasper of (A)
<taxonomicName id="4C34E7D1A232FFB9FD0D4702FC93F8D1" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[697,853,1803,1827]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="112" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A232FFB9FD0D4702FC93F8D1" box="[697,853,1803,1827]" italics="true" pageId="9" pageNumber="112">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A232FFB9FCE84702FC6BF8D1" bold="true" box="[860,941,1803,1827]" pageId="9" pageNumber="112">
<taxonomicNameLabel id="A273FD3BA232FFB9FCE84702FC6BF8D1" box="[860,941,1803,1827]" pageId="9" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male holotype, NMMB-P36013, 429 mm TL; and (B)
<taxonomicName id="4C34E7D1A232FFB9FEA84726FDE2F8B5" authority=", EBFS" authorityName="EBFS" box="[284,548,1839,1863]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="112" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A232FFB9FEA84726FE1FF8B5" box="[284,473,1839,1863]" italics="true" pageId="9" pageNumber="112">Okamejei hollandi</emphasis>
, EBFS
</taxonomicName>
adult male, uncatalogued specimen, 432 mm TL. ax, axial cartilage; at12, accessory terminal cartilage 1 and 2; dm, dorsal marginal cartilage; dt13, dorsal terminal cartilage 13; vm, ventral marginal cartilage; vt, ventral terminal cartilage.
</paragraph>
</caption>
<caption id="DF4BCCDAA231FFBAFF2C4419FC26FBBF" ID-DOI="http://doi.org/10.5281/zenodo.7895062" ID-Zenodo-Dep="7895062" httpUri="https://zenodo.org/record/7895062/files/figure.png" pageId="10" pageNumber="113" startId="10.[152,255,1040,1064]" targetBox="[303,1284,181,1017]" targetPageId="10" targetType="figure">
<paragraph id="8B8B9C52A231FFBAFF2C4419FC26FBBF" blockId="10.[151,1436,1040,1101]" pageId="10" pageNumber="113">
<emphasis id="B9404040A231FFBAFF2C4419FEDFFBDA" bold="true" box="[152,281,1040,1064]" pageId="10" pageNumber="113">FIGURE 8.</emphasis>
Ventral terminal cartilage from left clasper of
<taxonomicName id="4C34E7D1A231FFBAFD424418FC57FBDA" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[758,913,1041,1065]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="113" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A231FFBAFD424418FC57FBDA" box="[758,913,1041,1065]" italics="true" pageId="10" pageNumber="113">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A231FFBAFC234418FC2FFBDB" bold="true" box="[919,1001,1041,1065]" pageId="10" pageNumber="113">
<taxonomicNameLabel id="A273FD3BA231FFBAFC234418FC2FFBDB" box="[919,1001,1041,1065]" pageId="10" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male paratype, ASIZP0081116, 405 mm TL. (A) Dorsal view; (B) ventral view; (C) proximal tip viewed from left side.
</paragraph>
</caption>
<caption id="DF4BCCDAA231FFBAFF2C4746FD05F821" ID-DOI="http://doi.org/10.5281/zenodo.7895064" ID-Zenodo-Dep="7895064" httpUri="https://zenodo.org/record/7895064/files/figure.png" pageId="10" pageNumber="113" startId="10.[152,255,1871,1895]" targetBox="[312,1275,1125,1848]" targetPageId="10" targetType="figure">
<paragraph id="8B8B9C52A231FFBAFF2C4746FD05F821" blockId="10.[151,1437,1871,2003]" pageId="10" pageNumber="113">
<emphasis id="B9404040A231FFBAFF2C4746FEDEF895" bold="true" box="[152,280,1871,1895]" pageId="10" pageNumber="113">FIGURE 9.</emphasis>
<taxonomicName id="4C34E7D1A231FFBAFEAA4746FE71F895" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[286,439,1871,1895]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="10" pageNumber="113" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A231FFBAFEAA4746FE71F895" box="[286,439,1871,1895]" italics="true" pageId="10" pageNumber="113">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A231FFBAFE094746FDCBF895" bold="true" box="[445,525,1871,1895]" pageId="10" pageNumber="113">
<taxonomicNameLabel id="A273FD3BA231FFBAFE094746FDCBF895" box="[445,525,1871,1895]" pageId="10" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male holotype, NMMB-P36013, 429 mm TL, X-radiograph of neurocranium, snout and pectoral area. ac, antorbital cartilage; af, anterior fontanelle; cr, neurocranium; ms, mesopterygium; mt, metapterygium; nc, nasal capsule; pf, posterior fontanelle; poc, preorbital canal foramen, postp, postorbital process; pr, propterygium; prep, preorbital process; rb, rostral base; rc, rostral cartilage.
</paragraph>
</caption>
<caption id="DF4BCCDAA230FFBBFF2C45E7FE4FF9D8" ID-DOI="http://doi.org/10.5281/zenodo.7895066" ID-Zenodo-Dep="7895066" httpUri="https://zenodo.org/record/7895066/files/figure.png" pageId="11" pageNumber="114" startId="11.[152,255,1518,1542]" targetBox="[158,1368,214,1490]" targetPageId="11" targetType="figure">
<paragraph id="8B8B9C52A230FFBBFF2C45E7FE4FF9D8" blockId="11.[151,1436,1518,1578]" pageId="11" pageNumber="114">
<emphasis id="B9404040A230FFBBFF2C45E7FEE2F9F4" bold="true" box="[152,292,1518,1542]" pageId="11" pageNumber="114">FIGURE 10.</emphasis>
Fresh coloration of
<taxonomicName id="4C34E7D1A230FFBBFE5F45E7FD2BF9F4" authority=", EBFS" authorityName="EBFS" box="[491,749,1518,1542]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="11" pageNumber="114" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A230FFBBFE5F45E7FD63F9F4" box="[491,677,1518,1542]" italics="true" pageId="11" pageNumber="114">Okamejei hollandi</emphasis>
, EBFS
</taxonomicName>
adult male, uncatalogued specimen, 432 mm TL: top, dorsal surface; bottom, ventral surface.
</paragraph>
</caption>
<paragraph id="8B8B9C52A230FFBCFF734666FD56FE97" blockId="11.[151,1437,1647,2034]" lastBlockId="12.[151,1437,151,897]" lastPageId="12" lastPageNumber="115" pageId="11" pageNumber="114">
Clasper skeleton highly calcified, consists of basal, axial and terminal cartilages. Basal cartilages contain beta cartilage (β), basipterygium (bpv), and intermediate segments 1 (b1) and 2 (b2) (
<figureCitation id="130F80D7A230FFBBFB9D469AFBA9F95F" box="[1065,1135,1683,1709]" captionStart="FIGURE 6" captionStartId="8.[152,255,1797,1821]" captionTargetBox="[290,1297,1093,1773]" captionTargetId="figure-167@8.[290,1297,1093,1773]" captionTargetPageId="8" captionText="FIGURE 6. Okamejei pictasp.nov., adult male holotype,NMMB-P36013,429 mmTL,X-radiograph of pelvic girdleandclasper basal cartilages (ventral view). ax, axial cartilage; β, beta cartilage; b12, intermediate segments 1 and 2; bpv, basipterygium; ip, iliac process; lp, lateral process; lpp, lateral prepelvic process of obturator foramina; pvp, pelvic propterygium." figureDoi="http://doi.org/10.5281/zenodo.7895058" httpUri="https://zenodo.org/record/7895058/files/figure.png" pageId="11" pageNumber="114">Fig. 6</figureCitation>
); β cartilage long, situated dorsally of posterior half of b1 and the whole b2 cartilages; bpv slightly arched outwards, linked with pelvic girdle anteriorly and with b1 posteriorly; b1 and b2 short, length about half of bpv, both rod-like. Pelvic girdle with large, slightly arched puboischiadic bar; iliac region enlarged, with two obturator foramina (of) on each side; iliac process (ip) slightly curved inwards; lateral prepelvic process (lpp) long and narrow; lateral process (lp) very short, with rounded tips. Terminal cartilages comprise nine components: axial cartilage (ax), three dorsal terminal cartilages (dt13), two accessory terminal cartilages (at1, at2), ventral marginal cartilage (vm), dorsal marginal cartilage (dm), and ventral terminal cartilage (vt) (
<figureCitation id="130F80D7A230FFBBFDC54786FD0BF85B" box="[625,717,1935,1961]" captionStart="FIGURE 7" captionStartId="9.[152,255,1803,1827]" captionTargetBox="[347,1241,849,1777]" captionTargetId="figure-348@9.[344,1244,849,1780]" captionTargetPageId="9" captionText="FIGURE 7. Terminal skeleton of left clasper of (A) Okamejei picta sp. nov., adult male holotype, NMMB-P36013, 429 mm TL; and (B) Okamejei hollandi, EBFS adult male, uncatalogued specimen, 432 mm TL. ax, axial cartilage; at12, accessory terminal cartilage 1 and 2; dm, dorsal marginal cartilage; dt13, dorsal terminal cartilage 13; vm, ventral marginal cartilage; vt, ventral terminal cartilage." figureDoi="http://doi.org/10.5281/zenodo.7895060" httpUri="https://zenodo.org/record/7895060/files/figure.png" pageId="11" pageNumber="114">Fig. 7A</figureCitation>
); ax longest, slender, with a rounded tip; at1 Y-shaped, deeply forked, distal tip paddle-shaped, attaching proximally to vm and forming st externally; at2 slender, hook-like, with a flattened tip, alongside at1, joining the tip of at1 distally, and forming sk externally; dm long, slightly concave, with distal end very weakly bifurcated, both lateral and medial tips broadly rounded; dt1 moderate, shield-like, slightly concave proximally, covering ax, with a blunt tip; dt2 moderately short, attached to dm proximally, and dt3 distally; dt3 very slender, with pointed tips, proximally attaching dt2, distally attaching vt; vm long and forked, wide posteriorly, attached to ax and dm, distal tip forming ps externally; vt positioned at outer aspect of clasper glans, J-shaped, bifurcated proximally, broadly widened distally, tip spoon-shaped, forming sh and dk externally (
<figureCitation id="130F80D7A237FFBCFAFD412EFA4AFEB3" box="[1353,1420,295,321]" captionStart="FIGURE 8" captionStartId="10.[152,255,1040,1064]" captionTargetBox="[303,1284,181,1017]" captionTargetId="figure-31@10.[303,1284,181,1017]" captionTargetPageId="10" captionText="FIGURE 8. Ventral terminal cartilage from left clasper of Okamejei picta sp. nov., adult male paratype, ASIZP0081116, 405 mm TL. (A) Dorsal view; (B) ventral view; (C) proximal tip viewed from left side." figureDoi="http://doi.org/10.5281/zenodo.7895062" httpUri="https://zenodo.org/record/7895062/files/figure.png" pageId="12" pageNumber="115">Fig. 8</figureCitation>
); not showing externally as component funnel.
</paragraph>
<paragraph id="8B8B9C52A237FFBCFF734166FD63FDCF" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="115">
Neurocranium (
<figureCitation id="130F80D7A237FFBCFEC84166FE05FE7B" box="[380,451,367,393]" captionStart="FIGURE 9" captionStartId="10.[152,255,1871,1895]" captionTargetBox="[312,1275,1125,1848]" captionTargetId="figure-72@10.[312,1275,1125,1848]" captionTargetPageId="10" captionText="FIGURE 9. Okamejei picta sp. nov., adult male holotype, NMMB-P36013, 429 mm TL, X-radiograph of neurocranium, snout and pectoral area. ac, antorbital cartilage; af, anterior fontanelle; cr, neurocranium; ms, mesopterygium; mt, metapterygium; nc, nasal capsule; pf, posterior fontanelle; poc, preorbital canal foramen, postp, postorbital process; pr, propterygium; prep, preorbital process; rb, rostral base; rc, rostral cartilage." figureDoi="http://doi.org/10.5281/zenodo.7895064" httpUri="https://zenodo.org/record/7895064/files/figure.png" pageId="12" pageNumber="115">Fig. 9</figureCitation>
) with a bullet-shaped anterior fontanelle (af); posterior fontanelle (pf) bottle-shaped, shorter than af; rostral cartilage (rc) long, length 59.0% (43.9%, note that the rostral cartilage is slightly broken in the
<typeStatus id="548F22F0A237FFBCFF7441BEFEE4FE23" box="[192,290,439,465]" pageId="12" pageNumber="115" type="paratype">paratype</typeStatus>
) of dorsal head length; rostral base (rb) moderately narrow, its width about one quarter of rc; antorbital cartilages (ac) thin, arched, attaching nasal capsule (nc) and the second segments of propterygium (pr); nc ovalshaped, preorbital canal foramen (poc) situated proximally to the leading edge of nc; preorbital process (prep) and postorbital process (postp) scarcely developed.
</paragraph>
<paragraph id="8B8B9C52A237FFBCFF73424EFEA9FD03" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="115">
Distribution of ventral sensory pores overall W-shaped, denser on head, but sparsely distributed on other parts; pores in 12 rows at rostral cartilage, nasal curtain, area between gill silts, and pectoral fins before their maximum width; sensory pores present on anterior to middle edge of abdomen, absent from pelvic fins; pores generally small (slightly larger in
<typeStatus id="548F22F0A237FFBCFED242BAFE00FD3F" box="[358,454,691,717]" pageId="12" pageNumber="115" type="paratype">paratype</typeStatus>
than
<typeStatus id="548F22F0A237FFBCFDBD42BAFDA9FD3F" box="[521,623,691,717]" pageId="12" pageNumber="115" type="holotype">holotype</typeStatus>
), those on area between gill gilts and on abdomen edge slightly larger, others rather small.
</paragraph>
<paragraph id="8B8B9C52A237FFBCFF7342F2FB11FC73" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="115">
Meristics:Also provided in
<tableCitation id="C6B6A9E9A237FFBCFE4342F2FD8DFCE7" box="[503,587,763,789]" captionStart="TABLE 2" captionStartId="12.[152,245,944,968]" captionTargetPageId="12" captionText="TABLE 2. Comparisons of selected characters of O. picta sp. nov. and the similar species. Data of the holotypes of O. hollandi and O. mengae were obtained from Ishihara (1986) and Jeong et al. (2007), respectively." httpUri="http://table.plazi.org/id/DF4BCCDAA237FFBCFF2C43B9FB6FFC1E" pageId="12" pageNumber="115" tableUuid="DF4BCCDAA237FFBCFF2C43B9FB6FFC1E">Table 2</tableCitation>
. Tooth rows in upper jaw 40 (41); lower jaw 40 (40). Pectoral-fin propterygial radials 29 (28); mesopterygial radials 10 (8); metapterygial radials 37 (37); total basal radials 76 (73). Pelvic-fin radials 1 (1) + 19 (20). Trunk centra 31 (27); predorsal caudal centra 44 (43); predorsal centra 75 (70); centra between origins of dorsal fins 16 (16); diplospondylous centra 95 (96); total centra ca. 126 (123).
</paragraph>
<caption id="DF4BCCDAA237FFBCFF2C43B9FB6FFC1E" ID-Table-UUID="DF4BCCDAA237FFBCFF2C43B9FB6FFC1E" httpUri="http://table.plazi.org/id/DF4BCCDAA237FFBCFF2C43B9FB6FFC1E" pageId="12" pageNumber="115" startId="12.[152,245,944,968]" targetBox="[163,1421,1020,1689]" targetIsTable="true" targetPageId="12" targetType="table">
<paragraph id="8B8B9C52A237FFBCFF2C43B9FB6FFC1E" blockId="12.[151,1436,944,1005]" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFF2C43B9FEC9FC3A" bold="true" box="[152,271,944,968]" pageId="12" pageNumber="115">TABLE 2.</emphasis>
Comparisons of selected characters of
<taxonomicName id="4C34E7D1A237FFBCFD0C43B8FCC8FC3A" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[696,782,945,968]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A237FFBCFD0C43B8FCC8FC3A" box="[696,782,945,968]" italics="true" pageId="12" pageNumber="115">O. picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A237FFBCFCA243B8FCAAFC3B" bold="true" box="[790,876,945,969]" pageId="12" pageNumber="115">
<taxonomicNameLabel id="A273FD3BA237FFBCFCA243B8FCAAFC3B" box="[790,876,945,969]" pageId="12" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and the similar species. Data of the holotypes of
<taxonomicName id="4C34E7D1A237FFBCFA3643B8FF36FC1E" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A237FFBCFA3643B8FF36FC1E" italics="true" pageId="12" pageNumber="115">O. hollandi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34E7D1A237FFBCFE9043DCFE51FC1E" box="[292,407,981,1004]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A237FFBCFE9043DCFE51FC1E" box="[292,407,981,1004]" italics="true" pageId="12" pageNumber="115">O. mengae</emphasis>
</taxonomicName>
were obtained from Ishihara (1986) and
<bibRefCitation id="EFA5E1A3A237FFBCFCF943DCFBDEFC1F" author="Jeong, C. - H. &amp; Nakabo, T. &amp; Wu, H. - L." box="[845,1048,980,1005]" pageId="12" pageNumber="115" pagination="57 - 65" refId="ref7491" refString="Jeong, C. - H., Nakabo, T. &amp; Wu, H. - L. (2007) A new species of skate (Chondrichthyes: Rajidae), Okamejei mengae from the South China Sea. Korean Journal of Ichthyology, 19 (1), 57 - 65." type="journal article" year="2007">
Jeong
<emphasis id="B9404040A237FFBCFC2643DCFC0EFC1E" box="[914,968,980,1004]" italics="true" pageId="12" pageNumber="115">et al.</emphasis>
(2007)
</bibRefCitation>
, respectively.
</paragraph>
</caption>
<paragraph id="8B8B9C52A237FFBCFF1743F4FAA8F96B" pageId="12" pageNumber="115">
<table id="F9346EF2A237004FFF1743F5FA4BF96B" box="[163,1421,1020,1689]" gridcols="4" gridrows="17" pageId="12" pageNumber="115">
<tr id="35049E10A237004FFF1743F5FA4BFBE7" box="[163,1421,1020,1045]" gridrow="0" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF1743F5FD56FBE7" box="[163,656,1020,1045]" gridcol="0" gridrow="0" pageId="12" pageNumber="115">Characters</th>
<th id="76D5F76CA237004FFD0F43F5FC0AFBE7" box="[699,972,1020,1045]" gridcol="1" gridrow="0" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFD4C43F4FC96FBE6" box="[760,848,1021,1045]" italics="true" pageId="12" pageNumber="115">
<taxonomicName id="4C34E7D1A237FFBCFD4C43F4FC8CFBE7" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[760,842,1021,1045]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A237FFBCFD4C43F4FC8CFBE7" bold="true" box="[760,842,1021,1045]" italics="true" pageId="12" pageNumber="115">O. picta</emphasis>
</taxonomicName>
.
</emphasis>
(n=2)
</th>
<th id="76D5F76CA237004FFC4F43F5FB7CFBE7" box="[1019,1210,1020,1045]" gridcol="2" gridrow="0" pageId="12" pageNumber="115">
<taxonomicName id="4C34E7D1A237FFBCFC4B43F4FBB3FBE6" box="[1023,1141,1020,1045]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A237FFBCFC4B43F4FBB3FBE6" bold="true" box="[1023,1141,1020,1045]" italics="true" pageId="12" pageNumber="115">O. hollandi</emphasis>
</taxonomicName>
(n=9)
</th>
<th id="76D5F76CA237004FFB6A43F5FA4BFBE7" box="[1246,1421,1020,1045]" gridcol="3" gridrow="0" pageId="12" pageNumber="115">
<taxonomicName id="4C34E7D1A237FFBCFB6A43F4FA89FBE6" box="[1246,1359,1021,1045]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="12" pageNumber="115" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A237FFBCFB6A43F4FA89FBE6" bold="true" box="[1246,1359,1021,1045]" italics="true" pageId="12" pageNumber="115">O. mengae</emphasis>
</taxonomicName>
(n=1)
</th>
</tr>
<tr id="35049E10A237004FFF17442DFA4BFBCE" box="[163,1421,1060,1084]" gridrow="1" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF17442DFA4BFBCE" box="[163,1421,1060,1084]" colspan="4" colspanRight="3" gridcol="0" gridrow="1" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFF17442DFE8AFBCE" bold="true" box="[163,332,1060,1084]" pageId="12" pageNumber="115">Morphometrics</emphasis>
</th>
</tr>
<tr id="35049E10A237004FFF174444FA4BFB97" box="[163,1421,1101,1125]" gridrow="2" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174444FD56FB97" box="[163,656,1101,1125]" gridcol="0" gridrow="2" pageId="12" pageNumber="115">Disc length (%TL)</th>
<td id="76D5F76CA237004FFD0F4444FC0AFB97" box="[699,972,1101,1125]" gridcol="1" gridrow="2" pageId="12" pageNumber="115">45.047.7</td>
<td id="76D5F76CA237004FFC4F4444FB7CFB97" box="[1019,1210,1101,1125]" gridcol="2" gridrow="2" pageId="12" pageNumber="115">51.655.8</td>
<td id="76D5F76CA237004FFB6A4444FA4BFB97" box="[1246,1421,1101,1125]" gridcol="3" gridrow="2" pageId="12" pageNumber="115">55.7</td>
</tr>
<tr id="35049E10A237004FFF17447CFA4BFB7C" box="[163,1421,1141,1166]" gridrow="3" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF17447CFD56FB7C" box="[163,656,1141,1166]" gridcol="0" gridrow="3" pageId="12" pageNumber="115">Distance between cloaca to caudal-fin tip (%TL)</th>
<td id="76D5F76CA237004FFD0F447CFC0AFB7C" box="[699,972,1141,1166]" gridcol="1" gridrow="3" pageId="12" pageNumber="115">53.655.1</td>
<td id="76D5F76CA237004FFC4F447CFB7CFB7C" box="[1019,1210,1141,1166]" gridcol="2" gridrow="3" pageId="12" pageNumber="115">47.853.4</td>
<td id="76D5F76CA237004FFB6A447CFA4BFB7C" box="[1246,1421,1141,1166]" gridcol="3" gridrow="3" pageId="12" pageNumber="115">48.5</td>
</tr>
<tr id="35049E10A237004FFF174494FA4BFB47" box="[163,1421,1181,1205]" gridrow="4" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174494FA4BFB47" box="[163,1421,1181,1205]" colspan="4" colspanRight="3" gridcol="0" gridrow="4" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFF174494FF36FB47" bold="true" box="[163,240,1181,1205]" pageId="12" pageNumber="115">Counts</emphasis>
</th>
</tr>
<tr id="35049E10A237004FFF1744CFFA4BFB2C" box="[163,1421,1222,1246]" gridrow="5" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF1744CFFD56FB2C" box="[163,656,1222,1246]" gridcol="0" gridrow="5" pageId="12" pageNumber="115">Trunk centra</th>
<td id="76D5F76CA237004FFD0F44CFFC0AFB2C" box="[699,972,1222,1246]" gridcol="1" gridrow="5" pageId="12" pageNumber="115">31</td>
<td id="76D5F76CA237004FFC4F44CFFB7CFB2C" box="[1019,1210,1222,1246]" gridcol="2" gridrow="5" pageId="12" pageNumber="115">2830</td>
<td id="76D5F76CA237004FFB6A44CFFA4BFB2C" box="[1246,1421,1222,1246]" gridcol="3" gridrow="5" pageId="12" pageNumber="115">23</td>
</tr>
<tr id="35049E10A237004FFF1744E7FA4BFAF4" box="[163,1421,1262,1286]" gridrow="6" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF1744E7FD56FAF4" box="[163,656,1262,1286]" gridcol="0" gridrow="6" pageId="12" pageNumber="115">Total basal radials</th>
<td id="76D5F76CA237004FFD0F44E7FC0AFAF4" box="[699,972,1262,1286]" gridcol="1" gridrow="6" pageId="12" pageNumber="115">7376</td>
<td id="76D5F76CA237004FFC4F44E7FB7CFAF4" box="[1019,1210,1262,1286]" gridcol="2" gridrow="6" pageId="12" pageNumber="115">8286</td>
<td id="76D5F76CA237004FFB6A44E7FA4BFAF4" box="[1246,1421,1262,1286]" gridcol="3" gridrow="6" pageId="12" pageNumber="115">96</td>
</tr>
<tr id="35049E10A237004FFF17451FFA4BFADC" box="[163,1421,1302,1326]" gridrow="7" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF17451FFA4BFADC" box="[163,1421,1302,1326]" colspan="4" colspanRight="3" gridcol="0" gridrow="7" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFF17451FFE5DFADC" bold="true" box="[163,411,1302,1326]" pageId="12" pageNumber="115">Clasper characteristics</emphasis>
</th>
</tr>
<tr id="35049E10A237004FFF174536FA4BFAA5" box="[163,1421,1343,1367]" gridrow="8" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174536FD56FAA5" box="[163,656,1343,1367]" gridcol="0" gridrow="8" pageId="12" pageNumber="115">Shape of ventral terminal cartilage</th>
<td id="76D5F76CA237004FFD0F4536FC0AFAA5" box="[699,972,1343,1367]" gridcol="1" gridrow="8" pageId="12" pageNumber="115">J-shaped</td>
<td id="76D5F76CA237004FFC4F4536FB7CFAA5" box="[1019,1210,1343,1367]" gridcol="2" gridrow="8" pageId="12" pageNumber="115">Slightly curved</td>
<td id="76D5F76CA237004FFB6A4536FA4BFAA5" box="[1246,1421,1343,1367]" gridcol="3" gridrow="8" pageId="12" pageNumber="115">Unknown</td>
</tr>
<tr id="35049E10A237004FFF17456EFA4BFA8D" box="[163,1421,1383,1407]" gridrow="9" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF17456EFD56FA8D" box="[163,656,1383,1407]" gridcol="0" gridrow="9" pageId="12" pageNumber="115">Proximal end of ventral terminal cartilage</th>
<td id="76D5F76CA237004FFD0F456EFC0AFA8D" box="[699,972,1383,1407]" gridcol="1" gridrow="9" pageId="12" pageNumber="115">Bifurcated</td>
<td id="76D5F76CA237004FFC4F456EFB7CFA8D" box="[1019,1210,1383,1407]" gridcol="2" gridrow="9" pageId="12" pageNumber="115">Blunt</td>
<td id="76D5F76CA237004FFB6A456EFA4BFA8D" box="[1246,1421,1383,1407]" gridcol="3" gridrow="9" pageId="12" pageNumber="115">Unknown</td>
</tr>
<tr id="35049E10A237004FFF174586FA4BFA55" box="[163,1421,1423,1447]" gridrow="10" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174586FD56FA55" box="[163,656,1423,1447]" gridcol="0" gridrow="10" pageId="12" pageNumber="115">Component funnel</th>
<td id="76D5F76CA237004FFD0F4586FC0AFA55" box="[699,972,1423,1447]" gridcol="1" gridrow="10" pageId="12" pageNumber="115">Absent</td>
<td id="76D5F76CA237004FFC4F4586FB7CFA55" box="[1019,1210,1423,1447]" gridcol="2" gridrow="10" pageId="12" pageNumber="115">Present</td>
<td id="76D5F76CA237004FFB6A4586FA4BFA55" box="[1246,1421,1423,1447]" gridcol="3" gridrow="10" pageId="12" pageNumber="115">Unknown</td>
</tr>
<tr id="35049E10A237004FFF1745BEFA4BFA3D" box="[163,1421,1463,1487]" gridrow="11" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF1745BEFA4BFA3D" box="[163,1421,1463,1487]" colspan="4" colspanRight="3" gridcol="0" gridrow="11" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFF1745BEFD8AFA3D" bold="true" box="[163,588,1463,1487]" pageId="12" pageNumber="115">Coloration patterns (on the dorsal disc)</emphasis>
</th>
</tr>
<tr id="35049E10A237004FFF1745E9FA4BFA0A" box="[163,1421,1504,1528]" gridrow="12" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF1745E9FD56FA0A" box="[163,656,1504,1528]" gridcol="0" gridrow="12" pageId="12" pageNumber="115">Fresh coloration</th>
<td id="76D5F76CA237004FFD0F45E9FC0AFA0A" box="[699,972,1504,1528]" gridcol="1" gridrow="12" pageId="12" pageNumber="115">Yellowish brown</td>
<td id="76D5F76CA237004FFC4F45E9FB7CFA0A" box="[1019,1210,1504,1528]" gridcol="2" gridrow="12" pageId="12" pageNumber="115">Dark brown</td>
<td id="76D5F76CA237004FFB6A45E9FA4BFA0A" box="[1246,1421,1504,1528]" gridcol="3" gridrow="12" pageId="12" pageNumber="115">Unknown</td>
</tr>
<tr id="35049E10A237004FFF174601FA4BF9D2" box="[163,1421,1544,1568]" gridrow="13" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174601FD56F9D2" box="[163,656,1544,1568]" gridcol="0" gridrow="13" pageId="12" pageNumber="115">Shape of the black spots</th>
<td id="76D5F76CA237004FFD0F4601FC0AF9D2" box="[699,972,1544,1568]" gridcol="1" gridrow="13" pageId="12" pageNumber="115">circular to irregular shaped</td>
<td id="76D5F76CA237004FFC4F4601FB7CF9D2" box="[1019,1210,1544,1568]" gridcol="2" gridrow="13" pageId="12" pageNumber="115">circular</td>
<td id="76D5F76CA237004FFB6A4601FA4BF9D2" box="[1246,1421,1544,1568]" gridcol="3" gridrow="13" pageId="12" pageNumber="115">Unknown</td>
</tr>
<tr id="35049E10A237004FFF174639FA4BF9BA" box="[163,1421,1584,1608]" gridrow="14" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174639FD56F9BA" box="[163,656,1584,1608]" gridcol="0" gridrow="14" pageId="12" pageNumber="115">Blotches</th>
<td id="76D5F76CA237004FFD0F4639FC0AF9BA" box="[699,972,1584,1608]" gridcol="1" gridrow="14" pageId="12" pageNumber="115">indistinct</td>
<td id="76D5F76CA237004FFC4F4639FB7CF9BA" box="[1019,1210,1584,1608]" gridcol="2" gridrow="14" pageId="12" pageNumber="115">distinct</td>
<td id="76D5F76CA237004FFB6A4639FA4BF9BA" box="[1246,1421,1584,1608]" gridcol="3" gridrow="14" pageId="12" pageNumber="115">absent</td>
</tr>
<tr id="35049E10A237004FFF174651FA4BF982" box="[163,1421,1624,1648]" gridrow="15" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174651FD56F982" box="[163,656,1624,1648]" gridcol="0" gridrow="15" pageId="12" pageNumber="115">Posterior ocellus</th>
<td id="76D5F76CA237004FFD0F4651FC0AF982" box="[699,972,1624,1648]" gridcol="1" gridrow="15" pageId="12" pageNumber="115">absent</td>
<td id="76D5F76CA237004FFC4F4651FB7CF982" box="[1019,1210,1624,1648]" gridcol="2" gridrow="15" pageId="12" pageNumber="115">sometimes present</td>
<td id="76D5F76CA237004FFB6A4651FA4BF982" box="[1246,1421,1624,1648]" gridcol="3" gridrow="15" pageId="12" pageNumber="115">absent</td>
</tr>
<tr id="35049E10A237004FFF174689FA4BF96B" box="[163,1421,1664,1689]" gridrow="16" pageId="12" pageNumber="115">
<th id="76D5F76CA237004FFF174689FD56F96B" box="[163,656,1664,1689]" gridcol="0" gridrow="16" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFF174689FE28F96A" bold="true" box="[163,494,1664,1688]" pageId="12" pageNumber="115">Colorations on the ventral disc</emphasis>
</th>
<td id="76D5F76CA237004FFD0F4689FC0AF96B" box="[699,972,1664,1689]" gridcol="1" gridrow="16" pageId="12" pageNumber="115">white</td>
<td id="76D5F76CA237004FFC4F4689FB7CF96B" box="[1019,1210,1664,1689]" gridcol="2" gridrow="16" pageId="12" pageNumber="115">pale to dark brown</td>
<td id="76D5F76CA237004FFB6A4689FA4BF96B" box="[1246,1421,1664,1689]" gridcol="3" gridrow="16" pageId="12" pageNumber="115">pale brown</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8B9C52A237FFBCFF7346D7FA9BF807" blockId="12.[151,1436,1758,2037]" pageId="12" pageNumber="115">
<emphasis id="B9404040A237FFBCFF7346D7FE8BF90A" bold="true" box="[199,333,1758,1784]" pageId="12" pageNumber="115">Coloration.</emphasis>
When fresh (
<figureCitation id="130F80D7A237FFBCFE5646D6FD8DF90B" box="[482,587,1759,1785]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="5.[152,255,1456,1480]" captionStartId-1="6.[152,255,1608,1632]" captionTargetBox-0="[151,1436,181,1432]" captionTargetBox-1="[151,1436,181,1584]" captionTargetId-0="figure-17@5.[151,1436,181,1433]" captionTargetId-1="figure-31@6.[151,1436,181,1584]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 1. Fresh coloration of Okamejei picta sp. nov., NMMB-P36013, 429 mm TL, adult male holotype: top, dorsal surface; bottom, ventral surface." captionText-1="FIGURE 2. Fresh coloration of Okamejei picta sp. nov., ASIZP0081116, 405 mm TL, adult male paratype: top, dorsal surface; bottom, ventral surface." figureDoi-0="http://doi.org/10.5281/zenodo.7895046" figureDoi-1="http://doi.org/10.5281/zenodo.7895048" httpUri-0="https://zenodo.org/record/7895046/files/figure.png" httpUri-1="https://zenodo.org/record/7895048/files/figure.png" pageId="12" pageNumber="115">Figs. 12</figureCitation>
), dorsal surface of disc, posterior pelvic-fin lobes, claspers and tail yellowish brown, densely covered by small black irregular spots; scattered larger blotches but indistinct; no prominent pair of pectoral markings and posterior ocellus; pectoral-fin margins translucent without spots or blotches; rostral cartilage uniformly pale brown, except for the posterior part with a few black speckles; dorsal fins and caudal fin translucent; eyelid brownish to translucent, eye dark; thorns pale, contrasted against darker dorsal surface of tail; tail strongly bicolored laterally, yellow dorsolaterally, pale ventralaterally; ventral surface of disc and tail uniformly whitish, pectoral fins slightly translucent; sensory pores dark edged, some surrounded by blotches. Coloration of female and juvenile unknown. In preservative, dorsal surface and sensory pores darker in general, others similar to fresh coloration.
</paragraph>
</subSubSection>
<caption id="DF4BCCDAA236FFBDFF2C44AEFC26FB11" ID-DOI="http://doi.org/10.5281/zenodo.7895068" ID-Zenodo-Dep="7895068" httpUri="https://zenodo.org/record/7895068/files/figure.png" pageId="13" pageNumber="116" startId="13.[152,255,1191,1215]" targetBox="[359,1228,181,1167]" targetPageId="13" targetType="figure">
<paragraph id="8B8B9C52A236FFBDFF2C44AEFC26FB11" blockId="13.[151,1436,1191,1251]" pageId="13" pageNumber="116">
<emphasis id="B9404040A236FFBDFF2C44AEFEE2FB4D" bold="true" box="[152,292,1191,1215]" pageId="13" pageNumber="116">FIGURE 11.</emphasis>
Ventral terminal cartilage from left clasper of
<taxonomicName id="4C34E7D1A236FFBDFD4E44AEFBC6FB4D" authority=", EBFS" authorityName="EBFS" box="[762,1024,1191,1215]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A236FFBDFD4E44AEFC70FB4D" box="[762,950,1191,1215]" italics="true" pageId="13" pageNumber="116">Okamejei hollandi</emphasis>
, EBFS
</taxonomicName>
adult male, uncatalogued specimen, 432 mm TL. (A) Dorsal view; (B) ventral view; (C) proximal tip viewed from left side.
</paragraph>
</caption>
<subSubSection id="C32ECFD9A236FFBDFF734527FA52FABB" box="[199,1428,1326,1353]" pageId="13" pageNumber="116" type="distribution">
<paragraph id="8B8B9C52A236FFBDFF734527FA52FABB" blockId="13.[151,1437,1326,2022]" box="[199,1428,1326,1353]" pageId="13" pageNumber="116">
<emphasis id="B9404040A236FFBDFF734527FE99FABA" bold="true" box="[199,351,1326,1352]" pageId="13" pageNumber="116">Distribution.</emphasis>
Known only from deep waters off western Dong-sha Atoll, South
<collectingCountry id="F323DCC2A236FFBDFBF54526FB45FABB" box="[1089,1155,1327,1353]" name="China" pageId="13" pageNumber="116">China</collectingCountry>
Sea, at depth ca.
<specimenCount id="9D3257DBA236FFBDFAF24526FA52FABB" box="[1350,1428,1327,1353]" pageId="13" pageNumber="116" type="generic">
<quantity id="4CCC31B7A236FFBDFAF24526FA56FABB" box="[1350,1424,1327,1353]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="116" unit="m" value="300.0">300 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C32ECFD9A236FFBDFF73455BFD96FA7D" pageId="13" pageNumber="116" type="description">
<paragraph id="8B8B9C52A236FFBDFF73455BFD96FA7D" blockId="13.[151,1437,1326,2022]" pageId="13" pageNumber="116">
<emphasis id="B9404040A236FFBDFF73455BFF38FA9E" bold="true" box="[199,254,1362,1388]" pageId="13" pageNumber="116">Size.</emphasis>
The
<typeStatus id="548F22F0A236FFBDFE83455BFEB4FA9E" box="[311,370,1362,1388]" pageId="13" pageNumber="116">types</typeStatus>
are both mature males (to
<quantity id="4CCC31B7A236FFBDFD2E455BFD31FA9E" box="[666,759,1362,1388]" metricMagnitude="-1" metricUnit="m" metricValue="4.29" pageId="13" pageNumber="116" unit="mm" value="429.0">429 mm</quantity>
TL) and no information on females, juveniles and egg cases. Females presumably attain larger sizes.
</paragraph>
</subSubSection>
<subSubSection id="C32ECFD9A236FFBDFF734591FB52FA27" pageId="13" pageNumber="116" type="etymology">
<paragraph id="8B8B9C52A236FFBDFF734591FB52FA27" blockId="13.[151,1437,1326,2022]" pageId="13" pageNumber="116">
<emphasis id="B9404040A236FFBDFF734591FE88FA40" bold="true" box="[199,334,1432,1458]" pageId="13" pageNumber="116">Etymology.</emphasis>
Derived from the Latin
<emphasis id="B9404040A236FFBDFDEF4590FD58FA40" box="[603,670,1433,1458]" italics="true" pageId="13" pageNumber="116">pictus</emphasis>
, which means painted or colored, referring to the densely distributed black spots on the contrasting yellowish brown dorsal disc. Vernacular: Fine-spotted Skate.
</paragraph>
</subSubSection>
<subSubSection id="C32ECFD9A236FFBEFF7345D6FBA3FB79" lastPageId="14" lastPageNumber="117" pageId="13" pageNumber="116" type="discussion">
<paragraph id="8B8B9C52A236FFBDFF7345D6FA48F990" blockId="13.[151,1437,1326,2022]" pageId="13" pageNumber="116">
<emphasis id="B9404040A236FFBDFF7345D6FEFCFA0B" bold="true" box="[199,314,1503,1529]" pageId="13" pageNumber="116">Remarks.</emphasis>
Among the genus
<taxonomicName id="4C34E7D1A236FFBDFDBB45D6FDBFFA0B" authorityName="Ishiyama" authorityYear="1958" box="[527,633,1503,1529]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="genus">
<emphasis id="B9404040A236FFBDFDBB45D6FDBFFA0B" box="[527,633,1503,1529]" italics="true" pageId="13" pageNumber="116">Okamejei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C34E7D1A236FFBDFD3C45D6FD24FA0A" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[648,738,1503,1529]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A236FFBDFD3C45D6FD24FA0A" box="[648,738,1503,1529]" italics="true" pageId="13" pageNumber="116">O. picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A236FFBDFD5F45D6FC83FA0B" bold="true" box="[747,837,1503,1529]" pageId="13" pageNumber="116">
<taxonomicNameLabel id="A273FD3BA236FFBDFD5F45D6FC83FA0B" box="[747,837,1503,1529]" pageId="13" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles two other western Pacific species, viz.,
<taxonomicName id="4C34E7D1A236FFBDFA3545D6FF35F9EE" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A236FFBDFA3545D6FF35F9EE" italics="true" pageId="13" pageNumber="116">O. hollandi</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C34E7D1A236FFBDFE84460BFE6EF9EE" box="[304,424,1538,1564]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A236FFBDFE84460BFE6EF9EE" box="[304,424,1538,1564]" italics="true" pageId="13" pageNumber="116">O. mengae</emphasis>
</taxonomicName>
. Both species share the characters of densely distributed black spots on the dorsal disc, and a W-shaped distribution of ventral sensory pores (
<bibRefCitation id="EFA5E1A3A236FFBDFD7E462CFC4DF9CD" author="Jeong, C. - H. &amp; Nakabo, T. &amp; Wu, H. - L." box="[714,907,1573,1600]" pageId="13" pageNumber="116" pagination="57 - 65" refId="ref7491" refString="Jeong, C. - H., Nakabo, T. &amp; Wu, H. - L. (2007) A new species of skate (Chondrichthyes: Rajidae), Okamejei mengae from the South China Sea. Korean Journal of Ichthyology, 19 (1), 57 - 65." type="journal article" year="2007">
Jeong
<emphasis id="B9404040A236FFBDFCA7462FFC85F9CD" box="[787,835,1573,1599]" italics="true" pageId="13" pageNumber="116">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="EFA5E1A3A236FFBDFC21462CFBAAF9CD" author="Misawa, R. &amp; Babaran, R. P. &amp; Motomura, H." box="[917,1132,1573,1600]" pageId="13" pageNumber="116" pagination="161 - 176" refId="ref8281" refString="Misawa, R., Babaran, R. P. &amp; Motomura, H. (2022) Okamejei panayensis sp. nov., a new skate (Rajiformes: Rajidae) from the Philippines. Ichthyological Research, 70, pages 161 - 176. https: // doi. org / 10.1007 / s 10228 - 022 - 00874 - 1" type="journal article" year="2022">
Misawa
<emphasis id="B9404040A236FFBDFC47462FFBE2F9CD" box="[1011,1060,1573,1599]" italics="true" pageId="13" pageNumber="116">et al</emphasis>
. 2022
</bibRefCitation>
). However, the new species is readily distinguished from the two species by coloration patterns, morphometrics and vertebral counts (
<tableCitation id="C6B6A9E9A236FFBDFA9E4641FA46F990" box="[1322,1408,1608,1634]" captionStart="TABLE 2" captionStartId="12.[152,245,944,968]" captionTargetPageId="12" captionText="TABLE 2. Comparisons of selected characters of O. picta sp. nov. and the similar species. Data of the holotypes of O. hollandi and O. mengae were obtained from Ishihara (1986) and Jeong et al. (2007), respectively." httpUri="http://table.plazi.org/id/DF4BCCDAA237FFBCFF2C43B9FB6FFC1E" pageId="13" pageNumber="116" tableUuid="DF4BCCDAA237FFBCFF2C43B9FB6FFC1E">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8B8B9C52A236FFBEFF734662FD62FDE3" blockId="13.[151,1437,1326,2022]" lastBlockId="14.[151,1437,150,1164]" lastPageId="14" lastPageNumber="117" pageId="13" pageNumber="116">
The coloration and patterns of skates are sensitive to environmental influences, that the same species may have different colorations, and sympatric species may exhibit overlapping coloration patterns (
<bibRefCitation id="EFA5E1A3A236FFBDFB6E4686FA51F95B" author="Weigmann, S." box="[1242,1431,1679,1705]" pageId="13" pageNumber="116" pagination="1176 - 1181" refId="ref8582" refString="Weigmann, S. (2017) Reply to Borsa (2017): Comment on ' Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity by Weigmann (2016) '. Journal of Fish Biology, 90 (4), 1176 - 1181. https: // doi. org / 10.1111 / jfb. 13234" type="journal article" year="2017">Weigmann 2017</bibRefCitation>
;
<bibRefCitation id="EFA5E1A3A236FFBDFF2346BBFEB0F93E" author="Misawa, R. &amp; Narimatsu, Y. &amp; Endo, H. &amp; Kai, Y." box="[151,374,1714,1740]" pageId="13" pageNumber="116" pagination="24 - 36" refId="ref8209" refString="Misawa, R., Narimatsu, Y., Endo, H. &amp; Kai, Y. (2019) Population structure of the ocellate spot skate (Okamejei kenojei) inferred from variations in mitochondrial DNA (mtDNA) sequences and from morphological characters of regional populations. Fishery Bulletin, 117, 24 - 36. https: // doi. org / 10.7755 / FB. 117.1.4" type="journal article" year="2019">
Misawa
<emphasis id="B9404040A236FFBDFF4D46BAFEF5F93E" box="[249,307,1714,1740]" italics="true" pageId="13" pageNumber="116">et al.</emphasis>
2019
</bibRefCitation>
;
<bibRefCitation id="EFA5E1A3A236FFBDFE3046BBFD41F93E" author="Gabbanelli, V. &amp; Naylor, G. &amp; Weigmann, S. &amp; Yang, L. &amp; Vazquez, D. M. &amp; Last, P. &amp; Diaz de Astarloa, J. M. &amp; Mabragana, E." box="[388,647,1714,1740]" pageId="13" pageNumber="116" pagination="76" refId="ref7324" refString="Gabbanelli, V., Naylor, G., Weigmann, S., Yang, L., Vazquez, D. M., Last, P., Diaz de Astarloa, J. M. &amp; Mabragana, E. (2022) Morphological and molecular evidence reveals Zearaja brevicaudata (Marini, 1933) is a senior synonym of Dipturus lamillai Concha, Caira, Ebert &amp; Pompert 2019. Zoological Studies, 61, 76." type="journal article" year="2022">
Gabbanelli
<emphasis id="B9404040A236FFBDFDB346BAFD87F93E" box="[519,577,1714,1740]" italics="true" pageId="13" pageNumber="116">et al.</emphasis>
2022
</bibRefCitation>
). Thus, cautions should be taken when using coloration as diagnostic character in description of new species. Nevertheless,
<taxonomicName id="4C34E7D1A236FFBDFCA346DCFCB0F91D" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[791,886,1749,1775]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A236FFBDFCA346DCFCB0F91D" box="[791,886,1749,1775]" italics="true" pageId="13" pageNumber="116">O. picta</emphasis>
</taxonomicName>
shows notable difference in several coloration patterns from
<taxonomicName id="4C34E7D1A236FFBDFE8046F0FE74F8E0" box="[308,434,1784,1811]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A236FFBDFE8046F0FE74F8E0" box="[308,434,1784,1811]" italics="true" pageId="13" pageNumber="116">O. hollandi</emphasis>
</taxonomicName>
(
<figureCitation id="130F80D7A236FFBDFE7646F1FDD0F8E0" box="[450,534,1784,1810]" captionStart="FIGURE 10" captionStartId="11.[152,255,1518,1542]" captionTargetBox="[158,1368,214,1490]" captionTargetId="figure-17@11.[151,1436,181,1494]" captionTargetPageId="11" captionText="FIGURE 10. Fresh coloration of Okamejei hollandi, EBFS adult male, uncatalogued specimen, 432 mm TL: top, dorsal surface; bottom, ventral surface." figureDoi="http://doi.org/10.5281/zenodo.7895066" httpUri="https://zenodo.org/record/7895066/files/figure.png" pageId="13" pageNumber="116">Fig. 10</figureCitation>
): a yellowish brown dorsal disc (vs. grayish to dark brown in
<taxonomicName id="4C34E7D1A236FFBDFB7246F0FA82F8E0" box="[1222,1348,1784,1811]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A236FFBDFB7246F0FA82F8E0" box="[1222,1348,1784,1811]" italics="true" pageId="13" pageNumber="116">O. hollandi</emphasis>
</taxonomicName>
); shape of the black spots on the dorsal disc circular to irregular (vs. uniformly circular); blotches indistinct (vs. distinct); posterior ocellus absent (vs. sometimes present); ventral disc mostly white (vs. pale to dark brown). By comparing the morphometrics,
<taxonomicName id="4C34E7D1A236FFBDFEC8476BFE1EF88E" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[380,472,1890,1916]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A236FFBDFEC8476BFE1EF88E" box="[380,472,1890,1916]" italics="true" pageId="13" pageNumber="116">O. picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A236FFBDFE55476BFDFBF88E" bold="true" box="[481,573,1890,1916]" pageId="13" pageNumber="116">
<taxonomicNameLabel id="A273FD3BA236FFBDFE55476BFDFBF88E" box="[481,573,1890,1916]" pageId="13" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C34E7D1A236FFBDFD63476BFC9EF88E" box="[727,856,1890,1916]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="13" pageNumber="116" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A236FFBDFD63476BFC9EF88E" box="[727,856,1890,1916]" italics="true" pageId="13" pageNumber="116">O. hollandi</emphasis>
</taxonomicName>
in having shorter disc (45.047.7, vs. 51.655.8% TL); a larger head (dorsal-head length 24.524.8, vs. 19.321.5% TL); a longer distance between cloaca to first dorsal-fin origin (32.132.3, vs. 25.129.2% TL); a longer distance between cloaca to second dorsal-fin origin (41.141.9, vs. 34.139.6% TL); a longer tail (distance between cloaca to caudal-fin tip 53.655.1, vs 47.853.4% TL); space between nostrils relatively narrow, its length 23.523.6% (vs. 24.125.7% ventral-head length); a longer anterior pelvic lobe (12.813.0, vs. 10.312.1% TL); a longer posterior pelvic lobe (17.618.5, vs. 14.517.4% TL); distance between first dorsal-fin origin and caudal-fin tip small (20.9, vs. 22.225.7% TL); distance between second dorsal-fin origin and caudal-fin tip small (11.412.2, vs. 13.214.5% TL); and a shorter caudal fin (5.35.7, vs. 5.88.0% TL). In meristics,
<taxonomicName id="4C34E7D1A235FFBEFE43412DFD97FECF" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[503,593,292,318]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A235FFBEFE43412DFD97FECF" box="[503,593,292,318]" italics="true" pageId="14" pageNumber="117">O. picta</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C34E7D1A235FFBEFD56412DFCA7FECF" box="[738,865,291,318]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A235FFBEFD56412DFCA7FECF" box="[738,865,291,318]" italics="true" pageId="14" pageNumber="117">O. hollandi</emphasis>
</taxonomicName>
in having more diplospondylous centra (9596, vs. 6683); more total centra (123126, vs. 95112); relatively few propterygials (2829, vs. 2932) and fewer total basal radials (7376, vs. 8286). When comparing clasper skeleton, the vt of
<taxonomicName id="4C34E7D1A235FFBEFC4B4163FB9FFE76" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[1023,1113,362,388]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A235FFBEFC4B4163FB9FFE76" box="[1023,1113,362,388]" italics="true" pageId="14" pageNumber="117">O. picta</emphasis>
</taxonomicName>
is J-shaped, with a bifurcate proximal tip, while the vt of
<taxonomicName id="4C34E7D1A235FFBEFE6C4184FD9EFE55" box="[472,600,397,423]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A235FFBEFE6C4184FD9EFE55" box="[472,600,397,423]" italics="true" pageId="14" pageNumber="117">O. hollandi</emphasis>
</taxonomicName>
is weakly-curved in shape, with a blunt proximal tip (
<figureCitation id="130F80D7A235FFBEFB024184FAC1FE55" box="[1206,1287,397,423]" captionStart="FIGURE 11" captionStartId="13.[152,255,1191,1215]" captionTargetBox="[359,1228,181,1167]" captionTargetId="figure-17@13.[359,1228,181,1167]" captionTargetPageId="13" captionText="FIGURE 11. Ventral terminal cartilage from left clasper of Okamejei hollandi, EBFS adult male, uncatalogued specimen, 432 mm TL. (A) Dorsal view; (B) ventral view; (C) proximal tip viewed from left side." figureDoi="http://doi.org/10.5281/zenodo.7895068" httpUri="https://zenodo.org/record/7895068/files/figure.png" pageId="14" pageNumber="117">Fig. 11</figureCitation>
). In
<taxonomicName id="4C34E7D1A235FFBEFA8F4184FA53FE55" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[1339,1429,397,423]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A235FFBEFA8F4184FA53FE55" box="[1339,1429,397,423]" italics="true" pageId="14" pageNumber="117">O. picta</emphasis>
</taxonomicName>
, the component funnel at the tip of clasper is not evident, but it is prominent in
<taxonomicName id="4C34E7D1A235FFBEFC4D41B8FBB1FE38" box="[1017,1143,432,459]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A235FFBEFC4D41B8FBB1FE38" box="[1017,1143,432,459]" italics="true" pageId="14" pageNumber="117">O. hollandi</emphasis>
</taxonomicName>
. Furthermore,
<taxonomicName id="4C34E7D1A235FFBEFAAA41B8FA5AFE38" box="[1310,1436,432,459]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A235FFBEFAAA41B8FA5AFE38" box="[1310,1436,432,459]" italics="true" pageId="14" pageNumber="117">O. hollandi</emphasis>
</taxonomicName>
is a shallow water species, typically found in coastal waters not deeper than
<quantity id="4CCC31B7A235FFBEFC4541DDFBFCFE1C" box="[1009,1082,468,494]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="117" unit="m" value="100.0">100 m</quantity>
, while the
<specimenCount id="9D3257DBA235FFBEFB0141DDFA9DFE1F" box="[1205,1371,467,494]" pageId="14" pageNumber="117" type="generic">two specimens</specimenCount>
of
<taxonomicName id="4C34E7D1A235FFBEFA3541DDFF09FDE2" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A235FFBEFA3541DDFF09FDE2" italics="true" pageId="14" pageNumber="117">O. picta</emphasis>
</taxonomicName>
were captured in deepwaters (ca.
<quantity id="4CCC31B7A235FFBEFDF941FEFD5EFDE3" box="[589,664,503,529]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="117" unit="m" value="300.0">300 m</quantity>
).
</paragraph>
<paragraph id="8B8B9C52A235FFBEFF734213FE0AFCD8" blockId="14.[151,1437,150,1164]" pageId="14" pageNumber="117">
<taxonomicName id="4C34E7D1A235FFBEFF734213FEB5FDC6" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[199,371,538,564]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A235FFBEFF734213FEB5FDC6" box="[199,371,538,564]" italics="true" pageId="14" pageNumber="117">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A235FFBEFECF4213FE13FDC6" bold="true" box="[379,469,538,564]" pageId="14" pageNumber="117">
<taxonomicNameLabel id="A273FD3BA235FFBEFECF4213FE13FDC6" box="[379,469,538,564]" pageId="14" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also similar to
<taxonomicName id="4C34E7D1A235FFBEFD154213FCDAFDC6" box="[673,796,538,564]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A235FFBEFD154213FCDAFDC6" box="[673,796,538,564]" italics="true" pageId="14" pageNumber="117">O. mengae</emphasis>
</taxonomicName>
known only from the
<typeStatus id="548F22F0A235FFBEFBAF4213FB46FDC6" box="[1051,1152,538,564]" pageId="14" pageNumber="117" type="holotype">holotype</typeStatus>
, for which the validity is often questioned (e.g.
<bibRefCitation id="EFA5E1A3A235FFBEFE3B4237FD86FDA5" author="Last P. R. &amp; Seret B. &amp; Stehmann M. F. W. &amp; Weigmann S." box="[399,576,573,600]" pageId="14" pageNumber="117" pagination="204 - 236" refId="ref7926" refString="Last P. R., Seret B., Stehmann M. F. W. &amp; Weigmann S. (2016) Skates. Family Rajidae. In: Last, P. R., Naylor, G., Seret, B., White, W. T., de Carvalho, M. &amp; Stehmann, M. (Eds.), Rays of the World. CSIRO publishing, Clayton North, pp. 204 - 236. https: // doi. org / 10.1071 / 9780643109148" type="book chapter" year="2016">
Last
<emphasis id="B9404040A235FFBEFE724237FE3BFDA5" box="[454,509,573,599]" italics="true" pageId="14" pageNumber="117">et al.</emphasis>
2016
</bibRefCitation>
;
<bibRefCitation id="EFA5E1A3A235FFBEFDFE4234FCE2FDA5" author="Misawa, R. &amp; Babaran, R. P. &amp; Motomura, H." box="[586,804,573,599]" pageId="14" pageNumber="117" pagination="161 - 176" refId="ref8281" refString="Misawa, R., Babaran, R. P. &amp; Motomura, H. (2022) Okamejei panayensis sp. nov., a new skate (Rajiformes: Rajidae) from the Philippines. Ichthyological Research, 70, pages 161 - 176. https: // doi. org / 10.1007 / s 10228 - 022 - 00874 - 1" type="journal article" year="2022">
Misawa
<emphasis id="B9404040A235FFBEFD1D4237FD27FDA5" box="[681,737,573,599]" italics="true" pageId="14" pageNumber="117">et al.</emphasis>
2022
</bibRefCitation>
). Nevertheless, the new species is readily separated from
<taxonomicName id="4C34E7D1A235FFBEFF234268FEC9FD88" box="[151,271,609,635]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A235FFBEFF234268FEC9FD88" box="[151,271,609,635]" italics="true" pageId="14" pageNumber="117">O. mengae</emphasis>
</taxonomicName>
by having a combination of characters (data obtained from
<bibRefCitation id="EFA5E1A3A235FFBEFC104268FBA3FD88" author="Jeong, C. - H. &amp; Nakabo, T. &amp; Wu, H. - L." box="[932,1125,608,635]" pageId="14" pageNumber="117" pagination="57 - 65" refId="ref7491" refString="Jeong, C. - H., Nakabo, T. &amp; Wu, H. - L. (2007) A new species of skate (Chondrichthyes: Rajidae), Okamejei mengae from the South China Sea. Korean Journal of Ichthyology, 19 (1), 57 - 65." type="journal article" year="2007">
Jeong
<emphasis id="B9404040A235FFBEFC5F4268FBDDFD88" box="[1003,1051,608,634]" italics="true" pageId="14" pageNumber="117">et al</emphasis>
. 2007
</bibRefCitation>
): blotches on the dorsal disc indistinct but present (vs. absent); a shorter disc (45.047.7, vs. 55.7% TL); a shorter snout (10.914.5, vs. 17.6% TL); a longer tail, distance between cloaca to caudal-fin tip 53.655.1 (vs. 48.5% TL); a much longer distance from second dorsal-fin origin to caudal-fin tip (11.412.2, vs. 5.8% TL); more trunk centra (31 vs. 23); and fewer total basal radials (7376, vs. 96). Further comparisons on clasper morphology and skeleton are needed when adult males of
<taxonomicName id="4C34E7D1A235FFBEFF024318FEE9FCD8" box="[182,303,785,811]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="mengae">
<emphasis id="B9404040A235FFBEFF024318FEE9FCD8" box="[182,303,785,811]" italics="true" pageId="14" pageNumber="117">O. mengae</emphasis>
</taxonomicName>
are available.
</paragraph>
<paragraph id="8B8B9C52A235FFBEFF73433AFBA3FB79" blockId="14.[151,1437,150,1164]" pageId="14" pageNumber="117">
The presence of a component funnel at the tip of clasper was generally considered a diagnostic character of the genus (e.g.
<bibRefCitation id="EFA5E1A3A235FFBEFEAC435EFDCBFC83" author="Last, P. R. &amp; Yearsley, G. K." box="[280,525,855,881]" pageId="14" pageNumber="117" pagination="1627 - 1641" refId="ref7782" refString="Last, P. R. &amp; Yearsley, G. K. (2002) Zoogeography and relationships of Australasian skates (Chondrichthyes: Rajidae). Journal of Biogeography, 29 (12), 1627 - 1641." type="journal article" year="2002">Last &amp; Yearsley 2002</bibRefCitation>
;
<bibRefCitation id="EFA5E1A3A235FFBEFDA3435EFCCEFC83" author="Last, P. R. &amp; Gledhill, D. C." box="[535,776,855,881]" pageId="14" pageNumber="117" pagination="119 - 134" refId="ref7693" refString="Last, P. R. &amp; Gledhill, D. C. (2008) Two new skates of the genus Okamejei (Rajoidei: Rajidae) from the south-east Indian Ocean. In: Last, P. R., White, W. T., Pogonoski, J. J. &amp; Gledhill, D. C. (Eds.), Descriptions of new Australian skates. CSIRO Marine and Atmospheric Research, Clayton North, pp. 119 - 134." type="book chapter" year="2008">Last &amp; Gledhill 2008</bibRefCitation>
;
<bibRefCitation id="EFA5E1A3A235FFBEFCA0435EFC03FC83" author="Last P. R. &amp; Fahmi &amp; Ishihara, H." box="[788,965,855,881]" pageId="14" pageNumber="117" pagination="89 - 100" refId="ref7839" refString="Last P. R., Fahmi &amp; Ishihara, H. (2010) Okamejei cairae sp. nov. (Rajoidei: Rajidae), a new skate from the South China Sea. In: Last, P. R., White, W. T. &amp; Pogonoski, J. J. (Eds.), Descriptions of New Sharks and Rays from Borneo. CSIRO Marine and Atmospheric Research, Clayton North, pp. 89 - 100." type="book chapter" year="2010">
Last
<emphasis id="B9404040A235FFBEFCFE435EFC44FC83" box="[842,898,855,881]" italics="true" pageId="14" pageNumber="117">et al.</emphasis>
2010
</bibRefCitation>
;
<bibRefCitation id="EFA5E1A3A235FFBEFC7B435EFB6FFC83" author="Misawa, R. &amp; Babaran, R. P. &amp; Motomura, H." box="[975,1193,855,881]" pageId="14" pageNumber="117" pagination="161 - 176" refId="ref8281" refString="Misawa, R., Babaran, R. P. &amp; Motomura, H. (2022) Okamejei panayensis sp. nov., a new skate (Rajiformes: Rajidae) from the Philippines. Ichthyological Research, 70, pages 161 - 176. https: // doi. org / 10.1007 / s 10228 - 022 - 00874 - 1" type="journal article" year="2022">
Misawa
<emphasis id="B9404040A235FFBEFB9A435EFBA0FC83" box="[1070,1126,855,881]" italics="true" pageId="14" pageNumber="117">et al.</emphasis>
2022
</bibRefCitation>
), but
<bibRefCitation id="EFA5E1A3A235FFBEFB51435EFF24FC67" author="Weigmann, S. &amp; Stehmann, M. F. W. &amp; Thiel, R." pageId="14" pageNumber="117" pagination="18 - 29" refId="ref8451" refString="Weigmann, S., Stehmann, M. F. W. &amp; Thiel, R. (2015) Okamejei ornata n. sp., a new deep-water skate (Elasmobranchii, Rajidae) from the northwestern Indian Ocean off Socotra Islands. Deep-Sea Research II: Topical Studies in Oceanography, 115, 18 - 29. https: // doi. org / 10.1016 / j. dsr 2.2013.09.005" type="journal article" year="2015">
Weigmann
<emphasis id="B9404040A235FFBEFAD1435EFA5BFC83" box="[1381,1437,855,881]" italics="true" pageId="14" pageNumber="117">et al.</emphasis>
(2015)
</bibRefCitation>
excluded it from the diagnosis. In the description of
<taxonomicName id="4C34E7D1A235FFBEFCF44373FB0CFC67" authority=", Weigmann et al. (2015)" authorityName="Weigmann" authorityYear="2015" box="[832,1226,890,917]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="ornata">
<emphasis id="B9404040A235FFBEFCF44373FC68FC66" box="[832,942,890,916]" italics="true" pageId="14" pageNumber="117">O. ornata</emphasis>
,
<bibRefCitation id="EFA5E1A3A235FFBEFC094373FB0CFC67" author="Weigmann, S. &amp; Stehmann, M. F. W. &amp; Thiel, R." box="[957,1226,890,917]" pageId="14" pageNumber="117" pagination="18 - 29" refId="ref8451" refString="Weigmann, S., Stehmann, M. F. W. &amp; Thiel, R. (2015) Okamejei ornata n. sp., a new deep-water skate (Elasmobranchii, Rajidae) from the northwestern Indian Ocean off Socotra Islands. Deep-Sea Research II: Topical Studies in Oceanography, 115, 18 - 29. https: // doi. org / 10.1016 / j. dsr 2.2013.09.005" type="journal article" year="2015">
Weigmann
<emphasis id="B9404040A235FFBEFB8A4372FBB1FC66" box="[1086,1143,890,916]" italics="true" pageId="14" pageNumber="117">et al.</emphasis>
(2015)
</bibRefCitation>
</taxonomicName>
found out that the component funnel was not evident, but the species share typical characters of genus
<taxonomicName id="4C34E7D1A235FFBEFBFF4394FB70FC45" authorityName="Ishiyama" authorityYear="1958" box="[1099,1206,925,951]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis id="B9404040A235FFBEFBFF4394FB70FC45" box="[1099,1206,925,951]" italics="true" pageId="14" pageNumber="117">Okamejei</emphasis>
</taxonomicName>
. Our results support
<bibRefCitation id="EFA5E1A3A235FFBEFF2343C9FE64FC29" author="Weigmann, S. &amp; Stehmann, M. F. W. &amp; Thiel, R." box="[151,418,960,987]" pageId="14" pageNumber="117" pagination="18 - 29" refId="ref8451" refString="Weigmann, S., Stehmann, M. F. W. &amp; Thiel, R. (2015) Okamejei ornata n. sp., a new deep-water skate (Elasmobranchii, Rajidae) from the northwestern Indian Ocean off Socotra Islands. Deep-Sea Research II: Topical Studies in Oceanography, 115, 18 - 29. https: // doi. org / 10.1016 / j. dsr 2.2013.09.005" type="journal article" year="2015">
Weigmann
<emphasis id="B9404040A235FFBEFEA343C8FE96FC28" box="[279,336,960,986]" italics="true" pageId="14" pageNumber="117">et al.</emphasis>
(2015)
</bibRefCitation>
that the presence of the component funnel may not be a diagnostic character of the genus.
<taxonomicName id="4C34E7D1A235FFBEFF2343EAFE85FC0F" authority="Ng &amp; Ho &amp; Joung &amp; Liu, 2023" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[151,323,995,1021]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta" status="sp. nov.">
<emphasis id="B9404040A235FFBEFF2343EAFE85FC0F" box="[151,323,995,1021]" italics="true" pageId="14" pageNumber="117">Okamejei picta</emphasis>
</taxonomicName>
<emphasis id="B9404040A235FFBEFEF843EDFE61FC0C" bold="true" box="[332,423,996,1022]" pageId="14" pageNumber="117">
<taxonomicNameLabel id="A273FD3BA235FFBEFEF843EDFE61FC0C" box="[332,423,996,1022]" pageId="14" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, shares all the diagnostic characters of
<taxonomicName id="4C34E7D1A235FFBEFCDF43EAFC10FC0F" authorityName="Ishiyama" authorityYear="1958" box="[875,982,995,1021]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis id="B9404040A235FFBEFCDF43EAFC10FC0F" box="[875,982,995,1021]" italics="true" pageId="14" pageNumber="117">Okamejei</emphasis>
</taxonomicName>
(defined in introduction) except for the absence of a component funnel, so the assignment of the new species to genus
<taxonomicName id="4C34E7D1A235FFBEFBA9440EFB4EFBD3" authorityName="Ishiyama" authorityYear="1958" box="[1053,1160,1031,1057]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis id="B9404040A235FFBEFBA9440EFB4EFBD3" box="[1053,1160,1031,1057]" italics="true" pageId="14" pageNumber="117">Okamejei</emphasis>
</taxonomicName>
is undoubted.
<taxonomicName id="4C34E7D1A235FFBEFA85440EFF09FBB6" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A235FFBEFA85440EFF09FBB6" italics="true" pageId="14" pageNumber="117">Okamejei picta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C34E7D1A235FFBEFEBF4423FEBDFBB6" box="[267,379,1066,1092]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="ornata">
<emphasis id="B9404040A235FFBEFEBF4423FEBDFBB6" box="[267,379,1066,1092]" italics="true" pageId="14" pageNumber="117">O. ornata</emphasis>
</taxonomicName>
are the only two species without the component funnel in the claspers, but
<taxonomicName id="4C34E7D1A235FFBEFB5F4423FA8EFBB6" authorityName="Ng &amp; Ho &amp; Joung &amp; Liu" authorityYear="2023" box="[1259,1352,1066,1092]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="picta">
<emphasis id="B9404040A235FFBEFB5F4423FA8EFBB6" box="[1259,1352,1066,1092]" italics="true" pageId="14" pageNumber="117">O. picta</emphasis>
</taxonomicName>
can be readily distinguished from
<taxonomicName id="4C34E7D1A235FFBEFE7C4444FDF0FB95" box="[456,566,1101,1127]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="ornata">
<emphasis id="B9404040A235FFBEFE7C4444FDF0FB95" box="[456,566,1101,1127]" italics="true" pageId="14" pageNumber="117">O. ornata</emphasis>
</taxonomicName>
by having densely distributed black spots on the dorsal disc (vs. only patchy distributed blotches present) and a bifurcated proximal tip of vt (vs. tip not bifurcated).
</paragraph>
</subSubSection>
<subSubSection id="C32ECFD9A235FFBEFF2344BBFDE1FA1E" pageId="14" pageNumber="117" type="materials_examined">
<paragraph id="8B8B9C52A235FFBEFF2344BBFDE1FA1E" blockId="14.[151,1437,1202,1517]" pageId="14" pageNumber="117">
<emphasis id="B9404040A235FFBEFF2344BBFD73FB3E" bold="true" box="[151,693,1202,1229]" pageId="14" pageNumber="117">
Comparative materials.
<taxonomicName id="4C34E7D1A235FFBEFE7844BBFD6DFB3F" box="[460,683,1202,1229]" class="Elasmobranchii" family="Rajidae" genus="Okamejei" kingdom="Animalia" order="Rajiformes" pageId="14" pageNumber="117" phylum="Chordata" rank="species" species="hollandi">
<emphasis id="B9404040A235FFBEFE7844BBFD6DFB3F" bold="true" box="[460,683,1202,1229]" italics="true" pageId="14" pageNumber="117">Okamejei hollandi</emphasis>
</taxonomicName>
:
</emphasis>
<materialsCitation id="3B5C960FA235FFBEFD7244BBFDFDFB02" ID-GBIF-Occurrence="4100388308" collectingDate="2010-12-31" collectionCode="NMMB-P" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCode="NMMB-P15670" specimenCount="1">
<specimenCode id="DB923429A235FFBEFD7244BBFCBBFB3F" box="[710,893,1202,1229]" collectionCode="NMMB-P" pageId="14" pageNumber="117">NMMB-P15670</specimenCode>
(1, 476),
<collectorName id="26C1F984A235FFBEFBBC44BAFBBBFB3F" box="[1032,1149,1203,1229]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<location id="8EEBCA89A235FFBEFA8444BBFA5EFB3E" LSID="urn:lsid:plazi:treatment:039D2D44A23AFFBEFF2345AFFDE1FA1E:8EEBCA89A235FFBEFA8444BBFA5EFB3E" box="[1328,1432,1202,1228]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</location>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFE8144DFFE4CFB02" box="[309,394,1238,1264]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFE2D44DFFDFDFB02" box="[409,571,1238,1265]" pageId="14" pageNumber="117" value="2010-12-31">
<collectingDate id="EFCE437AA235FFBEFE2D44DFFDFDFB02" box="[409,571,1238,1265]" pageId="14" pageNumber="117" value="2010-12-31">31 Dec. 2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFDFD44DFFE5EFAE6" ID-GBIF-Occurrence="4100388310" collectingDate="2010-08-18" collectionCode="NMMB-P" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCode="NMMB-P15678" specimenCount="1" stateProvince="Pingtung">
<specimenCode id="DB923429A235FFBEFDFD44DFFCC7FB03" box="[585,769,1238,1265]" collectionCode="NMMB-P" pageId="14" pageNumber="117">NMMB-P15678</specimenCode>
(1, 415),
<collectorName id="26C1F984A235FFBEFCCE44DEFC29FB03" box="[890,1007,1239,1265]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<collectingRegion id="49F052B0A235FFBEFB2544DFFB3FFB02" box="[1169,1273,1238,1264]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFF2344F3FF2AFAE6" box="[151,236,1274,1300]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFF4D44F3FE5EFAE6" box="[249,408,1274,1301]" pageId="14" pageNumber="117" value="2010-08-18">
<collectingDate id="EFCE437AA235FFBEFF4D44F3FE5EFAE6" box="[249,408,1274,1301]" pageId="14" pageNumber="117" value="2010-08-18">18 Aug. 2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFE1044F3FF15FACA" ID-GBIF-Occurrence="4100388305" collectingDate="2010-03-03" collectionCode="NMMB-P" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCode="NMMB-P15680" specimenCount="1" stateProvince="Pingtung">
<specimenCode id="DB923429A235FFBEFE1044F3FD9DFAE7" box="[420,603,1274,1301]" collectionCode="NMMB-P" pageId="14" pageNumber="117">NMMB-P15680</specimenCode>
(1, 460),
<collectorName id="26C1F984A235FFBEFD7A44F2FC85FAE7" box="[718,835,1275,1301]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<collectingRegion id="49F052B0A235FFBEFC6B44F3FB81FAE6" box="[991,1095,1274,1300]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFB4444F3FA83FAE6" box="[1264,1349,1274,1300]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFAE544F3FF15FACA" pageId="14" pageNumber="117" value="2010-03-03">
<collectingDate id="EFCE437AA235FFBEFAE544F3FF15FACA" pageId="14" pageNumber="117" value="2010-03-03">3 Mar. 2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFF544517FAEBFACA" ID-GBIF-Occurrence="4100388302" box="[224,1325,1310,1337]" collectingDate="2010-03-03" collectionCode="NMMB-P" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCode="NMMB-P15682" specimenCount="1" stateProvince="Pingtung">
<specimenCode id="DB923429A235FFBEFF544517FE51FACB" box="[224,407,1310,1337]" collectionCode="NMMB-P" pageId="14" pageNumber="117">NMMB-P15682</specimenCode>
(1, 460),
<collectorName id="26C1F984A235FFBEFDBB4516FD42FACB" box="[527,644,1311,1337]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<collectingRegion id="49F052B0A235FFBEFC904517FC4AFACA" box="[804,908,1310,1336]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFB8C4517FB4BFACA" box="[1080,1165,1310,1336]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFB2F4517FAEBFACA" box="[1179,1325,1310,1337]" pageId="14" pageNumber="117" value="2010-03-03">
<collectingDate id="EFCE437AA235FFBEFB2F4517FAEBFACA" box="[1179,1325,1310,1337]" pageId="14" pageNumber="117" value="2010-03-03">3 Mar. 2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFA8E4516FBB0FAAE" ID-GBIF-Occurrence="4100388307" collectingDate="2010-05-09" collectionCode="NMMBP" country="Taiwan" location="Kaohsiung" pageId="14" pageNumber="117" specimenCode="NMMBP16393" specimenCount="1" stateProvince="Kaohsiung">
<specimenCode id="DB923429A235FFBEFA8E4516FF2BFAAE" collectionCode="NMMBP" country="Taiwan" httpUri="http://grbio.org/cool/kt9t-t1e8" name="National Museum of Marine Biology and Aquarium" pageId="14" pageNumber="117">NMMBP16393</specimenCode>
(1, 350), Ke-tzu-liao fishing port,
<collectingRegion id="49F052B0A235FFBEFDD9454BFD2FFAAE" box="[621,745,1346,1372]" country="Taiwan" name="Kaohsiung" pageId="14" pageNumber="117">Kaohsiung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFC39454BFC24FAAE" box="[909,994,1346,1372]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFC58454BFBB0FAAE" box="[1004,1142,1346,1373]" pageId="14" pageNumber="117" value="2010-05-09">
<collectingDate id="EFCE437AA235FFBEFC58454BFBB0FAAE" box="[1004,1142,1346,1373]" pageId="14" pageNumber="117" value="2010-05-09">9 May 2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFBCB454BFC52FA72" ID-GBIF-Occurrence="4100388309" collectingDate="2010-07-10" collectionCode="NMMB-P" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCode="NMMB-P16757" specimenCount="1" stateProvince="Pingtung">
<specimenCode id="DB923429A235FFBEFBCB454BFAF1FAAF" box="[1151,1335,1346,1373]" collectionCode="NMMB-P" pageId="14" pageNumber="117">NMMB-P16757</specimenCode>
(1, 426),
<collectorName id="26C1F984A235FFBEFF23456EFECAFA73" box="[151,268,1383,1409]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<collectingRegion id="49F052B0A235FFBEFE2B456FFDC1FA72" box="[415,519,1382,1408]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFD1E456FFD39FA72" box="[682,767,1382,1408]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFCBD456FFC52FA72" box="[777,916,1382,1408]" pageId="14" pageNumber="117" value="2010-07-10">
<collectingDate id="EFCE437AA235FFBEFCBD456FFC52FA72" box="[777,916,1382,1408]" pageId="14" pageNumber="117" value="2010-07-10">10 Jul. 2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFC29456FFD26FA56" ID-GBIF-Occurrence="4100388306" collectingDate="2012-08-09" collectionCode="NMMB-P" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCode="NMMB-P17093" specimenCount="1" stateProvince="Pingtung">
<specimenCode id="DB923429A235FFBEFC29456FFB93FA72" box="[925,1109,1382,1408]" collectionCode="NMMB-P" pageId="14" pageNumber="117">NMMB-P17093</specimenCode>
(1, 462.5),
<collectorName id="26C1F984A235FFBEFB60456EFA8FFA73" box="[1236,1353,1383,1409]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<collectingRegion id="49F052B0A235FFBEFF614583FEFBFA56" box="[213,317,1418,1444]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFE5D4583FDF8FA56" box="[489,574,1418,1444]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFDF84583FD26FA56" box="[588,736,1418,1445]" pageId="14" pageNumber="117" value="2012-08-09">
<collectingDate id="EFCE437AA235FFBEFDF84583FD26FA56" box="[588,736,1418,1445]" pageId="14" pageNumber="117" value="2012-08-09">9 Aug. 2012</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFD594583FDF4FA3A" ID-GBIF-Occurrence="4100388301" collectingDate="2012-08-06" collectionCode="NMMB-P" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCode="NMMB-P17262" specimenCount="1" stateProvince="Pingtung">
<specimenCode id="DB923429A235FFBEFD594583FC62FA56" box="[749,932,1418,1444]" collectionCode="NMMB-P" pageId="14" pageNumber="117">NMMB-P17262</specimenCode>
(1, 480),
<collectorName id="26C1F984A235FFBEFBAF4582FB56FA57" box="[1051,1168,1419,1445]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<collectingRegion id="49F052B0A235FFBEFA844583FA5EFA56" box="[1328,1432,1418,1444]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFE8245A7FE4DFA3A" box="[310,395,1454,1480]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFE2F45A7FDF4FA3A" box="[411,562,1454,1481]" pageId="14" pageNumber="117" value="2012-08-06">
<collectingDate id="EFCE437AA235FFBEFE2F45A7FDF4FA3A" box="[411,562,1454,1481]" pageId="14" pageNumber="117" value="2012-08-06">6 Aug. 2012</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B5C960FA235FFBEFDF545A7FDE1FA1E" ID-GBIF-Occurrence="4100388303" collectingDate="2022-01-10" collectionCode="EBFS" collectorName="Donggang" country="Taiwan" location="Pingtung" pageId="14" pageNumber="117" specimenCount="1" stateProvince="Pingtung">
<collectionCode id="ED250497A235FFBEFDF545A7FD42FA3A" box="[577,644,1454,1480]" pageId="14" pageNumber="117">EBFS</collectionCode>
uncatalogued specimen (1, 432),
<collectorName id="26C1F984A235FFBEFBA245A6FB4DFA3B" box="[1046,1163,1455,1481]" pageId="14" pageNumber="117">Donggang</collectorName>
fishing port,
<collectingRegion id="49F052B0A235FFBEFA8445A7FA5EFA3A" box="[1328,1432,1454,1480]" country="Taiwan" name="Pingtung" pageId="14" pageNumber="117">Pingtung</collectingRegion>
, southwestern
<collectingCountry id="F323DCC2A235FFBEFE8545DBFE40FA1E" box="[305,390,1490,1516]" name="Taiwan" pageId="14" pageNumber="117">Taiwan</collectingCountry>
,
<date id="FF8ABA92A235FFBEFE2545DBFDE2FA1E" box="[401,548,1490,1517]" pageId="14" pageNumber="117" value="2022-01-10">
<collectingDate id="EFCE437AA235FFBEFE2545DBFDE2FA1E" box="[401,548,1490,1517]" pageId="14" pageNumber="117" value="2022-01-10">10 Jan. 2022</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>