treatments-xml/data/03/ED/87/03ED87D86B630937FE8D9872ACD8FBF7.xml

<document id="978F688AE92B0E7B0C16BF53EAA9B408" ID-DOI="10.5281/zenodo.5301164" ID-ISSN="0374-1036" ID-Zenodo-Dep="5301164" ID-ZooBank="402537AD-557B-4F95-8EB1-EC67EAEE0DDB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630099701985" checkinUser="marcus" docAuthor="Vendl, Tomáš, VONDRÁýEK, Dominik, Kubá, VítČzslav &amp; Šípek, Petr" docDate="2014" docId="03ED87D86B630937FE8D9872ACD8FBF7" docLanguage="en" docName="ActaEntMusNatPra.2014.54.2.571-604.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Euselates cineraceus " docType="treatment" docVersion="4" lastPageNumber="592" masterDocId="FFD4FFA06B6F0922FFF99D6EAE2CFFE4" masterDocTitle="Immature stages of Taenioderini (Coleoptera: Scarabaeidae: Cetoniinae): a report of hidden morphological diversity" masterLastPageNumber="604" masterPageNumber="571" pageNumber="583" updateTime="1699149442422" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="34D24CA2811DBFC9E8BFD2BA336BF513">Immature stages of Taenioderini (Coleoptera: Scarabaeidae: Cetoniinae): a report of hidden morphological diversity</mods:title>
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<mods:namePart id="8C3DCB684B1A8F45AE5D21654E1BD16D">Vendl, Tomáš</mods:namePart>
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<mods:namePart id="4535684D78AA2EAA2C1B276B366E29C2">VONDRÁýEK, Dominik</mods:namePart>
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<mods:namePart id="206F3A3600560250F9354030FBF483AC">Kubá, VítČzslav</mods:namePart>
<mods:affiliation id="171C94A3A4D1D7892467C19EAA0B25B1">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Poþernice, Czech Republic</mods:affiliation>
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<mods:namePart id="9C751E6F623E58E854EAC74FC82F1443">Šípek, Petr</mods:namePart>
<mods:affiliation id="851C9B46E4F0E27EB37525865E2A055B">Department of Zoology, Faculty of Science, Charles University in Prague, Viniþná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<heading id="D0B381A26B63092EFE8D9872ADA6FAD7" bold="true" box="[372,906,1308,1331]" centered="true" fontSize="8" level="2" pageId="12" pageNumber="583" reason="3">
<taxonomicName id="4C444D4D6B63092EFE8D9872ADA6FAD7" authority="(Gory &amp; Percheron, 1833)" authorityName="" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[372,906,1308,1331]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="583" phylum="Arthropoda" rank="species" species="cineraceus">
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<emphasis id="B930EADC6B63092EFE8D9872AC77FAD7" bold="true" box="[372,603,1308,1331]" italics="true" pageId="12" pageNumber="583">Euselates cineraceus</emphasis>
(Gory &amp; Percheron, 1833)
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<subSubSection id="C35E65456B63092EFEAC9854AD85FAA9" box="[341,937,1338,1357]" pageId="12" pageNumber="583" type="description">
<paragraph id="8BFB36CE6B63092EFEAC9854AD85FAA9" blockId="12.[341,937,1338,1357]" box="[341,937,1338,1357]" pageId="12" pageNumber="583">
(
<figureCitation id="137F2A4B6B63092EFEA29854AF8EFAA9" box="[347,418,1338,1357]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="12" pageNumber="583">Figs 2D</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFE529854AFE4FAA9" box="[427,456,1338,1357]" captionStart="Fig" captionStartId="6.[151,182,1208,1227]" captionTargetBox="[165,1084,224,1187]" captionTargetId="figure-10@6.[151,1120,216,1187]" captionTargetPageId="6" captionText="Fig. 3. Third instar larvae of Taenioderini. A–H – cranium. A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars: Figs 3A–H = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5301170" httpUri="https://zenodo.org/record/5301170/files/figure.png" pageId="12" pageNumber="583">3D</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFE289854AFC1FAA9" box="[465,493,1338,1357]" captionStart="Fig" captionStartId="7.[113,144,1144,1163]" captionTargetBox="[122,1055,222,1118]" captionTargetId="figure-11@7.[113,1086,219,1118]" captionTargetPageId="7" captionText="Fig. 4. Third instar larvae of Taenioderini. A–H – epipharynx. A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320612" httpUri="https://zenodo.org/record/5320612/files/figure.png" pageId="12" pageNumber="583">4D</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFE0F9854AC3FFAA9" box="[502,531,1338,1357]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="12" pageNumber="583">5D</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFDE29854AC75FAA9" box="[539,601,1338,1357]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="12" pageNumber="583">6J–L,S</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFD9B9854AC86FAA9" box="[610,682,1338,1357]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="12" pageNumber="583">8D–F,Q</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFD4A9854ACCDFAA9" box="[691,737,1338,1357]" captionStart="Fig" captionStartId="17.[114,145,1497,1516]" captionTargetBox="[169,1030,223,1459]" captionTargetId="figure-11@17.[152,1052,214,1485]" captionTargetPageId="17" captionText="Fig. 9. Third instar larvae of Taenioderini.A–C – right mandible (ventral, medial and dorsal aspect), D – pretarsus, E–L– antenna (dorsal and ventral aspect), M–T – metathoracic leg: A–D – Taeniodera sp. E,M – Chalcothea neglecta. F,N – Coilodera diardi. G,O – C. penicillata. H,P – Euselates cineraceus. I,Q – E. laoticus. J,R – Meroloba suturalis. K,S – Plectrone tristis. L, T – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301172" httpUri="https://zenodo.org/record/5301172/files/figure.png" pageId="12" pageNumber="583">9H,P</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFD139854AD3DFAA9" box="[746,785,1338,1357]" captionStart="Fig" captionStartId="19.[827,858,926,945]" captionTargetBox="[117,1089,233,1185]" captionTargetId="figure-150@19.[113,1085,219,1210]" captionTargetPageId="19" captionText="Fig. 10. Third instar larvae of Taenioderini. A–H – ultimate abdominal segment, ventral aspect. A – Chalcothea ne- glecta. B – Coilodera diardi. C – C. penicillata. D – Eusela- tes cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taenio- dera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320626" httpUri="https://zenodo.org/record/5320626/files/figure.png" pageId="12" pageNumber="583">10D</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFCE39854AD6CFAA9" box="[794,832,1338,1357]" captionStart="Fig" captionStartId="20.[845,878,904,923]" captionTargetBox="[157,1127,224,1163]" captionTargetId="figure-162@20.[151,1119,219,1170]" captionTargetPageId="20" captionText="Fig. 11. Third instar larvae of Taenioderini. A–H – ultima- te abdominal segment, lateral aspect: A – Chalcothea neglecta. B – Coilodera diardi. C – C. peni- cillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320630" httpUri="https://zenodo.org/record/5320630/files/figure.png" pageId="12" pageNumber="583">11D</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFCB09854AD44FAA9" box="[841,872,1338,1357]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="12" pageNumber="583">12J</figureCitation>
; 
<figureCitation id="137F2A4B6B63092EFC8B9854AD8EFAA9" box="[882,930,1338,1357]" captionStart="Fig" captionStartId="25.[113,144,1172,1191]" captionTargetBox="[113,1088,219,1150]" captionTargetId="figure-11@25.[113,1088,219,1150]" captionTargetPageId="25" captionText="Fig. 13. Third instar larvae of Taenioderini, SEM images. A–E – ligula: A – Chalcothea neglecta. B– Coilodera diardi, C – Euselates cineraceus. D – Meroloba suturalis. E – Taeniodera sp. F–L – mandibular stridulatory area: F – Chalcothea neglecta. G – Coilodera diardi. H – C. penicillata. I – E. cineraceus. J – E. laoticus. K – Meroloba suturalis. L – Taeniodera sp." figureDoi="http://doi.org/10.5281/zenodo.5320632" httpUri="https://zenodo.org/record/5320632/files/figure.png" pageId="12" pageNumber="583">13C,I</figureCitation>
)
</paragraph>
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<paragraph id="8BFB36CE6B63092EFF6E980AACADFA48" blockId="12.[151,1127,1380,1452]" pageId="12" pageNumber="583">
<emphasis id="B930EADC6B63092EFF6E980AAF60FA93" bold="true" box="[151,332,1380,1399]" pageId="12" pageNumber="583">Material examined.</emphasis>
14 third instar larvae reared from adults reared from ¿eld collected larvae at: 
<emphasis id="B930EADC6B63092EFBEA980AAF44FA76" bold="true" pageId="12" pageNumber="583">
MALAY- SIA: 
<collectingRegion id="4980F82C6B63092EFF319811AF2CFA74" box="[200,256,1407,1426]" country="Malaysia" name="Perak" pageId="12" pageNumber="583">PERAK</collectingRegion>
SULTANATE:
</emphasis>
Cameron 
<collectingRegion id="4980F82C6B63092EFE3A9811AC31FA76" box="[451,541,1407,1426]" country="Czech Republic" name="Vysocina" pageId="12" pageNumber="583">Highlands</collectingRegion>
, road between Tapah and Tanah Rata, Batu (= mile 19), 4°22.121ƍN 101°20.012ƍE, 
<quantity id="4CBC9B2B6B63092EFEEE98F7AF54FA48" box="[279,376,1433,1452]" metricMagnitude="2" metricUnit="m" metricValue="6.3" metricValueMax="6.6" metricValueMin="6.0" pageId="12" pageNumber="583" unit="m" value="630.0" valueMax="660.0" valueMin="600.0">600–660 m</quantity>
a.s.l., 
<date id="FFFA100E6B63092EFE5698F7AC21FA48" box="[431,525,1433,1452]" pageId="12" pageNumber="583" value="2010-03-12">12.iii.2010</date>
, P. Šípek leg.
</paragraph>
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<paragraph id="8BFB36CE6B63092FFF6E98ABACA0FED7" blockId="12.[151,1127,1477,1596]" lastBlockId="13.[113,1090,219,1491]" lastPageId="13" lastPageNumber="584" pageId="12" pageNumber="583">
<emphasis id="B930EADC6B63092EFF6E98ABAC12FA38" bold="true" box="[151,574,1477,1500]" pageId="12" pageNumber="583">
Description of third instar larva. 
<emphasis id="B930EADC6B63092EFDFF98ABAC12FA38" bold="true" box="[518,574,1477,1500]" italics="true" pageId="12" pageNumber="583">Body</emphasis>
</emphasis>
(
<figureCitation id="137F2A4B6B63092EFDB298ABACB3FA38" box="[587,671,1477,1500]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="12" pageNumber="583">Fig. 2D</figureCitation>
). Length 38.0–49.0 mm, dorsoventral interval of abdominal segments slightly larger than in thoracic segments (the abdominal segment VIII is the thickest). Chaetotaxy sparse, with three distinct 
<typeStatus id="54FF886C6B63092EFCE99B6BAD64F9F8" box="[784,840,1541,1564]" pageId="12" pageNumber="583">types</typeStatus>
of setae: Short or medium long hair-like setae on dorsal sublobes and the venter of the ultimate segment; long, stiff or slightly flattened setae on legs, ventral part of body and the dorsa of abdominal segments VII and VIII and oblanceolate* long setae on the ultimate abdominal segment; the last 
<specimenCount id="9D42FD476B62092FFC229D92AA6DFEF7" box="[987,1089,252,275]" pageId="13" pageNumber="584" type="generic" typeStatus="types">two types</specimenCount>
of setae, dark brown or black, the ¿rst pale brown.
</paragraph>
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<emphasis id="B930EADC6B62092FFF699C52AF09FEB7" bold="true" box="[144,293,316,339]" italics="true" pageId="13" pageNumber="584">Head capsule</emphasis>
(
<figureCitation id="137F2A4B6B62092FFECD9C52AFA6FEB7" box="[308,394,316,339]" captionStart="Fig" captionStartId="6.[151,182,1208,1227]" captionTargetBox="[165,1084,224,1187]" captionTargetId="figure-10@6.[151,1120,216,1187]" captionTargetPageId="6" captionText="Fig. 3. Third instar larvae of Taenioderini. A–H – cranium. A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars: Figs 3A–H = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5301170" httpUri="https://zenodo.org/record/5301170/files/figure.png" pageId="13" pageNumber="584">Fig. 3D</figureCitation>
). Maximum width 3.8–4.0 mm, glossy, yellowish or pale brown. Frontal sutures bisinuated. Epicranial insertions of antennal muscles indistinct. Cranial chaetotaxy summarized in 
<tableCitation id="C6C603756B62092FFE6E9C12AFC5FE77" box="[407,489,380,403]" captionStart="Table 1" captionStartId="28.[112,159,424,443]" captionText="Table 1. Cranial chaetotaxy of larvae of Taenioderini" pageId="13" pageNumber="584">Table 1</tableCitation>
. Anterior and exterior frontal setae with minute single seta or absent.
</paragraph>
<paragraph id="8BFB36CE6B62092FFF699CD2AA6EFE17" blockId="13.[113,1090,219,1491]" pageId="13" pageNumber="584">
<emphasis id="B930EADC6B62092FFF699CD2AED7FE37" bold="true" box="[144,251,444,467]" italics="true" pageId="13" pageNumber="584">Antennae</emphasis>
(
<figureCitation id="137F2A4B6B62092FFEF19CD2AF4DFE37" box="[264,353,444,467]" captionStart="Fig" captionStartId="6.[151,182,1208,1227]" captionTargetBox="[165,1084,224,1187]" captionTargetId="figure-10@6.[151,1120,216,1187]" captionTargetPageId="6" captionText="Fig. 3. Third instar larvae of Taenioderini. A–H – cranium. A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars: Figs 3A–H = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5301170" httpUri="https://zenodo.org/record/5301170/files/figure.png" pageId="13" pageNumber="584">Figs 3D</figureCitation>
; 
<figureCitation id="137F2A4B6B62092FFE959CD2AFBCFE37" box="[364,400,444,467]" captionStart="Fig" captionStartId="17.[114,145,1497,1516]" captionTargetBox="[169,1030,223,1459]" captionTargetId="figure-11@17.[152,1052,214,1485]" captionTargetPageId="17" captionText="Fig. 9. Third instar larvae of Taenioderini.A–C – right mandible (ventral, medial and dorsal aspect), D – pretarsus, E–L– antenna (dorsal and ventral aspect), M–T – metathoracic leg: A–D – Taeniodera sp. E,M – Chalcothea neglecta. F,N – Coilodera diardi. G,O – C. penicillata. H,P – Euselates cineraceus. I,Q – E. laoticus. J,R – Meroloba suturalis. K,S – Plectrone tristis. L, T – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301172" httpUri="https://zenodo.org/record/5301172/files/figure.png" pageId="13" pageNumber="584">9H</figureCitation>
). Relative length of antennomeres I–IV (an I–IV): an I&gt; an IV&gt; an II&gt; an III), ultimate antennomere with four to eight dorsal and 7–12 ventral sensory spots.
</paragraph>
<paragraph id="8BFB36CE6B62092FFF699C92AF8EFD57" blockId="13.[113,1090,219,1491]" pageId="13" pageNumber="584">
<emphasis id="B930EADC6B62092FFF699C92AF3DFDF7" bold="true" box="[144,273,508,531]" italics="true" pageId="13" pageNumber="584">Epipharynx</emphasis>
(
<figureCitation id="137F2A4B6B62092FFEE79C92AF5DFDF7" box="[286,369,508,531]" captionStart="Fig" captionStartId="7.[113,144,1144,1163]" captionTargetBox="[122,1055,222,1118]" captionTargetId="figure-11@7.[113,1086,219,1118]" captionTargetPageId="7" captionText="Fig. 4. Third instar larvae of Taenioderini. A–H – epipharynx. A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320612" httpUri="https://zenodo.org/record/5320612/files/figure.png" pageId="13" pageNumber="584">Fig. 4D</figureCitation>
). Haptomerum: Zygum convex, more or less protruding, with arcuate row of 16–18 stout conical setae and few setae proximad to the row. Sensilla of zygum grouped in single group distad to the row. Acroparia: Lateral lobes of epipharynx with three to ¿ve long setae, medial lobe with eight setae. Acanthoparia with ¿ve to nine setae surrounded by moderately swollen base (or tubercle); the size of the tubercles as well as the setae increasing towards apex of epipharynx.
</paragraph>
<paragraph id="8BFB36CE6B62092FFF699FD2ADA3FCB7" blockId="13.[113,1090,219,1491]" pageId="13" pageNumber="584">Chaetoparia: Asymmetric, right side composed of approximately 40–60 setae in three to ¿ve longitudinal rows and medial ¿eld (proximad to plate-shaped sclerite). Left side with approximately 30–40 setae in two to three rows. Both parts of chaetoparia with medial, more or less regular row of prominent long setae. Pedium large. Dexiotorma straight, right pternotorma well developed. Laeotorma absent, left pternotorma well developed.</paragraph>
<paragraph id="8BFB36CE6B62092FFF699E32ACA3FC57" blockId="13.[113,1090,219,1491]" pageId="13" pageNumber="584">Haptolachus: Sense cone conical, almost pointed. Plate-like sclerite large, occupying almost the entire area between sense cone and the regular rows on both sides of chaetoparia, however, it can be less distinct in some specimens.</paragraph>
<paragraph id="8BFB36CE6B62092FFF699ED2AD6FFBB7" blockId="13.[113,1090,219,1491]" pageId="13" pageNumber="584">
<emphasis id="B930EADC6B62092FFF699ED2AF28FC37" bold="true" box="[144,260,956,979]" italics="true" pageId="13" pageNumber="584">Mandibles</emphasis>
(
<figureCitation id="137F2A4B6B62092FFEE89ED2AF53FC37" box="[273,383,956,979]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="13" pageNumber="584">Figs 6J–L</figureCitation>
; 
<figureCitation id="137F2A4B6B62092FFE709ED2AFE4FC37" box="[393,456,956,979]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="13" pageNumber="584">8D–F</figureCitation>
; 
<figureCitation id="137F2A4B6B62092FFE2A9ED2AFD5FC37" box="[467,505,956,979]" captionStart="Fig" captionStartId="25.[113,144,1172,1191]" captionTargetBox="[113,1088,219,1150]" captionTargetId="figure-11@25.[113,1088,219,1150]" captionTargetPageId="25" captionText="Fig. 13. Third instar larvae of Taenioderini, SEM images. A–E – ligula: A – Chalcothea neglecta. B– Coilodera diardi, C – Euselates cineraceus. D – Meroloba suturalis. E – Taeniodera sp. F–L – mandibular stridulatory area: F – Chalcothea neglecta. G – Coilodera diardi. H – C. penicillata. I – E. cineraceus. J – E. laoticus. K – Meroloba suturalis. L – Taeniodera sp." figureDoi="http://doi.org/10.5281/zenodo.5320632" httpUri="https://zenodo.org/record/5320632/files/figure.png" pageId="13" pageNumber="584">13I</figureCitation>
). Stridulatory area well developed, with approximately 20–25 ridges (ridges in the proximal half less distinct and narrow). Scrobis with three to ¿ve lateral setae. Longitudinal furrow absent. Apical half of mandibles in dorsal aspect with two lateral setae and mediolateral depression (proximad to the base of the third scissorial tooth). Lateral outline of both mandibles without prominent external tooth.
</paragraph>
<paragraph id="8BFB36CE6B62092FFF699932AF9CFA37" blockId="13.[113,1090,219,1491]" pageId="13" pageNumber="584">
<emphasis id="B930EADC6B62092FFF699932AECAFB97" bold="true" box="[144,230,1116,1139]" italics="true" pageId="13" pageNumber="584">Maxilla</emphasis>
(
<figureCitation id="137F2A4B6B62092FFF0A9932AF67FB97" box="[243,331,1116,1139]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="13" pageNumber="584">Figs 5D</figureCitation>
; 
<figureCitation id="137F2A4B6B62092FFEAF9932AF52FB97" box="[342,382,1116,1139]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="13" pageNumber="584">12J</figureCitation>
). Dorsal surface of cardo and labacoparia with four to eight, 15–23 setae respectively. Ventral surface of cardo and labacoparia with two to ¿ve and 5–14 stiff setae respectively. Dorsal surface of stipes with 11–19 slender hair-like setae and one anterolateral stout seta. Maxillary stridulatory apparatus on stipes with more or less regular row of six to eight spine-like stridulatory teeth (in dorsal aspect, 
<figureCitation id="137F2A4B6B62092FFD6A99B2ACC6FB17" box="[659,746,1244,1267]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="13" pageNumber="584">Fig. 12J</figureCitation>
) and one or two blunt tubercles. Ventral surface of stipes with single proximal and single distal stout seta. Galear portion of mala in dorsal aspect beside large falcate uncus with four to ¿ve large stiff and one to three medium-sized hair-like setae, respectively. Lacinial part of mala with around 13–19 mostly very long and stiff setae. Lacinial apex with two unci (subapical uncus however small and reduced) and with conical, stout setae. Vental surface of mala with two longitudinal rows of two to four (seven) setae; distal setae of the exterior row long and stiff, distal setae of the interior row shorter and stout.
</paragraph>
<footnote id="E85F2AC06B62092FFF809B66AF46F9D2" pageId="13" pageNumber="584">
<paragraph id="8BFB36CE6B62092FFF809B66AF46F9D2" blockId="13.[121,1081,1544,1590]" pageId="13" pageNumber="584">* The extraordinary oblanceolate setae were present only in third instar larvae, ¿rst and second instar larvae have ‘normal’ cetoniinae pilosity.</paragraph>
</footnote>
<caption id="DF3B66466B61092CFF6E9880AD68F9D3" ID-DOI="http://doi.org/10.5281/zenodo.5320618" ID-Zenodo-Dep="5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="14" pageNumber="585" startId="14.[151,182,1518,1537]" targetBox="[222,1083,233,1473]" targetPageId="14">
<paragraph id="8BFB36CE6B61092CFF6E9880AD68F9D3" blockId="14.[151,1127,1518,1591]" pageId="14" pageNumber="585">
Fig. 6. Third instar larvae of 
<taxonomicName id="4C444D4D6B61092CFE749880AFD7F9E5" authorityName="Krikken" authorityYear="1984" box="[397,507,1518,1537]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="603" phylum="Arthropoda" rank="tribe" tribe="Taenioderini">Taenioderini</taxonomicName>
.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – 
<emphasis id="B930EADC6B61092CFEC89B67AFC4F9F8" box="[305,488,1545,1564]" italics="true" pageId="14" pageNumber="585">
<taxonomicName id="4C444D4D6B61092CFEC89B67AFC8F9F8" authorityName="Ritsema" authorityYear="1882" box="[305,484,1545,1564]" class="Insecta" family="Cetoniidae" genus="Chalcothea" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="585" phylum="Arthropoda" rank="species" species="neglecta">Chalcothea neglecta</taxonomicName>
.
</emphasis>
D–F,Q – 
<emphasis id="B930EADC6B61092CFDC39B67ACFDF9F8" box="[570,721,1545,1564]" italics="true" pageId="14" pageNumber="585">
<taxonomicName id="4C444D4D6B61092CFDC39B67ACE0F9F8" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[570,716,1545,1564]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="585" phylum="Arthropoda" rank="species" species="diardi">Coilodera diardi</taxonomicName>
.
</emphasis>
G–I,R – 
<emphasis id="B930EADC6B61092CFCE69B67ADB6F9F8" box="[799,922,1545,1564]" italics="true" pageId="14" pageNumber="585">
<taxonomicName id="4C444D4D6B61092CFCE69B67ADBAF9F8" authorityName="Hope" authorityYear="1831" box="[799,918,1545,1564]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="585" phylum="Arthropoda" rank="species" species="penicillata">C. penicillata</taxonomicName>
.
</emphasis>
J–L,S – 
<taxonomicName id="4C444D4D6B61092CFC1D9B67AEFCF9D3" authorityName="" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="585" phylum="Arthropoda" rank="species" species="cineraceus">
<emphasis id="B930EADC6B61092CFC1D9B67AEFCF9D3" italics="true" pageId="14" pageNumber="585">Euselates cineraceus</emphasis>
</taxonomicName>
. M–O,T – 
<emphasis id="B930EADC6B61092CFED69B4AAFBDF9D3" box="[303,401,1572,1591]" italics="true" pageId="14" pageNumber="585">
<taxonomicName id="4C444D4D6B61092CFED69B4AAFA1F9D3" authorityName="Miksiu" authorityYear="1974" box="[303,397,1572,1591]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="585" phylum="Arthropoda" rank="species" species="laoticus">E. laoticus</taxonomicName>
.
</emphasis>
Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm.
</paragraph>
</caption>
<caption id="DF3B66466B60092DFF88988DACB7F9A2" ID-DOI="http://doi.org/10.5281/zenodo.5320622" ID-Zenodo-Dep="5320622" httpUri="https://zenodo.org/record/5320622/files/figure.png" pageId="15" pageNumber="586" startId="15.[113,144,1507,1526]" targetBox="[163,1037,231,1485]" targetPageId="15">
<paragraph id="8BFB36CE6B60092DFF88988DACB7F9A2" blockId="15.[113,1089,1507,1606]" pageId="15" pageNumber="586">
Fig. 7. Third instar larvae of 
<taxonomicName id="4C444D4D6B60092DFE8D988DAFCEFA12" authorityName="Krikken" authorityYear="1984" box="[372,482,1507,1526]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="603" phylum="Arthropoda" rank="tribe" tribe="Taenioderini">Taenioderini</taxonomicName>
. A–I – left mandible (dorsal, medial and ventral aspect), J–O – right mandible (ventral, medial and dorsal aspect), P–R – thoracic spiracle, S–T – pretarsus.A–C,P – 
<taxonomicName id="4C444D4D6B60092DFC609890AA17F9F5" baseAuthorityName="Snellen van Vollenhoven" baseAuthorityYear="1858" box="[921,1083,1534,1553]" class="Insecta" family="Cetoniidae" genus="Meroloba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="586" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B930EADC6B60092DFC609890AA17F9F5" box="[921,1083,1534,1553]" italics="true" pageId="15" pageNumber="586">Meroloba suturalis</emphasis>
</taxonomicName>
. D–E,Q – 
<taxonomicName id="4C444D4D6B60092DFF3A9B76AF67F9CF" baseAuthorityName="Westwood" baseAuthorityYear="1842" box="[195,331,1560,1579]" class="Insecta" family="Cetoniidae" genus="Plectrone" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="586" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B930EADC6B60092DFF3A9B76AF67F9CF" box="[195,331,1560,1579]" italics="true" pageId="15" pageNumber="586">Plectrone tristis</emphasis>
</taxonomicName>
. G–I,R – 
<taxonomicName id="4C444D4D6B60092DFE669B76AC00F9CF" box="[415,556,1560,1579]" class="Insecta" family="Cetoniidae" genus="Taeniodera" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="586" phylum="Arthropoda" rank="species" species="undefined-j">
<emphasis id="B930EADC6B60092DFE669B76AFD3F9CF" box="[415,511,1560,1579]" italics="true" pageId="15" pageNumber="586">Taeniodera</emphasis>
sp. J
</taxonomicName>
–L,S – 
<taxonomicName id="4C444D4D6B60092DFD909B76AD35F9CF" authorityName="Ritsema" authorityYear="1882" box="[617,793,1560,1579]" class="Insecta" family="Cetoniidae" genus="Chalcothea" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="586" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="B930EADC6B60092DFD909B76AD35F9CF" box="[617,793,1560,1579]" italics="true" pageId="15" pageNumber="586">Chalcothea neglecta</emphasis>
</taxonomicName>
. M–O,T – 
<taxonomicName id="4C444D4D6B60092DFC819B76AA2BF9CF" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[888,1031,1560,1579]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="586" phylum="Arthropoda" rank="species" species="diardi">
<emphasis id="B930EADC6B60092DFC819B76AA2BF9CF" box="[888,1031,1560,1579]" italics="true" pageId="15" pageNumber="586">Coilodera diardi</emphasis>
</taxonomicName>
. Scale bars: Figs A–O,S = 1 mm; Figs P–R = 0.7 mm; Fig. T = 0.1 mm.
</paragraph>
</caption>
<caption id="DF3B66466B7F0932FF6E9893AD6DF9A2" ID-DOI="http://doi.org/10.5281/zenodo.5320624" ID-Zenodo-Dep="5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="16" pageNumber="587" startId="16.[151,182,1533,1552]" targetBox="[188,1060,222,1486]" targetPageId="16">
<paragraph id="8BFB36CE6B7F0932FF6E9893AD6DF9A2" blockId="16.[151,1128,1533,1606]" pageId="16" pageNumber="587">
Fig. 8. Third instar larvae of 
<taxonomicName id="4C444D4D6B7F0932FE7C9893AFDFF9F4" authorityName="Krikken" authorityYear="1984" box="[389,499,1533,1552]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="603" phylum="Arthropoda" rank="tribe" tribe="Taenioderini">Taenioderini</taxonomicName>
.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – 
<emphasis id="B930EADC6B7F0932FF119B76AF84F9CF" box="[232,424,1560,1579]" italics="true" pageId="16" pageNumber="587">
<taxonomicName id="4C444D4D6B7F0932FF119B76AF88F9CF" authorityName="Hope" authorityYear="1831" box="[232,420,1560,1579]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="penicillata">Coilodera penicillata</taxonomicName>
.
</emphasis>
D–F,Q – 
<emphasis id="B930EADC6B7F0932FE079B76AC95F9CF" box="[510,697,1560,1579]" italics="true" pageId="16" pageNumber="587">
<taxonomicName id="4C444D4D6B7F0932FE079B76AC99F9CF" authorityName="" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[510,693,1560,1579]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="cineraceus">Euselates cineraceus</taxonomicName>
.
</emphasis>
G–I,R – 
<taxonomicName id="4C444D4D6B7F0932FCF29B76AD44F9CF" authorityName="Miksiu" authorityYear="1974" box="[779,872,1560,1579]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="B930EADC6B7F0932FCF29B76AD44F9CF" box="[779,872,1560,1579]" italics="true" pageId="16" pageNumber="587">E. laoticus</emphasis>
</taxonomicName>
. J–L,S – 
<taxonomicName id="4C444D4D6B7F0932FC459B76AA4DF9CF" baseAuthorityName="Snellen van Vollenhoven" baseAuthorityYear="1858" box="[956,1121,1560,1579]" class="Insecta" family="Cetoniidae" genus="Meroloba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B930EADC6B7F0932FC459B76AA4DF9CF" box="[956,1121,1560,1579]" italics="true" pageId="16" pageNumber="587">Meroloba suturalis</emphasis>
</taxonomicName>
. M–O,T – 
<taxonomicName id="4C444D4D6B7F0932FF129B5DAF58F9A2" baseAuthorityName="Westwood" baseAuthorityYear="1842" box="[235,372,1587,1606]" class="Insecta" family="Cetoniidae" genus="Plectrone" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="587" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B930EADC6B7F0932FF129B5DAF58F9A2" box="[235,372,1587,1606]" italics="true" pageId="16" pageNumber="587">Plectrone tristis</emphasis>
</taxonomicName>
. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm.
</paragraph>
</caption>
<caption id="DF3B66466B7E0933FF8B98B7ACDEF9D8" ID-DOI="http://doi.org/10.5281/zenodo.5301172" ID-Zenodo-Dep="5301172" httpUri="https://zenodo.org/record/5301172/files/figure.png" pageId="17" pageNumber="588" startId="17.[114,145,1497,1516]" targetBox="[169,1030,223,1459]" targetPageId="17">
<paragraph id="8BFB36CE6B7E0933FF8B98B7ACDEF9D8" blockId="17.[114,1090,1497,1596]" pageId="17" pageNumber="588">
Fig. 9. Third instar larvae of 
<taxonomicName id="4C444D4D6B7E0933FE9F98B7AFF8FA08" authorityName="Krikken" authorityYear="1984" box="[358,468,1497,1516]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="603" phylum="Arthropoda" rank="tribe" tribe="Taenioderini">Taenioderini</taxonomicName>
.A–C – right mandible (ventral, medial and dorsal aspect), D – pretarsus, E–L– antenna (dorsal and ventral aspect), M–T – metathoracic leg: A–D – 
<taxonomicName id="4C444D4D6B7E0933FD05989AADA3F9E3" box="[764,911,1524,1543]" class="Insecta" family="Cetoniidae" genus="Taeniodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="undefined-e">
<emphasis id="B930EADC6B7E0933FD05989AAD70F9E3" box="[764,860,1524,1543]" italics="true" pageId="17" pageNumber="588">Taeniodera</emphasis>
sp. E
</taxonomicName>
,M – 
<taxonomicName id="4C444D4D6B7E0933FC44989AAE8AF9C6" authorityName="Ritsema" authorityYear="1882" class="Insecta" family="Cetoniidae" genus="Chalcothea" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="B930EADC6B7E0933FC44989AAE8AF9C6" italics="true" pageId="17" pageNumber="588">Chalcothea neglecta</emphasis>
</taxonomicName>
. F,N – 
<taxonomicName id="4C444D4D6B7E0933FF1D9B61AF5FF9C6" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[228,371,1551,1570]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="diardi">
<emphasis id="B930EADC6B7E0933FF1D9B61AF5FF9C6" box="[228,371,1551,1570]" italics="true" pageId="17" pageNumber="588">Coilodera diardi</emphasis>
</taxonomicName>
. G,O – 
<taxonomicName id="4C444D4D6B7E0933FE4F9B61AC06F9C6" authorityName="Hope" authorityYear="1831" box="[438,554,1551,1570]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B930EADC6B7E0933FE4F9B61AC06F9C6" box="[438,554,1551,1570]" italics="true" pageId="17" pageNumber="588">C. penicillata</emphasis>
</taxonomicName>
. H,P – 
<taxonomicName id="4C444D4D6B7E0933FD909B61AD30F9C6" authorityName="" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[617,796,1551,1570]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="cineraceus">
<emphasis id="B930EADC6B7E0933FD909B61AD30F9C6" box="[617,796,1551,1570]" italics="true" pageId="17" pageNumber="588">Euselates cineraceus</emphasis>
</taxonomicName>
. I,Q – 
<taxonomicName id="4C444D4D6B7E0933FCA19B61AD9FF9C6" authorityName="Miksiu" authorityYear="1974" box="[856,947,1551,1570]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="B930EADC6B7E0933FCA19B61AD9FF9C6" box="[856,947,1551,1570]" italics="true" pageId="17" pageNumber="588">E. laoticus</emphasis>
</taxonomicName>
. J,R – 
<taxonomicName id="4C444D4D6B7E0933FC179B61AE91F9D8" baseAuthorityName="Snellen van Vollenhoven" baseAuthorityYear="1858" class="Insecta" family="Cetoniidae" genus="Meroloba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B930EADC6B7E0933FC179B61AE91F9D8" italics="true" pageId="17" pageNumber="588">Meroloba suturalis</emphasis>
</taxonomicName>
. K,S – 
<taxonomicName id="4C444D4D6B7E0933FF049B47AFAAF9D8" baseAuthorityName="Westwood" baseAuthorityYear="1842" box="[253,390,1577,1596]" class="Insecta" family="Cetoniidae" genus="Plectrone" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B930EADC6B7E0933FF049B47AFAAF9D8" box="[253,390,1577,1596]" italics="true" pageId="17" pageNumber="588">Plectrone tristis</emphasis>
</taxonomicName>
. L, T – 
<taxonomicName id="4C444D4D6B7E0933FE309B47AC64F9D8" box="[457,584,1577,1596]" class="Insecta" family="Cetoniidae" genus="Taeniodera" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="588" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B930EADC6B7E0933FE309B47AC05F9D8" box="[457,553,1577,1596]" italics="true" pageId="17" pageNumber="588">Taeniodera</emphasis>
sp.
</taxonomicName>
Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BFB36CE6B7D0930FF4C9DB2AC3FFED7" blockId="18.[151,1127,220,1043]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF4C9DB2AFF4FF17" bold="true" box="[181,472,220,243]" italics="true" pageId="18" pageNumber="589">Hypopharyngeal sclerome</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FE119DB2AC6CFF17" box="[488,576,220,243]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="18" pageNumber="589">Fig. 5D</figureCitation>
). Four to ¿ve tegumentary expansions (phoba-like processes) present on the left lateral lobe of hypopharynx, tegumentary expansions on right medial portion of scleroma present.
</paragraph>
<paragraph id="8BFB36CE6B7D0930FF4C9C52ACDBFE57" blockId="18.[151,1127,220,1043]" pageId="18" pageNumber="589">
<taxonomicName id="4C444D4D6B7D0930FF4C9C52AED2FEB7" authorityName="Bloch" authorityYear="1782" box="[181,254,316,339]" class="Cestoda" family="Diphyllobothriidae" genus="Ligula" kingdom="Animalia" order="Diphyllobothriidea" pageId="18" pageNumber="589" phylum="Platyhelminthes" rank="genus">
<emphasis id="B930EADC6B7D0930FF4C9C52AED2FEB7" bold="true" box="[181,254,316,339]" italics="true" pageId="18" pageNumber="589">Ligula</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A4B6B7D0930FEF59C52AF48FEB7" box="[268,356,316,339]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="18" pageNumber="589">Figs 5D</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE969C52AFB2FEB7" box="[367,414,316,339]" captionStart="Fig" captionStartId="25.[113,144,1172,1191]" captionTargetBox="[113,1088,219,1150]" captionTargetId="figure-11@25.[113,1088,219,1150]" captionTargetPageId="25" captionText="Fig. 13. Third instar larvae of Taenioderini, SEM images. A–E – ligula: A – Chalcothea neglecta. B– Coilodera diardi, C – Euselates cineraceus. D – Meroloba suturalis. E – Taeniodera sp. F–L – mandibular stridulatory area: F – Chalcothea neglecta. G – Coilodera diardi. H – C. penicillata. I – E. cineraceus. J – E. laoticus. K – Meroloba suturalis. L – Taeniodera sp." figureDoi="http://doi.org/10.5281/zenodo.5320632" httpUri="https://zenodo.org/record/5320632/files/figure.png" pageId="18" pageNumber="589">13C</figureCitation>
). Dorsal surface with two lateral groups of nine hair-like setae, two paramedial setae at the apical margin and central group of setae and sensilla. This group is composed of two paramedial rows of three stout and conical setae, and transverse, basomedial row with four conical and four to six campaniform setae.
</paragraph>
<paragraph id="8BFB36CE6B7D0930FF4C9CD2AD6FFD57" blockId="18.[151,1127,220,1043]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF4C9CD2AF2AFE37" bold="true" box="[181,262,444,467]" italics="true" pageId="18" pageNumber="589">Thorax</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FEEC9CD2AF42FE37" box="[277,366,444,467]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="18" pageNumber="589">Figs 2D</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE839CD2AFB5FE37" box="[378,409,444,467]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="18" pageNumber="589">6S</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE5D9CD2AFEBFE37" box="[420,455,444,467]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="18" pageNumber="589">8Q</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE2A9CD2AFDEFE37" box="[467,498,444,467]" captionStart="Fig" captionStartId="17.[114,145,1497,1516]" captionTargetBox="[169,1030,223,1459]" captionTargetId="figure-11@17.[152,1052,214,1485]" captionTargetPageId="17" captionText="Fig. 9. Third instar larvae of Taenioderini.A–C – right mandible (ventral, medial and dorsal aspect), D – pretarsus, E–L– antenna (dorsal and ventral aspect), M–T – metathoracic leg: A–D – Taeniodera sp. E,M – Chalcothea neglecta. F,N – Coilodera diardi. G,O – C. penicillata. H,P – Euselates cineraceus. I,Q – E. laoticus. J,R – Meroloba suturalis. K,S – Plectrone tristis. L, T – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301172" httpUri="https://zenodo.org/record/5301172/files/figure.png" pageId="18" pageNumber="589">9P</figureCitation>
). Prothoracic sclerite with seven to ten setae on the anteroventral margin and another two setae in the posterio-dorsal area. Each sublobe of prothorax dorsally with one row of mostly medium-long or long setae interspersed with few short setae. Thoracic spiracle (
<figureCitation id="137F2A4B6B7D0930FE999F72AF9CFDD7" box="[352,432,540,563]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="18" pageNumber="589">Fig. 6S</figureCitation>
) 0.7 × 
<quantity id="4CBC9B2B6B7D0930FDF99F72AC7EFDD7" box="[512,594,540,563]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="589" unit="mm" value="0.4">0.4 mm</quantity>
(height × width), bullar opening narrow, arms of respiratory plate almost concealed. Respiratory plate with approximately 20–40 holes across diameter. Venter of each thoracic segment with four to six prominent dark setae (usually one pair of long and several medium-long or short setae present). Similar setae also on coxa. Pretarsus (
<figureCitation id="137F2A4B6B7D0930FEF19FF2AF77FD57" box="[264,347,668,691]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="18" pageNumber="589">Fig. 8Q</figureCitation>
) conical with falcate tip and two apical setae.
</paragraph>
<paragraph id="8BFB36CE6B7D0930FF4C9FD2AF6CFC97" blockId="18.[151,1127,220,1043]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF4C9FD2AF0CFD37" bold="true" box="[181,288,700,723]" italics="true" pageId="18" pageNumber="589">Abdomen</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FED49FD2AFABFD37" box="[301,391,700,723]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="18" pageNumber="589">Figs 2D</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE689FD2AFEFFD37" box="[401,451,700,723]" captionStart="Fig" captionStartId="19.[827,858,926,945]" captionTargetBox="[117,1089,233,1185]" captionTargetId="figure-150@19.[113,1085,219,1210]" captionTargetPageId="19" captionText="Fig. 10. Third instar larvae of Taenioderini. A–H – ultimate abdominal segment, ventral aspect. A – Chalcothea ne- glecta. B – Coilodera diardi. C – C. penicillata. D – Eusela- tes cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taenio- dera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320626" httpUri="https://zenodo.org/record/5320626/files/figure.png" pageId="18" pageNumber="589">10D</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE349FD2AFD2FD37" box="[461,510,700,723]" captionStart="Fig" captionStartId="20.[845,878,904,923]" captionTargetBox="[157,1127,224,1163]" captionTargetId="figure-162@20.[151,1119,219,1170]" captionTargetPageId="20" captionText="Fig. 11. Third instar larvae of Taenioderini. A–H – ultima- te abdominal segment, lateral aspect: A – Chalcothea neglecta. B – Coilodera diardi. C – C. peni- cillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320630" httpUri="https://zenodo.org/record/5320630/files/figure.png" pageId="18" pageNumber="589">11D</figureCitation>
). Each dorsal sublobe of abdominal segments I–VIII with one row of mostly medium long setae, however, short and long setae also present. Dorsum of abdominal segment VII–VIII, as well as venter, and spiracular and pleural areas of all abdominal segments with several prominent dark stiff setae. Spiracles on abdominal segments I–V elliptical, similar to thoracic spiracle, abdominal spiracles on segments VI–VIII almost circular.
</paragraph>
<paragraph id="8BFB36CE6B7D0930FF4C9E12AA3EFBF7" blockId="18.[151,1127,220,1043]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF4C9E12AED0FC77" bold="true" box="[181,252,892,915]" italics="true" pageId="18" pageNumber="589">Raster</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FEF39E12AF5CFC77" box="[266,368,892,915]" captionStart="Fig" captionStartId="19.[827,858,926,945]" captionTargetBox="[117,1089,233,1185]" captionTargetId="figure-150@19.[113,1085,219,1210]" captionTargetPageId="19" captionText="Fig. 10. Third instar larvae of Taenioderini. A–H – ultimate abdominal segment, ventral aspect. A – Chalcothea ne- glecta. B – Coilodera diardi. C – C. penicillata. D – Eusela- tes cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taenio- dera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320626" httpUri="https://zenodo.org/record/5320626/files/figure.png" pageId="18" pageNumber="589">Figs 10D</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE839E12AF86FC77" box="[378,426,892,915]" captionStart="Fig" captionStartId="20.[845,878,904,923]" captionTargetBox="[157,1127,224,1163]" captionTargetId="figure-162@20.[151,1119,219,1170]" captionTargetPageId="20" captionText="Fig. 11. Third instar larvae of Taenioderini. A–H – ultima- te abdominal segment, lateral aspect: A – Chalcothea neglecta. B – Coilodera diardi. C – C. peni- cillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320630" httpUri="https://zenodo.org/record/5320630/files/figure.png" pageId="18" pageNumber="589">11D</figureCitation>
). Palidium monostichous (however with few pali scattered around the main row), approximately 25 pali arranged in single elongate U-shaped row. Septula opened posteriorly, about three times longer than broad. Tegilla fused, with numerous short, stout, apically recurved setae, ventral and dorsal anal lip with dense rows of similar setae. The ultimate abdominal segment with sparse but prominent dark oblanceoate setae.
</paragraph>
<paragraph id="8BFB36CE6B7D0930FE289952AD9CFB89" blockId="18.[334,944,1082,1133]" pageId="18" pageNumber="589">
<taxonomicName id="4C444D4D6B7D0930FE289952ACF2FBB7" authority="Miksi" authorityName="Miksiu" authorityYear="1974" box="[465,734,1084,1107]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="589" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="B930EADC6B7D0930FE289952ACF2FBB7" bold="true" box="[465,734,1084,1107]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FE289952ACBBFBB7" bold="true" box="[465,663,1084,1107]" italics="true" pageId="18" pageNumber="589">Euselates laoticus</emphasis>
Mikši
</emphasis>
</taxonomicName>
ü, 
<emphasis id="B930EADC6B7D0930FD0E9952AD00FBB7" bold="true" box="[759,812,1084,1107]" pageId="18" pageNumber="589">1974</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FEAD9934AFB6FB89" box="[340,410,1114,1133]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="18" pageNumber="589">Figs 2E</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE5B9934AF91FB89" box="[418,445,1114,1133]" captionStart="Fig" captionStartId="6.[151,182,1208,1227]" captionTargetBox="[165,1084,224,1187]" captionTargetId="figure-10@6.[151,1120,216,1187]" captionTargetPageId="6" captionText="Fig. 3. Third instar larvae of Taenioderini. A–H – cranium. A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars: Figs 3A–H = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5301170" httpUri="https://zenodo.org/record/5301170/files/figure.png" pageId="18" pageNumber="589">3E</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE3C9934AFCCFB89" box="[453,480,1114,1133]" captionStart="Fig" captionStartId="7.[113,144,1144,1163]" captionTargetBox="[122,1055,222,1118]" captionTargetId="figure-11@7.[113,1086,219,1118]" captionTargetPageId="7" captionText="Fig. 4. Third instar larvae of Taenioderini. A–H – epipharynx. A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320612" httpUri="https://zenodo.org/record/5320612/files/figure.png" pageId="18" pageNumber="589">4E</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE119934AC2EFB89" box="[488,514,1114,1133]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="18" pageNumber="589">5E</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FDF29934AC7BFB89" box="[523,599,1114,1133]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="18" pageNumber="589">6M–O,T</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FDA69934AC8DFB89" box="[607,673,1114,1133]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="18" pageNumber="589">8G–I,R</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FD539934ACFFFB89" box="[682,723,1114,1133]" captionStart="Fig" captionStartId="17.[114,145,1497,1516]" captionTargetBox="[169,1030,223,1459]" captionTargetId="figure-11@17.[152,1052,214,1485]" captionTargetPageId="17" captionText="Fig. 9. Third instar larvae of Taenioderini.A–C – right mandible (ventral, medial and dorsal aspect), D – pretarsus, E–L– antenna (dorsal and ventral aspect), M–T – metathoracic leg: A–D – Taeniodera sp. E,M – Chalcothea neglecta. F,N – Coilodera diardi. G,O – C. penicillata. H,P – Euselates cineraceus. I,Q – E. laoticus. J,R – Meroloba suturalis. K,S – Plectrone tristis. L, T – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301172" httpUri="https://zenodo.org/record/5301172/files/figure.png" pageId="18" pageNumber="589">9I,Q</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FD259934AD2DFB89" box="[732,769,1114,1133]" captionStart="Fig" captionStartId="19.[827,858,926,945]" captionTargetBox="[117,1089,233,1185]" captionTargetId="figure-150@19.[113,1085,219,1210]" captionTargetPageId="19" captionText="Fig. 10. Third instar larvae of Taenioderini. A–H – ultimate abdominal segment, ventral aspect. A – Chalcothea ne- glecta. B – Coilodera diardi. C – C. penicillata. D – Eusela- tes cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taenio- dera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320626" httpUri="https://zenodo.org/record/5320626/files/figure.png" pageId="18" pageNumber="589">10E</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FCF09934AD01FB89" box="[777,813,1114,1133]" captionStart="Fig" captionStartId="20.[845,878,904,923]" captionTargetBox="[157,1127,224,1163]" captionTargetId="figure-162@20.[151,1119,219,1170]" captionTargetPageId="20" captionText="Fig. 11. Third instar larvae of Taenioderini. A–H – ultima- te abdominal segment, lateral aspect: A – Chalcothea neglecta. B – Coilodera diardi. C – C. peni- cillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320630" httpUri="https://zenodo.org/record/5320630/files/figure.png" pageId="18" pageNumber="589">11E</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FCCF9934ADAEFB89" box="[822,898,1114,1133]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="18" pageNumber="589">12F,K,N</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FC729934AD85FB89" box="[907,937,1114,1133]" captionStart="Fig" captionStartId="25.[113,144,1172,1191]" captionTargetBox="[113,1088,219,1150]" captionTargetId="figure-11@25.[113,1088,219,1150]" captionTargetPageId="25" captionText="Fig. 13. Third instar larvae of Taenioderini, SEM images. A–E – ligula: A – Chalcothea neglecta. B– Coilodera diardi, C – Euselates cineraceus. D – Meroloba suturalis. E – Taeniodera sp. F–L – mandibular stridulatory area: F – Chalcothea neglecta. G – Coilodera diardi. H – C. penicillata. I – E. cineraceus. J – E. laoticus. K – Meroloba suturalis. L – Taeniodera sp." figureDoi="http://doi.org/10.5281/zenodo.5320632" httpUri="https://zenodo.org/record/5320632/files/figure.png" pageId="18" pageNumber="589">13J</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E65456B7D0930FF6E99EAAA61FB56" pageId="18" pageNumber="589" type="materials_examined">
<paragraph id="8BFB36CE6B7D0930FF6E99EAAA61FB56" blockId="18.[151,1126,1156,1203]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF6E99EAAF6BFB73" bold="true" box="[151,327,1156,1175]" pageId="18" pageNumber="589">Material examined.</emphasis>
2 third instar larvae reared from eggs laid by adults collected in: 
<emphasis id="B930EADC6B7D0930FC9999EAAA4AFB73" bold="true" box="[864,1126,1156,1175]" pageId="18" pageNumber="589">
<collectingCountry id="F353765E6B7D0930FC9999EAADB1FB73" box="[864,925,1156,1175]" name="Laos" pageId="18" pageNumber="589">LAOS</collectingCountry>
: 
<collectionCode id="ED55AE0B6B7D0930FC5C99EAAA73FB71" box="[933,1119,1156,1175]" pageId="18" pageNumber="589">HOUA PHAN PROVINCE</collectionCode>
:
</emphasis>
20°13ƍ09–19ƎN 103°59ƍ54Ǝ–104°00ƍ03ƎE, Mount Phou Pane, 
<quantity id="4CBC9B2B6B7D0930FD5099F1AD0CFB56" box="[681,800,1183,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.51" metricValueMin="1.48" pageId="18" pageNumber="589" unit="m" value="1495.0" valueMax="1510.0" valueMin="1480.0">1480–1510 m</quantity>
a.s.l., 
<date id="FFFA100E6B7D0930FCAE99F1ADE3FB56" box="[855,975,1183,1202]" pageId="18" pageNumber="589" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">1.–16.vi.2009</date>
, 
<collectionCode id="ED55AE0B6B7D0930FC2E99F1ADCAFB56" box="[983,998,1183,1202]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="18" pageNumber="589" type="Museum">V</collectionCode>
. KubáĖ leg.
</paragraph>
</subSubSection>
<subSubSection id="C35E65456B7D0937FF6E99A4ACD8FBF7" lastPageId="21" lastPageNumber="592" pageId="18" pageNumber="589" type="description">
<paragraph id="8BFB36CE6B7D0930FF6E99A4AD5CFA85" blockId="18.[151,1127,1226,1569]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF6E99A4AC67FB05" bold="true" box="[151,587,1226,1249]" pageId="18" pageNumber="589">
Description of third instar larva. 
<emphasis id="B930EADC6B7D0930FDEA99A4AC67FB05" bold="true" box="[531,587,1226,1249]" italics="true" pageId="18" pageNumber="589">Body</emphasis>
</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FDA399A4AC82FB05" box="[602,686,1226,1249]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="18" pageNumber="589">Fig. 2E</figureCitation>
). Length 27.0–34.0 mm, dorsoventral interval of body segments gradually increasing toward the penultimate segment; penultimate and ultimate segment distinctly thicker than the previous segments. Chaetotaxy more or less sparse, however with numerous long or extremely long setae, giving the larva ‘hairy appearance’, the longest setae present on the ultimate abdominal segment.
</paragraph>
<paragraph id="8BFB36CE6B7D0930FF4C9804AFB9FA05" blockId="18.[151,1127,1226,1569]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF4C9804AF61FA65" bold="true" box="[181,333,1386,1409]" italics="true" pageId="18" pageNumber="589">Head capsule</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FEA79804AF9FFA65" box="[350,435,1386,1409]" captionStart="Fig" captionStartId="6.[151,182,1208,1227]" captionTargetBox="[165,1084,224,1187]" captionTargetId="figure-10@6.[151,1120,216,1187]" captionTargetPageId="6" captionText="Fig. 3. Third instar larvae of Taenioderini. A–H – cranium. A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars: Figs 3A–H = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5301170" httpUri="https://zenodo.org/record/5301170/files/figure.png" pageId="18" pageNumber="589">Fig. 3E</figureCitation>
). Width 2.9 (n = 1), brow with pale spots, clypeus and labrum yellow. Frontal sutures bisinuated. Epicranial insertions of antennal muscles visible as pale depressions. Cranial chaetotaxy summarized in 
<tableCitation id="C6C603756B7D0930FD6398C4ACC7FA25" box="[666,747,1450,1473]" captionStart="Table 1" captionStartId="28.[112,159,424,443]" captionText="Table 1. Cranial chaetotaxy of larvae of Taenioderini" pageId="18" pageNumber="589">Table 1</tableCitation>
. Anterior and exterior frontal setae with minute single seta.
</paragraph>
<paragraph id="8BFB36CE6B7D0930FF4C9884AA4BF9C5" blockId="18.[151,1127,1226,1569]" pageId="18" pageNumber="589">
<emphasis id="B930EADC6B7D0930FF4C9884AF0CF9E5" bold="true" box="[181,288,1514,1537]" italics="true" pageId="18" pageNumber="589">Antennae</emphasis>
(
<figureCitation id="137F2A4B6B7D0930FED69884AFA9F9E5" box="[303,389,1514,1537]" captionStart="Fig" captionStartId="6.[151,182,1208,1227]" captionTargetBox="[165,1084,224,1187]" captionTargetId="figure-10@6.[151,1120,216,1187]" captionTargetPageId="6" captionText="Fig. 3. Third instar larvae of Taenioderini. A–H – cranium. A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars: Figs 3A–H = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5301170" httpUri="https://zenodo.org/record/5301170/files/figure.png" pageId="18" pageNumber="589">Figs 3E</figureCitation>
; 
<figureCitation id="137F2A4B6B7D0930FE6B9884AF86F9E5" box="[402,426,1514,1537]" captionStart="Fig" captionStartId="17.[114,145,1497,1516]" captionTargetBox="[169,1030,223,1459]" captionTargetId="figure-11@17.[152,1052,214,1485]" captionTargetPageId="17" captionText="Fig. 9. Third instar larvae of Taenioderini.A–C – right mandible (ventral, medial and dorsal aspect), D – pretarsus, E–L– antenna (dorsal and ventral aspect), M–T – metathoracic leg: A–D – Taeniodera sp. E,M – Chalcothea neglecta. F,N – Coilodera diardi. G,O – C. penicillata. H,P – Euselates cineraceus. I,Q – E. laoticus. J,R – Meroloba suturalis. K,S – Plectrone tristis. L, T – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301172" httpUri="https://zenodo.org/record/5301172/files/figure.png" pageId="18" pageNumber="589">9I</figureCitation>
). Relative length of antennomeres I–IV (an I–IV): an I&gt; an IV&gt; an II&gt; an III), ultimate antennomere with eight dorsal and nine ventral sensory spots (n = 1).
</paragraph>
<caption id="DF3B66466B7C0931FCC29EF0AA07FB45" ID-DOI="http://doi.org/10.5281/zenodo.5320626" ID-Zenodo-Dep="5320626" httpUri="https://zenodo.org/record/5320626/files/figure.png" pageId="19" pageNumber="590" startId="19.[827,858,926,945]" targetBox="[117,1089,233,1185]" targetPageId="19">
<paragraph id="8BFB36CE6B7C0931FCC29EF0AA07FB45" blockId="19.[827,1089,926,1185]" pageId="19" pageNumber="590">
Fig. 10. Third instar larvae of 
<taxonomicName id="4C444D4D6B7C0931FCC29ED7AD87FC28" authorityName="Krikken" authorityYear="1984" box="[827,939,953,972]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="603" phylum="Arthropoda" rank="tribe" tribe="Taenioderini">Taenioderini</taxonomicName>
. A–H – ultimate abdominal segment, ventral aspect. A – 
<taxonomicName id="4C444D4D6B7C0931FC499E80AD5CFBF8" authorityName="Ritsema" authorityYear="1882" class="Insecta" family="Cetoniidae" genus="Chalcothea" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="B930EADC6B7C0931FC499E80AD5CFBF8" italics="true" pageId="19" pageNumber="590">Chalcothea neglecta</emphasis>
</taxonomicName>
. B – 
<taxonomicName id="4C444D4D6B7C0931FC5E9967AA17FBF8" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[935,1083,1033,1052]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="diardi">
<emphasis id="B930EADC6B7C0931FC5E9967AA17FBF8" box="[935,1083,1033,1052]" italics="true" pageId="19" pageNumber="590">Coilodera diardi</emphasis>
</taxonomicName>
. C – 
<taxonomicName id="4C444D4D6B7C0931FCA7994AADFDFBD3" authorityName="Hope" authorityYear="1831" box="[862,977,1060,1079]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B930EADC6B7C0931FCA7994AADFDFBD3" box="[862,977,1060,1079]" italics="true" pageId="19" pageNumber="590">C. penicillata</emphasis>
</taxonomicName>
. D – 
<taxonomicName id="4C444D4D6B7C0931FC06994AAD98FBB5" authorityName="" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="cineraceus">
<emphasis id="B930EADC6B7C0931FC06994AAD98FBB5" italics="true" pageId="19" pageNumber="590">Euselates cineraceus</emphasis>
</taxonomicName>
. E – 
<taxonomicName id="4C444D4D6B7C0931FC199950AA10FBB5" authorityName="Miksiu" authorityYear="1974" box="[992,1084,1086,1105]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="B930EADC6B7C0931FC199950AA10FBB5" box="[992,1084,1086,1105]" italics="true" pageId="19" pageNumber="590">E. laoticus</emphasis>
</taxonomicName>
. F – 
<taxonomicName id="4C444D4D6B7C0931FC9F9937AA0CFB88" baseAuthorityName="Snellen van Vollenhoven" baseAuthorityYear="1858" box="[870,1056,1113,1132]" class="Insecta" family="Cetoniidae" genus="Meroloba" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B930EADC6B7C0931FC9F9937AA0CFB88" box="[870,1056,1113,1132]" italics="true" pageId="19" pageNumber="590">Meroloba suturalis</emphasis>
</taxonomicName>
. G – 
<taxonomicName id="4C444D4D6B7C0931FCB2991AADFFFB63" baseAuthorityName="Westwood" baseAuthorityYear="1842" box="[843,979,1140,1159]" class="Insecta" family="Cetoniidae" genus="Plectrone" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B930EADC6B7C0931FCB2991AADFFFB63" box="[843,979,1140,1159]" italics="true" pageId="19" pageNumber="590">Plectrone tristis</emphasis>
</taxonomicName>
. H – 
<taxonomicName id="4C444D4D6B7C0931FBF9991AADADFB45" class="Insecta" family="Cetoniidae" genus="Taeniodera" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="590" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B930EADC6B7C0931FBF9991AAD4EFB45" italics="true" pageId="19" pageNumber="590">Taeniodera</emphasis>
sp.
</taxonomicName>
Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BFB36CE6B7C0931FF699981AD63FA42" blockId="19.[113,1089,1263,1606]" pageId="19" pageNumber="590">
<emphasis id="B930EADC6B7C0931FF699981AF3EFAE2" bold="true" box="[144,274,1263,1286]" italics="true" pageId="19" pageNumber="590">Epipharynx</emphasis>
(
<figureCitation id="137F2A4B6B7C0931FED99981AF5BFAE2" box="[288,375,1263,1286]" captionStart="Fig" captionStartId="7.[113,144,1144,1163]" captionTargetBox="[122,1055,222,1118]" captionTargetId="figure-11@7.[113,1086,219,1118]" captionTargetPageId="7" captionText="Fig. 4. Third instar larvae of Taenioderini. A–H – epipharynx. A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320612" httpUri="https://zenodo.org/record/5320612/files/figure.png" pageId="19" pageNumber="590">Figs 4E</figureCitation>
; 
<figureCitation id="137F2A4B6B7C0931FE7B9981AF83FAE2" box="[386,431,1263,1286]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="19" pageNumber="590">12F</figureCitation>
). Haptomerum: Zygum convex, more or less protruding, with arcuate row of approximately 14 stout conical setae and few setae proximad to the row. Sensilla of zygum grouped in single group on more or less prominent cone. Acroparia: Lateral lobes of epipharynx with four to ¿ve long setae, medial lobe with eight setae. Acanthoparia with ¿ve to six setae surrounded by moderately swollen base (or tubercle); the size of the tubercles as well as the setae increasing towards apex of epipharynx.
</paragraph>
<paragraph id="8BFB36CE6B7C0931FF6998C1AF88F9A2" blockId="19.[113,1089,1263,1606]" pageId="19" pageNumber="590">Chaetoparia: Asymmetric, right side composed of approximately 45 setae in irregular longitudinal rows and medial ¿eld (proximad to plate-shaped sclerite). Left side with approximately 30 setae in three irregular rows. Pedium large. Dexiotorma straight, right pternotorma absent (only slightly indicated). Laeotorma strongly reduced, however present, left pternotorma well developed.</paragraph>
<caption id="DF3B66466B7B0936FCB49EE6ADB4FB6F" ID-DOI="http://doi.org/10.5281/zenodo.5320630" ID-Zenodo-Dep="5320630" httpUri="https://zenodo.org/record/5320630/files/figure.png" pageId="20" pageNumber="591" startId="20.[845,878,904,923]" targetBox="[157,1127,224,1163]" targetPageId="20">
<paragraph id="8BFB36CE6B7B0936FCB49EE6ADB4FB6F" blockId="20.[845,1127,904,1163]" pageId="20" pageNumber="591">
Fig. 11. Third instar larvae of 
<taxonomicName id="4C444D4D6B7B0936FCB69ECDADE7FC52" authorityName="Krikken" authorityYear="1984" box="[847,971,931,950]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="603" phylum="Arthropoda" rank="tribe" tribe="Taenioderini">Taenioderini</taxonomicName>
. A–H – ultimate abdominal segment, lateral aspect: A – 
<taxonomicName id="4C444D4D6B7B0936FC489EB6AA4DFC0F" authorityName="Ritsema" authorityYear="1882" box="[945,1121,984,1003]" class="Insecta" family="Cetoniidae" genus="Chalcothea" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="B930EADC6B7B0936FC489EB6AA4DFC0F" box="[945,1121,984,1003]" italics="true" pageId="20" pageNumber="591">Chalcothea neglecta</emphasis>
</taxonomicName>
. B – 
<taxonomicName id="4C444D4D6B7B0936FC969E9DADD6FBE2" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[879,1018,1011,1030]" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="diardi">
<emphasis id="B930EADC6B7B0936FC969E9DADD6FBE2" box="[879,1018,1011,1030]" italics="true" pageId="20" pageNumber="591">Coilodera diardi</emphasis>
</taxonomicName>
. C – 
<taxonomicName id="4C444D4D6B7B0936FBDD9E9DADA9FBC4" authorityName="Hope" authorityYear="1831" class="Insecta" family="Cetoniidae" genus="Coilodera" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="penicillata">
<emphasis id="B930EADC6B7B0936FBDD9E9DADA9FBC4" italics="true" pageId="20" pageNumber="591">C. penicillata</emphasis>
</taxonomicName>
. D – 
<taxonomicName id="4C444D4D6B7B0936FC499963AA4DFBC4" authorityName="" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[944,1121,1037,1056]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="cineraceus">
<emphasis id="B930EADC6B7B0936FC499963AA4DFBC4" box="[944,1121,1037,1056]" italics="true" pageId="20" pageNumber="591">Euselates cineraceus</emphasis>
</taxonomicName>
. E – 
<taxonomicName id="4C444D4D6B7B0936FC819946ADF5FBDF" authorityName="Miksiu" authorityYear="1974" box="[888,985,1064,1083]" class="Insecta" family="Cetoniidae" genus="Euselates" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="laoticus">
<emphasis id="B930EADC6B7B0936FC819946ADF5FBDF" box="[888,985,1064,1083]" italics="true" pageId="20" pageNumber="591">E. laoticus</emphasis>
</taxonomicName>
. F – 
<taxonomicName id="4C444D4D6B7B0936FBE99946ADB7FBB2" baseAuthorityName="Snellen van Vollenhoven" baseAuthorityYear="1858" class="Insecta" family="Cetoniidae" genus="Meroloba" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B930EADC6B7B0936FBE99946ADB7FBB2" italics="true" pageId="20" pageNumber="591">Meroloba suturalis</emphasis>
</taxonomicName>
. G – 
<taxonomicName id="4C444D4D6B7B0936FC2A992DAA4DFBB2" baseAuthorityName="Westwood" baseAuthorityYear="1842" box="[979,1121,1091,1110]" class="Insecta" family="Cetoniidae" genus="Plectrone" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="B930EADC6B7B0936FC2A992DAA4DFBB2" box="[979,1121,1091,1110]" italics="true" pageId="20" pageNumber="591">Plectrone tristis</emphasis>
</taxonomicName>
. H – 
<taxonomicName id="4C444D4D6B7B0936FC839933AA2DFB94" box="[890,1025,1117,1136]" class="Insecta" family="Cetoniidae" genus="Taeniodera" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="591" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B930EADC6B7B0936FC839933ADF2FB94" box="[890,990,1117,1136]" italics="true" pageId="20" pageNumber="591">Taeniodera</emphasis>
sp.
</taxonomicName>
Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BFB36CE6B7B0936FF4C99BFAD50FB0C" blockId="20.[151,1127,1233,1608]" box="[181,892,1233,1256]" pageId="20" pageNumber="591">Haptolachus: Sense cone low, cob-like. Plate-like sclerite reduced.</paragraph>
<paragraph id="8BFB36CE6B7B0936FF4C999FAF1EFA6C" blockId="20.[151,1127,1233,1608]" pageId="20" pageNumber="591">
<emphasis id="B930EADC6B7B0936FF4C999FAF04FAEC" bold="true" box="[181,296,1265,1288]" italics="true" pageId="20" pageNumber="591">Mandibles</emphasis>
(
<figureCitation id="137F2A4B6B7B0936FECC999FAF9FFAEC" box="[309,435,1265,1288]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="20" pageNumber="591">Figs 6M–O</figureCitation>
; 
<figureCitation id="137F2A4B6B7B0936FE42999FAFD9FAEC" box="[443,501,1265,1288]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="20" pageNumber="591">8G–I</figureCitation>
; 
<figureCitation id="137F2A4B6B7B0936FE07999FAC0AFAEC" box="[510,550,1265,1288]" captionStart="Fig" captionStartId="25.[113,144,1172,1191]" captionTargetBox="[113,1088,219,1150]" captionTargetId="figure-11@25.[113,1088,219,1150]" captionTargetPageId="25" captionText="Fig. 13. Third instar larvae of Taenioderini, SEM images. A–E – ligula: A – Chalcothea neglecta. B– Coilodera diardi, C – Euselates cineraceus. D – Meroloba suturalis. E – Taeniodera sp. F–L – mandibular stridulatory area: F – Chalcothea neglecta. G – Coilodera diardi. H – C. penicillata. I – E. cineraceus. J – E. laoticus. K – Meroloba suturalis. L – Taeniodera sp." figureDoi="http://doi.org/10.5281/zenodo.5320632" httpUri="https://zenodo.org/record/5320632/files/figure.png" pageId="20" pageNumber="591">13J</figureCitation>
). Stridulatory area well developed, with approximately 20–25 ridges. Scrobis with two to ¿ve lateral setae. Longitudinal furrow absent. Apical half of mandibles in dorsal aspect with two lateral setae and mediolateral depression (proximad to the base of the third scissorial tooth). Lateral outline of both mandibles without prominent external tooth.
</paragraph>
<paragraph id="8BFB36CE6B7B0937FF4C98FFAF82FEF7" blockId="20.[151,1127,1233,1608]" lastBlockId="21.[113,1089,220,1043]" lastPageId="21" lastPageNumber="592" pageId="20" pageNumber="591">
<emphasis id="B930EADC6B7B0936FF4C98FFAF21FA4C" bold="true" box="[181,269,1425,1448]" italics="true" pageId="20" pageNumber="591">Maxilla</emphasis>
(
<figureCitation id="137F2A4B6B7B0936FEE498FFAF5AFA4C" box="[285,374,1425,1448]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="20" pageNumber="591">Figs 5E</figureCitation>
; 
<figureCitation id="137F2A4B6B7B0936FE7D98FFAF9AFA4C" box="[388,438,1425,1448]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="20" pageNumber="591">12K</figureCitation>
). Dorsal surface of cardo and labacoparia with four to six and 10 setae, respectively. Dorsal surface of stipes with 13–14 slender hair-like setae and one anterolateral stout seta. Maxillary stridulatory apparatus on stipes with four to ¿ve spinelike stridulatory teeth (in dorsal aspect, 
<figureCitation id="137F2A4B6B7B0936FDB3989FAC80F9EC" box="[586,684,1521,1544]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="20" pageNumber="591">Fig. 12K</figureCitation>
) and two blunt tubercles. Galear portion of mala in dorsal aspect beside large falcate uncus with ¿ve large stiff and three to four medium-sized hair-like setae, respectively. Lacinial part of mala with around 15 mostly long and stiff setae. Lacinial apex with two unci (subapical uncus however small and reduced) and with conical, stout setae.
</paragraph>
<paragraph id="8BFB36CE6B7A0937FF699C72AFC8FE97" blockId="21.[113,1089,220,1043]" pageId="21" pageNumber="592">
<emphasis id="B930EADC6B7A0937FF699C72AF84FED7" bold="true" box="[144,424,284,307]" italics="true" pageId="21" pageNumber="592">Hypopharyngeal sclerome</emphasis>
(
<figureCitation id="137F2A4B6B7A0937FE4A9C72AC2AFED7" box="[435,518,284,307]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="21" pageNumber="592">Figs 5E</figureCitation>
; 
<figureCitation id="137F2A4B6B7A0937FDF69C72AC6CFED7" box="[527,576,284,307]" captionStart="Fig" captionStartId="24.[151,182,1445,1464]" captionTargetBox="[152,1125,220,1379]" captionTargetId="figure-10@24.[151,1126,219,1380]" captionTargetPageId="24" captionText="Fig. 12. Third instar larvae of Taenioderini, SEM images.A–C – epipharynx:A – Chalcothea neglecta. B – Coilodera diardi. C– Taeniodera sp. D–F – sense cone (haptolachus, epipharynx): D – Chalcothea neglecta. E – Coilodera diardi. F– Euselates laoticus. G–M – maxillar stridulatory teeth: G – Chalcothea neglecta. H – Coilodera diardi. I – C. penicillata. J – E. cineraceus. K – E. laoticus. L – Meroloba suturalis.M – Taeniodera sp. N,O – hypopharyngeal scleroma: N – E. laoticus. O – Meroloba suturalis." figureDoi="http://doi.org/10.5281/zenodo.5301176" httpUri="https://zenodo.org/record/5301176/files/figure.png" pageId="21" pageNumber="592">12N</figureCitation>
). Four to ¿ve tegumentary expansions (phoba-like processes) present on the left lateral lobe of hypopharynx, tegumentary expansions on right medial portion of scleroma absent.
</paragraph>
<paragraph id="8BFB36CE6B7A0937FF699C12AC6DFE17" blockId="21.[113,1089,220,1043]" pageId="21" pageNumber="592">
<taxonomicName id="4C444D4D6B7A0937FF699C12AEF4FE77" authorityName="Bloch" authorityYear="1782" box="[144,216,380,403]" class="Cestoda" family="Diphyllobothriidae" genus="Ligula" kingdom="Animalia" order="Diphyllobothriidea" pageId="21" pageNumber="592" phylum="Platyhelminthes" rank="genus">
<emphasis id="B930EADC6B7A0937FF699C12AEF4FE77" bold="true" box="[144,216,380,403]" italics="true" pageId="21" pageNumber="592">Ligula</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A4B6B7A0937FF1D9C12AF18FE77" box="[228,308,380,403]" captionStart="Fig" captionStartId="8.[151,182,1046,1065]" captionTargetBox="[168,1126,222,1022]" captionTargetId="figure-10@8.[153,1130,217,1023]" captionTargetPageId="8" captionText="Fig. 5. Third instar larvae of Taenioderini. A–H – maxillo-labial complex.A – Chalcothea neglecta. B – Coilodera diardi. C – C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320616" httpUri="https://zenodo.org/record/5320616/files/figure.png" pageId="21" pageNumber="592">Fig. 5E</figureCitation>
). Dorsal surface with two lateral groups of nine hair-like setae, two paramedial setae at the apical margin and central group of setae and sensilla. This group is composed of two paramedial rows of three stout and conical setae, and transverse, basomedial row with three to conical and six campaniform setae.
</paragraph>
<paragraph id="8BFB36CE6B7A0937FF699C92AC27FD17" blockId="21.[113,1089,220,1043]" pageId="21" pageNumber="592">
<emphasis id="B930EADC6B7A0937FF699C92AECFFDF7" bold="true" box="[144,227,508,531]" italics="true" pageId="21" pageNumber="592">Thorax</emphasis>
(
<figureCitation id="137F2A4B6B7A0937FF089C92AF66FDF7" box="[241,330,508,531]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="21" pageNumber="592">Figs 2E</figureCitation>
; 
<figureCitation id="137F2A4B6B7A0937FEAC9C92AF59FDF7" box="[341,373,508,531]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="21" pageNumber="592">6T</figureCitation>
; 
<figureCitation id="137F2A4B6B7A0937FE799C92AF8EFDF7" box="[384,418,508,531]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="21" pageNumber="592">8R</figureCitation>
; 
<figureCitation id="137F2A4B6B7A0937FE569C92AFFFFDF7" box="[431,467,508,531]" captionStart="Fig" captionStartId="17.[114,145,1497,1516]" captionTargetBox="[169,1030,223,1459]" captionTargetId="figure-11@17.[152,1052,214,1485]" captionTargetPageId="17" captionText="Fig. 9. Third instar larvae of Taenioderini.A–C – right mandible (ventral, medial and dorsal aspect), D – pretarsus, E–L– antenna (dorsal and ventral aspect), M–T – metathoracic leg: A–D – Taeniodera sp. E,M – Chalcothea neglecta. F,N – Coilodera diardi. G,O – C. penicillata. H,P – Euselates cineraceus. I,Q – E. laoticus. J,R – Meroloba suturalis. K,S – Plectrone tristis. L, T – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5301172" httpUri="https://zenodo.org/record/5301172/files/figure.png" pageId="21" pageNumber="592">9Q</figureCitation>
). Prothoracic sclerite with anterior row of approximately 20 hair-like setae and posterior row of four to ¿ve setae. Each sublobe of prothorax dorsally with one row of mostly medium-long or long setae interspersed with few short setae. Thoracic spiracle (
<figureCitation id="137F2A4B6B7A0937FEC49F32AFBCFD97" box="[317,400,604,627]" captionStart="Fig" captionStartId="14.[151,182,1518,1537]" captionTargetBox="[222,1083,233,1473]" captionTargetId="figure-10@14.[194,1084,219,1490]" captionTargetPageId="14" captionText="Fig. 6. Third instar larvae of Taenioderini.A–O – left mandible (dorsal, medial and ventral aspect), P–T – thoracic spiracle. A–C,P – Chalcothea neglecta. D–F,Q – Coilodera diardi. G–I,R – C. penicillata. J–L,S – Euselates cineraceus. M–O,T – E. laoticus. Scale bars: Figs A–O = 1mm; Figs P–T = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320618" httpUri="https://zenodo.org/record/5320618/files/figure.png" pageId="21" pageNumber="592">Fig. 6T</figureCitation>
) 0.6 × 
<quantity id="4CBC9B2B6B7A0937FE249F32AC1CFD97" box="[477,560,604,627]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="21" pageNumber="592" unit="mm" value="0.3">0.3 mm</quantity>
(height × width), bullar opening broad, distance between the arms of respiratory plate equals the widest diameter of respiratory plate. Respiratory plate with approximately 15 holes across diameter. Venter of each thoracic segment with row of numerous long hair-like setae. Pretarsus (
<figureCitation id="137F2A4B6B7A0937FCD49FD2ADAEFD37" box="[813,898,700,723]" captionStart="Fig" captionStartId="16.[151,182,1533,1552]" captionTargetBox="[188,1060,222,1486]" captionTargetId="figure-10@16.[151,1092,219,1492]" captionTargetPageId="16" captionText="Fig. 8. Third instar larvae of Taenioderini.A–O – right mandible (ventral, medial and dorsal aspect), P–T – pretarsus. A–C,P – Coilodera penicillata. D–F,Q – Euselates cineraceus. G–I,R – E. laoticus. J–L,S – Meroloba suturalis. M–O,T – Plectrone tristis. Scale bars: Figs A–P,T = 1 mm; Figs Q–S = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5320624" httpUri="https://zenodo.org/record/5320624/files/figure.png" pageId="21" pageNumber="592">Fig. 8R</figureCitation>
) cylindrical with small falcate tip and two apical setae.
</paragraph>
<paragraph id="8BFB36CE6B7A0937FF699F92AD09FC97" blockId="21.[113,1089,220,1043]" pageId="21" pageNumber="592">
<emphasis id="B930EADC6B7A0937FF699F92AED5FCF7" bold="true" box="[144,249,764,787]" italics="true" pageId="21" pageNumber="592">Abdomen</emphasis>
(
<figureCitation id="137F2A4B6B7A0937FEF19F92AF4CFCF7" box="[264,352,764,787]" captionStart="Fig" captionStartId="4.[151,182,1243,1262]" captionTargetBox="[170,1087,218,1216]" captionTargetId="figure-90@4.[157,1091,213,1221]" captionTargetPageId="4" captionText="Fig. 2. Third instar larvae of Taenioderini.A–H – habitus.A – Chalcothea neglecta. B – Coilodera diardi. C –C. penicillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis.H – Taeniodera sp. Figures not to scale." figureDoi="http://doi.org/10.5281/zenodo.5301168" httpUri="https://zenodo.org/record/5301168/files/figure.png" pageId="21" pageNumber="592">Figs 2E</figureCitation>
; 
<figureCitation id="137F2A4B6B7A0937FE929F92AFB5FCF7" box="[363,409,764,787]" captionStart="Fig" captionStartId="19.[827,858,926,945]" captionTargetBox="[117,1089,233,1185]" captionTargetId="figure-150@19.[113,1085,219,1210]" captionTargetPageId="19" captionText="Fig. 10. Third instar larvae of Taenioderini. A–H – ultimate abdominal segment, ventral aspect. A – Chalcothea ne- glecta. B – Coilodera diardi. C – C. penicillata. D – Eusela- tes cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taenio- dera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320626" httpUri="https://zenodo.org/record/5320626/files/figure.png" pageId="21" pageNumber="592">10E</figureCitation>
; 
<figureCitation id="137F2A4B6B7A0937FE5C9F92AFFEFCF7" box="[421,466,764,787]" captionStart="Fig" captionStartId="20.[845,878,904,923]" captionTargetBox="[157,1127,224,1163]" captionTargetId="figure-162@20.[151,1119,219,1170]" captionTargetPageId="20" captionText="Fig. 11. Third instar larvae of Taenioderini. A–H – ultima- te abdominal segment, lateral aspect: A – Chalcothea neglecta. B – Coilodera diardi. C – C. peni- cillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320630" httpUri="https://zenodo.org/record/5320630/files/figure.png" pageId="21" pageNumber="592">11E</figureCitation>
). Each dorsal sublobe of abdominal segments I–VIII with one or two rows short and long hair-like setae. Setae of ventral and lateral parts of body extremely long. Spiracles on abdominal segments I–VII similar to the thoracic spiracle in shape and size, spiracle of abdominal segment distinctly smaller.
</paragraph>
<paragraph id="8BFB36CE6B7A0937FF699E12ACD8FBF7" blockId="21.[113,1089,220,1043]" pageId="21" pageNumber="592">
<emphasis id="B930EADC6B7A0937FF699E12AEFBFC77" bold="true" box="[144,215,892,915]" italics="true" pageId="21" pageNumber="592">Raster</emphasis>
(
<figureCitation id="137F2A4B6B7A0937FF1C9E12AF65FC77" box="[229,329,892,915]" captionStart="Fig" captionStartId="19.[827,858,926,945]" captionTargetBox="[117,1089,233,1185]" captionTargetId="figure-150@19.[113,1085,219,1210]" captionTargetPageId="19" captionText="Fig. 10. Third instar larvae of Taenioderini. A–H – ultimate abdominal segment, ventral aspect. A – Chalcothea ne- glecta. B – Coilodera diardi. C – C. penicillata. D – Eusela- tes cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taenio- dera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320626" httpUri="https://zenodo.org/record/5320626/files/figure.png" pageId="21" pageNumber="592">Figs 10E</figureCitation>
; 
<figureCitation id="137F2A4B6B7A0937FEAA9E12AFACFC77" box="[339,384,892,915]" captionStart="Fig" captionStartId="20.[845,878,904,923]" captionTargetBox="[157,1127,224,1163]" captionTargetId="figure-162@20.[151,1119,219,1170]" captionTargetPageId="20" captionText="Fig. 11. Third instar larvae of Taenioderini. A–H – ultima- te abdominal segment, lateral aspect: A – Chalcothea neglecta. B – Coilodera diardi. C – C. peni- cillata. D – Euselates cineraceus. E – E. laoticus. F – Meroloba suturalis. G – Plectrone tristis. H – Taeniodera sp. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5320630" httpUri="https://zenodo.org/record/5320630/files/figure.png" pageId="21" pageNumber="592">11E</figureCitation>
). Palidium monostichous (however with few scattered pali around the main row), approximately 20–25 pali arranged in single U-shaped row. Septula opened posteriorly, about two times longer than broad. Tegilla fused, with approximately 15 hamate setae. Hamate setae also present on the ventral anal lip. The ultimate abdominal segment with numerous extremely long hair-like setae especially on sides.
</paragraph>
</subSubSection>
</treatment>
</document>