409 lines
48 KiB
XML
409 lines
48 KiB
XML
<document id="91FE7E8BDCD24540A17BC3B4A0A8CC0E" ID-CLB-Dataset="20639" ID-DOI="10.11646/zootaxa.5099.4.5" ID-GBIF-Dataset="f0a97603-c958-4814-a1a2-44a2622b286f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6092195" ID-ZooBank="C2244780-3616-4188-A23E-0291BD463F03" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644912521413" checkinUser="plazi" docAuthor="Archibald, S. Bruce, Cannings, Robert A. & Greenwalt, Dale E." docDate="2022" docId="03ED5075EA22FFF33CCC7E2EC42AFE6D" docLanguage="en" docName="zootaxa.5099.4.5.pdf" docOrigin="Zootaxa 5099 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kishenehna Archibald & Cannings & Greenwalt 2022, new genus" docType="treatment" docVersion="7" lastPageNumber="498" masterDocId="FFD4280DEA23FFF13C5B7F09C701FFC0" masterDocTitle="Kishenehna prima, a new genus and species of darner dragonfly (Odonata, Aeshnidae, Gomphaeschninae) from the early middle Eocene Kishenehn Formation of Montana, USA" masterLastPageNumber="500" masterPageNumber="496" pageNumber="497" updateTime="1699312088466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="ACBF34375091DDA2BCEB8A2E096627EE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DF3DC9FA5972AFFA4B59552FE71B2279">
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<mods:title id="CA3981DAAF984D78EE2A3A6D1A9D5E87">Kishenehna prima, a new genus and species of darner dragonfly (Odonata, Aeshnidae, Gomphaeschninae) from the early middle Eocene Kishenehn Formation of Montana, USA</mods:title>
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</mods:titleInfo>
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<mods:name id="93F90851FC8109ADB9A0B811EB6B83C5" type="personal">
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<mods:roleTerm id="FEB3AF00E848E2C4D932370177FAE73E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7F9ACDC5B6CA2CBB41DB4D44CA42BB3B">Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier id="870FC99BE28EC013782CBCAF59DF63CB" type="ORCID">0000-0001-9491-8186</mods:nameIdentifier>
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<mods:affiliation id="2158262D29C46473F33AC12C337A406C">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada. RCannings @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0001 - 9491 - 8186</mods:affiliation>
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<mods:nameIdentifier id="8C2D076075726CA1539555B66C1902F7" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name id="CDA71FDF260745BCDE56056768C47275" type="personal">
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<mods:role id="14A9ACC20B398C91CF2E3C25121FDE9B">
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<mods:roleTerm id="BC56E6A3519AFA1031076AF451787991">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BA3D41D1E86AEAED7F291918B0F234B5">Cannings, Robert A.</mods:namePart>
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<mods:nameIdentifier id="B557B066E279E1EDEAF72887BCC0CC72" type="ORCID">0000-0001-9491-8186</mods:nameIdentifier>
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<mods:affiliation id="0208DA671D2C277BC4BB296CCFAD6677">Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada. RCannings @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0001 - 9491 - 8186</mods:affiliation>
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<mods:nameIdentifier id="45ACE93F08F89478300495A6494F84C8" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="F7CB2EA23B1DFC270E5A7435340EE287">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4B95199204699664262B666F5134EEAF">Greenwalt, Dale E.</mods:namePart>
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<mods:nameIdentifier id="E863C474F4AAFAC733B4048874FF5799" type="ORCID">0000-0002-9811-6356</mods:nameIdentifier>
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<mods:affiliation id="BD470C8993BC1863AD3AAB52C953FF40">Department of Paleobiology, NMNH, Smithsonian Institution, P. O. Box 37012 MRC 121, Washington, DC, 20013 - 7012, USA. greenwaltde @ gmail. com; https: // orcid. org / 0000 - 0002 - 9811 - 6356</mods:affiliation>
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<mods:nameIdentifier id="3976275B5C1A6FD77EA742E7123C3A67" type="email">greenwaltde@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="C4758FDBD48B02BCEFDC037747D8FC42">text</mods:typeOfResource>
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<mods:title id="45ED07A334DEE5A1EBF3B1965B1B6EB1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7D9D05A2E7A953BE68BC781DFD188AA6">
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<mods:date id="1EBA4833D973C01A5792A172F954CE6B">2022</mods:date>
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<mods:number id="E0FBB00E3222FF5E0FE596022AEABE84">2022-02-15</mods:number>
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<mods:number id="615827345ABDED3FCDCF1018E36AC610">5099</mods:number>
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<mods:start id="EB1DB1C1F682BB5DD9ECB0B60336A1C9">496</mods:start>
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<mods:classification id="428FF44E39EE1042DFE0722CE0BE2228">journal article</mods:classification>
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<mods:identifier id="59BCDF90372B647A15615A631FB98591" type="DOI">10.11646/zootaxa.5099.4.5</mods:identifier>
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<treatment id="03ED5075EA22FFF33CCC7E2EC42AFE6D" ID-DOI="http://doi.org/10.5281/zenodo.6311081" ID-GBIF-Taxon="193376308" ID-Zenodo-Dep="6311081" LSID="urn:lsid:plazi:treatment:03ED5075EA22FFF33CCC7E2EC42AFE6D" httpUri="http://treatment.plazi.org/id/03ED5075EA22FFF33CCC7E2EC42AFE6D" lastPageId="2" lastPageNumber="498" pageId="1" pageNumber="497">
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<subSubSection id="C35EB2E8EA22FFF03CCC7E2EC5D5FE81" box="[151,724,295,322]" pageId="1" pageNumber="497" type="nomenclature">
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<paragraph id="8BFBE163EA22FFF03CCC7E2EC5D5FE81" blockId="1.[151,724,295,322]" box="[151,724,295,322]" pageId="1" pageNumber="497">
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<heading id="D0B3560FEA22FFF03CCC7E2EC5D5FE81" bold="true" box="[151,724,295,322]" fontSize="11" level="1" pageId="1" pageNumber="497" reason="1">
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<emphasis id="B9303D71EA22FFF03CCC7E2EC5D5FE81" bold="true" box="[151,724,295,322]" pageId="1" pageNumber="497">
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<taxonomicName id="4C449AE0EA22FFF03CCC7E2EC629FE82" authority="Archibald & Cannings" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[151,296,295,322]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9303D71EA22FFF03CCC7E2EC629FE82" bold="true" box="[151,296,295,322]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A203800AEA22FFF03D6B7E2EC6B0FE81" box="[304,433,295,321]" pageId="1" pageNumber="497" rank="genus">new genus</taxonomicNameLabel>
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Archibald & Cannings
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35EB2E8EA22FFF03CCC7E66C6EFFA5D" pageId="1" pageNumber="497" type="diagnosis">
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<paragraph id="8BFBE163EA22FFF03CCC7E66C521FD32" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
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<emphasis id="B9303D71EA22FFF03CCC7E66C613FE49" bold="true" box="[151,274,367,393]" pageId="1" pageNumber="497">Diagnosis.</emphasis>
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The female hind wing of
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<taxonomicName id="4C449AE0EA22FFF03E1D7E66C5CBFE49" authority="Archibald & Cannings & Greenwalt, 2022" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[582,714,367,393]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9303D71EA22FFF03E1D7E66C5CBFE49" box="[582,714,367,393]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
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</taxonomicName>
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<emphasis id="B9303D71EA22FFF03E8F7E66C427FE49" bold="true" box="[724,806,367,393]" pageId="1" pageNumber="497">
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<taxonomicNameLabel id="A203800AEA22FFF03E8F7E66C427FE49" box="[724,806,367,393]" pageId="1" pageNumber="497" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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is most like those of
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<taxonomicName id="4C449AE0EA22FFF0387C7E66C29CFE49" authority="Wighton & Wilson." authorityName="Wighton & Wilson." box="[1063,1437,367,393]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF0387C7E66C3B0FE49" box="[1063,1201,367,393]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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Wighton & Wilson.
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</taxonomicName>
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<taxonomicName id="4C449AE0EA22FFF03CCC7E9AC61AFE6D" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[151,283,403,429]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF03CCC7E9AC61AFE6D" box="[151,283,403,429]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
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</taxonomicName>
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is excluded from
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<taxonomicName id="4C449AE0EA22FFF03DB97E9AC559FE6D" authorityName="Leach" authorityYear="1815" box="[482,600,403,429]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="subFamily" subFamily="Aeshninae">Aeshninae</taxonomicName>
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<emphasis id="B9303D71EA22FFF03E057E9DC59DFE6D" box="[606,668,404,429]" italics="true" pageId="1" pageNumber="497">sensu</emphasis>
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<bibRefCitation id="EFD59C92EA22FFF03EF87E9DC4A5FE6D" author="von Ellenrieder, N." box="[675,932,403,430]" pageId="1" pageNumber="497" pagination="437 - 467" refId="ref3433" refString="von Ellenrieder, N. (2002) A phylogenetic analysis of the extant Aeshnidae (Odonata: Anisoptera). Systematic Entomology, 27, 437 - 467. https: // doi. org / 10.1046 / j. 1365 - 3113.2002.00190. x" type="journal article" year="2002">von Ellenrieder (2002)</bibRefCitation>
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, by MA, RP3-4 parallel where one cell width becomes two (her character state 13.1), Mspl lacking concave bend in distal portion (her character state 14.2); and differs from all remaining
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<taxonomicName id="4C449AE0EA22FFF03DE27ED2C52EFE35" authorityName="Leach" authorityYear="1815" box="[441,559,475,501]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
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other than
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<taxonomicName id="4C449AE0EA22FFF03EF07ED2C434FE35" box="[683,821,475,501]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF03EF07ED2C434FE35" box="[683,821,475,501]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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by a combination of an unbranched IRP2 (shared by all
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<taxonomicName id="4C449AE0EA22FFF03CCC7EF6C620FDD9" box="[151,289,511,537]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF03CCC7EF6C620FDD9" box="[151,289,511,537]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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); RP2 curved basal to pterostigma (shared by all
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<taxonomicName id="4C449AE0EA22FFF03F1F7EF6C4CFFDD9" box="[836,974,511,537]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF03F1F7EF6C4CFFDD9" box="[836,974,511,537]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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); radial, medial planates, MP–CuA space narrow, each one cell wide (shared by most
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<taxonomicName id="4C449AE0EA22FFF03ECF7D2AC41FFDFD" box="[660,798,547,573]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF03ECF7D2AC41FFDFD" box="[660,798,547,573]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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: last row at margin in some, a few adventitiously more basally in some specimens;
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<taxonomicName id="4C449AE0EA22FFF03D807D4EC44BFDA1" authority="Wighton & Wilson" authorityName="Wighton & Wilson" authorityYear="1986" box="[475,842,583,610]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="quadrata">
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<emphasis id="B9303D71EA22FFF03D807D4EC6EDFDA1" box="[475,492,583,609]" italics="true" pageId="1" pageNumber="497">A</emphasis>
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.
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<emphasis id="B9303D71EA22FFF03DA67D4EC565FDA1" box="[509,612,583,609]" italics="true" pageId="1" pageNumber="497">quadrata</emphasis>
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Wighton & Wilson
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</taxonomicName>
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: MA–CuA terminus poorly known by preservation;
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<taxonomicName id="4C449AE0EA22FFF03CCC7D62C6F6FD45" authority="Wighton & Wilson" authorityName="Wighton & Wilson" authorityYear="1986" box="[151,503,619,646]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="marklae">
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<emphasis id="B9303D71EA22FFF03CCC7D62C7A9FD45" box="[151,168,619,645]" italics="true" pageId="1" pageNumber="497">A</emphasis>
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.
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<emphasis id="B9303D71EA22FFF03CE37D62C615FD45" box="[184,276,619,645]" italics="true" pageId="1" pageNumber="497">marklae</emphasis>
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Wighton & Wilson
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</taxonomicName>
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: medial planate two cells wide for five? rows to margin); radial planate expanded slightly in middle (all
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<taxonomicName id="4C449AE0EA22FFF03DC17D86C525FD69" box="[410,548,655,681]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF03DC17D86C525FD69" box="[410,548,655,681]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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: slightly or not expanded); without crossveins in supratriangle (shared by all
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<taxonomicName id="4C449AE0EA22FFF03CCC7DBAC620FD0D" box="[151,289,691,717]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03CCC7DBAC620FD0D" box="[151,289,691,717]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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except
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<taxonomicName id="4C449AE0EA22FFF03D277DBAC6F8FD0D" authorityName="Wighton & Wilson" authorityYear="1986" box="[380,505,691,718]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="marklae">
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<emphasis id="B9303D71EA22FFF03D277DBAC68CFD0D" box="[380,397,691,717]" italics="true" pageId="1" pageNumber="497">A</emphasis>
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.
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<emphasis id="B9303D71EA22FFF03DC67DBAC6F8FD0D" box="[413,505,691,717]" italics="true" pageId="1" pageNumber="497">marklae</emphasis>
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</taxonomicName>
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, 1 crossvein), subtriangle (shared by all
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<taxonomicName id="4C449AE0EA22FFF03F967DBAC356FD0D" box="[973,1111,691,717]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF03F967DBAC356FD0D" box="[973,1111,691,717]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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); no cordulegastrid gap (
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<taxonomicName id="4C449AE0EA22FFF039217DBAC608FD31" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
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<emphasis id="B9303D71EA22FFF039217DBAC608FD31" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
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</taxonomicName>
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: see Discussion, below).
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</paragraph>
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<paragraph id="8BFBE163EA22FFF03C9C7DF2C4E3FC9D" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
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Hind wings of
|
||
<taxonomicName id="4C449AE0EA22FFF03D2F7DF2C6F9FCD5" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[372,504,763,789]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03D2F7DF2C6F9FCD5" box="[372,504,763,789]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
are separated most easily from those of
|
||
<taxonomicName id="4C449AE0EA22FFF03F977DF2C357FCD5" box="[972,1110,763,789]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03F977DF2C357FCD5" box="[972,1110,763,789]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
|
||
</taxonomicName>
|
||
by anal loop with four cells [
|
||
<taxonomicName id="4C449AE0EA22FFF03CFA7C16C62AFCF9" box="[161,299,799,825]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03CFA7C16C62AFCF9" box="[161,299,799,825]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
|
||
</taxonomicName>
|
||
: six]; by RP2–IRP2 space maximum of three cells wide from level of base of pterostigma to cell row at margin [
|
||
<taxonomicName id="4C449AE0EA22FFF03D497C4AC69DFC9D" box="[274,412,835,861]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03D497C4AC69DFC9D" box="[274,412,835,861]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
|
||
</taxonomicName>
|
||
: two cells wide (except a few cell rows at margin)].
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C7C6EC67CFC65" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">The following genera are only known from partial/damaged fossil wings; all character states discussed above cannot be evaluated.</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C7CA6C249FBD1" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
<taxonomicName id="4C449AE0EA22FFF03C9C7CA6C575FC09" authority="Jarzembowski & Nel" authorityName="Jarzembowski & Nel" authorityYear="1996" box="[199,628,943,969]" class="Insecta" family="Aeshnidae" genus="Cretalloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03C9C7CA6C67EFC09" box="[199,383,943,969]" italics="true" pageId="1" pageNumber="497">Cretalloaeschna</emphasis>
|
||
Jarzembowski & Nel
|
||
</taxonomicName>
|
||
(based on a partial forewing, see
|
||
<bibRefCitation id="EFD59C92EA22FFF03FA97CA6C221FC0A" author="Jarzembowski, E. A. & Nel, A." box="[1010,1312,943,970]" pageId="1" pageNumber="497" pagination="97 - 101" refId="ref3224" refString="Jarzembowski, E. A. & Nel, A. (1996) A new genus and species of hawker dragonfly (Aeshnidae) from the Lower Cretaceous of southern England. Cretaceous Research, 17, 97 - 101. https: // doi. org / 10.1006 / cres. 1996.0007" type="journal article" year="1996">Jarzembowski & Nel 1996</bibRefCitation>
|
||
) differs by two rows of cells in RP2–IRP2 space [
|
||
<taxonomicName id="4C449AE0EA22FFF03E0D7CDAC5DBFC2D" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[598,730,979,1005]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03E0D7CDAC5DBFC2D" box="[598,730,979,1005]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: three cells wide distal to level of mid-pterostigma (except at margin)]; one row in RP3-4–MA space but two close to margin [
|
||
<taxonomicName id="4C449AE0EA22FFF03F377CFEC4F1FBD1" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[876,1008,1015,1041]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03F377CFEC4F1FBD1" box="[876,1008,1015,1041]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: two cells wide more basally].
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C7B12C3DEFBBD" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
<taxonomicName id="4C449AE0EA22FFF03C9C7B12C57EFBF6" authority="Zheng et al." authorityName="Zheng" box="[199,639,1051,1078]" class="Insecta" family="Gomphaeschnidae" genus="Cretagomphaeschnaoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03C9C7B12C6E8FBF5" box="[199,489,1051,1077]" italics="true" pageId="1" pageNumber="497">Cretagomphaeschnaoides</emphasis>
|
||
Zheng
|
||
<emphasis id="B9303D71EA22FFF03E1F7B15C579FBF5" box="[580,632,1051,1077]" italics="true" pageId="1" pageNumber="497">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
(based on a fragmentary forewing, see
|
||
<bibRefCitation id="EFD59C92EA22FFF038167B12C21CFBF6" author="Zheng, D. & Jarzembowski, E. A. & Su-Chin Chang, S. - C. & Wang, B." box="[1101,1309,1051,1078]" pageId="1" pageNumber="497" pagination="629 - 632" refId="ref3615" refString="Zheng, D., Jarzembowski, E. A., Su-Chin Chang, S. - C. & Wang, B. (2016) A new true dragonfly (Odonata, Anisoptera, Gomphaeschnaoidini) from mid-Cretaceous Burmese amber. Proceedings of the Geologists' Association, 127, 629 - 632. https: // doi. org / 10.1016 / j. pgeola. 2016.07.006" type="journal article" year="2016">
|
||
Zheng
|
||
<emphasis id="B9303D71EA22FFF038C57B15C3D3FBF5" box="[1182,1234,1051,1077]" italics="true" pageId="1" pageNumber="497">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
) differs by estimated wing length
|
||
<quantity id="4CBC4C86EA22FFF03DC07B36C516FB9A" box="[411,535,1087,1114]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="1" pageNumber="497" unit="mm" value="28.0" valueMax="29.0" valueMin="27.0">27–29 mm</quantity>
|
||
[
|
||
<taxonomicName id="4C449AE0EA22FFF03E737B36C5ADFB99" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[552,684,1087,1113]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03E737B36C5ADFB99" box="[552,684,1087,1113]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="4CBC4C86EA22FFF03EE67B36C412FB99" box="[701,787,1086,1113]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="497" unit="mm" value="39.0">39 mm</quantity>
|
||
]; discoidal triangle three-celled [
|
||
<taxonomicName id="4C449AE0EA22FFF038D07B36C20EFB99" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[1163,1295,1087,1113]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF038D07B36C20EFB99" box="[1163,1295,1087,1113]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: two]; three aligned antenodal crossveins present immediately basal of nodus [
|
||
<taxonomicName id="4C449AE0EA22FFF03F267B6AC300FBBD" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[893,1025,1123,1149]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03F267B6AC300FBBD" box="[893,1025,1123,1149]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: these not aligned].
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C7B8EC340FB29" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
<taxonomicName id="4C449AE0EA22FFF03C9C7B8EC6F5FB62" authority="Zheng et al." authorityName="Zheng" box="[199,500,1159,1186]" class="Insecta" family="Gomphaeschnidae" genus="Kachinaeshna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03C9C7B8EC666FB61" box="[199,359,1159,1185]" italics="true" pageId="1" pageNumber="497">Kachinaeshna</emphasis>
|
||
Zheng
|
||
<emphasis id="B9303D71EA22FFF03DE77B81C6ECFB61" box="[444,493,1159,1185]" italics="true" pageId="1" pageNumber="497">et al</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
(based on most of a forewing, portion of a hind wing, see
|
||
<bibRefCitation id="EFD59C92EA22FFF038247B8EC247FB61" author="Zheng, D. R. & Nel, A. & Wang, B." box="[1151,1350,1159,1186]" pageId="1" pageNumber="497" pagination="138 - 144" refId="ref3685" refString="Zheng, D. R., Nel, A. & Wang, B. (2019) New gomphaeschnid dragonflies (Odonata: Anisoptera: Aeshnoptera) from mid-Cretaceous Burmese amber. Cretaceous Research, 100, 138 - 144. https: // doi. org / 10.1016 / j. cretres. 2019.03.027" type="journal article" year="2019">
|
||
Zheng
|
||
<emphasis id="B9303D71EA22FFF038967B81C3FFFB61" box="[1229,1278,1159,1185]" italics="true" pageId="1" pageNumber="497">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
) differs by MP–CuA space dividing to two cells width near margin [
|
||
<taxonomicName id="4C449AE0EA22FFF03F607BA2C4BEFB05" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[827,959,1195,1221]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03F607BA2C4BEFB05" box="[827,959,1195,1221]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: one to margin]; IRP1 origin at brace vein [
|
||
<taxonomicName id="4C449AE0EA22FFF03CFA7BC6C624FB29" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[161,293,1231,1257]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03CFA7BC6C624FB29" box="[161,293,1231,1257]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: 1.5 cells basal to this]; forewing
|
||
<quantity id="4CBC4C86EA22FFF03EF97BC6C407FB2A" box="[674,774,1231,1258]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="1" pageNumber="497" unit="mm" value="31.1">31.1 mm</quantity>
|
||
long [
|
||
<taxonomicName id="4C449AE0EA22FFF03F157BC6C4D3FB29" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[846,978,1231,1257]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03F157BC6C4D3FB29" box="[846,978,1231,1257]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="4CBC4C86EA22FFF03FBA7BC6C337FB29" box="[993,1078,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="497" unit="mm" value="36.0">36 mm</quantity>
|
||
].
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C7BFAC268FAF1" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
UAPC 6190 (unnamed base of hind wing from Paleocene of
|
||
<collectingRegion id="49802F81EA22FFF03F3B7BFAC4B9FACD" box="[864,952,1267,1293]" country="Canada" name="Alberta" pageId="1" pageNumber="497">Alberta</collectingRegion>
|
||
:
|
||
<bibRefCitation id="EFD59C92EA22FFF03F9A7BFAC3D1FACE" author="Wighton, D. C. & Wilson, M. V. H." box="[961,1232,1267,1294]" pageId="1" pageNumber="497" pagination="505 - 522" refId="ref3550" refString="Wighton, D. C. & Wilson, M. V. H. (1986) The Gomphaeschninae (Odonata: Aeshnidae): new fossil genus, reconstructed phylogeny, and geographical history. Systematic Entomology, 11, 505 - 522. https: // doi. org / 10.1111 / j. 1365 - 3113.1986. tb 00539. x" type="journal article" year="1986">Wighton & Wilson 1986</bibRefCitation>
|
||
) by anal loop with ten cells [
|
||
<taxonomicName id="4C449AE0EA22FFF03D5F7A1EC689FAF1" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[260,392,1303,1329]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03D5F7A1EC689FAF1" box="[260,392,1303,1329]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: four]; triangle with four cells [
|
||
<taxonomicName id="4C449AE0EA22FFF03EB77A1EC471FAF1" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[748,880,1303,1329]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03EB77A1EC471FAF1" box="[748,880,1303,1329]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: two]; subtriangle with crossveins [without].
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C7A32C6EFFA5D" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
<taxonomicName id="4C449AE0EA22FFF03C9C7A32C58AFA95" authority="Archibald & Cannings (2019" authorityName="Archibald & Cannings" authorityYear="2019" box="[199,651,1339,1365]" class="Insecta" family="Aeshnidae" genus="Idemlinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03C9C7A32C636FA95" box="[199,311,1339,1365]" italics="true" pageId="1" pageNumber="497">Idemlinea</emphasis>
|
||
<bibRefCitation id="EFD59C92EA22FFF03D647A32C58AFA95" author="Archibald, S. B. & Cannings, R. A." box="[319,651,1339,1365]" pageId="1" pageNumber="497" pagination="783 - 816" refId="ref2515" refString="Archibald, S. B. & Cannings, R. A. (2019) Fossil dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands, western North America. The Canadian Entomologist, 151, 783 - 816. https: // doi. org / 10.4039 / tce. 2019.61" type="journal article" year="2019">Archibald & Cannings (2019</bibRefCitation>
|
||
</taxonomicName>
|
||
, based on basal portion of a hind wing) differs most notably by basal portions of MP, CuA parallel to hind margin [curves toward it]; supratriangle about 4.5 times longer than wide [
|
||
<taxonomicName id="4C449AE0EA22FFF03CFA7A8AC624FA5D" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[161,293,1411,1437]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03CFA7A8AC624FA5D" box="[161,293,1411,1437]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
: about six times].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB2E8EA22FFF03C9C7AAEC41EFA02" box="[199,799,1447,1474]" pageId="1" pageNumber="497" type="type_taxon">
|
||
<paragraph id="8BFBE163EA22FFF03C9C7AAEC41EFA02" blockId="1.[151,1437,367,2013]" box="[199,799,1447,1474]" pageId="1" pageNumber="497">
|
||
<emphasis id="B9303D71EA22FFF03C9C7AAEC663FA01" bold="true" box="[199,354,1447,1473]" pageId="1" pageNumber="497">
|
||
<typeStatus id="54FF5FC1EA22FFF03C9C7AAEC603FA01" box="[199,258,1447,1473]" pageId="1" pageNumber="497">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C449AE0EA22FFF03D327AAEC536FA01" authority="Archibald & Cannings & Greenwalt, 2022" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[361,567,1447,1473]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="prima" status="sp. nov.">
|
||
<emphasis id="B9303D71EA22FFF03D327AAEC536FA01" box="[361,567,1447,1473]" italics="true" pageId="1" pageNumber="497">Kishenehna prima</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9303D71EA22FFF03E657AAEC57CFA01" bold="true" box="[574,637,1447,1473]" pageId="1" pageNumber="497">
|
||
<taxonomicNameLabel id="A203800AEA22FFF03E657AAEC57CFA01" box="[574,637,1447,1473]" pageId="1" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by monotypy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB2E8EA22FFF03C9C7AC2C5AEFA25" box="[199,687,1483,1509]" pageId="1" pageNumber="497" type="description">
|
||
<paragraph id="8BFBE163EA22FFF03C9C7AC2C5AEFA25" blockId="1.[151,1437,367,2013]" box="[199,687,1483,1509]" pageId="1" pageNumber="497">
|
||
<emphasis id="B9303D71EA22FFF03C9C7AC2C658FA25" bold="true" box="[199,345,1483,1509]" pageId="1" pageNumber="497">Description.</emphasis>
|
||
As for its only species, below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB2E8EA22FFF03C9C7AE6C5D7F991" pageId="1" pageNumber="497" type="etymology">
|
||
<paragraph id="8BFBE163EA22FFF03C9C7AE6C663F9ED" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
<emphasis id="B9303D71EA22FFF03C9C7AE6C64FF9C9" bold="true" box="[199,334,1519,1545]" pageId="1" pageNumber="497">Etymology.</emphasis>
|
||
The genus name
|
||
<taxonomicName id="4C449AE0EA22FFF03E447AE6C5A2F9C9" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[543,675,1519,1545]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03E447AE6C5A2F9C9" box="[543,675,1519,1545]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
is the Latinized form of “Kishenehn”, referring to the formation. Gender, feminine.
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C793EC5D7F991" blockId="1.[151,1437,367,2013]" box="[199,726,1591,1617]" pageId="1" pageNumber="497">
|
||
<emphasis id="B9303D71EA22FFF03C9C793EC67FF991" bold="true" box="[199,382,1591,1617]" pageId="1" pageNumber="497">Range and age.</emphasis>
|
||
As for its only species, below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35EB2E8EA22FFF33C9C7952C42AFE6D" lastPageId="2" lastPageNumber="498" pageId="1" pageNumber="497" type="discussion">
|
||
<paragraph id="8BFBE163EA22FFF03C9C7952C527F8C6" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
<emphasis id="B9303D71EA22FFF03C9C7952C64DF9B5" bold="true" box="[199,332,1627,1653]" pageId="1" pageNumber="497">Discussion.</emphasis>
|
||
We assign
|
||
<taxonomicName id="4C449AE0EA22FFF03D957952C553F9B5" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[462,594,1627,1653]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03D957952C553F9B5" box="[462,594,1627,1653]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
to the
|
||
<taxonomicName id="4C449AE0EA22FFF03EC47952C414F9B5" authorityName="Leach" authorityYear="1815" box="[671,789,1627,1653]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
|
||
by its distinct radial and medial planates and well-developed hind wing anal loop. We exclude the subfamily
|
||
<taxonomicName id="4C449AE0EA22FFF03EA37976C46FF959" authorityName="Leach" authorityYear="1815" box="[760,878,1663,1689]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="subFamily" subFamily="Aeshninae">Aeshninae</taxonomicName>
|
||
as in the genus diagnosis.
|
||
<taxonomicName id="4C449AE0EA22FFF038F87976C273F959" box="[1187,1394,1663,1689]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="subFamily" subFamily="Gomphaeschninae">Gomphaeschninae</taxonomicName>
|
||
are not strongly defined by wing characters, none of which are found to be consistent throughout the taxon, which
|
||
<bibRefCitation id="EFD59C92EA22FFF0392979ADC664F921" author="von Ellenrieder, N." pageId="1" pageNumber="497" pagination="437 - 467" refId="ref3433" refString="von Ellenrieder, N. (2002) A phylogenetic analysis of the extant Aeshnidae (Odonata: Anisoptera). Systematic Entomology, 27, 437 - 467. https: // doi. org / 10.1046 / j. 1365 - 3113.2002.00190. x" type="journal article" year="2002">von Ellenrieder (2002)</bibRefCitation>
|
||
defined by male secondary genitalia. We associate the new genus with the
|
||
<taxonomicName id="4C449AE0EA22FFF038F079CEC27BF921" box="[1195,1402,1735,1761]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="subFamily" subFamily="Gomphaeschninae">Gomphaeschninae</taxonomicName>
|
||
by its strong similarity to
|
||
<taxonomicName id="4C449AE0EA22FFF03DCE79E2C51EF8C5" box="[405,543,1771,1797]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03DCE79E2C51EF8C5" box="[405,543,1771,1797]" italics="true" pageId="1" pageNumber="497">Alloaeschna</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA22FFF03C9C7806C644F81D" blockId="1.[151,1437,367,2013]" pageId="1" pageNumber="497">
|
||
<taxonomicName id="4C449AE0EA22FFF03C9C7806C64AF8E9" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[199,331,1807,1833]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA22FFF03C9C7806C64AF8E9" box="[199,331,1807,1833]" italics="true" pageId="1" pageNumber="497">Kishenehna</emphasis>
|
||
</taxonomicName>
|
||
lacks a cordulegastrid gap, considered an important trait in associating wings of many
|
||
<taxonomicName id="4C449AE0EA22FFF039697806C609F88D" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="subFamily" subFamily="Gomphaeschninae">Gomphaeschninae</taxonomicName>
|
||
, although it is not present in all assigned to it (
|
||
<bibRefCitation id="EFD59C92EA22FFF03F7E783AC4FDF88D" author="Bechly, G. & Nel, A. & Martinez-Delclos, X. & Jarzembowski, E. A. & Coram, R. & Martill, D." box="[805,1020,1843,1870]" pageId="1" pageNumber="497" pagination="1 - 219" refId="ref2571" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D. et al. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of numerous new taxa (Insecta: Odonata: Anisoptera). Neue Palaontologilsche Abhandlungen, 4, 1 - 219, 48 pls." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B9303D71EA22FFF03F25783DC4B0F88D" box="[894,945,1843,1869]" italics="true" pageId="1" pageNumber="497">et al</emphasis>
|
||
. 2001
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD59C92EA22FFF03852783DC3FDF88D" author="von Ellenrieder, N." box="[1033,1276,1843,1870]" pageId="1" pageNumber="497" pagination="437 - 467" refId="ref3433" refString="von Ellenrieder, N. (2002) A phylogenetic analysis of the extant Aeshnidae (Odonata: Anisoptera). Systematic Entomology, 27, 437 - 467. https: // doi. org / 10.1046 / j. 1365 - 3113.2002.00190. x" type="journal article" year="2002">von Ellenrieder 2002</bibRefCitation>
|
||
), is weakly to very weakly expressed in some, and is present in a species of the Cretaceous
|
||
<taxonomicName id="4C449AE0EA22FFF03FAD785EC299F8B1" authority="(Bechly et al. 2001)" baseAuthorityName="Bechly" baseAuthorityYear="2001" box="[1014,1432,1879,1906]" class="Insecta" family="Liupanshaniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="family">
|
||
Liupanshaniidae (
|
||
<bibRefCitation id="EFD59C92EA22FFF038E6785EC291F8B1" author="Bechly, G. & Nel, A. & Martinez-Delclos, X. & Jarzembowski, E. A. & Coram, R. & Martill, D." box="[1213,1424,1879,1906]" pageId="1" pageNumber="497" pagination="1 - 219" refId="ref2571" refString="Bechly, G., Nel, A., Martinez-Delclos, X., Jarzembowski, E. A., Coram, R., Martill, D. et al. (2001) A revision and phylogenetic study of Mesozoic Aeshnoptera, with description of numerous new taxa (Insecta: Odonata: Anisoptera). Neue Palaontologilsche Abhandlungen, 4, 1 - 219, 48 pls." type="journal article" year="2001">
|
||
Bechly
|
||
<emphasis id="B9303D71EA22FFF0394E7851C247F8B1" box="[1301,1350,1879,1905]" italics="true" pageId="1" pageNumber="497">et al</emphasis>
|
||
. 2001
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The forewing of
|
||
<taxonomicName id="4C449AE0EA22FFF03D0C7872C551F855" authorityName="Wighton & Wilson" authorityYear="1986" box="[343,592,1915,1941]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="quadrata">
|
||
<emphasis id="B9303D71EA22FFF03D0C7872C551F855" box="[343,592,1915,1941]" italics="true" pageId="1" pageNumber="497">Alloaeschna quadrata</emphasis>
|
||
</taxonomicName>
|
||
lacks it (hind wing unknown), and its presence appears variable in
|
||
<taxonomicName id="4C449AE0EA22FFF039177872C512F87A" authority="Wighton & Wilson. It" authorityName="Wighton & Wilson. It" authorityYear="1986" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="497" phylum="Arthropoda" rank="species" species="paskapooensis">
|
||
<emphasis id="B9303D71EA22FFF039177872C25CF855" box="[1356,1373,1915,1941]" italics="true" pageId="1" pageNumber="497">A</emphasis>
|
||
.
|
||
<emphasis id="B9303D71EA22FFF039377875C612F879" italics="true" pageId="1" pageNumber="497">paskapooensis</emphasis>
|
||
Wighton & Wilson. It
|
||
</taxonomicName>
|
||
is arguably absent in the hind wing specimen UAPC 6183 (
|
||
<bibRefCitation id="EFD59C92EA22FFF038987896C7D3F81D" author="Wighton, D. C. & Wilson, M. V. H." pageId="1" pageNumber="497" pagination="505 - 522" refId="ref3550" refString="Wighton, D. C. & Wilson, M. V. H. (1986) The Gomphaeschninae (Odonata: Aeshnidae): new fossil genus, reconstructed phylogeny, and geographical history. Systematic Entomology, 11, 505 - 522. https: // doi. org / 10.1111 / j. 1365 - 3113.1986. tb 00539. x" type="journal article" year="1986">Wighton & Wilson 1986</bibRefCitation>
|
||
, figure 8).
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA21FFF33C9C7F9EC419FE81" blockId="2.[151,1437,151,429]" pageId="2" pageNumber="498">
|
||
The radial and medial planates and MP–CuA spaces a single cell wide, the oblique vein O as the first crossvein distal to the origin of RP2, and the hypertriangle and subtriangle without crossveins are conditions found in numerous
|
||
<taxonomicName id="4C449AE0EA21FFF33C9D7FD6C699FF39" box="[198,408,223,249]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="497" phylum="Arthropoda" rank="subFamily" subFamily="Gomphaeschninae">Gomphaeschninae</taxonomicName>
|
||
, sometimes together, as in,
|
||
<emphasis id="B9303D71EA21FFF33E947FD6C5F1FF38" box="[719,752,223,248]" italics="true" pageId="2" pageNumber="498">e.g</emphasis>
|
||
.,
|
||
<emphasis id="B9303D71EA21FFF33F5D7FD6C348FF39" box="[774,1097,223,249]" italics="true" pageId="2" pageNumber="498">Gomphaeschnoides obliquus</emphasis>
|
||
Wighton, see
|
||
<bibRefCitation id="EFD59C92EA21FFF338B07FD6C299FF39" author="Wighton, D. C." box="[1259,1432,223,249]" pageId="2" pageNumber="498" pagination="311 - 314" refId="ref3508" refString="Wighton, D. C. (1987) Gomphaeschna obliqua spec. nov., a new species of Gomphaeschninae from the Lower Cretaceous of northeastern Brazil (Anisoptera: Aeshnidae). Odonatologica, 16, 311 - 314." type="journal article" year="1987">Wighton (1987</bibRefCitation>
|
||
, figure 2, as
|
||
<taxonomicName id="4C449AE0EA21FFF33D477E0AC529FEDD" baseAuthorityName="Wighton" baseAuthorityYear="1987" box="[284,552,259,285]" class="Insecta" family="Aeshnidae" genus="Gomphaeschna" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="498" phylum="Arthropoda" rank="species" species="obliqua">
|
||
<emphasis id="B9303D71EA21FFF33D477E0AC529FEDD" box="[284,552,259,285]" italics="true" pageId="2" pageNumber="498">Gomphaeschna obliqua</emphasis>
|
||
</taxonomicName>
|
||
), which also has its triangle crossed by a single crossvein, but which differs in other ways,
|
||
<emphasis id="B9303D71EA21FFF33D447E2EC641FE80" box="[287,320,295,320]" italics="true" pageId="2" pageNumber="498">e.g</emphasis>
|
||
., by bearing a distinct cordulegastrid gap.
|
||
</paragraph>
|
||
<paragraph id="8BFBE163EA21FFF33C9C7E42C42AFE6D" blockId="2.[151,1437,151,429]" pageId="2" pageNumber="498">
|
||
<taxonomicName id="4C449AE0EA21FFF33C9C7E42C650FEA5" box="[199,337,331,357]" class="Insecta" family="Aeshnidae" genus="Alloaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="498" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9303D71EA21FFF33C9C7E42C650FEA5" box="[199,337,331,357]" italics="true" pageId="2" pageNumber="498">Alloaeschna</emphasis>
|
||
</taxonomicName>
|
||
fossils are known from the middle or late Paleocene Tiffanian North American Land Mammal Age (
|
||
<bibRefCitation id="EFD59C92EA21FFF33CC57E66C6C1FE49" author="Hoffman, G. L. & Stockey, R. A." box="[158,448,367,393]" pageId="2" pageNumber="498" pagination="2073 - 2084" refId="ref3161" refString="Hoffman, G. L. & Stockey, R. A. (1999) Geological setting and paleobotany of the Joffre Bridge Roadcut fossil locality (Late Paleocene), Red Deer Valley, Alberta. Canadian Journal of Earth Sciences, 36, 2073 - 2084. https: // doi. org / 10.1139 / e 99 - 095" type="journal article" year="1999">Hoffman & Stockey 1999</bibRefCitation>
|
||
) of the Paskapoo Formation of
|
||
<collectingRegion id="49802F81EA21FFF33F407E66C473FE49" box="[795,882,367,393]" country="Canada" name="Alberta" pageId="2" pageNumber="498">Alberta</collectingRegion>
|
||
,
|
||
<collectingCountry id="F353A1F3EA21FFF33F277E66C4D2FE49" box="[892,979,367,393]" name="Canada" pageId="2" pageNumber="498">Canada</collectingCountry>
|
||
, about 10–16 million years older than the Coal Creek Member and some 440 kilometers to the north.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |