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<mods:title id="FF525FE55250964749409189B42846D7">On the generic placement of the narrow-range endemic ‘ Helix ’ arguineguinensis Seddon &amp; Aparicio, 1998 from Gran Canaria (Canary Islands)</mods:title>
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<treatment id="038E4F18AC1BFFA8FF2C4412FBA169AF" ID-DOI="http://doi.org/10.5281/zenodo.6114361" ID-GBIF-Taxon="119635276" ID-Zenodo-Dep="6114361" LSID="urn:lsid:plazi:treatment:038E4F18AC1BFFA8FF2C4412FBA169AF" httpUri="http://treatment.plazi.org/id/038E4F18AC1BFFA8FF2C4412FBA169AF" lastPageId="3" lastPageNumber="299" pageId="1" pageNumber="297">
<subSubSection id="C33DAD85AC1BFFAAFF2C4412FE396B04" box="[151,486,322,348]" pageId="1" pageNumber="297" type="nomenclature">
<paragraph id="8B98FE0EAC1BFFAAFF2C4412FE396B04" blockId="1.[151,486,322,348]" box="[151,486,322,348]" pageId="1" pageNumber="297">
<heading id="D0D04962AC1BFFAAFF2C4412FE396B04" bold="true" box="[151,486,322,348]" fontSize="11" level="1" pageId="1" pageNumber="297" reason="1">
<emphasis id="B953221CAC1BFFAAFF2C4412FE396B04" bold="true" box="[151,486,322,348]" pageId="1" pageNumber="297">
Subgenus
<taxonomicName id="4C27858DAC1BFFAAFEAC4412FE416B04" box="[279,414,322,348]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="B953221CAC1BFFAAFEAC4412FE416B04" bold="true" box="[279,414,322,348]" italics="true" pageId="1" pageNumber="297">Monilearia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2609F67AC1BFFAAFE1E4412FE396B04" box="[421,486,322,348]" pageId="1" pageNumber="297" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C33DAD85AC1BFFAAFF2C44D8FA856DA9" pageId="1" pageNumber="297" type="reference_group">
<paragraph id="8B98FE0EAC1BFFAAFF2C44D8FC586BC7" blockId="1.[151,903,392,415]" box="[151,903,392,415]" pageId="1" pageNumber="297">
<taxonomicName id="4C27858DAC1BFFAAFF2C44D8FC586BC7" authority="Seddon &amp; Aparicio, 1998" authorityName="Seddon &amp; Aparicio" authorityYear="1998" box="[151,903,392,415]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="species" species="arguineguinensis" subGenus="Monilearia">
<emphasis id="B953221CAC1BFFAAFF2C44D8FED06BC7" box="[151,271,392,415]" italics="true" pageId="1" pageNumber="297">Monilearia</emphasis>
(
<emphasis id="B953221CAC1BFFAAFEA544D8FE496BC7" box="[286,406,392,415]" italics="true" pageId="1" pageNumber="297">Monilearia</emphasis>
)
<emphasis id="B953221CAC1BFFAAFE1D44D8FD836BC7" box="[422,604,392,415]" italics="true" pageId="1" pageNumber="297">arguineguinensis</emphasis>
(
<bibRefCitation id="EFB683FFAC1BFFAAFDD144D8FCA16BC7" author="Seddon" box="[618,894,392,415]" pageId="1" pageNumber="297" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998">Seddon &amp; Aparicio, 1998</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8B98FE0EAC1BFFAAFF2C449AFA8C685C" blockId="1.[151,1435,458,516]" pageId="1" pageNumber="297">
<emphasis id="B953221CAC1BFFAAFF2C449AFECC6BB9" bold="true" box="[151,275,458,481]" pageId="1" pageNumber="297">Synonyms.</emphasis>
<taxonomicName id="4C27858DAC1BFFAAFEA0449AFE006BB9" box="[283,479,458,481]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="species" species="argonautula">
<emphasis id="B953221CAC1BFFAAFEA0449AFE006BB9" box="[283,479,458,481]" italics="true" pageId="1" pageNumber="297">Helix argonautula</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC1BFFAAFE5C449AFCE16BB9" authority="Mousson, 1874" authorityName="Mousson" authorityYear="1874" box="[487,830,458,481]" pageId="1" pageNumber="297" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC1BFFAAFDAD449AFD4E6BB9" box="[534,657,458,481]" italics="true" pageId="1" pageNumber="297">canariensis</emphasis>
<bibRefCitation id="EFB683FFAC1BFFAAFD22449AFCE16BB9" author="Mousson" box="[665,830,458,481]" pageId="1" pageNumber="297" refString="Mousson, A. (1874) Bemerkungen uber die von Hrn. Dr. von Fritsch und Dr. Rein aus West-Marocco 1872 zuruckgebrachten Land- und Susswasser-Mollusken. (Schluss). Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 1, 81 - 107, pls. 4 - 6." type="journal article" year="1874">Mousson, 1874</bibRefCitation>
</taxonomicName>
(non
<taxonomicName id="4C27858DAC1BFFAAFC3B449AFAB26BB9" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[896,1389,458,481]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="species" species="canariensis" subGenus="Xerophila">
<emphasis id="B953221CAC1BFFAAFC3B449AFC656BB9" box="[896,954,458,481]" italics="true" pageId="1" pageNumber="297">Helix</emphasis>
(
<emphasis id="B953221CAC1BFFAAFC71449AFBEB6BB9" box="[970,1076,458,481]" italics="true" pageId="1" pageNumber="297">Xerophila</emphasis>
)
<emphasis id="B953221CAC1BFFAAFBFE449AFB1F6BB9" box="[1093,1216,458,481]" italics="true" pageId="1" pageNumber="297">canariensis</emphasis>
<bibRefCitation id="EFB683FFAC1BFFAAFB73449AFAB26BB9" author="Mousson" box="[1224,1389,458,481]" pageId="1" pageNumber="297" refString="Mousson, A. (1872) Revision de la faune malacologique des iles Canaries. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, 25, 1 - 176, pls. I - VI." type="journal article" year="1872">Mousson, 1872</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C27858DAC1BFFAAFF2C44BDFE7E685C" box="[151,417,493,516]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="species" species="lactea" subGenus="Macularia">
<emphasis id="B953221CAC1BFFAAFF2C44BDFF0E685C" box="[151,209,493,516]" italics="true" pageId="1" pageNumber="297">Helix</emphasis>
(
<emphasis id="B953221CAC1BFFAAFF5B44BDFE8E685C" box="[224,337,493,516]" italics="true" pageId="1" pageNumber="297">Macularia</emphasis>
)
<emphasis id="B953221CAC1BFFAAFEDB44BDFE7E685C" box="[352,417,493,516]" italics="true" pageId="1" pageNumber="297">lactea</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC1BFFAAFE1344BDFD20685C" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[424,767,493,516]" pageId="1" pageNumber="297" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC1BFFAAFE6D44BDFD8E685C" box="[470,593,493,516]" italics="true" pageId="1" pageNumber="297">canariensis</emphasis>
<bibRefCitation id="EFB683FFAC1BFFAAFDEC44BDFD20685C" author="Mousson" box="[599,767,493,516]" pageId="1" pageNumber="297" refString="Mousson, A. (1872) Revision de la faune malacologique des iles Canaries. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, 25, 1 - 176, pls. I - VI." type="journal article" year="1872">Mousson, 1872</bibRefCitation>
</taxonomicName>
). For further synonyms, see
<bibRefCitation id="EFB683FFAC1BFFAAFB9444BDFA90685C" author="Seddon" box="[1071,1359,493,516]" pageId="1" pageNumber="297" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998">Seddon &amp; Aparicio (1998)</bibRefCitation>
.
</paragraph>
<paragraph id="8B98FE0EAC1BFFAAFF2C4217FA856DA9" blockId="1.[151,1436,1863,2033]" pageId="1" pageNumber="297">
<emphasis id="B953221CAC1BFFAAFF2C4217FECC6D05" bold="true" box="[151,275,1863,1885]" pageId="1" pageNumber="297">FIGURE 1.</emphasis>
Schematic representations of the generalized penial appendage of orthurethran gastropods (A) and the vagianal appendage of genera in Cochlicellini:
<taxonomicName id="4C27858DAC1BFFAAFDB04234FD2B6D21" authority="Hartmann, 1842" authorityName="Hartmann" authorityYear="1842" box="[523,756,1892,1914]" class="Gastropoda" family="Hygromiidae" genus="Obelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="genus">
<emphasis id="B953221CAC1BFFAAFDB04234FD906D22" box="[523,591,1892,1914]" italics="true" pageId="1" pageNumber="297">Obelus</emphasis>
Hartmann, 1842
</taxonomicName>
(B),
<taxonomicName id="4C27858DAC1BFFAAFC924234FBEE6D21" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[809,1073,1892,1914]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="genus">
<emphasis id="B953221CAC1BFFAAFC924234FC4A6D22" box="[809,917,1892,1914]" italics="true" pageId="1" pageNumber="297">Monilearia</emphasis>
<bibRefCitation id="EFB683FFAC1BFFAAFC274234FBEE6D21" author="Mousson" box="[924,1073,1892,1913]" pageId="1" pageNumber="297" refString="Mousson, A. (1872) Revision de la faune malacologique des iles Canaries. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, 25, 1 - 176, pls. I - VI." type="journal article" year="1872">Mousson, 1872</bibRefCitation>
</taxonomicName>
(C),
<taxonomicName id="4C27858DAC1BFFAAFBDD4234FAB26D21" authority="Ferussac, 1821" authorityName="Ferussac" authorityYear="1821" box="[1126,1389,1892,1914]" class="Gastropoda" family="Hygromiidae" genus="Cochlicella" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="genus">
<emphasis id="B953221CAC1BFFAAFBDD4234FB0A6D22" box="[1126,1237,1892,1914]" italics="true" pageId="1" pageNumber="297">Cochlicella</emphasis>
<bibRefCitation id="EFB683FFAC1BFFAAFB674234FAB26D21" author="Ferussac" box="[1244,1389,1892,1913]" pageId="1" pageNumber="297" refString="Ferussac, A. E. J. P. J. F. d'Audebard de (1821 - 1822) Tableaux systematiques des animaux mollusques classes en familles naturelles, dans lesquels on a etabli la concordance de tous les systemes; suivis d'un prodrome general pour tous les mollusques terrestres ou fluviatiles, vivants ou fossiles. Bertrand &amp; Sowerby, Paris &amp; Londres, XLVII + 1 + 27 + 110 + 1 pp." type="book" year="1821">Férussac, 1821</bibRefCitation>
</taxonomicName>
(D), and
<taxonomicName id="4C27858DAC1BFFAAFF7842D1FDBC6DCF" authority="Schileyko &amp; Menkhorst, 1997" authorityName="Schileyko &amp; Menkhorst" authorityYear="1997" box="[195,611,1921,1943]" class="Gastropoda" family="Hygromiidae" genus="Prietocella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="genus">
<emphasis id="B953221CAC1BFFAAFF7842D1FEF16DCF" box="[195,302,1921,1943]" italics="true" pageId="1" pageNumber="297">Prietocella</emphasis>
<bibRefCitation id="EFB683FFAC1BFFAAFE8D42D2FDBC6DCF" author="Schileyko" box="[310,611,1922,1943]" pageId="1" pageNumber="297" refString="Schileyko, A. A. &amp; Menkhorst, H. P. M. G. (1997) Composition and phylogenetic relations of the Cochlicellidae (Gastropoda, Pulmonata). Ruthenica, 7, 51 - 60." type="journal article" year="1997">Schileyko &amp; Menkhorst, 1997</bibRefCitation>
</taxonomicName>
(E) (after
<bibRefCitation id="EFB683FFAC1BFFAAFD6A42D2FC506DCF" author="Ibanez" box="[721,911,1922,1943]" pageId="1" pageNumber="297" refString="Ibanez, M., Alonso, M. R., Groh, K. &amp; Hutterer, R. (2003) The Genus Obelus Hartmann, 1842 (Gastropoda, Pulmonata, Helicoidea) and its phylogenetic relationships. Zoologischer Anzeiger, 242, 157 - 167. http: // dx. doi. org / 10.1078 / 0044 - 5231 - 00094" type="journal article" year="2003">Ibáñez et al., 2003</bibRefCitation>
), and genital system (F) of
<taxonomicName id="4C27858DAC1BFFAAFB1042D1FDBD6DEC" authority="Seddon &amp; Aparicio, 1998" authorityName="Seddon &amp; Aparicio" authorityYear="1998" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="297" phylum="Mollusca" rank="species" species="arguineguinensis" subGenus="Monilearia">
<emphasis id="B953221CAC1BFFAAFB1042D1FAC86DCF" box="[1195,1303,1921,1943]" italics="true" pageId="1" pageNumber="297">Monilearia</emphasis>
(
<emphasis id="B953221CAC1BFFAAFA9342D1FA4B6DCF" box="[1320,1428,1921,1943]" italics="true" pageId="1" pageNumber="297">Monilearia</emphasis>
)
<emphasis id="B953221CAC1BFFAAFF2C42CFFEE46DEC" box="[151,315,1951,1972]" italics="true" pageId="1" pageNumber="297">arguineguinensis</emphasis>
(
<bibRefCitation id="EFB683FFAC1BFFAAFEF642CFFD846DEC" author="Seddon" box="[333,603,1951,1972]" pageId="1" pageNumber="297" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998">Seddon &amp; Aparicio, 1998</bibRefCitation>
)
</taxonomicName>
. Abbreviations: A1–A5, sections of generalized penial appendage of orthurethran gastropods and sections of vaginal appendage of Cochlicellini; ag, albumen gland; at, genital atrium; bc, bursa copulatrix; ep, epiphallus; f, flagellum; hd, hermaphroditic duct; p, penis; rm, retractor muscle; sov, spermoviduct; v, vagina; vd, vas deferens.
</paragraph>
</subSubSection>
<caption id="DF58AE86AC18FFA9FF2C4031FD3E6FB6" httpUri="https://zenodo.org/record/241985/files/figure.png" pageId="2" pageNumber="298" targetBox="[151,1436,191,1356]" targetPageId="2">
<paragraph id="8B98FE0EAC18FFA9FF2C4031FD3E6FB6" blockId="2.[151,1436,1377,1518]" pageId="2" pageNumber="298">
<emphasis id="B953221CAC18FFA9FF2C4031FED36F2F" bold="true" box="[151,268,1377,1399]" pageId="2" pageNumber="298">FIGURE 2.</emphasis>
Shells of
<taxonomicName id="4C27858DAC18FFA9FED74031FBCE6F2F" authority="Seddon &amp; Aparicio, 1998" authorityName="Seddon &amp; Aparicio" authorityYear="1998" box="[364,1041,1377,1399]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="298" phylum="Mollusca" rank="species" species="arguineguinensis" subGenus="Monilearia">
<emphasis id="B953221CAC18FFA9FED74031FE076F2F" box="[364,472,1377,1399]" italics="true" pageId="2" pageNumber="298">Monilearia</emphasis>
(
<emphasis id="B953221CAC18FFA9FE5E4031FD8E6F2F" box="[485,593,1377,1399]" italics="true" pageId="2" pageNumber="298">Monilearia</emphasis>
)
<emphasis id="B953221CAC18FFA9FDE54032FCDD6F2F" box="[606,770,1378,1399]" italics="true" pageId="2" pageNumber="298">arguineguinensis</emphasis>
(Seddon &amp; Aparicio, 1998)
</taxonomicName>
. A) Spain, Canary Islands, Gran Canaria, settlement E of Argüineguin, 27°45’38’’ N, 15°40’04’’ W, 50 m a.s.l., leg. E. Schlegel (ZMH 92264), B) Spain, Canary Islands, Gran Canaria, coll. Melvill-Tomlin, ex Wollaston (NMW 1955.158.02113), C) Spain, Canary Islands, Gran Canaria, Argüineguin, syntype coll. Mousson, ex Wollaston (ZMZ 505021). Photographs: A) F. Walther, B) Copyright Amgueddfa Cymru - National Museum Wales C) Copyright CSCF Neuchâtel, E. Neubert. Scale bar: 5 mm.
</paragraph>
</caption>
<subSubSection id="C33DAD85AC18FFA9FF7C434AFB046CE3" pageId="2" pageNumber="298" type="materials_examined">
<paragraph id="8B98FE0EAC18FFA9FF7C434AFB046CE3" blockId="2.[151,1437,1562,2001]" pageId="2" pageNumber="298">
<emphasis id="B953221CAC18FFA9FF7C434AFE786C69" bold="true" box="[199,423,1562,1585]" pageId="2" pageNumber="298">Material examined.</emphasis>
<collectingCountry id="F330BE9EAC18FFA9FE14434AFE316C69" box="[431,494,1562,1585]" name="Spain" pageId="2" pageNumber="298">Spain</collectingCountry>
, Canary Islands, Gran Canaria, settlement E of Argüineguin,
<geoCoordinate id="EE1398C9AC18FFA9FB30434AFAC36C69" box="[1163,1308,1562,1585]" direction="north" orientation="latitude" pageId="2" pageNumber="298" precision="15" value="27.760555">27°45’38’’ N</geoCoordinate>
,
<geoCoordinate id="EE1398C9AC18FFA9FA93434AFF6E6C0B" direction="west" orientation="longitude" pageId="2" pageNumber="298" precision="15" value="-15.667778">15°40’04’’ W</geoCoordinate>
,
<quantity id="4CDF53EBAC18FFA9FF00436CFF2D6C0B" box="[187,242,1596,1619]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="298" unit="m" value="50.0">50 m</quantity>
a.s.l.,
<date id="FF99D8CEAC18FFA9FE83436CFE086C0B" box="[312,471,1596,1619]" pageId="2" pageNumber="298" value="2015-02">February 2015</date>
, E. Schlegel leg. (1 shell, MN GEO-045 and 2 preserved specimens,
<collectionCode id="ED3666CBAC18FFA9FB07436CFB276C0B" box="[1212,1272,1596,1619]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="298">ZMH</collectionCode>
92264);
<collectingCountry id="F330BE9EAC18FFA9FAE3436CFA486C0B" box="[1368,1431,1596,1619]" name="Spain" pageId="2" pageNumber="298">Spain</collectingCountry>
, Canary Islands, Gran Canaria, coll. Melvill-Tomlin, ex Wollaston (1 shell,
<collectionCode id="ED3666CBAC18FFA9FC5C430FFBF46C2E" box="[999,1067,1631,1654]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="298">NMW</collectionCode>
1955.158.02113);
<collectingCountry id="F330BE9EAC18FFA9FB44430FFAE16C2E" box="[1279,1342,1631,1654]" name="Spain" pageId="2" pageNumber="298">Spain</collectingCountry>
, Canary Islands, Gran Canaria, Argüineguin,
<typeStatus id="549C40ACAC18FFA9FD9E43D2FDA76CC1" box="[549,632,1666,1689]" pageId="2" pageNumber="298" type="syntype">syntype</typeStatus>
coll. Mousson, ex Wollaston (4 shells,
<typeStatus id="549C40ACAC18FFA9FB9F43D2FB5E6CC1" box="[1060,1153,1666,1689]" pageId="2" pageNumber="298" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C27858DAC18FFA9FB1343D2FAB26CC1" box="[1192,1389,1666,1689]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="298" phylum="Mollusca" rank="species" species="argonautula">
<emphasis id="B953221CAC18FFA9FB1343D2FAB26CC1" box="[1192,1389,1666,1689]" italics="true" pageId="2" pageNumber="298">Helix argonautula</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC18FFA9FACF43D2FE636CE3" authority="Mousson, 1874" authorityName="Mousson" authorityYear="1874" pageId="2" pageNumber="298" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC18FFA9FF2C43F4FECD6CE3" box="[151,274,1700,1723]" italics="true" pageId="2" pageNumber="298">canariensis</emphasis>
<bibRefCitation id="EFB683FFAC18FFA9FEA343F4FE636CE3" author="Mousson" box="[280,444,1700,1723]" pageId="2" pageNumber="298" refString="Mousson, A. (1874) Bemerkungen uber die von Hrn. Dr. von Fritsch und Dr. Rein aus West-Marocco 1872 zuruckgebrachten Land- und Susswasser-Mollusken. (Schluss). Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 1, 81 - 107, pls. 4 - 6." type="journal article" year="1874">Mousson, 1874</bibRefCitation>
</taxonomicName>
and of ‘
<taxonomicName id="4C27858DAC18FFA9FDAD43F4FD8F6CE3" box="[534,592,1700,1723]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="298" phylum="Mollusca" rank="genus">
<emphasis id="B953221CAC18FFA9FDAD43F4FD8F6CE3" box="[534,592,1700,1723]" italics="true" pageId="2" pageNumber="298">Helix</emphasis>
</taxonomicName>
’
<taxonomicName id="4C27858DAC18FFA9FDE443F4FBEE6CE3" authority="Seddon &amp; Aparicio, 1998" authorityName="Seddon &amp; Aparicio" authorityYear="1998" box="[607,1073,1700,1723]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="298" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC18FFA9FDE443F4FCCA6CE3" box="[607,789,1700,1723]" italics="true" pageId="2" pageNumber="298">arguineguinensis</emphasis>
<bibRefCitation id="EFB683FFAC18FFA9FCA643F4FBEE6CE3" author="Seddon" box="[797,1073,1700,1723]" pageId="2" pageNumber="298" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998">Seddon &amp; Aparicio, 1998</bibRefCitation>
</taxonomicName>
, ZMZ 505021).
</paragraph>
</subSubSection>
<subSubSection id="C33DAD85AC18FFA8FF7C4397FE456B19" lastPageId="3" lastPageNumber="299" pageId="2" pageNumber="298" type="description">
<paragraph id="8B98FE0EAC18FFA9FF7C4397FA436D89" blockId="2.[151,1437,1562,2001]" pageId="2" pageNumber="298">
<emphasis id="B953221CAC18FFA9FF7C4397FE8D6C86" bold="true" box="[199,338,1735,1758]" pageId="2" pageNumber="298">Description.</emphasis>
A detailed description of the shell (
<figureCitation id="131CE28BAC18FFA9FD764397FCD36C86" box="[717,780,1735,1758]" captionStart="FIGURE 2" captionStartId="2.[151,244,1377,1399]" captionTargetBox="[151,1436,191,1356]" captionTargetId="figure@2.[151,1436,191,1356]" captionTargetPageId="2" captionText="FIGURE 2. Shells of Monilearia (Monilearia) arguineguinensis (Seddon &amp; Aparicio, 1998). A) Spain, Canary Islands, Gran Canaria, settlement E of Argüineguin, 27 ° 45 ’ 38 ’’ N, 15 ° 40 ’ 04 ’’ W, 50 m a. s. l., leg. E. Schlegel (ZMH 92264), B) Spain, Canary Islands, Gran Canaria, coll. Melvill-Tomlin, ex Wollaston (NMW 1955.158.02113), C) Spain, Canary Islands, Gran Canaria, Argüineguin, syntype coll. Mousson, ex Wollaston (ZMZ 505021). Photographs: A) F. Walther, B) Copyright Amgueddfa Cymru - National Museum Wales C) Copyright CSCF Neuchâtel, E. Neubert. Scale bar: 5 mm." httpUri="https://zenodo.org/record/241985/files/figure.png" pageId="2" pageNumber="298">Fig. 2</figureCitation>
A) is given in
<bibRefCitation id="EFB683FFAC18FFA9FC184397FB836C86" author="Wollaston" box="[931,1116,1735,1758]" pageId="2" pageNumber="298" refString="Wollaston, T. V. (1878) Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena. Reeve, London, XI + 1 + 588 + 16 pp." type="book" year="1878">Wollaston (1878)</bibRefCitation>
or
<bibRefCitation id="EFB683FFAC18FFA9FBC44397FA436C86" author="Seddon" box="[1151,1436,1735,1758]" pageId="2" pageNumber="298" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998">Seddon &amp; Aparicio (1998)</bibRefCitation>
and therefore is not repeated here. Genitalia (Fig. 1F): Right ommatophoran retractor independent of genitalia. Atrium short (
<quantity id="4CDF53EBAC18FFA9FF64425CFEE56D7B" box="[223,314,1804,1827]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="298" unit="mm" value="0.9">0.9 mm</quantity>
), about as long as vagina (
<quantity id="4CDF53EBAC18FFA9FDCB425CFD166D7B" box="[624,713,1804,1827]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="298" unit="mm" value="0.8">0.8 mm</quantity>
) and free oviduct (
<quantity id="4CDF53EBAC18FFA9FC1E425CFBDF6D7B" box="[933,1024,1804,1827]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="298" unit="mm" value="0.8">0.8 mm</quantity>
), the latter about as wide as vagina. Spermoviduct (4.0 mm) with a light touch of dark pigment, albumen gland withish, tongue-shaped (
<quantity id="4CDF53EBAC18FFA9FA8A427FFA506D1E" box="[1329,1423,1839,1862]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="298" unit="mm" value="2.8">2.8 mm</quantity>
), hermaphroditic duct light-brownish, slightly undulating. Bursa copulatrix complex adhering to spermoviduct, with a long duct (
<quantity id="4CDF53EBAC18FFA9FEB34225FE826DD4" box="[264,349,1909,1932]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="298" unit="mm" value="3.2">3.2 mm</quantity>
) and an oval bursa (
<quantity id="4CDF53EBAC18FFA9FD8F4225FD546DD4" box="[564,651,1909,1932]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="298" unit="mm" value="1.1">1.1 mm</quantity>
). Penis very short (
<quantity id="4CDF53EBAC18FFA9FCEE4225FC736DD4" box="[853,940,1909,1932]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="298" unit="mm" value="0.6">0.6 mm</quantity>
), inflated, internally with a plug-shaped penial papilla with apical pore; epiphallus (
<quantity id="4CDF53EBAC18FFA9FDA742C7FDAE6DF6" box="[540,625,1943,1966]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="298" unit="mm" value="6.2">6.2 mm</quantity>
) about 10 times as long as penis, retractor muscle inserting in its distal fifth (in relation to the position of the hermaphroditic gland). Flagellum (
<quantity id="4CDF53EBAC18FFA9FC3B42EAFC076D89" box="[896,984,1978,2001]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="298" unit="mm" value="2.1">2.1 mm</quantity>
), thin, about 1/3 the length of epiphallus.
</paragraph>
<paragraph id="8B98FE0EAC19FFA8FF2C45C8FE456B19" blockId="3.[151,1437,152,1015]" pageId="3" pageNumber="299">
Vaginal appendage well developed, with a hook-shaped, broad A1 section (
<quantity id="4CDF53EBAC19FFA8FC0745C8FBCB6AF7" box="[956,1044,152,175]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="299" unit="mm" value="2.9">2.9 mm</quantity>
); A2 section short (
<quantity id="4CDF53EBAC19FFA8FB5A45C8FAE66AF7" box="[1249,1337,152,175]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="299" unit="mm" value="0.4">0.4 mm</quantity>
), slightly bent and well muscularized; A4 section
<quantity id="4CDF53EBAC19FFA8FD8145EDFD536A8C" box="[570,652,189,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="299" unit="mm" value="2.8">2.8 mm</quantity>
long and thin, the branched A5 section (
<quantity id="4CDF53EBAC19FFA8FB8845EDFB586A8C" box="[1075,1159,189,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="299" unit="mm" value="2.8">2.8 mm</quantity>
) split into three thin ducts that are about as long as the A4 section. Internal wall of genital atrium equipped with several rugose, longitudinal pilasters that can be traced into the vagina and the vaginal appendage, in the latter two stronger and several weaker pilasters are discernible.
</paragraph>
</subSubSection>
<subSubSection id="C33DAD85AC19FFA8FF7C441DFBA169AF" pageId="3" pageNumber="299" type="discussion">
<paragraph id="8B98FE0EAC19FFA8FF7C441DFBA169AF" blockId="3.[151,1437,152,1015]" pageId="3" pageNumber="299">
<emphasis id="B953221CAC19FFA8FF7C441DFEEA6B3C" bold="true" box="[199,309,333,356]" pageId="3" pageNumber="299">Remarks.</emphasis>
The structure of the vaginal appendage of
<taxonomicName id="4C27858DAC19FFA8FCB0441DFC396B3C" box="[779,998,333,356]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC19FFA8FCB0441DFCFE6B3C" box="[779,801,333,356]" italics="true" pageId="3" pageNumber="299">M</emphasis>
.
<emphasis id="B953221CAC19FFA8FC8B441DFC396B3C" box="[816,998,333,356]" italics="true" pageId="3" pageNumber="299">arguineguinensis</emphasis>
</taxonomicName>
(Fig. 1) corresponds to that observed in other species assigned to the genus (
<bibRefCitation id="EFB683FFAC19FFA8FD924420FD6E6BDF" author="Hesse" box="[553,689,368,391]" pageId="3" pageNumber="299" refString="Hesse, P. (1911) Zur Kenntnis der Anatomie von Monilearia phalerata W. B. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft, 43, 161 - 165." type="journal article" year="1911">Hesse, 1911</bibRefCitation>
;
<bibRefCitation id="EFB683FFAC19FFA8FD7B4420FC886BDF" author="Odhner" box="[704,855,368,391]" pageId="3" pageNumber="299" refString="Odhner, N. H. (1932) Beitrage zur Malakozoologie der Kanarischen Inseln. Lamellibranchien, Cephalopoden, Gastropoden. Arkiv for Zoologi, 23 A, 1 - 116, pls. 1 - 2." type="journal article" year="1932">Odhner, 1932</bibRefCitation>
;
<bibRefCitation id="EFB683FFAC19FFA8FCDF4420FB6D6BDF" author="Schileyko" box="[868,1202,368,391]" pageId="3" pageNumber="299" refString="Schileyko, A. A. &amp; Menkhorst, H. P. M. G. (1997) Composition and phylogenetic relations of the Cochlicellidae (Gastropoda, Pulmonata). Ruthenica, 7, 51 - 60." type="journal article" year="1997">Schileyko &amp; Menkhorst, 1997</bibRefCitation>
;
<bibRefCitation id="EFB683FFAC19FFA8FB7B4420FA4E6BDF" author="Ibanez" box="[1216,1425,368,391]" pageId="3" pageNumber="299" refString="Ibanez, M., Groh, K., Alonso, M. R., Castillo, C. &amp; Yanes, Y. (2006) The subgenus Monilearia (Lyrula) Wollaston, 1878 (Gastropoda: Helicoidea: Cochlicellidae) from Lanzarote and Fuerteventura (Canary Islands), with the description of Monilearia (Lyrula) tubaeformis sp. nov. Zootaxa, 1320, 29 - 41." type="journal article" year="2006">Ibáñez et al., 2006</bibRefCitation>
).
<bibRefCitation id="EFB683FFAC19FFA8FF2C44C2FEAB6BF1" box="[151,372,402,425]" pageId="3" pageNumber="299" refString="Ibanez, M., Groh, K., Alonso, M. R., Castillo, C. &amp; Yanes, Y. (2006) The subgenus Monilearia (Lyrula) Wollaston, 1878 (Gastropoda: Helicoidea: Cochlicellidae) from Lanzarote and Fuerteventura (Canary Islands), with the description of Monilearia (Lyrula) tubaeformis sp. nov. Zootaxa, 1320, 29 - 41." type="journal article">Ibáñez et al. (2006)</bibRefCitation>
included three species in
<taxonomicName id="4C27858DAC19FFA8FD1944C2FBE46BF1" authority="Wollaston, 1878" authorityName="Wollaston" authorityYear="1878" box="[674,1083,402,425]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="subGenus" subGenus="Lyrula">
<emphasis id="B953221CAC19FFA8FD1944C2FCC56BF1" box="[674,794,402,425]" italics="true" pageId="3" pageNumber="299">Monilearia</emphasis>
(
<emphasis id="B953221CAC19FFA8FC9544C2FCAB6BF1" box="[814,884,402,425]" italics="true" pageId="3" pageNumber="299">Lyrula</emphasis>
)
<bibRefCitation id="EFB683FFAC19FFA8FC3244C2FBE46BF1" author="Wollaston" box="[905,1083,402,425]" pageId="3" pageNumber="299" refString="Wollaston, T. V. (1878) Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena. Reeve, London, XI + 1 + 588 + 16 pp." type="book" year="1878">Wollaston, 1878</bibRefCitation>
</taxonomicName>
on the basis of a similar shell sculpture consisting of a decussate ornamentation of slightly undulating radial riblets and numerous (sometimes stronger) spiral lines. The shell of
<taxonomicName id="4C27858DAC19FFA8FDB24488FD3C6BB7" box="[521,739,472,495]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC19FFA8FDB24488FDC06BB7" box="[521,543,472,495]" italics="true" pageId="3" pageNumber="299">M</emphasis>
.
<emphasis id="B953221CAC19FFA8FD964488FD3C6BB7" box="[557,739,472,495]" italics="true" pageId="3" pageNumber="299">arguineguinensis</emphasis>
</taxonomicName>
(
<figureCitation id="131CE28BAC19FFA8FD484488FCEC6BB7" box="[755,819,472,495]" captionStart="FIGURE 2" captionStartId="2.[151,244,1377,1399]" captionTargetBox="[151,1436,191,1356]" captionTargetId="figure@2.[151,1436,191,1356]" captionTargetPageId="2" captionText="FIGURE 2. Shells of Monilearia (Monilearia) arguineguinensis (Seddon &amp; Aparicio, 1998). A) Spain, Canary Islands, Gran Canaria, settlement E of Argüineguin, 27 ° 45 ’ 38 ’’ N, 15 ° 40 ’ 04 ’’ W, 50 m a. s. l., leg. E. Schlegel (ZMH 92264), B) Spain, Canary Islands, Gran Canaria, coll. Melvill-Tomlin, ex Wollaston (NMW 1955.158.02113), C) Spain, Canary Islands, Gran Canaria, Argüineguin, syntype coll. Mousson, ex Wollaston (ZMZ 505021). Photographs: A) F. Walther, B) Copyright Amgueddfa Cymru - National Museum Wales C) Copyright CSCF Neuchâtel, E. Neubert. Scale bar: 5 mm." httpUri="https://zenodo.org/record/241985/files/figure.png" pageId="3" pageNumber="299">Fig. 2</figureCitation>
) has a well-developed radial ornamentation consisting of numerous, irregular riblets, but spiral lines are absent. Therefore,
<taxonomicName id="4C27858DAC19FFA8FCC644AAFB826849" box="[893,1117,506,529]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC19FFA8FCC644AAFC4C6849" box="[893,915,506,529]" italics="true" pageId="3" pageNumber="299">M</emphasis>
.
<emphasis id="B953221CAC19FFA8FC1D44AAFB826849" box="[934,1117,506,529]" italics="true" pageId="3" pageNumber="299">arguineguinensis</emphasis>
</taxonomicName>
is included in the nominate subgenus here.
<bibRefCitation id="EFB683FFAC19FFA8FEF0474DFD11686C" author="Seddon" box="[331,718,541,564]" pageId="3" pageNumber="299" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998" yearSuffix="p">Seddon &amp; Aparicio (1998, p. 19)</bibRefCitation>
clearly intended the name ‘
<taxonomicName id="4C27858DAC19FFA8FBA7474DFB89686C" box="[1052,1110,541,564]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="genus">
<emphasis id="B953221CAC19FFA8FBA7474DFB89686C" box="[1052,1110,541,564]" italics="true" pageId="3" pageNumber="299">Helix</emphasis>
</taxonomicName>
’
<taxonomicName id="4C27858DAC19FFA8FBD6474DFAFC686C" box="[1133,1315,541,564]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC19FFA8FBD6474DFAFC686C" box="[1133,1315,541,564]" italics="true" pageId="3" pageNumber="299">arguineguinensis</emphasis>
</taxonomicName>
as a new replacement name for the variety from the Canary Islands introduced by
<bibRefCitation id="EFB683FFAC19FFA8FC1B4710FB8D680F" author="Mousson" box="[928,1106,576,599]" pageId="3" pageNumber="299" refString="Mousson, A. (1874) Bemerkungen uber die von Hrn. Dr. von Fritsch und Dr. Rein aus West-Marocco 1872 zuruckgebrachten Land- und Susswasser-Mollusken. (Schluss). Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 1, 81 - 107, pls. 4 - 6." type="journal article" year="1874">Mousson (1874)</bibRefCitation>
, i.e. for
<taxonomicName id="4C27858DAC19FFA8FB104710FAB1680F" box="[1195,1390,576,599]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="argonautula">
<emphasis id="B953221CAC19FFA8FB104710FAB1680F" box="[1195,1390,576,599]" italics="true" pageId="3" pageNumber="299">Helix argonautula</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC19FFA8FACE4710FECD6821" pageId="3" pageNumber="299" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC19FFA8FF2C4732FECD6821" box="[151,274,610,633]" italics="true" pageId="3" pageNumber="299">canariensis</emphasis>
</taxonomicName>
(not
<taxonomicName id="4C27858DAC19FFA8FEF14732FD6F6821" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[330,688,610,633]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="canariensis">
<emphasis id="B953221CAC19FFA8FEF14732FDDB6821" box="[330,516,610,633]" italics="true" pageId="3" pageNumber="299">Helix canariensis</emphasis>
<bibRefCitation id="EFB683FFAC19FFA8FDB74732FD6F6821" author="Mousson" box="[524,688,610,633]" pageId="3" pageNumber="299" refString="Mousson, A. (1872) Revision de la faune malacologique des iles Canaries. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, 25, 1 - 176, pls. I - VI." type="journal article" year="1872">Mousson, 1872</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="4C27858DAC19FFA8FD684732FC8A6821" box="[723,853,610,633]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="lactea">
<emphasis id="B953221CAC19FFA8FD684732FC8A6821" box="[723,853,610,633]" italics="true" pageId="3" pageNumber="299">Helix lactea</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC19FFA8FCE74732FB6C6821" authority="Mousson, 1872" authorityName="Mousson" authorityYear="1872" box="[860,1203,610,633]" pageId="3" pageNumber="299" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC19FFA8FC324732FBDB6821" box="[905,1028,610,633]" italics="true" pageId="3" pageNumber="299">canariensis</emphasis>
<bibRefCitation id="EFB683FFAC19FFA8FBB04732FB6C6821" author="Mousson" box="[1035,1203,610,633]" pageId="3" pageNumber="299" refString="Mousson, A. (1872) Revision de la faune malacologique des iles Canaries. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, 25, 1 - 176, pls. I - VI." type="journal article" year="1872">Mousson, 1872</bibRefCitation>
</taxonomicName>
). Although
<bibRefCitation id="EFB683FFAC19FFA8FA8A4732FE7968C4" author="Seddon" pageId="3" pageNumber="299" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998" yearSuffix="p">Seddon &amp; Aparicion (1998, p. 21)</bibRefCitation>
attributed the name ‘
<taxonomicName id="4C27858DAC19FFA8FD1D47D5FCFE68C4" box="[678,801,645,668]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="canariensis">
<emphasis id="B953221CAC19FFA8FD1D47D5FCFE68C4" box="[678,801,645,668]" italics="true" pageId="3" pageNumber="299">canariensis</emphasis>
</taxonomicName>
’ erroneously to
<bibRefCitation id="EFB683FFAC19FFA8FC5D47D5FB7768C4" author="Wollaston" box="[998,1192,645,668]" pageId="3" pageNumber="299" refString="Wollaston, T. V. (1878) Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena. Reeve, London, XI + 1 + 588 + 16 pp." type="book" year="1878">Wollaston (1878)</bibRefCitation>
, the reference to the combination used by
<bibRefCitation id="EFB683FFAC19FFA8FEC147F8FDEA68E7" author="Wollaston" box="[378,565,680,703]" pageId="3" pageNumber="299" refString="Wollaston, T. V. (1878) Testacea Atlantica or the land and freshwater shells of the Azores, Madeiras, Salvages, Canaries, Cape Verdes, and Saint Helena. Reeve, London, XI + 1 + 588 + 16 pp." type="book" year="1878">Wollaston (1878)</bibRefCitation>
,
<taxonomicName id="4C27858DAC19FFA8FDFB47F8FCF968E7" box="[576,806,680,703]" class="Gastropoda" family="Architectonicidae" genus="Disculus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="argonautula">
<emphasis id="B953221CAC19FFA8FDFB47F8FCF968E7" box="[576,806,680,703]" italics="true" pageId="3" pageNumber="299">Disculus argonautula</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC19FFA8FC9747F8FB4D68E7" authority="Mousson, 1874" authorityName="Mousson" authorityYear="1874" box="[812,1170,680,703]" pageId="3" pageNumber="299" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC19FFA8FCE247F8FC0B68E7" box="[857,980,680,703]" italics="true" pageId="3" pageNumber="299">canariensis</emphasis>
(
<bibRefCitation id="EFB683FFAC19FFA8FC5947F8FB5468E7" author="Mousson" box="[994,1163,680,703]" pageId="3" pageNumber="299" refString="Mousson, A. (1874) Bemerkungen uber die von Hrn. Dr. von Fritsch und Dr. Rein aus West-Marocco 1872 zuruckgebrachten Land- und Susswasser-Mollusken. (Schluss). Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 1, 81 - 107, pls. 4 - 6." type="journal article" year="1874">Mousson, 1874</bibRefCitation>
)
</taxonomicName>
, in the synonymy of ‘
<emphasis id="B953221CAC19FFA8FAC247F8FA4B68E7" box="[1401,1428,680,703]" italics="true" pageId="3" pageNumber="299">H.</emphasis>
’
<taxonomicName id="4C27858DAC19FFA8FF2C479AFE9168B9" box="[151,334,714,737]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC19FFA8FF2C479AFE9168B9" box="[151,334,714,737]" italics="true" pageId="3" pageNumber="299">arguineguinensis</emphasis>
</taxonomicName>
makes it clear, which name the authors replaced by the new name. According to the ICZN code Art. 72.7 both nominal taxa have the same name-bearing
<typeStatus id="549C40ACAC19FFA8FD6C47BDFCDA695C" box="[727,773,749,772]" pageId="3" pageNumber="299">type</typeStatus>
despite any contrary designation.
<bibRefCitation id="EFB683FFAC19FFA8FBC147BDFA43695C" author="Seddon" box="[1146,1436,749,772]" pageId="3" pageNumber="299" refString="Seddon, M. B. &amp; Aparicio, M. - T. (1998) Problematic taxa from Morocco - the status of &quot; Helix argonautula &quot; Webb &amp; Berthelot, 1833, &quot; Helix renati &quot; Dautzenberg, 1894 and Xeroleuca antoinei Pallary, 1936. Journal of Conchology, 36 (2), 19 - 29. http: // dx. doi. org / 10.1093 / mollus / eym 030" type="journal article" year="1998">Seddon &amp; Aparicio (1998)</bibRefCitation>
designated a shell from the Mellvill-Tomlin collection (NMW 1955.158.0 2113, see also
<figureCitation id="131CE28BAC19FFA8FBF74640FB54697F" box="[1100,1163,784,807]" captionStart="FIGURE 2" captionStartId="2.[151,244,1377,1399]" captionTargetBox="[151,1436,191,1356]" captionTargetId="figure@2.[151,1436,191,1356]" captionTargetPageId="2" captionText="FIGURE 2. Shells of Monilearia (Monilearia) arguineguinensis (Seddon &amp; Aparicio, 1998). A) Spain, Canary Islands, Gran Canaria, settlement E of Argüineguin, 27 ° 45 ’ 38 ’’ N, 15 ° 40 ’ 04 ’’ W, 50 m a. s. l., leg. E. Schlegel (ZMH 92264), B) Spain, Canary Islands, Gran Canaria, coll. Melvill-Tomlin, ex Wollaston (NMW 1955.158.02113), C) Spain, Canary Islands, Gran Canaria, Argüineguin, syntype coll. Mousson, ex Wollaston (ZMZ 505021). Photographs: A) F. Walther, B) Copyright Amgueddfa Cymru - National Museum Wales C) Copyright CSCF Neuchâtel, E. Neubert. Scale bar: 5 mm." httpUri="https://zenodo.org/record/241985/files/figure.png" pageId="3" pageNumber="299">Fig. 2</figureCitation>
B) as
<typeStatus id="549C40ACAC19FFA8FB724640FAF4697F" box="[1225,1323,784,807]" pageId="3" pageNumber="299" type="lectotype">lectotype</typeStatus>
of ‘
<taxonomicName id="4C27858DAC19FFA8FAE24640FA4C697F" box="[1369,1427,784,807]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="genus">
<emphasis id="B953221CAC19FFA8FAE24640FA4C697F" box="[1369,1427,784,807]" italics="true" pageId="3" pageNumber="299">Helix</emphasis>
</taxonomicName>
’
<taxonomicName id="4C27858DAC19FFA8FF2C4662FE916911" box="[151,334,818,841]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC19FFA8FF2C4662FE916911" box="[151,334,818,841]" italics="true" pageId="3" pageNumber="299">arguineguinensis</emphasis>
</taxonomicName>
. However, there are four shells labelled as
<taxonomicName id="4C27858DAC19FFA8FCAC4662FC656911" box="[791,954,818,841]" class="Gastropoda" family="Architectonicidae" genus="Disculus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="argonautula">
<emphasis id="B953221CAC19FFA8FCAC4662FC656911" box="[791,954,818,841]" italics="true" pageId="3" pageNumber="299">H. argonautula</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC19FFA8FC7A4662FBB66911" box="[961,1129,818,841]" pageId="3" pageNumber="299" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC19FFA8FC554662FBB66911" box="[1006,1129,818,841]" italics="true" pageId="3" pageNumber="299">canariensis</emphasis>
</taxonomicName>
in the Mousson collection at Zurich (ZMZ 505021, see also
<figureCitation id="131CE28BAC19FFA8FE5A4605FDC06934" box="[481,543,853,876]" captionStart="FIGURE 2" captionStartId="2.[151,244,1377,1399]" captionTargetBox="[151,1436,191,1356]" captionTargetId="figure@2.[151,1436,191,1356]" captionTargetPageId="2" captionText="FIGURE 2. Shells of Monilearia (Monilearia) arguineguinensis (Seddon &amp; Aparicio, 1998). A) Spain, Canary Islands, Gran Canaria, settlement E of Argüineguin, 27 ° 45 ’ 38 ’’ N, 15 ° 40 ’ 04 ’’ W, 50 m a. s. l., leg. E. Schlegel (ZMH 92264), B) Spain, Canary Islands, Gran Canaria, coll. Melvill-Tomlin, ex Wollaston (NMW 1955.158.02113), C) Spain, Canary Islands, Gran Canaria, Argüineguin, syntype coll. Mousson, ex Wollaston (ZMZ 505021). Photographs: A) F. Walther, B) Copyright Amgueddfa Cymru - National Museum Wales C) Copyright CSCF Neuchâtel, E. Neubert. Scale bar: 5 mm." httpUri="https://zenodo.org/record/241985/files/figure.png" pageId="3" pageNumber="299">Fig. 2</figureCitation>
C) which are
<typeStatus id="549C40ACAC19FFA8FD164605FCD56934" box="[685,778,853,876]" pageId="3" pageNumber="299" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C27858DAC19FFA8FC964605FC1C6934" box="[813,963,853,876]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="argonatula">
<emphasis id="B953221CAC19FFA8FC964605FC9F6934" box="[813,832,853,876]" italics="true" pageId="3" pageNumber="299">H</emphasis>
.
<emphasis id="B953221CAC19FFA8FCF64605FC1C6934" box="[845,963,853,876]" italics="true" pageId="3" pageNumber="299">argonatula</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC19FFA8FC734605FBAE6934" box="[968,1137,853,876]" pageId="3" pageNumber="299" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC19FFA8FC4D4605FBAE6934" box="[1014,1137,853,876]" italics="true" pageId="3" pageNumber="299">canariensis</emphasis>
</taxonomicName>
and according to ICZN Art. 72.7 also of ‘
<emphasis id="B953221CAC19FFA8FE884628FE9969D7" box="[307,326,888,911]" italics="true" pageId="3" pageNumber="299">H</emphasis>
.’
<taxonomicName id="4C27858DAC19FFA8FED94628FDC769D7" box="[354,536,888,911]" class="Gastropoda" family="Cochlicellidae" genus="Monilearia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="arguineguinensis">
<emphasis id="B953221CAC19FFA8FED94628FDC769D7" box="[354,536,888,911]" italics="true" pageId="3" pageNumber="299">arguineguinensis</emphasis>
</taxonomicName>
. There is no evidence that the specimen from the collection in Cardiff (NMW 1955.158.02113) was known to Mousson when introducing the name
<taxonomicName id="4C27858DAC19FFA8FC2046CAFBEC69E9" box="[923,1075,922,945]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="299" phylum="Mollusca" rank="species" species="argonatula">
<emphasis id="B953221CAC19FFA8FC2046CAFBEC69E9" box="[923,1075,922,945]" italics="true" pageId="3" pageNumber="299">H. argonatula</emphasis>
</taxonomicName>
<taxonomicName id="4C27858DAC19FFA8FB8646CAFB3669E9" box="[1085,1257,922,945]" pageId="3" pageNumber="299" rank="variety" variety="canariensis">
var.
<emphasis id="B953221CAC19FFA8FBD546CAFB3669E9" box="[1134,1257,922,945]" italics="true" pageId="3" pageNumber="299">canariensis</emphasis>
</taxonomicName>
and therefore it cannot be regarded as belonging to the
<typeStatus id="549C40ACAC19FFA8FDF946EDFD40698C" box="[578,671,957,980]" pageId="3" pageNumber="299" type="syntype">syntypes</typeStatus>
series. Hence, I regard its selection as
<typeStatus id="549C40ACAC19FFA8FBF346EDFB75698C" box="[1096,1194,957,980]" pageId="3" pageNumber="299" type="lectotype">lectotype</typeStatus>
as invalid (ICZN Art. 74.2) and only the shells in the Mousson collection at Zurich (ZMZ 505021) have
<typeStatus id="549C40ACAC19FFA8FBBE46B0FBEC69AF" box="[1029,1075,992,1015]" pageId="3" pageNumber="299">type</typeStatus>
status.
</paragraph>
</subSubSection>
</treatment>
</document>
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