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<document id="5F09379FC1CA3A5FC0015C23B9A6AB96" ID-DOI="10.5281/zenodo.5353724" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353724" ID-ZooBank="945E9711-35FE-466D-80BC-725AD484DE4B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212417419" checkinUser="felipe" docAuthor="Cano, Enio B." docDate="2020" docId="03ED87BFFFADFF86FF09FA6A6090B059" docLanguage="en" docName="0827.pdf" docOrigin="Insecta Mundi 2020 (827)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Phyllophaga (Phyllophaga) badbunnyi Cano 2020, new species" docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="FFD4FFC7FFAEFF8FFFC9FFD1623EB470" masterDocTitle="Two new species of Phyllophaga of the Schizorhina species group from cloud forests of Chiapas, México and Guatemala (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1699187864575" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="537AA648D020FA0F7EAB75989272CBE3">Two new species of Phyllophaga of the Schizorhina species group from cloud forests of Chiapas, México and Guatemala (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
</mods:titleInfo>
<mods:name id="2CFE18168BC77E114B28823B9F91AD99" type="personal">
<mods:role id="3EBF3F72291AA056A7020027DD1FD8BB">
<mods:roleTerm id="B7405CA1C5B2998724BED96E208520E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="367AFAFCB39CC3D2DC33C7F08EE35767">Cano, Enio B.</mods:namePart>
<mods:affiliation id="E11E08AD61C281AE7E7EEF6CF58ACD21">Escuela de Biología Facultad de Ciencias Químicas y Farmacia Universidad de San Carlos de Guatemala Edificio T- 10, segundo Nivel, Ciudad Universitaria, zona 12 Ciudad de Guatemala</mods:affiliation>
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<mods:titleInfo id="096B6F66AE3F21CFE5C2AD61268B732B">
<mods:title id="4ECB17742F5F6EECF4150CD4FB1BA515">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="4A4ECDFFA9BEAFE0618A44364F3EB5BD">
<mods:date id="84AABDCEDB2C82F1C4FFEF4A980FF3F6">2020</mods:date>
<mods:detail id="08FED852D5F8A3AF1F19EF939BC030E4" type="pubDate">
<mods:number id="0C17FE0B640C3BD3B763A91CA236E11A">2020-12-18</mods:number>
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<mods:detail id="F569E7652EB8C12F112496FF9DC704FA" type="volume">
<mods:number id="801580C0ACE39FB197C223806087C585">2020</mods:number>
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<mods:identifier id="982932541769CF302E30A14951DAC713" type="DOI">10.5281/zenodo.5353724</mods:identifier>
<mods:identifier id="8F9429D966E5AA1FEF0C5BA9DDE300F2" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03ED87BFFFADFF86FF09FA6A6090B059" ID-DOI="http://doi.org/10.5281/zenodo.5451662" ID-Zenodo-Dep="5451662" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059" httpUri="http://treatment.plazi.org/id/03ED87BFFFADFF86FF09FA6A6090B059" lastPageId="9" lastPageNumber="8" pageId="3" pageNumber="2">
<subSubSection id="C35E6522FFADFF8CFF09FA6A61AAB1A8" box="[192,916,1467,1496]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8BFB36A9FFADFF8CFF09FA6A61AAB1A8" blockId="3.[192,916,1467,1530]" box="[192,916,1467,1496]" pageId="3" pageNumber="2">
<heading id="D0B381C5FFADFF8CFF09FA6A61AAB1A8" box="[192,916,1467,1496]" fontSize="12" level="2" pageId="3" pageNumber="2" reason="2">
<emphasis id="B930EABBFFADFF8CFF09FA6A61AAB1A8" bold="true" box="[192,916,1467,1496]" pageId="3" pageNumber="2">
<taxonomicName id="4C444D2AFFADFF8CFF09FA6A60CCB1A8" authority="Cano" authorityName="Cano" authorityYear="2020" box="[192,754,1467,1496]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="badbunnyi" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="B930EABBFFADFF8CFF09FA6A6098B1A8" bold="true" box="[192,678,1467,1496]" italics="true" pageId="3" pageNumber="2">Phyllophaga (Phyllophaga) badbunnyi</emphasis>
Cano
</taxonomicName>
,
<taxonomicNameLabel id="A20357C0FFADFF8CFD36FA6D61AAB1A8" box="[767,916,1467,1496]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E6522FFADFF8CFF09FA316313B18A" box="[192,301,1504,1530]" pageId="3" pageNumber="2" type="description">
<paragraph id="8BFB36A9FFADFF8CFF09FA316313B18A" blockId="3.[192,916,1467,1530]" box="[192,301,1504,1530]" pageId="3" pageNumber="2">
(
<figureCitation id="137F2A2CFFADFF8CFF01FA31631BB18A" box="[200,293,1504,1530]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="4.[192,268,1620,1646]" captionStartId-1="6.[192,268,1607,1633]" captionStartId-2="7.[325,401,1474,1500]" captionStartId-3="8.[194,270,1512,1538]" captionTargetBox-0="[192,1440,230,1590]" captionTargetBox-1="[223,1406,221,1567]" captionTargetBox-2="[323,1312,217,1431]" captionTargetBox-3="[219,1427,222,1475]" captionTargetId-0="figure-14@4.[192,1440,213,1592]" captionTargetId-1="figure-14@6.[192,1440,211,1577]" captionTargetId-2="figure-175@7.[301,1331,213,1439]" captionTargetId-3="figure-166@8.[192,1440,213,1481]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Figure 1. Phyllophaga badbunnyi n. sp. A) Dorsal habitus. B) Lateral habitus." captionText-1="Figure 2. Genital capsule of Phyllophaga badbunnyi, n. sp. A) Lateral view. B) Dorsal view. C) Lateral view of aedeagus. Arrows indicate the finger-like dorsal projections of phallobase (sensu Morón 2003: 231)." captionText-2="Figure 3.Genital plates of Phyllophaga badbunnyi, n. sp. A) Superior and lateral plates. B) Inferior plates (the right broken)." captionText-3="Figure 4.Collecting localities of the new species. Circles: Phyllophaga badbunnyi n. sp. Cano. Square:Phyllophaga sechi n. sp. Cano." figureDoi-0="http://doi.org/10.5281/zenodo.5353726" figureDoi-1="http://doi.org/10.5281/zenodo.5353728" figureDoi-2="http://doi.org/10.5281/zenodo.5353732" figureDoi-3="http://doi.org/10.5281/zenodo.5353734" httpUri-0="https://zenodo.org/record/5353726/files/figure.png" httpUri-1="https://zenodo.org/record/5353728/files/figure.png" httpUri-2="https://zenodo.org/record/5353732/files/figure.png" httpUri-3="https://zenodo.org/record/5353734/files/figure.png" pageId="3" pageNumber="2">Fig. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E6522FFADFF8CFF29F9D86389B230" pageId="3" pageNumber="2" type="reference_group">
<paragraph id="8BFB36A9FFADFF8CFF29F9D86389B230" blockId="3.[224,1441,1545,1600]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB541187FFADFF8CFF29F9D86389B230" pageId="3" pageNumber="2">
<treatmentCitation id="0AE510B8FFADFF8CFF29F9D860CEB251" author="Moron MA &amp; Blas M." box="[224,752,1545,1569]" page="39" pageId="3" pageNumber="2" year="2006">
<taxonomicName id="4C444D2AFFADFF8CFF29F9D860CEB251" authority=": Moron and Blas (2006: 39" authorityName=": Moron and Blas" authorityYear="2006" box="[224,752,1545,1569]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="alvareztoroi">
<emphasis id="B930EABBFFADFF8CFF29F9D863E8B251" box="[224,470,1545,1569]" italics="true" pageId="3" pageNumber="2">Phyllophaga alvareztoroi</emphasis>
:
<bibRefCitation id="EFD54B58FFADFF8CFE2AF9D860CEB251" author="Moron MA &amp; Blas M." box="[483,752,1545,1569]" pageId="3" pageNumber="2" pagination="35 - 46" refId="ref7181" refString="Moron MA, Blas M. 2006. Dos especies nuevas de Phyllophaga del grupo &quot; Schizorhina &quot; (Coleoptera: Melolonthidae: Melolonthinae) de Chiapas, Mexico. Folia Entomologica Mexicana 45: 35 - 46." type="journal article" year="2006">Morón and Blas (2006: 39</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, in part);
<treatmentCitation id="0AE510B8FFADFF8CFC9EF9D866B4B251" author="Schuster JC &amp; Cano EB" box="[855,1162,1545,1569]" page="63" pageId="3" pageNumber="2" year="2006">
<bibRefCitation id="EFD54B58FFADFF8CFC9EF9D866B4B251" author="Schuster JC &amp; Cano EB" box="[855,1162,1545,1569]" pageId="3" pageNumber="2" pagination="57 - 70" refId="ref7365" refString="Schuster JC, Cano EB. 2006. What can Scarabaeoidea contribute to the knowledge of the biogeography of Guatemala? Coleopterists Society Monograph 5: 57 - 70." type="journal article" year="2006">Schuster and Cano (2006: 63)</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE510B8FFADFF8CFB5CF9D86364B230" author="Gomez B &amp; Moron MA" page="212" pageId="3" pageNumber="2" year="2010">
<bibRefCitation id="EFD54B58FFADFF8CFB5CF9D86364B230" author="Gomez B &amp; Moron MA" pageId="3" pageNumber="2" pagination="207 - 225" refId="ref6909" refString="Gomez B, Moron MA. 2010. Los escarabajos de bosques de niebla de Chiapas (Coleoptera: Melolonthidae). p. 207 - 225. In: Perez MA, Tejeda C, Silva E (eds.). Los bosques mesofilos de montana en Chiapas. Situacion actual, diversidad y conservacion. Universidad de Ciencias y Artes de Chiapas: Mexico." type="book chapter" year="2010">Gómez and Morón (2010: 212213</bibRefCitation>
</treatmentCitation>
, in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35E6522FFADFF8AFF09F98166C4B53B" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8BFB36A9FFADFF8CFF09F9816732B21B" blockId="3.[192,1292,1616,1644]" box="[192,1292,1616,1644]" pageId="3" pageNumber="2">
<emphasis id="B930EABBFFADFF8CFF09F981639DB21A" bold="true" box="[192,419,1616,1642]" pageId="3" pageNumber="2">Material examined.</emphasis>
<specimenCount id="9D42FD20FFADFF8CFE60F980607FB21B" box="[425,577,1617,1644]" pageId="3" pageNumber="2" type="generic">35 specimens</specimenCount>
, plus one not seen (species figured in
<bibRefCitation id="EFD54B58FFADFF8CFC2CF98066C1B21B" author="Moron MA &amp; Blas M." box="[997,1279,1617,1643]" pageId="3" pageNumber="2" pagination="35 - 46" refId="ref7181" refString="Moron MA, Blas M. 2006. Dos especies nuevas de Phyllophaga del grupo &quot; Schizorhina &quot; (Coleoptera: Melolonthidae: Melolonthinae) de Chiapas, Mexico. Folia Entomologica Mexicana 45: 35 - 46." type="journal article" year="2006">Morón and Blas 2006: 39</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB36A9FFADFF8CFF09F9AA6668B2C8" blockId="3.[191,1441,1659,1927]" pageId="3" pageNumber="2">
<emphasis id="B930EABBFFADFF8CFF09F9AA6357B2E5" bold="true" box="[192,361,1659,1685]" pageId="3" pageNumber="2">Type material.</emphasis>
<materialsCitation id="3B2C3CF4FFADFF8CFEA7F9AA6668B2C8" collectingDate="2002-06-14" collectionCode="UVG, USAC" collectorName="Col. E. Cano" country="Guatemala" latitude="14.546818" location="Reserva" longLatPrecision="1" longitude="-91.19364" municipality="Santa Barbara" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Suchitepequez" typeStatus="holotype">
<typeStatus id="54FF880BFFADFF8CFEA7F9AA63E8B2E5" box="[366,470,1659,1685]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F3537639FFADFF8CFDD6F9AA60F4B2E5" box="[543,714,1659,1685]" name="Guatemala" pageId="3" pageNumber="2">GUATEMALA</collectingCountry>
:
<collectingRegion id="4980F84BFFADFF8CFD1DF9AA6146B2E5" box="[724,888,1659,1685]" country="Guatemala" name="Suchitepequez" pageId="3" pageNumber="2">Suchitepéquez</collectingRegion>
,
<collectingMunicipality id="6B9FACD3FFADFF8CFC4BF9AA661FB2E5" box="[898,1057,1659,1685]" pageId="3" pageNumber="2">Santa Bárbara</collectingMunicipality>
, Fca. Panamá,
<location id="8E9B6072FFADFF8CFB0DF9AD6722B2E6" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059:8E9B6072FFADFF8CFB0DF9AD6722B2E6" box="[1220,1308,1660,1686]" country="Guatemala" latitude="14.546818" longLatPrecision="1" longitude="-91.19364" municipality="Santa Barbara" name="Reserva" pageId="3" pageNumber="2" stateProvince="Suchitepequez">Reserva</location>
<collectionCode id="ED55AE6CFFADFF8CFAEBF9AA675EB2E5" box="[1314,1376,1659,1685]" country="Guatemala" httpUri="http://grbio.org/cool/vx15-6kjy" name="Universidad del Valle" pageId="3" pageNumber="2">UVG</collectionCode>
, 1500 msnm,
<geoCoordinate id="EE70506EFFADFF8CFEDDF94C63B6B2C8" box="[276,392,1693,1720]" degrees="14.546818" direction="north" orientation="latitude" pageId="3" pageNumber="2" precision="1" value="14.546818">14.546818</geoCoordinate>
,
<geoCoordinate id="EE70506EFFADFF8CFE5AF94C6026B2C7" box="[403,536,1693,1719]" degrees="91.193639" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="1" value="-91.19364">91.193639</geoCoordinate>
. Bosque nuboso.
<date id="FFFA1069FFADFF8CFD10F94F6169B2C8" box="[729,855,1694,1720]" pageId="3" pageNumber="2" value="2002-06-14">
<collectingDate id="EFBEE981FFADFF8CFD10F94F6169B2C8" box="[729,855,1694,1720]" pageId="3" pageNumber="2" value="2002-06-14">14 VI 2002</collectingDate>
</date>
.
<collectorName id="26B1537FFFADFF8CFCA8F94C61D0B2C8" box="[865,1006,1693,1720]" pageId="3" pageNumber="2">Col. E. Cano</collectorName>
(
<collectionCode id="ED55AE6CFFADFF8CFC34F94F6675B2C8" box="[1021,1099,1694,1720]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="3" pageNumber="2">USAC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFB36A9FFADFF8CFEC9F91167A5B28D" blockId="3.[191,1441,1659,1927]" pageId="3" pageNumber="2">
<materialsCitation id="3B2C3CF4FFADFF8CFEC9F91167A5B28D" collectingDate="2007-03-15" collectingDateMax="2007-03-18" collectingDateMin="2007-03-15" collectionCode="UVG, USAC" country="Guatemala" elevation="1600" location="Santa Barbara" pageId="3" pageNumber="2" specimenCount="1" stateProvince="Suchitepequez" typeStatus="allotype">
<typeStatus id="54FF880BFFADFF8CFEC9F911635EB2AA" box="[256,352,1728,1754]" pageId="3" pageNumber="2" type="allotype">Allotype</typeStatus>
female.
<collectingCountry id="F3537639FFADFF8CFE75F9116059B2AA" box="[444,615,1728,1754]" name="Guatemala" pageId="3" pageNumber="2">GUATEMALA</collectingCountry>
:
<collectingRegion id="4980F84BFFADFF8CFDB9F911612AB2AA" box="[624,788,1728,1754]" country="Guatemala" name="Suchitepequez" pageId="3" pageNumber="2">Suchitepéquez</collectingRegion>
,
<location id="8E9B6072FFADFF8CFCD7F9116182B2AA" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059:8E9B6072FFADFF8CFCD7F9116182B2AA" box="[798,956,1728,1754]" country="Guatemala" name="Santa Barbara" pageId="3" pageNumber="2" stateProvince="Suchitepequez">Santa Bárbara</location>
, Res. Refugio Quetzal
<collectionCode id="ED55AE6CFFADFF8CFB7DF91166CCB2AA" box="[1204,1266,1728,1754]" country="Guatemala" httpUri="http://grbio.org/cool/vx15-6kjy" name="Universidad del Valle" pageId="3" pageNumber="2">UVG</collectionCode>
,
<date id="FFFA1069FFADFF8CFB35F91167A3B2AA" box="[1276,1437,1728,1755]" pageId="3" pageNumber="2" value="2007-03-15" valueMax="2007-03-18" valueMin="2007-03-15">
<collectingDate id="EFBEE981FFADFF8CFB35F91167A3B2AA" box="[1276,1437,1728,1755]" pageId="3" pageNumber="2" value="2007-03-15" valueMax="2007-03-18" valueMin="2007-03-15">15-18 III 2007</collectingDate>
</date>
.
<quantity id="4CBC9B4CFFADFF8CFF76F9336332B28C" box="[191,268,1762,1788]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="2" unit="m" value="1600.0">
<elevation id="0069D19AFFADFF8CFF76F9336332B28C" box="[191,268,1762,1788]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="2" unit="m" value="1600.0">1600m</elevation>
</quantity>
[
<quantity id="4CBC9B4CFFADFF8CFED0F9336350B28D" box="[281,366,1762,1789]" metricMagnitude="3" metricUnit="m" metricValue="1.373" pageId="3" pageNumber="2" unit="m" value="1373.0">
<elevation id="0069D19AFFADFF8CFED0F9336350B28D" box="[281,366,1762,1789]" metricMagnitude="3" metricUnit="m" metricValue="1.373" pageId="3" pageNumber="2" unit="m" value="1373.0">1373m</elevation>
</quantity>
]. Latitud 14.5417598494, Longitud 91.1972949818. [Cols.] Camposeco, Pérez, Cortéz (
<collectionCode id="ED55AE6CFFADFF8CFA8BF93267AEB28D" box="[1346,1424,1763,1789]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="3" pageNumber="2">USAC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFB36A9FFADFF8BFEC9F8D460B1B3F6" blockId="3.[191,1441,1659,1927]" lastBlockId="4.[192,1440,1698,1927]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<materialsCitation id="3B2C3CF4FFADFF8CFEC9F8D461EFB36F" box="[256,977,1797,1823]" collectingDate="2002-06-14" collectionCode="USAC" collectorName="Same" country="Guatemala" latitude="14.546818" location="Reserva" longLatPrecision="1" longitude="-91.19364" municipality="Santa Barbara" pageId="3" pageNumber="2" specimenCount="39" specimenCount-female="15" specimenCount-male="24" stateProvince="Suchitepequez" typeStatus="paratype">
<typeStatus id="54FF880BFFADFF8CFEC9F8D46352B36F" box="[256,364,1797,1823]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D42FD20FFADFF8CFEB2F8D4638CB36E" box="[379,434,1797,1823]" pageId="3" pageNumber="2" type="male">21 ♂</specimenCount>
,
<specimenCount id="9D42FD20FFADFF8CFE76F8D463CBB36E" box="[447,501,1797,1823]" pageId="3" pageNumber="2" type="female">13 ♀</specimenCount>
).
<collectorName id="26B1537FFFADFF8CFDC5F8D46076B36F" box="[524,584,1797,1823]" pageId="3" pageNumber="2">Same</collectorName>
data as holotype (
<specimenCount id="9D42FD20FFADFF8CFCDAF8D46102B36E" box="[787,828,1797,1823]" pageId="3" pageNumber="2" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D42FD20FFADFF8CFC80F8D4614FB36E" box="[841,881,1797,1823]" pageId="3" pageNumber="2" type="female">2 ♀</specimenCount>
,
<collectionCode id="ED55AE6CFFADFF8CFCB7F8D461F4B36F" box="[894,970,1797,1823]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="3" pageNumber="2">USAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2C3CF4FFADFF8CFC15F8D467A2B36F" box="[988,1436,1797,1823]" collectingDate="2007-03-15" collectingDateMax="2007-03-18" collectingDateMin="2007-03-15" collectionCode="UVGC" country="Guatemala" elevation="1600" isTailEnumeration="true" location="Santa Barbara" pageId="3" pageNumber="2" specimenCount="17" specimenCount-female="8" specimenCount-male="9" stateProvince="Suchitepequez" typeStatus="paratype">
same data as allotype (
<specimenCount id="9D42FD20FFADFF8CFB1FF8D4673EB36E" box="[1238,1280,1797,1823]" pageId="3" pageNumber="2" type="male">9 ♂</specimenCount>
,
<specimenCount id="9D42FD20FFADFF8CFAC4F8D4670BB36E" box="[1293,1333,1797,1823]" pageId="3" pageNumber="2" type="female">8 ♀</specimenCount>
,
<collectionCode id="ED55AE6CFFADFF8CFA8BF8D467AAB36F" box="[1346,1428,1797,1823]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="3" pageNumber="2">UVGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2C3CF4FFADFF8CFF09F8F9611EB331" box="[192,800,1831,1858]" collectingDate="2018-08" collectingDateMax="2007-03-18" collectingDateMin="2007-03-15" collectionCode="USAC" country="Guatemala" elevation="1600" location="Santa Barbara" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Suchitepequez" typeStatus="paratype">
same data as allotype except
<date id="FFFA1069FFADFF8CFDC6F8F96041B331" box="[527,639,1831,1858]" pageId="3" pageNumber="2" value="2018-08">
<collectingDate id="EFBEE981FFADFF8CFDC6F8F96041B331" box="[527,639,1831,1858]" pageId="3" pageNumber="2" value="2018-08">VIII 2018</collectingDate>
</date>
(
<specimenCount id="9D42FD20FFADFF8CFD5AF8F66083B331" box="[659,701,1831,1857]" pageId="3" pageNumber="2" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED55AE6CFFADFF8CFD04F8F66127B331" box="[717,793,1831,1857]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="3" pageNumber="2">USAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2C3CF4FFADFF8CFCE7F8F967A2B331" box="[814,1436,1831,1858]" collectingDate="2007-02-20" collectingDateMax="2007-03-18" collectingDateMin="2007-03-15" collectionCode="UVGC" country="Guatemala" elevation="1600" location="Santa Barbara" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Suchitepequez" typeStatus="paratype">
same data as allotype except
<date id="FFFA1069FFADFF8CFBB4F8F666CDB331" box="[1149,1267,1831,1858]" pageId="3" pageNumber="2" value="2007-02-20">
<collectingDate id="EFBEE981FFADFF8CFBB4F8F666CDB331" box="[1149,1267,1831,1858]" pageId="3" pageNumber="2" value="2007-02-20">20 II 2007</collectingDate>
</date>
(
<specimenCount id="9D42FD20FFADFF8CFAC1F8F6670CB331" box="[1288,1330,1831,1857]" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED55AE6CFFADFF8CFA8BF8F667AAB331" box="[1346,1428,1831,1857]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="3" pageNumber="2">UVGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2C3CF4FFADFF8CFF09F89B6667B314" box="[192,1113,1866,1892]" collectingDate="2013-05-11" collectingDateMax="2007-03-18" collectingDateMin="2007-03-15" collectionCode="V, UVGC" collectorName="Col. Monzon &amp; Camposeco" country="Guatemala" elevation="1600" location="Santa Barbara" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Suchitepequez" typeStatus="paratype">
same data as allotype except
<date id="FFFA1069FFADFF8CFDCDF89B6047B314" box="[516,633,1866,1892]" pageId="3" pageNumber="2" value="2013-05-11">
<collectingDate id="EFBEE981FFADFF8CFDCDF89B6047B314" box="[516,633,1866,1892]" pageId="3" pageNumber="2" value="2013-05-11">
11
<collectionCode id="ED55AE6CFFADFF8CFDEFF89B6004B314" box="[550,570,1866,1892]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="2" type="Museum">V</collectionCode>
2013
</collectingDate>
</date>
,
<collectorName id="26B1537FFFADFF8CFD4DF89B6127B314" box="[644,793,1866,1892]" pageId="3" pageNumber="2">Col. Monzón</collectorName>
y
<collectorName id="26B1537FFFADFF8CFCFAF89B6188B314" box="[819,950,1866,1892]" pageId="3" pageNumber="2">Camposeco</collectorName>
(
<specimenCount id="9D42FD20FFADFF8CFC00F89B61CFB313" box="[969,1009,1866,1892]" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED55AE6CFFADFF8CFC37F89B666EB314" box="[1022,1104,1866,1892]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="3" pageNumber="2">UVGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2C3CF4FFADFF8BFBAAF89B671AB2AE" collectingDate="2000-04-05" collectingDateMax="2008-05-10" collectingDateMin="2000-04-05" collectionCode="USAC, UVGC" collectorName="San Rafael Pie de la Cuesta &amp; El Bojonal &amp; Col. Monzon &amp; Camposeco" country="Guatemala" county="Finca Moca" elevation="1600" lastPageId="4" lastPageNumber="3" latitude="14.9459" location="Colector Jose Monzon" longLatPrecision="7" longitude="-91.8806" municipality="Volcan Atitlan" pageId="3" pageNumber="2" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Suchitepequez" typeStatus="paratype">
same data as allotype except
<date id="FFFA1069FFADFF8CFF08F8BD635AB3F6" box="[193,356,1900,1927]" pageId="3" pageNumber="2" value="2007-04-20" valueMax="2007-04-30" valueMin="2007-04-20">
<collectingDate id="EFBEE981FFADFF8CFF08F8BD635AB3F6" box="[193,356,1900,1927]" pageId="3" pageNumber="2" value="2007-04-20" valueMax="2007-04-30" valueMin="2007-04-20">2030 IV 2007</collectingDate>
</date>
(
<specimenCount id="9D42FD20FFADFF8CFEBAF8BD63A4B3F6" box="[371,410,1900,1926]" pageId="3" pageNumber="2" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D42FD20FFADFF8CFE6EF8BD63F0B3F6" box="[423,462,1900,1926]" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
,
<collectionCode id="ED55AE6CFFADFF8CFE12F8BD6019B3F6" box="[475,551,1900,1926]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="3" pageNumber="2">USAC</collectionCode>
).
<collectingCountry id="F3537639FFADFF8CFDF1F8BD60DDB3F6" box="[568,739,1900,1926]" name="Guatemala" pageId="3" pageNumber="2">GUATEMALA</collectingCountry>
:
<collectingRegion id="4980F84BFFADFF8CFD25F8BD61AEB3F6" box="[748,912,1900,1926]" country="Guatemala" name="Suchitepequez" pageId="3" pageNumber="2">Suchitepéquez</collectingRegion>
,
<collectingCounty id="629A4E25FFADFF8CFC53F8BD6620B3F6" box="[922,1054,1900,1926]" pageId="3" pageNumber="2">Finca Mocá</collectingCounty>
,
<collectingMunicipality id="6B9FACD3FFADFF8CFBE1F8BD66F5B3F6" box="[1064,1227,1900,1926]" pageId="3" pageNumber="2">Volcán Atitlán</collectingMunicipality>
,
<quantity id="4CBC9B4CFFADFF8CFB1CF8BD671AB3F6" box="[1237,1316,1900,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="2" unit="m" value="1600.0">
<elevation id="0069D19AFFADFF8CFB1CF8BD671AB3F6" box="[1237,1316,1900,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="2" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="FFFA1069FFADFF8CFAE7F8BD67A2B3F6" box="[1326,1436,1900,1927]" pageId="3" pageNumber="2" value="2000-04-05">
<collectingDate id="EFBEE981FFADFF8CFAE7F8BD67A2B3F6" box="[1326,1436,1900,1927]" pageId="3" pageNumber="2" value="2000-04-05">5 IV 2000</collectingDate>
</date>
,
<location id="8E9B6072FFAAFF8BFF09F9736382B2CD" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059:8E9B6072FFAAFF8BFF09F9736382B2CD" box="[192,444,1698,1725]" country="Guatemala" county="Finca Moca" latitude="14.9459" longLatPrecision="7" longitude="-91.8806" municipality="Volcan Atitlan" name="Colector Jose Monzon" pageId="4" pageNumber="3" stateProvince="Suchitepequez">Colector José Monzón</location>
(
<specimenCount id="9D42FD20FFAAFF8BFE19F97263C7B2CC" box="[464,505,1699,1725]" pageId="4" pageNumber="3" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED55AE6CFFAAFF8BFDCEF9726064B2CD" box="[519,602,1699,1725]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="4" pageNumber="3">UVGC</collectionCode>
).
<collectingCountry id="F3537639FFAAFF8BFDA4F9726126B2CD" box="[621,792,1699,1725]" name="Guatemala" pageId="4" pageNumber="3">GUATEMALA</collectingCountry>
:
<collectingRegion id="4980F84BFFAAFF8BFCEDF9726194B2CD" box="[804,938,1699,1725]" country="Guatemala" name="San Marcos" pageId="4" pageNumber="3">San Marcos</collectingRegion>
,
<collectorName id="26B1537FFFAAFF8BFC7FF97266D5B2CD" box="[950,1259,1698,1725]" pageId="4" pageNumber="3">
<location id="8E9B6072FFAAFF8BFC7FF97266D5B2CD" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059:8E9B6072FFAAFF8BFC7FF97266D5B2CD" box="[950,1259,1698,1725]" country="Guatemala" county="Finca Moca" latitude="14.9459" longLatPrecision="7" longitude="-91.8806" municipality="Volcan Atitlan" name="San Rafael Pie de la Cuesta" pageId="4" pageNumber="3" stateProvince="Suchitepequez">San Rafael Pié de la Cuesta</location>
</collectorName>
, camino
<location id="8E9B6072FFAAFF8BFA9DF972633FB2AE" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059:8E9B6072FFAAFF8BFA9DF972633FB2AE" country="Guatemala" county="Finca Moca" latitude="14.9459" longLatPrecision="7" longitude="-91.8806" municipality="Volcan Atitlan" name="Fraternidad" pageId="4" pageNumber="3" stateProvince="Suchitepequez">Fraternidad</location>
a
<collectorName id="26B1537FFFAAFF8BFED5F91563AAB2AE" box="[284,404,1732,1758]" pageId="4" pageNumber="3">
<location id="8E9B6072FFAAFF8BFED5F91563AAB2AE" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059:8E9B6072FFAAFF8BFED5F91563AAB2AE" box="[284,404,1732,1758]" country="Guatemala" county="Finca Moca" latitude="14.9459" longLatPrecision="7" longitude="-91.8806" municipality="Volcan Atitlan" name="El Bojonal" pageId="4" pageNumber="3" stateProvince="Suchitepequez">El Bojonal</location>
</collectorName>
,
<quantity id="4CBC9B4CFFAAFF8BFE56F91563D0B2AE" box="[415,494,1732,1758]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="3" unit="m" value="1600.0">
<elevation id="0069D19AFFAAFF8BFE56F91563D0B2AE" box="[415,494,1732,1758]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="3" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="FFFA1069FFAAFF8BFE33F9156051B2AE" box="[506,623,1732,1759]" pageId="4" pageNumber="3" value="2008-05-10">
<collectingDate id="EFBEE981FFAAFF8BFE33F9156051B2AE" box="[506,623,1732,1759]" pageId="4" pageNumber="3" value="2008-05-10">10 V 2008</collectingDate>
</date>
,
<geoCoordinate id="EE70506EFFAAFF8BFDB2F91560EAB2AF" box="[635,724,1732,1759]" degrees="14.9459" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="5" value="14.9459">14.9459</geoCoordinate>
,
<geoCoordinate id="EE70506EFFAAFF8BFD29F9156177B2AE" box="[736,841,1732,1758]" degrees="91.8806" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="5" value="-91.8806">91.8806</geoCoordinate>
,
<collectorName id="26B1537FFFAAFF8BFC9CF91561D4B2AE" box="[853,1002,1732,1758]" pageId="4" pageNumber="3">Col. Monzón</collectorName>
y
<collectorName id="26B1537FFFAAFF8BFBCCF91566B6B2AE" box="[1029,1160,1732,1758]" pageId="4" pageNumber="3">Camposeco</collectorName>
(
<specimenCount id="9D42FD20FFAAFF8BFB53F91566FDB2AE" box="[1178,1219,1732,1758]" pageId="4" pageNumber="3" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED55AE6CFFAAFF8BFB18F9156723B2AE" box="[1233,1309,1732,1758]" country="Australia" httpUri="http://grbio.org/cool/tsp0-m621" name="University of Western Australia" pageId="4" pageNumber="3">USAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2C3CF4FFAAFF8BFAF9F91467A2B370" collectingDate="2005-08-28" collectingDateMax="2005-08-31" collectingDateMin="2005-08-28" collectionCode="UVGC" collectorName="de Fraternidad-Bojonal &amp; Colector Jose Monzon Sierra" country="Guatemala" county="Finca Moca" elevation="1600" latitude="14.9459" location="Colector Jose Monzon" longLatPrecision="7" longitude="-91.8806" municipality="Volcan Atitlan" pageId="4" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Suchitepequez" typeStatus="paratype">
same data except camino
<collectorName id="26B1537FFFAAFF8BFEA7F9376047B370" box="[366,633,1766,1792]" pageId="4" pageNumber="3">de Fraternidad-Bojonal</collectorName>
,
<date id="FFFA1069FFAAFF8BFD4FF937617AB370" box="[646,836,1765,1792]" pageId="4" pageNumber="3" value="2005-08-28" valueMax="2005-08-31" valueMin="2005-08-28">
<collectingDate id="EFBEE981FFAAFF8BFD4FF937617AB370" box="[646,836,1765,1792]" pageId="4" pageNumber="3" value="2005-08-28" valueMax="2005-08-31" valueMin="2005-08-28">2831 VIII 2005</collectingDate>
</date>
,
<quantity id="4CBC9B4CFFAAFF8BFC98F937619EB370" box="[849,928,1766,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="3" unit="m" value="1600.0">
<elevation id="0069D19AFFAAFF8BFC98F937619EB370" box="[849,928,1766,1792]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="3" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<collectorName id="26B1537FFFAAFF8BFC64F93766C9B370" box="[941,1271,1766,1792]" pageId="4" pageNumber="3">Colector José Monzón Sierra</collectorName>
(
<specimenCount id="9D42FD20FFAAFF8BFAC3F937670DB28F" box="[1290,1331,1766,1792]" pageId="4" pageNumber="3" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED55AE6CFFAAFF8BFA8BF93767AAB370" box="[1346,1428,1766,1792]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="4" pageNumber="3">UVGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2C3CF4FFAAFF8BFF09F8D960B5B3F6" collectingDate="2001-03-20" collectingDateMax="2010-10" collectingDateMin="2001-03-20" collectionCode="UV, UVGC, V" collectorName="El Bojonal &amp; F. Camposeco &amp; Col." country="Mexico" county="Finca Moca" elevation="1600" latitude="15.557778" location="Galdamez" longLatPrecision="1" longitude="-92.428055" municipality="Metalarc" pageId="4" pageNumber="3" specimenCount="2" specimenCount-male="2" stateProvince="Suchitepequez" typeStatus="paratype">
same data except aldea
<collectorName id="26B1537FFFAAFF8BFE0DF8D66004B351" box="[452,570,1799,1825]" pageId="4" pageNumber="3">El Bojonal</collectorName>
,
<geoCoordinate id="EE70506EFFAAFF8BFD8CF8D66090B352" box="[581,686,1799,1826]" degrees="14" direction="north" minutes="57" orientation="latitude" pageId="4" pageNumber="3" precision="925" value="14.95">14° 57 N</geoCoordinate>
,
<geoCoordinate id="EE70506EFFAAFF8BFD71F8D6611AB352" box="[696,804,1799,1826]" degrees="91" direction="west" minutes="53" orientation="longitude" pageId="4" pageNumber="3" precision="925" value="-91.88333">91° 53 W</geoCoordinate>
,
<quantity id="4CBC9B4CFFAAFF8BFCE7F8D66143B351" box="[814,893,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="3" unit="m" value="1600.0">
<elevation id="0069D19AFFAAFF8BFCE7F8D66143B351" box="[814,893,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="3" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="FFFA1069FFAAFF8BFC4EF8D961E6B351" box="[903,984,1799,1826]" pageId="4" pageNumber="3" value="2010-10">
<collectingDate id="EFBEE981FFAAFF8BFC4EF8D961E6B351" box="[903,984,1799,1826]" pageId="4" pageNumber="3" value="2010-10">X 2010</collectingDate>
</date>
, bosque nuboso, luz
<collectionCode id="ED55AE6CFFAAFF8BFB76F8D666D9B351" box="[1215,1255,1799,1825]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="4" pageNumber="3">UV</collectionCode>
+
<collectingMunicipality id="6B9FACD3FFAAFF8BFB3FF8D66764B351" box="[1270,1370,1799,1825]" pageId="4" pageNumber="3">Metalarc</collectingMunicipality>
400w,
<collectorName id="26B1537FFFAAFF8BFF09F8F86363B333" box="[192,349,1833,1859]" pageId="4" pageNumber="3">F. Camposeco</collectorName>
(
<specimenCount id="9D42FD20FFAAFF8BFEB9F8F863A7B332" box="[368,409,1833,1859]" pageId="4" pageNumber="3" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED55AE6CFFAAFF8BFE6EF8F863C4B333" box="[423,506,1833,1859]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="4" pageNumber="3">UVGC</collectionCode>
).
<collectingCountry id="F3537639FFAAFF8BFDC4F8FA6042B337" box="[525,636,1835,1863]" name="Mexico" pageId="4" pageNumber="3">MÉXICO</collectingCountry>
:
<taxonomicName id="4C444D2AFFAAFF8BFD4EF8F86728B333" authority=", Siltepec, Reserva Cordon Pico, Barrio Las Moras" authorityName="Siltepec, Reserva Cordon Pico, Barrio Las Moras" box="[647,1302,1833,1859]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="chiapas">Chiapas, Siltepec, Reserva Cordón Pico, Barrio Las Moras</taxonomicName>
[
<geoCoordinate id="EE70506EFFAAFF8BFAEDF8F867A2B334" box="[1316,1436,1833,1860]" degrees="15.557778" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="15.557778">15.557778</geoCoordinate>
,
<geoCoordinate id="EE70506EFFAAFF8BFF08F89B6378B315" box="[193,326,1866,1893]" degrees="92.428056" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-92.428055">92.428056</geoCoordinate>
;
<quantity id="4CBC9B4CFFAAFF8BFE9BF89B639FB315" box="[338,417,1866,1893]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="3" unit="m" value="1750.0">
<elevation id="0069D19AFFAAFF8BFE9BF89B639FB315" box="[338,417,1866,1893]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="3" unit="m" value="1750.0">1750m</elevation>
</quantity>
; cloud forest],
<date id="FFFA1069FFAAFF8BFD82F89A60F5B315" box="[587,715,1867,1893]" pageId="4" pageNumber="3" value="2001-03-20">
<collectingDate id="EFBEE981FFAAFF8BFD82F89A60F5B315" box="[587,715,1867,1893]" pageId="4" pageNumber="3" value="2001-03-20">20 III 2001</collectingDate>
</date>
,
<collectorName id="26B1537FFFAAFF8BFD11F89B6139B314" box="[728,775,1866,1892]" pageId="4" pageNumber="3">Col.</collectorName>
<collectionCode id="ED55AE6CFFAAFF8BFCC7F89A611FB315" box="[782,801,1867,1893]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="3" type="Museum">V</collectionCode>
.
<location id="8E9B6072FFAAFF8BFCE4F89B61A0B314" LSID="urn:lsid:plazi:treatment:03ED87BFFFADFF86FF09FA6A6090B059:8E9B6072FFAAFF8BFCE4F89B61A0B314" box="[813,926,1866,1892]" country="Mexico" latitude="15.557778" longLatPrecision="1" longitude="-92.428055" municipality="Metalarc" name="Galdamez" pageId="4" pageNumber="3">Galdamez</location>
(
<specimenCount id="9D42FD20FFAAFF8BFC78F89A61E7B314" box="[945,985,1867,1893]" pageId="4" pageNumber="3" type="male">1 ♂</specimenCount>
, figured in
<bibRefCitation id="EFD54B58FFAAFF8BFB96F89A67A2B315" author="Moron MA &amp; Blas M." box="[1119,1436,1866,1893]" pageId="4" pageNumber="3" pagination="35 - 46" refId="ref7181" refString="Moron MA, Blas M. 2006. Dos especies nuevas de Phyllophaga del grupo &quot; Schizorhina &quot; (Coleoptera: Melolonthidae: Melolonthinae) de Chiapas, Mexico. Folia Entomologica Mexicana 45: 35 - 46." type="journal article" year="2006">Morón and Blas (2006: 39))</bibRefCitation>
, officially at MXAL-IEXA but not located
</materialsCitation>
.
</paragraph>
<caption id="DF3B6621FFAAFF8BFF09F9856627B21E" ID-DOI="http://doi.org/10.5281/zenodo.5353726" ID-Zenodo-Dep="5353726" box="[192,1049,1620,1647]" httpUri="https://zenodo.org/record/5353726/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,268,1620,1646]" targetBox="[192,1440,230,1590]" targetPageId="4">
<paragraph id="8BFB36A9FFAAFF8BFF09F9856627B21E" blockId="4.[192,1049,1620,1647]" box="[192,1049,1620,1647]" pageId="4" pageNumber="3">
<emphasis id="B930EABBFFAAFF8BFF09F9856316B21E" bold="true" box="[192,296,1620,1646]" pageId="4" pageNumber="3">Figure 1.</emphasis>
<taxonomicName id="4C444D2AFFAAFF8BFEE7F985600EB21E" authority="Cano, 2020" authorityName="Cano" authorityYear="2020" box="[302,560,1620,1646]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="badbunnyi" status="sp. nov.">
<emphasis id="B930EABBFFAAFF8BFEE7F985600EB21E" box="[302,560,1620,1646]" italics="true" pageId="4" pageNumber="3">Phyllophaga badbunnyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357C0FFAAFF8BFDFFF984604FB21F" box="[566,625,1621,1647]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B930EABBFFAAFF8BFDBEF98560ABB21E" bold="true" box="[631,661,1620,1646]" pageId="4" pageNumber="3">A)</emphasis>
Dorsal habitus.
<emphasis id="B930EABBFFAAFF8BFC83F9856158B21E" bold="true" box="[842,870,1620,1646]" pageId="4" pageNumber="3">B)</emphasis>
Lateral habitus.
</paragraph>
</caption>
<paragraph id="8BFB36A9FFABFF8AFF09FF3F66C4B53B" blockId="5.[191,1440,237,331]" pageId="5" pageNumber="4">
<emphasis id="B930EABBFFABFF8AFF09FF3F6365B577" bold="true" box="[192,347,237,264]" pageId="5" pageNumber="4">Type locality.</emphasis>
<collectingCountry id="F3537639FFABFF8AFE96FF3F63E5B578" box="[351,475,238,264]" name="Guatemala" pageId="5" pageNumber="4">Guatemala</collectingCountry>
,
<collectingRegion id="4980F84BFFABFF8AFE2CFF3F60B8B578" box="[485,646,238,264]" country="Guatemala" name="Suchitepequez" pageId="5" pageNumber="4">Suchitepéquez</collectingRegion>
Dept., Santa Bárbara, Reserva Refugio del Quetzal,
<quantity id="4CBC9B4CFFABFF8AFB0CFF3C672AB578" box="[1221,1300,237,264]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="4" unit="m" value="1500.0">1500m</quantity>
, coordinates
<geoCoordinate id="EE70506EFFABFF8AFF76FEDE630DB55A" box="[191,307,271,298]" degrees="14.546818" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="14.546818">14.546818</geoCoordinate>
,
<geoCoordinate id="EE70506EFFABFF8AFEF7FEDE63FCB559" box="[318,450,271,297]" degrees="91.193639" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-91.19364">91.193639</geoCoordinate>
, cloud forest on the southern slopes of Atitlán volcano. Some specimens (including
<typeStatus id="54FF880BFFABFF8AFAAAFEDE62CCB53B" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
) were collected before the territorial changes for the establishment of the Biological Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C35E6522FFABFF8AFF09FE8B6699B66D" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BFB36A9FFABFF8AFF09FE8B6699B66D" blockId="5.[192,1441,346,541]" pageId="5" pageNumber="4">
<emphasis id="B930EABBFFABFF8AFF09FE8B6302B504" bold="true" box="[192,316,346,372]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="4C444D2AFFABFF8AFE8DFE8B6076B504" authority="Cano, 2020" authorityName="Cano" authorityYear="2020" box="[324,584,346,372]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="badbunnyi" status="sp. nov.">
<emphasis id="B930EABBFFABFF8AFE8DFE8B6076B504" box="[324,584,346,372]" italics="true" pageId="5" pageNumber="4">Phyllophaga badbunnyi</emphasis>
</taxonomicName>
<emphasis id="B930EABBFFABFF8AFD86FE8A60E7B504" bold="true" box="[591,729,346,373]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="A20357C0FFABFF8AFD86FE8A60E7B504" box="[591,729,346,373]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="137F2A2CFFABFF8AFD20FE8B6178B505" box="[745,838,346,373]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="4.[192,268,1620,1646]" captionStartId-1="6.[192,268,1607,1633]" captionStartId-2="7.[325,401,1474,1500]" captionTargetBox-0="[192,1440,230,1590]" captionTargetBox-1="[223,1406,221,1567]" captionTargetBox-2="[323,1312,217,1431]" captionTargetId-0="figure-14@4.[192,1440,213,1592]" captionTargetId-1="figure-14@6.[192,1440,211,1577]" captionTargetId-2="figure-175@7.[301,1331,213,1439]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Figure 1. Phyllophaga badbunnyi n. sp. A) Dorsal habitus. B) Lateral habitus." captionText-1="Figure 2. Genital capsule of Phyllophaga badbunnyi, n. sp. A) Lateral view. B) Dorsal view. C) Lateral view of aedeagus. Arrows indicate the finger-like dorsal projections of phallobase (sensu Morón 2003: 231)." captionText-2="Figure 3.Genital plates of Phyllophaga badbunnyi, n. sp. A) Superior and lateral plates. B) Inferior plates (the right broken)." figureDoi-0="http://doi.org/10.5281/zenodo.5353726" figureDoi-1="http://doi.org/10.5281/zenodo.5353728" figureDoi-2="http://doi.org/10.5281/zenodo.5353732" httpUri-0="https://zenodo.org/record/5353726/files/figure.png" httpUri-1="https://zenodo.org/record/5353728/files/figure.png" httpUri-2="https://zenodo.org/record/5353732/files/figure.png" pageId="5" pageNumber="4">Fig. 13</figureCitation>
) is similar to
<taxonomicName id="4C444D2AFFABFF8AFC2EFE8B66C9B504" authorityName=": Moron and Blas" authorityYear="2006" box="[999,1271,346,372]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="alvareztoroi">
<emphasis id="B930EABBFFABFF8AFC2EFE8B66C9B504" box="[999,1271,346,372]" italics="true" pageId="5" pageNumber="4">Phyllophaga alvareztoroi</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A2CFFABFF8AFACEFE8B6764B505" box="[1287,1370,346,373]" captionStart="Figure 7" captionStartId="14.[192,268,1799,1825]" captionTargetBox="[202,1412,329,1683]" captionTargetId="figure-73@14.[192,1440,328,1712]" captionTargetPageId="14" captionText="Figure 7. Genital capsule of species of Phyllophaga of Schizorhina subgroup 2, from Chiapas, México and Guatemala. A) P. acacoyahuana, dorsolateral view. B) P. alvareztoroi, dorsal view, redrawn from Morón and Blas (2006). C) P. ginigra, lateral view. D) P. javepacuana, lateral view. Arrows indicate the finger-like dorsal projections of phallobase (sensu Morón 2003: 231)." figureDoi="http://doi.org/10.5281/zenodo.5353742" httpUri="https://zenodo.org/record/5353742/files/figure.png" pageId="5" pageNumber="4">Fig. 7B</figureCitation>
) from which it can be separated by dorsal keel of tectum barely marked, tectum separated from basal piece by a strong constriction, parameres (lower projections of phallobase) thinner apically and superior genital plates of females with upper margin markedly dentate. Although genital plates of female are similar to those of
<emphasis id="B930EABBFFABFF8AFB17FE6E631BB58B" italics="true" pageId="5" pageNumber="4">
<taxonomicName id="4C444D2AFFABFF8AFB17FE6E631FB58B" authorityName="Moron" authorityYear="2003" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="javepacuana">Phyllophaga javepacuana</taxonomicName>
,
</emphasis>
both, males and females of the new species can be separated by absence of a row of setae along sides of abdomen and absence of a strong sulcus on prepygidium, between the posterior spiracles.
</paragraph>
</subSubSection>
<subSubSection id="C35E6522FFABFF86FF09FDFD6065B6CD" lastPageId="9" lastPageNumber="8" pageId="5" pageNumber="4" type="description">
<paragraph id="8BFB36A9FFABFF8AFF09FDFD612BB6FA" blockId="5.[191,1442,556,1927]" pageId="5" pageNumber="4">
<emphasis id="B930EABBFFABFF8AFF09FDFD6361B636" bold="true" box="[192,351,556,582]" pageId="5" pageNumber="4">Description.</emphasis>
Dorsal coloration shiny black with some bluish reflections, epipleuron and adjacent elytral border reddish-brown, ventral coloration of abdomen and metasternum dark reddish-brown; legs reddish-brown. Total length of body:
<quantity id="4CBC9B4CFFABFF8AFEB9FDBE63DDB6FA" box="[368,483,623,650]" metricMagnitude="-2" metricUnit="m" metricValue="2.405" pageId="5" pageNumber="4" unit="mm" value="24.05">24.05 mm</quantity>
. Humeral width
<quantity id="4CBC9B4CFFABFF8AFD57FDBE612CB6FA" box="[670,786,623,650]" metricMagnitude="-2" metricUnit="m" metricValue="1.0619999999999998" pageId="5" pageNumber="4" unit="mm" value="10.62">10.62 mm</quantity>
.
</paragraph>
<paragraph id="8BFB36A9FFABFF8AFEC9FD40662DB04E" blockId="5.[191,1442,556,1927]" pageId="5" pageNumber="4">
<emphasis id="B930EABBFFABFF8AFEC9FD406374B6DB" bold="true" box="[256,330,657,683]" pageId="5" pageNumber="4">Head.</emphasis>
Clypeus short, 3.75 times wider than long, anterior border deeply and widely emarginate, anterior angles markedly rounded and moderately elevated; clypeal disc glabrous, concave towards lateral sides and slightly convex centrally, with circular dense deep punctures separated by less than a diameter, giving a strong rugose aspect. Frontoclypeal suture marked and bisinuate, less marked medially. Frons 3.55 times wider than long, uniformly convex, with deep circular punctures giving a slightly rugose aspect, more abundant at disc and smaller and sparse toward sides and anterolateral areas, each puncture with a very minute whitish, decumbent seta (visible only at great magnification); vertex smooth. Antennae 10 segmented; segments 3 and 4 subcylindrical of equal length; segments 5 and 6 shorter and wide, with a rounded projection; segment 7 slightly shorter than 6 and very wide, with an acute projection; wide (length in most authors) of antennal club as the length of segments 35 combined, with segment 8 wider than 9 or 10. Interocular distance
<quantity id="4CBC9B4CFFABFF8AFBAFFC6E66F3B7AA" box="[1126,1229,959,986]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="5" pageNumber="4" unit="mm" value="10.8">10.8 mm</quantity>
, width of each eye
<quantity id="4CBC9B4CFFABFF8AFF0BFC306322B78B" box="[194,284,993,1019]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="4" unit="mm" value="2.1">2.1 mm</quantity>
. Ocular canthus short and narrow with 811 erect setae. Labrum bilobate, with anterior border deeply emarginate, with many erect setae on lateral borders. Labium longitudinally widely and deeply excavate, disc bare, with abundant long setae toward the sides, anterior border emarginate.
</paragraph>
<paragraph id="8BFB36A9FFABFF8AFEC9FB976049B178" blockId="5.[191,1442,556,1927]" pageId="5" pageNumber="4">
<emphasis id="B930EABBFFABFF8AFEC9FB9763B6B010" bold="true" box="[256,392,1094,1120]" pageId="5" pageNumber="4">Prothorax.</emphasis>
Pronotum convex, 1.8 times wider than long. Anterior angles almost straight, slightly acute; posterior angles slightly obtuse. Lateral borders forming a wide angle at the central portion; marginal border narrow and crenulate with scarce sparse setae; posterior border marked by an impression and evanescent at middle in front of scutellum and crenulate at sides. Disc with circular and deep punctures, irregularly distributed giving a rugose aspect, each puncture with a decumbent, whitish setae; punctures denser towards the sides and scarce medially and near the posterior border.
</paragraph>
<paragraph id="8BFB36A9FFABFF8AFEC9FADE616DB2CD" blockId="5.[191,1442,556,1927]" pageId="5" pageNumber="4">
<emphasis id="B930EABBFFABFF8AFEC9FADE63A1B159" bold="true" box="[256,415,1295,1321]" pageId="5" pageNumber="4">Pterothorax.</emphasis>
Scutellum wider than long (1.7:1) with deep punctures toward anterolateral portions, each puncture with a decumbent, whitish seta; middle line barely marked; anterior border declivous. Elytra almost parallel and almost the same width as posterior margin of pronotum (at the level of humeri), slightly wider from anterior 1/3 to apex; moderately rugose-punctured; punctures rounded and each with a minute seta (only visible at great magnification); each elytron 2.6 times longer than wide. A row of minute decumbent white setae is present on punctures at most anterior portion, between epipleuron and humeral callus. Elytral epipleura with scattered short yellow-whitish setae; epipleural keel progressively effacing from middle of third abdominal segment to apex. Sutural costa elevated and well-marked, with scarce and disperse punctures, each puncture with a minute, decumbent seta; two discal costae longitudinally transverse and arising near humeral callus, proximal narrowly marked at anterior 2/3 and more widely marked at posterior 1/3 where fused with apical callus, the second very narrow and barely marked; lateral costa behind humeral callus, narrowly marked. Humeral callus prominent and rounded; apical callus wide, rounded, moderately prominent. Pterosternum densely puncturate-rugose, each puncture with a long, yellowish setae, giving a hairy aspect.
</paragraph>
<paragraph id="8BFB36A9FFABFF89FEC9F9156049B3F6" blockId="5.[191,1442,556,1927]" lastBlockId="6.[191,1440,1731,1926]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
<emphasis id="B930EABBFFABFF8AFEC9F91563BCB2AE" bold="true" box="[256,386,1732,1758]" pageId="5" pageNumber="4">Abdomen.</emphasis>
Prepygidium with a small excavation separated from fifth abdominal segment of venter, at level of posterior spiracle. Pygidium convex and shiny, puncturate-rugose, punctures wider at disc, each puncture with a whitish minute seta (only visible at great magnification); disc with a longitudinal, irregular and smooth tumosity; apical border with each puncture containing a long, yellowish seta. Abdominal segments 24 widely convex; sides with moderately abundant small and shallow punctures, each containing a minute decumbent whitish seta; setose punctures scarce medially (small setae on segment 2). Abominal segmets 34 with a shallow, longitudinal, lateral impression. Abdominal segment 5 widely convex, except at posterior transversal, almost smooth concavity and densely rugose and setose-puncturate posterolateral shallow and rounded concavity; setae of concavity whitish and long, forming a brush when seen laterally; anterior border of posterolateral concavity excavate at level of impression of fourth segment. Medial portion of fifth segment with minute rugose-punctures, denser posteriorly before transverse concavity. Anal plate narrow, medially concave, with minute and shallow rugose-punctures, surface with some long and short setae.
</paragraph>
<caption id="DF3B6621FFA8FF89FF09F996672CB2F3" ID-DOI="http://doi.org/10.5281/zenodo.5353728" ID-Zenodo-Dep="5353728" httpUri="https://zenodo.org/record/5353728/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,268,1607,1633]" targetBox="[223,1406,221,1567]" targetPageId="6">
<paragraph id="8BFB36A9FFA8FF89FF09F996672CB2F3" blockId="6.[192,1440,1607,1667]" pageId="6" pageNumber="5">
<emphasis id="B930EABBFFA8FF89FF09F9966314B211" bold="true" box="[192,298,1607,1633]" pageId="6" pageNumber="5">Figure 2.</emphasis>
Genital capsule of
<taxonomicName id="4C444D2AFFA8FF89FDC1F9966135B211" authority="Cano, 2020" authorityName="Cano" authorityYear="2020" box="[520,779,1607,1633]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="badbunnyi" status="sp. nov.">
<emphasis id="B930EABBFFA8FF89FDC1F9966135B211" box="[520,779,1607,1633]" italics="true" pageId="6" pageNumber="5">Phyllophaga badbunnyi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20357C0FFA8FF89FCD0F9996168B212" box="[793,854,1608,1634]" pageId="6" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B930EABBFFA8FF89FCA9F9966140B211" bold="true" box="[864,894,1607,1633]" pageId="6" pageNumber="5">A)</emphasis>
Lateral view.
<emphasis id="B930EABBFFA8FF89FBD4F9966606B211" bold="true" box="[1053,1080,1607,1633]" pageId="6" pageNumber="5">B)</emphasis>
Dorsal view.
<emphasis id="B930EABBFFA8FF89FB1CF99666CFB211" bold="true" box="[1237,1265,1607,1633]" pageId="6" pageNumber="5">C)</emphasis>
Lateral view of aedeagus. Arrows indicate the finger-like dorsal projections of phallobase (
<emphasis id="B930EABBFFA8FF89FBC8F9B86603B2F2" box="[1025,1085,1641,1666]" italics="true" pageId="6" pageNumber="5">sensu</emphasis>
<bibRefCitation id="EFD54B58FFA8FF89FB8AF9B8673BB2F3" author="Moron MA" box="[1091,1285,1641,1667]" pageId="6" pageNumber="5" pagination="213 - 302" refId="ref7116" refString="Moron MA. 2003. Revision of the Phyllophaga s. s. schizorhina species group (Coleoptera: Melolonthidae: Melolonthinae). The Canadian Entomologist 135: 213 - 302." type="journal article" year="2003">Morón 2003: 231</bibRefCitation>
).
</paragraph>
</caption>
<caption id="DF3B6621FFA9FF88FE8CFA136088B18E" ID-DOI="http://doi.org/10.5281/zenodo.5353732" ID-Zenodo-Dep="5353732" httpUri="https://zenodo.org/record/5353732/files/figure.png" pageId="7" pageNumber="6" startId="7.[325,401,1474,1500]" targetBox="[323,1312,217,1431]" targetPageId="7">
<paragraph id="8BFB36A9FFA9FF88FE8CFA136088B18E" blockId="7.[323,1308,1474,1534]" pageId="7" pageNumber="6">
<emphasis id="B930EABBFFA9FF88FE8CFA136392B1AC" bold="true" box="[325,428,1474,1500]" pageId="7" pageNumber="6">Figure 3.</emphasis>
Genital plates of
<taxonomicName id="4C444D2AFFA9FF88FDAEFA136156B1AC" authority="Cano, 2020" authorityName="Cano" authorityYear="2020" box="[615,872,1474,1500]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="badbunnyi" status="sp. nov.">
<emphasis id="B930EABBFFA9FF88FDAEFA136156B1AC" box="[615,872,1474,1500]" italics="true" pageId="7" pageNumber="6">Phyllophaga badbunnyi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20357C0FFA9FF88FCBDFA126190B1AD" box="[884,942,1475,1501]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="B930EABBFFA9FF88FC7BFA1361EEB1AC" bold="true" box="[946,976,1474,1500]" pageId="7" pageNumber="6">A)</emphasis>
Superior and lateral plates.
<emphasis id="B930EABBFFA9FF88FAC8FA136722B1AC" bold="true" box="[1281,1308,1474,1500]" pageId="7" pageNumber="6">B)</emphasis>
Inferior plates (the right broken).
</paragraph>
</caption>
<paragraph id="8BFB36A9FFA9FF87FEC9F9E2661DB370" blockId="7.[191,1442,1587,1927]" lastBlockId="8.[191,1442,1665,1927]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<emphasis id="B930EABBFFA9FF88FEC9F9E2637EB23D" bold="true" box="[256,320,1587,1613]" pageId="7" pageNumber="6">Legs.</emphasis>
Protibiae tridentate, shorter than protarsus (1:1.13); protibial spur straight, with acute and slightly downward curved apex; longer than second segment of protarsus (1.2:1). Middle of external face of mesotibiae with a well-marked, crenulate, oblique, transverse keel, with 9 thorny setae arising from base in each crenulation. Proximal 1/3 of mesotibiae with small and incomplete keel, with 23 thorny setae. Mesotibial spurs articulate, almost straight and with acute and slightly curved, apex; both spurs almost with the same length. Metatibiae shorter than metatarsus (1:1.26); middle of external face with a well-marked, crenulate, oblique, transverse keel, with 8 thorny setae arising from base in each crenulation. Proximal 1/3 with only two thorny setae and two crenulations marked. Metatibial spurs articulate; superior elongated and slender, slightly curved and with rounded apex, longer than first segment of metatarsus; inferior spur 1/3 length of superior, markedly curved and with apex acute, shorter than first segment of metatarsus. Pro- and meso-tarsal segments 14 with small spine on distal border; second protarsal segment shorter than first and same length as the third; first meso-tarsal segment longer and wider than second. Second metatarsal segment longer than first and segments 35; meta-tarsal segments 14 with row of ventral thorny setae, forming comb. All tarsal claws divided and equal; basal tooth ventrally almost as long as apical tooth; basal tooth closer to basal process than to apical tooth.
</paragraph>
<caption id="DF3B6621FFA6FF87FF0BFA396340B254" ID-DOI="http://doi.org/10.5281/zenodo.5353734" ID-Zenodo-Dep="5353734" httpUri="https://zenodo.org/record/5353734/files/figure.png" pageId="8" pageNumber="7" startId="8.[194,270,1512,1538]" targetBox="[219,1427,222,1475]" targetPageId="8">
<paragraph id="8BFB36A9FFA6FF87FF0BFA396340B254" blockId="8.[192,1439,1512,1573]" pageId="8" pageNumber="7">
<emphasis id="B930EABBFFA6FF87FF0BFA396316B272" bold="true" box="[194,296,1512,1538]" pageId="8" pageNumber="7">Figure 4.</emphasis>
Collecting localities of the new species. Circles:
<taxonomicName id="4C444D2AFFA6FF87FCFFFA396606B272" authority="Cano, 2020" authorityName="Cano" authorityYear="2020" box="[822,1080,1512,1538]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="badbunnyi" status="sp. nov.">
<emphasis id="B930EABBFFA6FF87FCFFFA396606B272" box="[822,1080,1512,1538]" italics="true" pageId="8" pageNumber="7">Phyllophaga badbunnyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357C0FFA6FF87FBF5FA386648B273" box="[1084,1142,1513,1539]" pageId="8" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
Cano. Square:
<taxonomicName id="4C444D2AFFA6FF87FAD0FA3962CAB254" authority="Cano, 2020" authorityName="Cano" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="sechi" status="sp. nov.">
<emphasis id="B930EABBFFA6FF87FAD0FA3962CAB254" italics="true" pageId="8" pageNumber="7">Phyllophaga sechi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357C0FFA6FF87FF32F9DA6308B255" box="[251,310,1547,1573]" pageId="8" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
Cano.
</paragraph>
</caption>
<paragraph id="8BFB36A9FFA6FF86FEC9F8D6671EB562" blockId="8.[191,1442,1665,1927]" lastBlockId="9.[191,1440,214,274]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="B930EABBFFA6FF87FEC9F8D663F5B351" bold="true" box="[256,459,1799,1825]" pageId="8" pageNumber="7">Genital capsule.</emphasis>
Parameres in lateral view narrow, elongate and free at apex; angulate before thin, apical 1/4; basally fused and with pair of superior projections in lateral view in form of “U”, with expanded apex (
<figureCitation id="137F2A2CFFA6FF87FABCF8F862D8B315" captionStart="Figure 2" captionStartId="6.[192,268,1607,1633]" captionTargetBox="[223,1406,221,1567]" captionTargetId="figure-14@6.[192,1440,211,1577]" captionTargetPageId="6" captionText="Figure 2. Genital capsule of Phyllophaga badbunnyi, n. sp. A) Lateral view. B) Dorsal view. C) Lateral view of aedeagus. Arrows indicate the finger-like dorsal projections of phallobase (sensu Morón 2003: 231)." figureDoi="http://doi.org/10.5281/zenodo.5353728" httpUri="https://zenodo.org/record/5353728/files/figure.png" pageId="8" pageNumber="7">Fig. 2A</figureCitation>
). Tectum (
<figureCitation id="137F2A2CFFA6FF87FE95F89B63F1B315" box="[348,463,1866,1893]" captionStart="Figure 2" captionStartId="6.[192,268,1607,1633]" captionTargetBox="[223,1406,221,1567]" captionTargetId="figure-14@6.[192,1440,211,1577]" captionTargetPageId="6" captionText="Figure 2. Genital capsule of Phyllophaga badbunnyi, n. sp. A) Lateral view. B) Dorsal view. C) Lateral view of aedeagus. Arrows indicate the finger-like dorsal projections of phallobase (sensu Morón 2003: 231)." figureDoi="http://doi.org/10.5281/zenodo.5353728" httpUri="https://zenodo.org/record/5353728/files/figure.png" pageId="8" pageNumber="7">Fig. 2AB</figureCitation>
) separated from basal piece by constriction; dorsally with weakly marked wide keel, not extended over phallobase. Basal 2/3 of aedeagus ventrally membranous and dorsally sclerotized and furrowed; asymmetrical and partially coiled near level of apical 2/3 to apex; apical 1/3 containing a dorsal membranous tubular projection (
<figureCitation id="137F2A2CFFA7FF86FE89FF2963A8B562" box="[320,406,248,274]" captionStart="Figure 2" captionStartId="6.[192,268,1607,1633]" captionTargetBox="[223,1406,221,1567]" captionTargetId="figure-14@6.[192,1440,211,1577]" captionTargetPageId="6" captionText="Figure 2. Genital capsule of Phyllophaga badbunnyi, n. sp. A) Lateral view. B) Dorsal view. C) Lateral view of aedeagus. Arrows indicate the finger-like dorsal projections of phallobase (sensu Morón 2003: 231)." figureDoi="http://doi.org/10.5281/zenodo.5353728" httpUri="https://zenodo.org/record/5353728/files/figure.png" pageId="9" pageNumber="8">Fig. 2C</figureCitation>
). Length of genital capsule from apex of parameres to end of basal piece:
<quantity id="4CBC9B4CFFA7FF86FB0DFF296722B562" box="[1220,1308,248,274]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="9" pageNumber="8" unit="mm" value="6.9">6.9 mm</quantity>
.
</paragraph>
<paragraph id="8BFB36A9FFA7FF86FF09FEF363DDB67A" blockId="9.[192,1441,290,522]" pageId="9" pageNumber="8">
<emphasis id="B930EABBFFA7FF86FF09FEF3630AB54C" bold="true" box="[192,308,290,316]" pageId="9" pageNumber="8">
<typeStatus id="54FF880BFFA7FF86FF09FEF36311B54C" box="[192,303,290,316]" pageId="9" pageNumber="8" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to
<typeStatus id="54FF880BFFA7FF86FDD4FEF36041B54C" box="[541,639,290,316]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
except as follows: pygidium scarcely convex, puncturate-rugose with longitudinal tumosity scarcely marked; fifth abdominal segment more plain; anal plate long and plain, puncturate-rugose. Metatibial spurs widened with rounded apex. Inferior genital plates rounded and separated, with few scattered minute setae (extreme of right plate broken in
<typeStatus id="54FF880BFFA7FF86FCADFE5961FFB5D3" box="[868,961,392,419]" pageId="9" pageNumber="8" type="allotype">allotype</typeStatus>
); lateral plates present and irregularly oval; superior genital plates united at middle forming a longitudinal keel, apical border roughly dentate and with long setae, latero-basally excavated forming an apical lateral projection (
<figureCitation id="137F2A2CFFA7FF86FC22FE1C6613B597" box="[1003,1069,461,487]" captionStart="Figure 3" captionStartId="7.[325,401,1474,1500]" captionTargetBox="[323,1312,217,1431]" captionTargetId="figure-175@7.[301,1331,213,1439]" captionTargetPageId="7" captionText="Figure 3.Genital plates of Phyllophaga badbunnyi, n. sp. A) Superior and lateral plates. B) Inferior plates (the right broken)." figureDoi="http://doi.org/10.5281/zenodo.5353732" httpUri="https://zenodo.org/record/5353732/files/figure.png" pageId="9" pageNumber="8">Fig. 3</figureCitation>
). Total length of body:
<quantity id="4CBC9B4CFFA7FF86FAFFFE1C67A3B597" box="[1334,1437,461,488]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="9" pageNumber="8" unit="mm" value="24.5">24.5 mm</quantity>
. Humeral width:
<quantity id="4CBC9B4CFFA7FF86FEB0FE3E63E1B67A" box="[377,479,495,522]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="8" unit="mm" value="11.1">11.1 mm</quantity>
.
</paragraph>
<paragraph id="8BFB36A9FFA7FF86FF09FDC86065B6CD" blockId="9.[191,1442,537,701]" pageId="9" pageNumber="8">
<emphasis id="B930EABBFFA7FF86FF09FDC863F5B643" bold="true" box="[192,459,537,563]" pageId="9" pageNumber="8">
Variation in
<typeStatus id="54FF880BFFA7FF86FE9BFDC863F8B643" box="[338,454,537,563]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
.
</emphasis>
Males similar to
<typeStatus id="54FF880BFFA7FF86FD41FDC860D4B643" box="[648,746,537,563]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
except as follows: old specimens without minute seta on frons, pronotum and elytra. Two female, teneral specimens with 10 long, yellowish setae on middle of frons and
<specimenCount id="9D42FD20FFA7FF86FABFFDED6332B607" pageId="9" pageNumber="8" type="female">one female</specimenCount>
with tumosity of pygidium almost smooth. Males: total body length
<quantity id="4CBC9B4CFFA7FF86FBDAFD8F66F1B608" box="[1043,1231,606,632]" metricMagnitude="-2" metricUnit="m" metricValue="2.586" metricValueMax="2.774" metricValueMin="2.398" pageId="9" pageNumber="8" unit="mm" value="25.86" valueMax="27.74" valueMin="23.98">23.9827.74 mm</quantity>
(µ = 26.33, n = 6), humeral width 10.5412.21 (µ = 11.34, n = 6). Females: total body length
<quantity id="4CBC9B4CFFA7FF86FC2DFDAE6646B6EA" box="[996,1144,639,666]" metricMagnitude="-2" metricUnit="m" metricValue="2.749" metricValueMax="2.9" metricValueMin="2.598" pageId="9" pageNumber="8" unit="mm" value="27.49" valueMax="29.0" valueMin="25.98">25.98-29 mm</quantity>
(µ = 27.32, n = 6), humeral width 11.0512.04 (µ = 11.67, n = 6).
</paragraph>
</subSubSection>
<subSubSection id="C35E6522FFA7FF86FF09FD1D61A6B75A" pageId="9" pageNumber="8" type="etymology">
<paragraph id="8BFB36A9FFA7FF86FF09FD1D61A6B75A" blockId="9.[192,1440,716,811]" pageId="9" pageNumber="8">
<emphasis id="B930EABBFFA7FF86FF09FD1D637AB696" bold="true" box="[192,324,716,742]" pageId="9" pageNumber="8">Etymology.</emphasis>
Named after the Puerto Rican urban singer and composer Benito Antonio Martínez, known as “Bad Bunny”. During the first weeks of the quarantine due to the SARS-CoV-2 I worked on the description of this new species listening to old and new works of the inspirational bunny.
</paragraph>
</subSubSection>
<subSubSection id="C35E6522FFA7FF86FF09FCEB6090B059" pageId="9" pageNumber="8" type="biology_ecology">
<paragraph id="8BFB36A9FFA7FF86FF09FCEB6748B7E9" blockId="9.[192,1442,826,922]" pageId="9" pageNumber="8">
<emphasis id="B930EABBFFA7FF86FF09FCEB63D0B724" bold="true" box="[192,494,826,852]" pageId="9" pageNumber="8">Ecology and distribution.</emphasis>
This species is known from cloud forests between 1500 and 1750 meters elevation, located at the southern slopes of the Sierra Madre de
<collectingRegion id="4980F84BFFA7FF86FCDDFC8D6150B706" box="[788,878,860,886]" country="Mexico" name="Chiapas" pageId="9" pageNumber="8">Chiapas</collectingRegion>
and Guatemalan western volcanoes (
<figureCitation id="137F2A2CFFA7FF86FAC6FC8D676FB707" box="[1295,1361,860,887]" captionStart="Figure 4" captionStartId="8.[194,270,1512,1538]" captionTargetBox="[219,1427,222,1475]" captionTargetId="figure-166@8.[192,1440,213,1481]" captionTargetPageId="8" captionText="Figure 4.Collecting localities of the new species. Circles: Phyllophaga badbunnyi n. sp. Cano. Square:Phyllophaga sechi n. sp. Cano." figureDoi="http://doi.org/10.5281/zenodo.5353734" httpUri="https://zenodo.org/record/5353734/files/figure.png" pageId="9" pageNumber="8">Fig. 4</figureCitation>
). It has been collected at lights in February (1), March (19), April (5), May (2), June (6), August (2) and October (1).
</paragraph>
<paragraph id="8BFB36A9FFA7FF86FF09FC786090B059" blockId="9.[192,1444,937,1066]" pageId="9" pageNumber="8">
<emphasis id="B930EABBFFA7FF86FF09FC7863C4B7B3" bold="true" box="[192,506,937,963]" pageId="9" pageNumber="8">Nomenclatural comments.</emphasis>
Although unseen and unlabeled, I included the Siltepec
<typeStatus id="54FF880BFFA7FF86FB46FC7866CEB7B3" box="[1167,1264,937,963]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
specimen of
<taxonomicName id="4C444D2AFFA7FF86FA47FC7B6039B795" authority="Moron and Blas" authorityName=": Moron and Blas" authorityYear="2006" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="alvareztoroi">
<emphasis id="B930EABBFFA7FF86FA47FC7B637CB795" italics="true" pageId="9" pageNumber="8">P. alvareztoroi</emphasis>
Morón and Blas
</taxonomicName>
, as a
<typeStatus id="54FF880BFFA7FF86FD85FC1D6093B796" box="[588,685,972,998]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C444D2AFFA7FF86FD11FC1D6154B795" authority="Cano, 2020" authorityName="Cano" authorityYear="2020" box="[728,874,971,997]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="badbunnyi" status="sp. nov.">
<emphasis id="B930EABBFFA7FF86FD11FC1D6154B795" box="[728,874,971,997]" italics="true" pageId="9" pageNumber="8">P. badbunnyi</emphasis>
</taxonomicName>
<emphasis id="B930EABBFFA7FF86FCBDFC1A61C1B795" bold="true" box="[884,1023,971,997]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A20357C0FFA7FF86FCBDFC1A61C1B795" box="[884,1023,971,997]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
according to article 72.4.1 of
<bibRefCitation id="EFD54B58FFA7FF86FA97FC1A6335B077" author="ICZN" pageId="9" pageNumber="8" refId="ref6977" refString="ICZN. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature; London. 306 p." type="book" year="1999">ICZN (1999)</bibRefCitation>
, because the figures 68 are signed by Morón himself, in
<bibRefCitation id="EFD54B58FFA7FF86FC45FC3C6687B077" author="Moron MA &amp; Blas M." box="[908,1209,1005,1031]" pageId="9" pageNumber="8" pagination="35 - 46" refId="ref7181" refString="Moron MA, Blas M. 2006. Dos especies nuevas de Phyllophaga del grupo &quot; Schizorhina &quot; (Coleoptera: Melolonthidae: Melolonthinae) de Chiapas, Mexico. Folia Entomologica Mexicana 45: 35 - 46." type="journal article" year="2006">Morón and Blas (2006: 39)</bibRefCitation>
, clearly indicate that it belongs to the new species here described.
</paragraph>
</subSubSection>
</treatment>
</document>