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<mods:title id="99986956E091C8D9526995C4CCF03FE1">Crinoids of the stalked family Hyocrinidae (Echinodermata) collected by the USNS Eltanin in the Southern Ocean</mods:title>
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<mods:namePart id="1EBD2699F1AB1BE1CA6547D409A11635">Roux, Michel</mods:namePart>
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<treatment id="03FFAD25FFC2FFE35483F958A883FE08" ID-DOI="http://doi.org/10.5281/zenodo.6095285" ID-GBIF-Taxon="119638578" ID-Zenodo-Dep="6095285" LSID="urn:lsid:plazi:treatment:03FFAD25FFC2FFE35483F958A883FE08" httpUri="http://treatment.plazi.org/id/03FFAD25FFC2FFE35483F958A883FE08" lastPageId="26" lastPageNumber="407" pageId="24" pageNumber="405">
<subSubSection id="C34C4FB8FFC2FFE15483F958A862F889" pageId="24" pageNumber="405" type="nomenclature">
<paragraph id="8BE91C33FFC2FFE15483F958A825F902" blockId="24.[151,777,1677,1736]" box="[151,777,1677,1703]" pageId="24" pageNumber="405">
<heading id="D0A1AB5FFFC2FFE15483F958A825F902" bold="true" box="[151,777,1677,1703]" fontSize="11" level="1" pageId="24" pageNumber="405" reason="1">
<taxonomicName id="4C5667B0FFC2FFE15483F958A825F902" ID-CoL="8GKH6" authority="Mironov &amp; Sorokina, 1998" authorityName="Mironov &amp; Sorokina" authorityYear="1998" box="[151,777,1677,1703]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="24" pageNumber="405" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC2FFE15483F958A825F902" bold="true" box="[151,777,1677,1703]" pageId="24" pageNumber="405">
<emphasis id="B922C021FFC2FFE15483F958AA9FF902" bold="true" box="[151,435,1677,1703]" italics="true" pageId="24" pageNumber="405">Thalassocrinus clausus</emphasis>
<bibRefCitation id="EFC761C2FFC2FFE155AEF958A825F902" author="Mironov" box="[442,777,1677,1703]" pageId="24" pageNumber="405" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina, 1998</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE91C33FFC2FFE15483F97AAA82F96D" blockId="24.[151,777,1677,1736]" box="[151,430,1711,1736]" pageId="24" pageNumber="405">
<figureCitation id="136D00B6FFC2FFE15483F97AAA29F96D" box="[151,261,1711,1736]" captionStart="FIGURE 17" captionStartId="25.[151,250,1032,1054]" captionTargetBox="[151,1436,589,1010]" captionTargetId="figure@25.[151,1436,589,1011]" captionTargetPageId="25" captionText="FIGURE 17. Proxistele and proximal mesistele columnals in Thalassocrinus clausus specimen 1005456. A: columnal of mid proxistele, B C: columnal of proximal mesistele; C: oblique view." httpUri="https://zenodo.org/record/243066/files/figure.png" pageId="24" pageNumber="405">Figure 17</figureCitation>
,
<tableCitation id="C6D42988FFC2FFE15505F97AAA86F96D" box="[273,426,1711,1736]" captionStart-0="TABLE 16" captionStart-1="TABLE 17" captionStartId-0="25.[151,239,1130,1153]" captionStartId-1="25.[151,239,1437,1460]" captionText-0="TABLE 16. Main quantitative characters in theca and proximal stalk in Thalassocrinus clausus specimen USNM 1005456. Wb: Br 1 width; Wr-b: radial lower width; Wr-t: radial upper width; Hc: aboral cup height; Hr-r: radial axial height; Hr-i: radial side height; Htg-a: tegmen height at anus; Htg-b: tegmen height at mouth; Dc: uppermost diameter of aboral cup; Drb: diameter at radial-basal sutures; Dp: proximalmost stalk diameter; Dm: minimum stalk diameter in mesistele. Values in mm." captionText-1="TABLE 17. Main morphological ratios in theca and proximal stalk in Thalassocrinus clausus specimen USNM 1005456. Abbreviations like in Table 16." pageId="24" pageNumber="405">Tables 1617</tableCitation>
.
</paragraph>
<paragraph id="8BE91C33FFC2FFE15483F92DA862F889" blockId="24.[151,1436,1784,1837]" pageId="24" pageNumber="405">
Synonymy:
<taxonomicName id="4C5667B0FFC2FFE15506F92DA868F8AB" authority="Mironov &amp; Sorokina, 1998: 36" authorityName="Mironov &amp; Sorokina" authorityPageNumber="36" authorityYear="1998" box="[274,836,1784,1806]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="24" pageNumber="405" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC2FFE15506F92DA92CF8AB" box="[274,512,1784,1806]" italics="true" pageId="24" pageNumber="405">Thalassocrinus clausus</emphasis>
<bibRefCitation id="EFC761C2FFC2FFE15613F92DA80CF8AB" author="Mironov" box="[519,800,1784,1806]" pageId="24" pageNumber="405" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina, 1998</bibRefCitation>
: 36
</taxonomicName>
39, fig. 11, pls. 5 (56), 6 (12), 10 (6); Roux 2002: 173 174, 184; Roux
<emphasis id="B922C021FFC2FFE1557EF8CDAAB1F889" box="[362,413,1814,1837]" italics="true" pageId="24" pageNumber="405">et al.</emphasis>
2002: 822;
<bibRefCitation id="EFC761C2FFC2FFE15602F8C2A9D3F889" author="Roux" box="[534,767,1814,1836]" pageId="24" pageNumber="405" refString="Roux, M. &amp; Lambert, P. (2011) Two new species of stalked crinoids from the north-eastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy. Zootaxa, 2825, 1 - 54." type="journal article" year="2011">Roux &amp; Lambert 2011</bibRefCitation>
: 48, 51.
</paragraph>
</subSubSection>
<subSubSection id="C34C4FB8FFC2FFE15483F88CAE5DF833" pageId="24" pageNumber="405" type="materials_examined">
<paragraph id="8BE91C33FFC2FFE15483F88CAE5DF833" blockId="24.[151,1437,1881,2014]" pageId="24" pageNumber="405">
<emphasis id="B922C021FFC2FFE15483F88CAAACF8D7" bold="true" box="[151,384,1881,1906]" pageId="24" pageNumber="405">Material examined.</emphasis>
<collectionCode id="ED4784F6FFC2FFE15592F88FAAFBF8D7" LSID="urn:lsid:biocol.org:col:34871" box="[390,471,1882,1906]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="24" pageNumber="405">USNM</collectionCode>
1005456 from South of Gambier Islands, SE Pacific basin, 42°01 à
<geoCoordinate id="EE627AF4FFC2FFE150CCF88CAE1CF8D7" box="[1240,1328,1881,1906]" direction="south" orientation="latitude" pageId="24" pageNumber="405" precision="925" value="-41.95">41°57S</geoCoordinate>
130°02 à
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, depth
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: one specimen with aboral cup and proximal arms, and three stalk fragments.
</paragraph>
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<subSubSection id="C34C4FB8FFC2FFE054D3F874AE51F8E4" lastPageId="25" lastPageNumber="406" pageId="24" pageNumber="405" type="description">
<paragraph id="8BE91C33FFC2FFE054D3F874A9F4FF5D" blockId="24.[151,1437,1881,2014]" lastBlockId="25.[151,1437,151,536]" lastPageId="25" lastPageNumber="406" pageId="24" pageNumber="405">
<emphasis id="B922C021FFC2FFE154D3F874AA75F81F" bold="true" box="[199,345,1953,1978]" pageId="24" pageNumber="405">Description.</emphasis>
Theca very similar to
<typeStatus id="54EDA291FFC2FFE1564AF874A9EFF81F" box="[606,707,1953,1978]" pageId="24" pageNumber="405" type="holotype">holotype</typeStatus>
, except cup higher than wide, orals without terminal finger-like projection, and anal cone slightly higher than oral cone. Measurements are given in
<tableCitation id="C6D42988FFC2FFE1502EF810AF89F87B" box="[1082,1189,1989,2014]" captionStart="TABLE 16" captionStartId="25.[151,239,1130,1153]" captionText="TABLE 16. Main quantitative characters in theca and proximal stalk in Thalassocrinus clausus specimen USNM 1005456. Wb: Br 1 width; Wr-b: radial lower width; Wr-t: radial upper width; Hc: aboral cup height; Hr-r: radial axial height; Hr-i: radial side height; Htg-a: tegmen height at anus; Htg-b: tegmen height at mouth; Dc: uppermost diameter of aboral cup; Drb: diameter at radial-basal sutures; Dp: proximalmost stalk diameter; Dm: minimum stalk diameter in mesistele. Values in mm." pageId="24" pageNumber="405">Tables 16</tableCitation>
and
<tableCitation id="C6D42988FFC2FFE150CEF813AFD5F87B" box="[1242,1273,1990,2014]" captionStart="TABLE 17" captionStartId="25.[151,239,1437,1460]" captionText="TABLE 17. Main morphological ratios in theca and proximal stalk in Thalassocrinus clausus specimen USNM 1005456. Abbreviations like in Table 16." pageId="24" pageNumber="405">17</tableCitation>
. Proximal arm pattern 1+2 3 4+5 with first pinnule on Br5, one arm missing, three arms broken after Br5, the fifth separated (length
<quantity id="4CAEB1D6FFC3FFE054FAFF69AA69FF71" box="[238,325,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="25" pageNumber="406" unit="mm" value="9.2">9.2 mm</quantity>
) with successive brachial pairs beyond Br6. A single row of H-shaped plates in proximal pinnules only, probably indicative of an immature specimen.
</paragraph>
<paragraph id="8BE91C33FFC3FFE054D3FED1AF12FDBD" blockId="25.[151,1437,151,536]" pageId="25" pageNumber="406">
Length of preserved stalk
<quantity id="4CAEB1D6FFC3FFE055EAFED1A946FEB9" box="[510,618,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="25" pageNumber="406" unit="mm" value="22.6">22.6 mm</quantity>
. Proxistele rough and hexalobate in cross section proximally, distally becoming smoother and hexagonal in cross section beyond columnal 25. Columnal diameter decreasing from
<quantity id="4CAEB1D6FFC3FFE0516DFEFDABEFFEC1" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="25" pageNumber="406" unit="mm" value="1.3">1.3 mm</quantity>
below basal ring to
<quantity id="4CAEB1D6FFC3FFE055B2FE99A926FEC1" box="[422,522,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="25" pageNumber="406" unit="mm" value="1.05">1.05 mm</quantity>
at a distance of
<quantity id="4CAEB1D6FFC3FFE056D4FE99A83AFEC1" box="[704,790,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="25" pageNumber="406" unit="mm" value="5.4">5.4 mm</quantity>
and up to
<quantity id="4CAEB1D6FFC3FFE05798FE99A8DCFEC1" box="[908,1008,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="25" pageNumber="406" unit="mm" value="0.95">0.95 mm</quantity>
at
<quantity id="4CAEB1D6FFC3FFE05005FE99AF5BFEC1" box="[1041,1143,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="25" pageNumber="406" unit="mm" value="14.5">14.5 mm</quantity>
, and then increasing up to 1.0 mm at distal end of preserved stalk. Columnal height &lt;
<quantity id="4CAEB1D6FFC3FFE05736FEA5A854FE22" box="[802,888,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="406" unit="mm" value="0.3">0.3 mm</quantity>
proximally, increasing to
<quantity id="4CAEB1D6FFC3FFE0508EFEBAAFC3FE22" box="[1178,1263,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="406" unit="mm" value="0.5">0.5 mm</quantity>
at a distance of 6.0 mm from aboral cup, 1.0 mm at
<quantity id="4CAEB1D6FFC3FFE0562BFE41A98BFE09" box="[575,679,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="25" pageNumber="406" unit="mm" value="14.5">14.5 mm</quantity>
, and
<quantity id="4CAEB1D6FFC3FFE056F3FE41A861FE09" box="[743,845,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="25" pageNumber="406" unit="mm" value="1.25">1.25 mm</quantity>
distally. Maximum ratio height to diameter
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distalmost columnal. Columnal symplexies of 6 crenular units of 1 crenula (
<figureCitation id="136D00B6FFC3FFE05044FE62AFB1FE75" box="[1104,1181,439,464]" captionStart="FIGURE 17" captionStartId="25.[151,250,1032,1054]" captionTargetBox="[151,1436,589,1010]" captionTargetId="figure@25.[151,1436,589,1011]" captionTargetPageId="25" captionText="FIGURE 17. Proxistele and proximal mesistele columnals in Thalassocrinus clausus specimen 1005456. A: columnal of mid proxistele, B C: columnal of proximal mesistele; C: oblique view." httpUri="https://zenodo.org/record/243066/files/figure.png" pageId="25" pageNumber="406">Fig.17</figureCitation>
). Axial canal poorly differentiated in proximal mesistele (
<figureCitation id="136D00B6FFC3FFE05658FE09A98DFE51" box="[588,673,476,501]" captionStart="FIGURE 17" captionStartId="25.[151,250,1032,1054]" captionTargetBox="[151,1436,589,1010]" captionTargetId="figure@25.[151,1436,589,1011]" captionTargetPageId="25" captionText="FIGURE 17. Proxistele and proximal mesistele columnals in Thalassocrinus clausus specimen 1005456. A: columnal of mid proxistele, B C: columnal of proximal mesistele; C: oblique view." httpUri="https://zenodo.org/record/243066/files/figure.png" pageId="25" pageNumber="406">Fig. 17</figureCitation>
A) and highly differentiated distally with annular facet of small crenulae and large meshed galleried stereom bordering a wide lumen (
<figureCitation id="136D00B6FFC3FFE057BEFE2AA8D6FDBD" box="[938,1018,511,536]" captionStart="FIGURE 17" captionStartId="25.[151,250,1032,1054]" captionTargetBox="[151,1436,589,1010]" captionTargetId="figure@25.[151,1436,589,1011]" captionTargetPageId="25" captionText="FIGURE 17. Proxistele and proximal mesistele columnals in Thalassocrinus clausus specimen 1005456. A: columnal of mid proxistele, B C: columnal of proximal mesistele; C: oblique view." httpUri="https://zenodo.org/record/243066/files/figure.png" pageId="25" pageNumber="406">Fig. 17</figureCitation>
BC).
</paragraph>
<caption id="DF294CBBFFC3FFE05483FBDDA814FB98" httpUri="https://zenodo.org/record/243066/files/figure.png" pageId="25" pageNumber="406" targetBox="[151,1436,589,1010]" targetPageId="25">
<paragraph id="8BE91C33FFC3FFE05483FBDDA814FB98" blockId="25.[151,1435,1032,1085]" pageId="25" pageNumber="406">
<emphasis id="B922C021FFC3FFE05483FBDDAA0CFBB8" bold="true" box="[151,288,1032,1054]" pageId="25" pageNumber="406">FIGURE 17.</emphasis>
Proxistele and proximal mesistele columnals in
<taxonomicName id="4C5667B0FFC3FFE05705FBDDA8D3FBBB" box="[785,1023,1032,1054]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC3FFE05705FBDDA8D3FBBB" box="[785,1023,1032,1054]" italics="true" pageId="25" pageNumber="406">Thalassocrinus clausus</emphasis>
</taxonomicName>
specimen 1005456. A: columnal of mid proxistele, BC: columnal of proximal mesistele; C: oblique view.
</paragraph>
</caption>
<caption id="DF294CBBFFC3FFE05483FBBFAA8DFAA7" pageId="25" pageNumber="406">
<paragraph id="8BE91C33FFC3FFE05483FBBFAA8DFAA7" blockId="25.[151,1436,1130,1282]" pageId="25" pageNumber="406">
<emphasis id="B922C021FFC3FFE05483FBBFAA30FB24" bold="true" box="[151,284,1130,1153]" pageId="25" pageNumber="406">TABLE 16.</emphasis>
Main quantitative characters in theca and proximal stalk in
<taxonomicName id="4C5667B0FFC3FFE057C7FBBFAFFEFB24" box="[979,1234,1130,1153]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC3FFE057C7FBBFAFFEFB24" box="[979,1234,1130,1153]" italics="true" pageId="25" pageNumber="406">Thalassocrinus clausus</emphasis>
</taxonomicName>
specimen USNM 1005456. Wb: Br1 width; Wr-b: radial lower width; Wr-t: radial upper width; Hc: aboral cup height; Hr-r: radial axial height; Hr-i: radial side height; Htg-a: tegmen height at anus; Htg-b: tegmen height at mouth; Dc: uppermost diameter of aboral cup; Drb: diameter at radial-basal sutures; Dp: proximalmost stalk diameter; Dm: minimum stalk diameter in mesistele. Values in mm.
</paragraph>
</caption>
<paragraph id="8BE91C33FFC3FFE05489FAC8AE51F8E4" blockId="25.[157,1373,1309,1373]" lastBlockId="25.[151,1437,1652,2037]" pageId="25" pageNumber="406">
Wb Wrt Hc Hrr Hri Htga Htgo Dc Drb Dp Dm 2.7 4.2 8.1 3.7 4.5 3.8 1.7 6.2 5.2 1.3 0.95 Hc/Dc Hri/Hc Hrr/Hc Htgo/Hc Hri/Wrt Wrt/ Wb Dc/Dp Dc/Drb Drb/Dp Dp/Dm 1.30 0.56 0.46 0.21 1.07 1.57 4.89 1.19 4.10 1.34
<emphasis id="B922C021FFC3FFE054D3F9A1AA17F928" bold="true" box="[199,315,1652,1677]" pageId="25" pageNumber="406">Remarks.</emphasis>
The Eltanin specimen is substantially smaller than the
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(Dp
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) and likely immature. Its mesistele symplexies differ in having short crenulae and large axial canal and claustrum (
<figureCitation id="136D00B6FFC3FFE0511CF94DAE72F914" box="[1288,1374,1688,1713]" captionStart="FIGURE 17" captionStartId="25.[151,250,1032,1054]" captionTargetBox="[151,1436,589,1010]" captionTargetId="figure@25.[151,1436,589,1011]" captionTargetPageId="25" captionText="FIGURE 17. Proxistele and proximal mesistele columnals in Thalassocrinus clausus specimen 1005456. A: columnal of mid proxistele, B C: columnal of proximal mesistele; C: oblique view." httpUri="https://zenodo.org/record/243066/files/figure.png" pageId="25" pageNumber="406">Fig. 17</figureCitation>
BC) corresponding to juvenile characters. New columnals appearing below basal ring (i.e., later during ontogeny) display symplexial pattern (
<figureCitation id="136D00B6FFC3FFE055CAF935A91FF95C" box="[478,563,1760,1785]" captionStart="FIGURE 17" captionStartId="25.[151,250,1032,1054]" captionTargetBox="[151,1436,589,1010]" captionTargetId="figure@25.[151,1436,589,1011]" captionTargetPageId="25" captionText="FIGURE 17. Proxistele and proximal mesistele columnals in Thalassocrinus clausus specimen 1005456. A: columnal of mid proxistele, B C: columnal of proximal mesistele; C: oblique view." httpUri="https://zenodo.org/record/243066/files/figure.png" pageId="25" pageNumber="406">Fig. 17</figureCitation>
A) similar to those of proxistele and mesistele columnals in the
<typeStatus id="54EDA291FFC3FFE0512EF935AEB0F95C" box="[1338,1436,1760,1785]" pageId="25" pageNumber="406" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFC761C2FFC3FFE0548AF8D1A9A5F8B8" author="Mironov" box="[158,649,1796,1821]" pageId="25" pageNumber="406" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina 1998, pl. 5-5 and 6-1</bibRefCitation>
). The species is now known from two specimens only. The
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was found in NW Pacific at a shallower depth (
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<geoCoordinate id="EE627AF4FFC3FFE0572DF8FDA890F8E5" box="[825,956,1832,1857]" direction="east" orientation="longitude" pageId="25" pageNumber="406" precision="92" value="161.52667">161°31.6 E</geoCoordinate>
at a depth between 1,292 and
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).
</paragraph>
</subSubSection>
<caption id="DF294CBBFFC3FFE05483FA48A960FA71" pageId="25" pageNumber="406">
<paragraph id="8BE91C33FFC3FFE05483FA48A960FA71" blockId="25.[151,1436,1437,1492]" pageId="25" pageNumber="406">
<emphasis id="B922C021FFC3FFE05483FA48AA32FA11" bold="true" box="[151,286,1437,1460]" pageId="25" pageNumber="406">TABLE 17.</emphasis>
Main morphological ratios in theca and proximal stalk in
<taxonomicName id="4C5667B0FFC3FFE057DBFA48AFE3FA11" box="[975,1231,1437,1460]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC3FFE057DBFA48AFE3FA11" box="[975,1231,1437,1460]" italics="true" pageId="25" pageNumber="406">Thalassocrinus clausus</emphasis>
</taxonomicName>
specimen USNM 1005456. Abbreviations like in Table 16.
</paragraph>
</caption>
<subSubSection id="C34C4FB8FFC3FFE35483F898A883FE08" lastPageId="26" lastPageNumber="407" pageId="25" pageNumber="406" type="reference_group">
<paragraph id="8BE91C33FFC3FFE35483F898A903FEE5" blockId="25.[151,1437,1652,2037]" lastBlockId="26.[151,1437,151,429]" lastPageId="26" lastPageNumber="407" pageId="25" pageNumber="406">
<taxonomicName id="4C5667B0FFC3FFE05483F898AA2AF8C0" box="[151,262,1868,1893]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="alvinae">
<emphasis id="B922C021FFC3FFE05483F898AA2AF8C0" box="[151,262,1868,1893]" italics="true" pageId="25" pageNumber="406">T. alvinae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5667B0FFC3FFE0552AF898AA80F8C0" box="[318,428,1868,1893]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC3FFE0552AF898AA80F8C0" box="[318,428,1868,1893]" italics="true" pageId="25" pageNumber="406">T. clausus</emphasis>
</taxonomicName>
share both the same anal cone feature (
<bibRefCitation id="EFC761C2FFC3FFE0577AF898AF5EF8C0" author="Roux" box="[878,1138,1868,1893]" pageId="25" pageNumber="406" refString="Roux, M. &amp; Lambert, P. (2011) Two new species of stalked crinoids from the north-eastern Pacific in the genera Gephyrocrinus and Ptilocrinus (Echinodermata, Crinoidea, Hyocrinidae). Effects of ontogeny and variability on hyocrinid taxonomy. Zootaxa, 2825, 1 - 54." type="journal article" year="2011">Roux &amp; Lambert 2011</bibRefCitation>
) and a large axial canal in the mesistele of juvenile specimens (see Roux 2002, fig. 2AE). They mainly differ in proximal arm morphology.
<taxonomicName id="4C5667B0FFC3FFE05483F840AA2FF808" box="[151,259,1940,1965]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="alvinae">
<emphasis id="B922C021FFC3FFE05483F840AA2FF808" box="[151,259,1940,1965]" italics="true" pageId="25" pageNumber="406">T. alvinae</emphasis>
</taxonomicName>
displays a greater Wr/Wb ratio than
<taxonomicName id="4C5667B0FFC3FFE056B2F840A83AF808" box="[678,790,1940,1965]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC3FFE056B2F840A83AF808" box="[678,790,1940,1965]" italics="true" pageId="25" pageNumber="406">T. clausus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC761C2FFC3FFE05730F841AF4EF808" author="Mironov" box="[804,1122,1940,1965]" pageId="25" pageNumber="406" refString="Mironov, A. N. &amp; Sorokina, O. A. (1998) Sea lilies of the order Hyocrinida (Echinodermata, Crinoidea). Zoologicheskie Issledovania, 2, 1 - 117. [in Russian]" type="journal article" year="1998">Mironov &amp; Sorokina (1998)</bibRefCitation>
attributed to a new species, i. e.,
<taxonomicName id="4C5667B0FFC3FFE054C4F86CAAA9F874" box="[208,389,1976,2001]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="25" pageNumber="406" phylum="Echinodermata" rank="species" species="depauperatus">
<emphasis id="B922C021FFC3FFE054C4F86CAAA9F874" box="[208,389,1976,2001]" italics="true" pageId="25" pageNumber="406">T. depauperatus</emphasis>
</taxonomicName>
, a poorly preserved specimen with broad proximal arms, anal cone substantially higher than oral cone and orals without finger-like terminal projection. It differs from Eltanin specimen in having larger size (Dp
<quantity id="4CAEB1D6FFC0FFE354DEFF42AA1FFF15" box="[202,307,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="26" pageNumber="407" unit="mm" value="1.65">1.65 mm</quantity>
), three interbasal sutures, aboral cup more elongated (Hc/Dc 1.38), stalk symplexies with 7 crenular units and proximal brachials without lateral wings. Such differences could be accounted for by intraspecific variation, and/or correspond to ontogenetic changes. Moreover,
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<emphasis id="B922C021FFC0FFE35773FF34AF37FF5D" box="[871,1051,223,248]" italics="true" pageId="26" pageNumber="407">T. depauperatus</emphasis>
</taxonomicName>
was collected at a slightly greater depth (
<quantity id="4CAEB1D6FFC0FFE354F0FED1AA6DFEB8" box="[228,321,260,285]" metricMagnitude="3" metricUnit="m" metricValue="5.02" pageId="26" pageNumber="407" unit="m" value="5020.0">5,020 m</quantity>
<quantity id="4CAEB1D6FFC0FFE3555BFED1AA80FEB9" box="[335,428,260,285]" metricMagnitude="3" metricUnit="m" metricValue="5.13" pageId="26" pageNumber="407" unit="m" value="5130.0">5,130 m</quantity>
) off Sunda Strait (
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<geoCoordinate id="EE627AF4FFC0FFE35716FED1A8A0FEB9" box="[770,908,260,285]" direction="east" orientation="longitude" pageId="26" pageNumber="407" precision="92" value="102.91">102°54.6 E</geoCoordinate>
). Additional material is required to know if the two species are distinct or synonym.
</paragraph>
<paragraph id="8BE91C33FFC0FFE354D3FE99A883FE08" blockId="26.[151,1437,151,429]" pageId="26" pageNumber="407">
<emphasis id="B922C021FFC0FFE354D3FE99AA75FEC0" bold="true" box="[199,345,332,357]" pageId="26" pageNumber="407">Occurrence.</emphasis>
North western Pacific and South Central Pacific, depth
<quantity id="4CAEB1D6FFC0FFE357C5FE99AF01FEC1" box="[977,1069,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.334" pageId="26" pageNumber="407" unit="m" value="1334.0">1,334 m</quantity>
to
<quantity id="4CAEB1D6FFC0FFE35046FE99AF82FEC1" box="[1106,1198,332,356]" metricMagnitude="3" metricUnit="m" metricValue="4.831" pageId="26" pageNumber="407" unit="m" value="4831.0">4,831 m</quantity>
(possibly
<quantity id="4CAEB1D6FFC0FFE35137FE99AE53FEC1" box="[1315,1407,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.2919999999999998" pageId="26" pageNumber="407" unit="m" value="1292.0">1,292 m</quantity>
to
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). This depth range could be extending to
<quantity id="4CAEB1D6FFC0FFE356D8FEBAA804FE22" box="[716,808,367,392]" metricMagnitude="3" metricUnit="m" metricValue="5.02" pageId="26" pageNumber="407" unit="m" value="5020.0">5,020 m</quantity>
(possibly
<quantity id="4CAEB1D6FFC0FFE3578AFEBAA8D0FE2D" box="[926,1020,367,392]" metricMagnitude="3" metricUnit="m" metricValue="5.13" pageId="26" pageNumber="407" unit="m" value="5130.0">5,130 m</quantity>
) and the geographical distribution to NW Indian Ocean if
<taxonomicName id="4C5667B0FFC0FFE35596FE40AAC3FE08" box="[386,495,404,429]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="26" pageNumber="407" phylum="Echinodermata" rank="species" species="clausus">
<emphasis id="B922C021FFC0FFE35596FE40AAC3FE08" box="[386,495,404,429]" italics="true" pageId="26" pageNumber="407">T. clausus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5667B0FFC0FFE35633FE40A9F0FE08" box="[551,732,404,429]" class="Crinoidea" family="Hyocrinidae" genus="Thalassocrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hyocrinida" pageId="26" pageNumber="407" phylum="Echinodermata" rank="species" species="depauperatus">
<emphasis id="B922C021FFC0FFE35633FE40A9F0FE08" box="[551,732,404,429]" italics="true" pageId="26" pageNumber="407">T. depauperatus</emphasis>
</taxonomicName>
become synonym.
</paragraph>
</subSubSection>
</treatment>
</document>