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<document id="E491EB6400D879629F0B9CC4EA87B2AB" ID-CLB-Dataset="7433" ID-DOI="10.5852/ejt.2021.744.1303" ID-GBIF-Dataset="f62caa9b-9794-41d0-8551-a6ccb1d2e092" ID-ISSN="2118-9773" ID-Zenodo-Dep="4673914" ID-ZooBank="BBB5D83C-54BE-43AB-B4B8-8B896B5909BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617944647591" checkinUser="plazi" docAuthor="Chatzimanolis, Stylianos &amp; Brunke, Adam J." docDate="2021" docId="752487D7FF8E0678FDDFFEB3FDC036D1" docLanguage="en" docName="ejt-744_67-82.pdf" docOrigin="European Journal of Taxonomy 744" docSource="http://dx.doi.org/10.5852/ejt.2021.744.1303" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ikaros apteros Chatzimanolis &amp; Brunke 2021, gen. et sp. nov." docType="treatment" docUuid="2FB02118-DAD8-4812-AFFE-2ADA3356F97B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="75" masterDocId="891DFFAFFF890670FFB8FFBBFFDC3525" masterDocTitle="A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position" masterLastPageNumber="82" masterPageNumber="67" pageNumber="74" updateTime="1698936220334" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ECCA44B2E237548F0DDFA415E6B80440">A new apterous rove beetle genus (Coleoptera: Staphylinidae) from the Northern Andes with an assessment of its phylogenetic position</mods:title>
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<mods:namePart id="7C008BA7E7609E7BFD7528C203B3A5B4">Chatzimanolis, Stylianos</mods:namePart>
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<mods:affiliation id="75DF4DE4EA57775B6F2F2B0822ED7A45">Department of Biology, Geology and Environmental Science, University of Tennessee at Chattanooga, 615 McCallie Ave. Dept 2653, Chattanooga, TN 37403, USA.</mods:affiliation>
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<mods:namePart id="B038B6507BB529CCCDD062E00811F2CF">Brunke, Adam J.</mods:namePart>
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<mods:affiliation id="D1014EB6E2D77A66D673171192F2140A">Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Avenue, K. W. Neatby Building, Ottawa, ON K 1 A 0 C 6, Canada.</mods:affiliation>
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<mods:date id="9F6DF154ADE45084924FF012D1E83630">2021</mods:date>
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<subSubSection id="B597654AFF8E0677FDDFFEB3FC103406" box="[615,972,264,291]" pageId="7" pageNumber="74" type="nomenclature">
<paragraph id="FD3236C1FF8E0677FDDFFEB3FC103406" blockId="7.[361,1227,264,361]" box="[615,972,264,291]" pageId="7" pageNumber="74">
<heading id="A67A81ADFF8E0677FDDFFEB3FC103406" box="[615,972,264,291]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="74" reason="2">
<taxonomicName id="3A8D4D42FF8E0677FDDFFEB3FCC93406" ID-CoL="8Y4VP" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[615,789,264,291]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="apteros" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF8E0677FDDFFEB3FCC93406" bold="true" box="[615,789,264,291]" italics="true" pageId="7" pageNumber="74">Ikaros apteros</emphasis>
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<taxonomicNameLabel id="D4CA57A8FF8E0677FCA4FEB2FC103406" box="[796,972,265,291]" pageId="7" pageNumber="74" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<figureCitation id="65B62A44FF8E0677FD64FEF4FCE0344C" box="[732,828,335,361]" captionStart="Fig" captionStartId="6.[189,232,1913,1939]" captionTargetBox="[201,1380,691,1837]" captionTargetId="figure-114@6.[191,1399,688,1877]" captionTargetPageId="6" captionText="Fig. 2. Habitus photographs of species of Ikaros gen. nov. A. I. apteros gen. et sp. nov. Holotype, ♂ (SDEI). B. I. paramo gen. et sp. nov. Paratype, ♂ (barcode: SM05500251, UTCI). C. I. polygonos gen. et sp. nov. Holotype, ♂ (barcode: SM0114364, MIZA). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4673920" httpUri="https://zenodo.org/record/4673920/files/figure.png" pageId="7" pageNumber="74">Figs 2A</figureCitation>
,
<figureCitation id="65B62A44FF8E0677FCF1FEF4FC84344C" box="[841,856,335,361]" captionStart="Fig" captionStartId="7.[189,232,1953,1979]" captionTargetBox="[189,1385,1175,1897]" captionTargetId="figure-174@7.[189,1398,1175,1910]" captionTargetPageId="7" captionText="Fig. 3.Aedeagus ofIkaros apteros gen. et sp. nov.A. Dorsal view.B. Lateral view.C. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673924" httpUri="https://zenodo.org/record/4673924/files/figure.png" pageId="7" pageNumber="74">3</figureCitation>
</paragraph>
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<paragraph id="FD3236C1FF8E0677FF05FE2BFEEA348F" blockId="7.[189,310,400,426]" box="[189,310,400,426]" pageId="7" pageNumber="74">
<heading id="A67A81ADFF8E0677FF05FE2BFEEA348F" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="7" pageNumber="74" reason="3">
<emphasis id="CFF9EAD3FF8E0677FF05FE2BFEEA348F" bold="true" box="[189,310,400,426]" pageId="7" pageNumber="74">Diagnosis</emphasis>
</heading>
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<paragraph id="FD3236C1FF8E0677FF05FE04FD203705" blockId="7.[188,1398,447,544]" pageId="7" pageNumber="74">
Among species of
<taxonomicName id="3A8D4D42FF8E0677FE2BFE04FE0134FC" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[403,477,447,473]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CFF9EAD3FF8E0677FE2BFE04FE0134FC" box="[403,477,447,473]" italics="true" pageId="7" pageNumber="74">Ikaros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF8E0677FE5CFE04FD9234FC" box="[484,590,447,473]" pageId="7" pageNumber="74" rank="genus">gen. nov.</taxonomicNameLabel>
that lack arch-like carina on terga 35,
<taxonomicName id="3A8D4D42FF8E0677FBABFE7BFB5D34FC" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[1043,1153,448,473]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="apteros" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF8E0677FBABFE7BFB5D34FC" box="[1043,1153,448,473]" italics="true" pageId="7" pageNumber="74">I. apteros</emphasis>
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<taxonomicNameLabel id="D4CA57A8FF8E0677FB30FE04FAE934FC" box="[1160,1333,447,473]" pageId="7" pageNumber="74" rank="species">gen. et sp. nov.</taxonomicNameLabel>
lacks any punctures on the pronotum, beyond the punctures along the median area, whereas
<taxonomicName id="3A8D4D42FF8E0677FB16FE58FAFD34D9" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[1198,1313,483,508]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="paramo" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF8E0677FB16FE58FAFD34D9" box="[1198,1313,483,508]" italics="true" pageId="7" pageNumber="74">I. paramo</emphasis>
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<taxonomicNameLabel id="D4CA57A8FF8E0677FA91FE58FECB3705" pageId="7" pageNumber="74" rank="species">gen. et sp. nov.</taxonomicNameLabel>
has punctures arranged in multiple rows.
</paragraph>
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<heading id="A67A81ADFF8E0677FF05FDFCFE983744" bold="true" box="[189,324,583,609]" fontSize="11" level="3" pageId="7" pageNumber="74" reason="3">
<emphasis id="CFF9EAD3FF8E0677FF05FDFCFE983744" bold="true" box="[189,324,583,609]" pageId="7" pageNumber="74">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF8E0677FF05FDCEFCF43797" blockId="7.[188,1398,629,691]" pageId="7" pageNumber="74">The specific epithet is derived from the Greek word ‘άπτερος’ (without wings) and refers to the lack of hind wings. It is treated here as a noun in apposition.</paragraph>
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<emphasis id="CFF9EAD3FF8E0677FF05FD61FEB037D1" bold="true" box="[189,364,730,756]" pageId="7" pageNumber="74">Type material</emphasis>
</heading>
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(here designated) COUNTRY UNKNOWN •
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; “Nov. Gran. [
<collectingCountry id="859A7651FF8E0677FD6BFC97FC543663" box="[723,904,811,838]" name="Colombia" pageId="7" pageNumber="74">Nova Granada</collectingCountry>
], L.F.S / coll.
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/
<taxonomicName id="3A8D4D42FF8E0677FB27FC97FA993663" authorityName="Sharp" authorityYear="1884" box="[1183,1349,812,838]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
ref.
<taxonomicName id="3A8D4D42FF8E0677FF05FCF4FE8A364C" authorityName="Kraatz" authorityYear="1857" box="[189,342,847,873]" class="Insecta" family="Staphylinidae" genus="Xanthopygus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="CFF9EAD3FF8E0677FF05FCF4FE8A364C" box="[189,342,847,873]" italics="true" pageId="7" pageNumber="74">Xanthopygus</emphasis>
</taxonomicName>
det. A. Brunke 2012 / HOLOTYPE
<taxonomicName id="3A8D4D42FF8E0677FC89FCF4FAEA364C" authority="Chatzimanolis and Brunke" authorityName="Chatzimanolis and Brunke" box="[817,1334,847,873]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="apteros">
<emphasis id="CFF9EAD3FF8E0677FC89FCF4FC03364C" box="[817,991,847,873]" italics="true" pageId="7" pageNumber="74">Ikaros apteros</emphasis>
Chatzimanolis and Brunke
</taxonomicName>
, des. Chatzimanolis and Brunke 2020”;
<collectionCode id="9B9CAE04FF8E0677FDECFCC9FD4536A9" box="[596,665,882,908]" pageId="7" pageNumber="74">SDEI</collectionCode>
.
</materialsCitation>
</paragraph>
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<materialsCitation id="4DE53C9CFF8E0677FF52FC0FFB583112" ID-GBIF-Occurrence="3069587306" collectingDate="2002-06-19" collectingDateMax="2002-06-21" collectingDateMin="2002-06-19" collectionCode="SEMC" collectorName="F. Guzman Leg." country="Colombia" elevation="2990" latitude="4.5166664" location="Chingaza Charrascalles" longLatPrecision="1306" longitude="-73.75" municipality="Colombia" pageId="7" pageNumber="74" specimenCount="1" specimenCount-female="1" stateProvince="Cundinamarca" typeStatus="paratype">
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<typeStatus id="22368863FF8E0677FF52FC0FFE8036EB" box="[234,348,948,974]" pageId="7" pageNumber="74" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="859A7651FF8E0677FF05FC6CFE8436D4" box="[189,344,983,1009]" name="Colombia" pageId="7" pageNumber="74">COLOMBIA</collectingCountry>
<specimenCount id="EB8BFD48FF8E0677FED7FC6CFE5A36D4" box="[367,390,983,1009]" pageId="7" pageNumber="74" type="female"></specimenCount>
; “
<collectingMunicipality id="1D56ACBBFF8E0677FE26FC6CFDC436D4" box="[414,536,983,1009]" pageId="7" pageNumber="74">Colombia</collectingMunicipality>
,
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, PNN
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,
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</quantity>
,
<location id="F852601AFF8E0677FF05FC41FEC23131" LSID="urn:lsid:plazi:treatment:752487D7FF8E0678FDDFFEB3FDC036D1:F852601AFF8E0677FF05FC41FEC23131" box="[189,286,1018,1044]" country="Colombia" latitude="4.5166664" longLatPrecision="1306" longitude="-73.75" municipality="Colombia" name="Winkler" pageId="7" pageNumber="74" stateProvince="Cundinamarca">Winkler</location>
,
<date id="89331001FF8E0677FE94FC41FE043131" box="[300,472,1018,1044]" pageId="7" pageNumber="74" value="2002-06-19" valueMax="2002-06-21" valueMin="2002-06-19">
<collectingDate id="9977E9E9FF8E0677FE94FC41FE043131" box="[300,472,1018,1044]" pageId="7" pageNumber="74" value="2002-06-19" valueMax="2002-06-21" valueMin="2002-06-19">1921.vi.2002</collectingDate>
</date>
,
<collectorName id="50785317FF8E0677FE5EFC41FD763131" box="[486,682,1018,1044]" pageId="7" pageNumber="74">F. Guzmán Leg.</collectorName>
, M3239 / SM0548722 [barcode label] / PARATYPE
<taxonomicName id="3A8D4D42FF8E0677FA97FC41FD8B3112" authority="Chatzimanolis and Brunke" authorityName="Chatzimanolis and Brunke" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="apteros">
<emphasis id="CFF9EAD3FF8E0677FA97FC41FEC83112" italics="true" pageId="7" pageNumber="74">Ikaros apteros</emphasis>
Chatzimanolis and Brunke
</taxonomicName>
, des. Chatzimanolis and Brunke 2020”;
<collectionCode id="9B9CAE04FF8E0677FB88FBA6FB583112" box="[1072,1156,1053,1079]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="7" pageNumber="74" type="Museum">SEMC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="A9F26649FF8E0677FF05F81AFE8F32FB" ID-DOI="http://doi.org/10.5281/zenodo.4673924" ID-Zenodo-Dep="4673924" httpUri="https://zenodo.org/record/4673924/files/figure.png" pageId="7" pageNumber="74" startId="7.[189,232,1953,1979]" targetBox="[189,1385,1175,1897]" targetPageId="7">
<paragraph id="FD3236C1FF8E0677FF05F81AFE8F32FB" blockId="7.[187,1398,1953,2014]" pageId="7" pageNumber="74">
<emphasis id="CFF9EAD3FF8E0677FF05F81AFED5329E" bold="true" box="[189,265,1953,1979]" pageId="7" pageNumber="74">Fig. 3.</emphasis>
Aedeagus of
<taxonomicName id="3A8D4D42FF8E0677FE18F81AFD9A329E" authority="Chatzimanolis &amp; Brunke, 2021" authorityName="Chatzimanolis &amp; Brunke" authorityYear="2021" box="[416,582,1953,1979]" class="Insecta" family="Staphylinidae" genus="Ikaros" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="74" phylum="Arthropoda" rank="species" species="apteros" status="gen. et sp. nov.">
<emphasis id="CFF9EAD3FF8E0677FE18F81AFD9A329E" box="[416,582,1953,1979]" italics="true" pageId="7" pageNumber="74">Ikaros apteros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D4CA57A8FF8E0677FDF7F81AFD26329E" box="[591,762,1953,1979]" pageId="7" pageNumber="74" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="CFF9EAD3FF8E0677FD45F81AFCCE329E" bold="true" box="[765,786,1953,1979]" pageId="7" pageNumber="74">A</emphasis>
. Dorsal view.
<emphasis id="CFF9EAD3FF8E0677FC0EF81AFC16329E" bold="true" box="[950,970,1953,1979]" pageId="7" pageNumber="74">B</emphasis>
. Lateral view.
<emphasis id="CFF9EAD3FF8E0677FBCBF81AFB54329E" bold="true" box="[1139,1160,1953,1979]" pageId="7" pageNumber="74">C</emphasis>
. Detail of paramere, ventral view.
</paragraph>
</caption>
<subSubSection id="B597654AFF810678FF05FEB3FB04363E" pageId="8" pageNumber="75" type="description">
<paragraph id="FD3236C1FF810678FF05FEB3FE923407" blockId="8.[189,334,264,290]" box="[189,334,264,290]" pageId="8" pageNumber="75">
<heading id="A67A81ADFF810678FF05FEB3FE923407" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="8" pageNumber="75" reason="3">
<emphasis id="CFF9EAD3FF810678FF05FEB3FE923407" bold="true" box="[189,334,264,290]" pageId="8" pageNumber="75">Description</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF810678FF05FE8CFB04363E" blockId="8.[187,1400,311,795]" pageId="8" pageNumber="75">
Forebody length
<quantity id="3A759B24FF810678FE36FE8CFE303477" box="[398,492,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="8" pageNumber="75" unit="mm" value="7.1">7.1 mm</quantity>
long. Coloration reddish brown with head and mouthparts slightly darker brown. Head transverse, HW/HL ratio = 1.18. Epicranium mostly impunctate, with few large punctures posteriorly, medially and around margin of head and eye; with faint polygon-shaped microsculpture. Labial palpus with palpomere 3 widest apically, subparallel-sided. Antennomeres with crown-like macrosetae shorter than length of antennomere. Pronotum longer than wide, PW/PL ratio = 0.91; surface of pronotum impunctate except two punctures on each side of median area and few punctures around margin; with faint polygon-shaped microsculpture. Elytra shorter than pronotum, EL/PL ratio = 0.75. Elytra with large, deep contiguous punctures and dense polygon-shaped microsculpture.Abdominal terga 35 without arch-like carina. Male secondary sexual structures with shallow but broad emargination on sternum 7; with deep, broad V-shaped emargination on sternum 8; borders of emargination on sternum 7 and 8 appearing shaved (with no setae). Aedeagus as in
<figureCitation id="65B62A44FF810678FCD7FD2CFC6B3794" box="[879,951,663,689]" captionStart="Fig" captionStartId="7.[189,232,1953,1979]" captionTargetBox="[189,1385,1175,1897]" captionTargetId="figure-174@7.[189,1398,1175,1910]" captionTargetPageId="7" captionText="Fig. 3.Aedeagus ofIkaros apteros gen. et sp. nov.A. Dorsal view.B. Lateral view.C. Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4673924" httpUri="https://zenodo.org/record/4673924/files/figure.png" pageId="8" pageNumber="75">Fig. 3</figureCitation>
; in dorsal view paramere longer than median lobe; converging to elongate, narrow tip; apex of paramere with small emargination; in lateral view paramere concave, becoming narrower. Median lobe in dorsal view narrowing to rounded apex; in lateral view median lobe becoming narrower near flattened apex, with no subapical tooth.
</paragraph>
</subSubSection>
<subSubSection id="B597654AFF810678FF05FCF9FDC036D1" pageId="8" pageNumber="75" type="distribution">
<paragraph id="FD3236C1FF810678FF05FCF9FE8A3679" blockId="8.[189,342,834,860]" box="[189,342,834,860]" pageId="8" pageNumber="75">
<heading id="A67A81ADFF810678FF05FCF9FE8A3679" bold="true" box="[189,342,834,860]" fontSize="11" level="3" pageId="8" pageNumber="75" reason="3">
<emphasis id="CFF9EAD3FF810678FF05FCF9FE8A3679" bold="true" box="[189,342,834,860]" pageId="8" pageNumber="75">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="FD3236C1FF810678FF05FCCAFDC036D1" blockId="8.[189,1400,880,1013]" pageId="8" pageNumber="75">
Known from the National Park Chingaza in the department of
<collectingRegion id="3F49F823FF810678FC10FCCAFB8A36AE" box="[936,1110,881,907]" country="Colombia" name="Cundinamarca" pageId="8" pageNumber="75">Cundinamarca</collectingRegion>
,
<collectingCountry id="859A7651FF810678FBDBFCCAFB0B36AE" box="[1123,1239,881,907]" name="Colombia" pageId="8" pageNumber="75">Colombia</collectingCountry>
(
<figureCitation id="65B62A44FF810678FB51FCCAFAEE36AE" box="[1257,1330,880,907]" captionStart="Fig" captionStartId="12.[191,234,1945,1971]" captionTargetBox="[189,1398,680,1912]" captionTargetId="figure-145@12.[189,1398,677,1912]" captionTargetPageId="12" captionText="Fig. 6. Distribution map of Ikaros apteros gen. et sp. nov. (square), I. paramo gen. et sp. nov. (circle) and I. polygonos gen. et sp. nov. (triangle)." figureDoi="http://doi.org/10.5281/zenodo.4673930" httpUri="https://zenodo.org/record/4673930/files/figure.png" pageId="8" pageNumber="75">Fig. 6</figureCitation>
). The
<typeStatus id="22368863FF810678FF06FC2FFEF9368B" box="[190,293,916,942]" pageId="8" pageNumber="75" type="holotype">holotype</typeStatus>
was collected in the Republic of
<collectingCountry id="859A7651FF810678FD7CFC2FFCB5368B" box="[708,873,916,942]" name="Colombia" pageId="8" pageNumber="75">New Granada</collectingCountry>
(18311858), which refers to a region that included primarily
<collectingCountry id="859A7651FF810678FE18FC0CFDC836F4" box="[416,532,951,977]" name="Colombia" pageId="8" pageNumber="75">Colombia</collectingCountry>
and
<collectingCountry id="859A7651FF810678FDE8FC03FD6C36F7" box="[592,688,952,978]" name="Panama" pageId="8" pageNumber="75">Panama</collectingCountry>
, and smaller areas from the countries of
<collectingCountry id="859A7651FF810678FB2EFC0CFB3D36F4" box="[1174,1249,951,977]" name="Brazil" pageId="8" pageNumber="75">Brazil</collectingCountry>
,
<collectingCountry id="859A7651FF810678FB56FC0CFAAE36F4" box="[1262,1394,951,977]" name="Costa Rica" pageId="8" pageNumber="75">Costa Rica</collectingCountry>
,
<collectingCountry id="859A7651FF810678FF06FC61FEFE36D1" box="[190,290,986,1012]" name="Ecuador" pageId="8" pageNumber="75">Ecuador</collectingCountry>
,
<collectingCountry id="859A7651FF810678FE95FC60FEBF36D0" box="[301,355,987,1013]" name="Peru" pageId="8" pageNumber="75">Peru</collectingCountry>
and
<collectingCountry id="859A7651FF810678FE24FC61FDC536D1" box="[412,537,986,1012]" name="Venezuela" pageId="8" pageNumber="75">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>