131 lines
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131 lines
16 KiB
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<document id="EFD69C815D7B75BD99EAFA42506E8176" ID-CLB-Dataset="20639" ID-DOI="10.11646/zootaxa.5099.4.5" ID-GBIF-Dataset="f0a97603-c958-4814-a1a2-44a2622b286f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6092195" ID-ZooBank="C2244780-3616-4188-A23E-0291BD463F03" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644912521413" checkinUser="plazi" docAuthor="Archibald, S. Bruce, Cannings, Robert A. & Greenwalt, Dale E." docDate="2022" docId="03ED5075EA20FFF23CCC7D87C647FB29" docLanguage="en" docName="zootaxa.5099.4.5.pdf" docOrigin="Zootaxa 5099 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aeshnidae Leach 1815" docType="treatment" docVersion="8" lastPageNumber="499" masterDocId="FFD4280DEA23FFF13C5B7F09C701FFC0" masterDocTitle="Kishenehna prima, a new genus and species of darner dragonfly (Odonata, Aeshnidae, Gomphaeschninae) from the early middle Eocene Kishenehn Formation of Montana, USA" masterLastPageNumber="500" masterPageNumber="496" pageNumber="499" updateTime="1699312088466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3A4E64D6E4D23DE7EA165EDB55EEC395">Kishenehna prima, a new genus and species of darner dragonfly (Odonata, Aeshnidae, Gomphaeschninae) from the early middle Eocene Kishenehn Formation of Montana, USA</mods:title>
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<mods:roleTerm id="C8235A2C522E42FCA0E6C8B04D1BE09F">Author</mods:roleTerm>
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<mods:namePart id="1C00873174554E575E3035638D9FDFDD">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="D3CFF34BE4BF67ECD6129341FD005179">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada. RCannings @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0001 - 9491 - 8186</mods:affiliation>
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<mods:nameIdentifier id="503A371D3373F33600128E1C329A0E42" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name id="B679C8581B9E6DE6208610C7BEB32E2A" type="personal">
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<mods:namePart id="8F45F64B94B432141998AE7B5EAB23F6">Cannings, Robert A.</mods:namePart>
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<mods:nameIdentifier id="FEC7A925CCB3D88E4D23766ADB5D8DA9" type="ORCID">0000-0001-9491-8186</mods:nameIdentifier>
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<mods:affiliation id="7D18A9C6A158887833FF40EB94F47C7B">Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada. RCannings @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0001 - 9491 - 8186</mods:affiliation>
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<mods:nameIdentifier id="0881ED694057B40086EB2D96B84D5025" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="67A775EDC5BE8C963A685F2CCD343CBB">Greenwalt, Dale E.</mods:namePart>
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<mods:nameIdentifier id="44ECF62BC81FA120FB4B0355C2A32AFE" type="ORCID">0000-0002-9811-6356</mods:nameIdentifier>
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<mods:affiliation id="E94C728C1583C876871AB69EDB4AE477">Department of Paleobiology, NMNH, Smithsonian Institution, P. O. Box 37012 MRC 121, Washington, DC, 20013 - 7012, USA. greenwaltde @ gmail. com; https: // orcid. org / 0000 - 0002 - 9811 - 6356</mods:affiliation>
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<mods:nameIdentifier id="0198C6352C57F9ECE0BC9A657F7A1161" type="email">greenwaltde@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="CA2A468D4E1E4FCDB132AB7BAA304118">text</mods:typeOfResource>
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<mods:title id="12FC949EAD2CA25995A7223D02689DA7">Zootaxa</mods:title>
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<mods:part id="B278F4FDBC147BA9C7045C01F38DF86E">
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<mods:date id="A755FDCCEDF5892D6A98CF3A93A24B39">2022</mods:date>
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<mods:number id="405A653F6B59F87B149A9530F61EE0B6">2022-02-15</mods:number>
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<mods:number id="B68E18E74145CE72363BC2D273746C05">5099</mods:number>
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<mods:identifier id="68D2EC3F083BC1B5C05EE6F5629539AF" type="DOI">10.11646/zootaxa.5099.4.5</mods:identifier>
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<treatment id="03ED5075EA20FFF23CCC7D87C647FB29" ID-DOI="http://doi.org/10.5281/zenodo.6311085" ID-GBIF-Taxon="193376307" ID-Zenodo-Dep="6311085" LSID="urn:lsid:plazi:treatment:03ED5075EA20FFF23CCC7D87C647FB29" httpUri="http://treatment.plazi.org/id/03ED5075EA20FFF23CCC7D87C647FB29" lastPageNumber="499" pageId="3" pageNumber="499">
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<subSubSection id="C35EB2E8EA20FFF23CCC7D87C532FD69" box="[151,563,654,681]" pageId="3" pageNumber="499" type="nomenclature">
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<paragraph id="8BFBE163EA20FFF23CCC7D87C532FD69" blockId="3.[151,563,654,681]" box="[151,563,654,681]" pageId="3" pageNumber="499">
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<heading id="D0B3560FEA20FFF23CCC7D87C532FD69" bold="true" box="[151,563,654,681]" fontSize="11" level="1" pageId="3" pageNumber="499" reason="1">
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<emphasis id="B9303D71EA20FFF23CCC7D87C532FD69" bold="true" box="[151,563,654,681]" pageId="3" pageNumber="499">
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<taxonomicName id="4C449AE0EA20FFF23CCC7D87C618FD68" ID-CoL="63Y" authorityName="Leach" authorityYear="1815" box="[151,281,654,680]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="499" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
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in the early Paleogene
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35EB2E8EA20FFF23CCC7DDEC42AFBF5" pageId="3" pageNumber="499" type="reference_group">
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<paragraph id="8BFBE163EA20FFF23CCC7DDEC569FC41" blockId="3.[151,1437,727,1257]" pageId="3" pageNumber="499">
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<taxonomicName id="4C449AE0EA20FFF23CCC7DDEC60CFD31" ID-CoL="63Y" authorityName="Leach" authorityYear="1815" box="[151,269,727,753]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="499" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
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comprise the majority of dragonflies described in the early Paleogene, see Table 1 of
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<bibRefCitation id="EFD59C92EA20FFF2388A7DDEC62BFCD5" author="Archibald, S. B. & Cannings, R. A." pageId="3" pageNumber="499" pagination="783 - 816" refId="ref2515" refString="Archibald, S. B. & Cannings, R. A. (2019) Fossil dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands, western North America. The Canadian Entomologist, 151, 783 - 816. https: // doi. org / 10.4039 / tce. 2019.61" type="journal article" year="2019">Archibald & Cannings (2019)</bibRefCitation>
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. They are still diverse and plentiful today, although they have been overtaken in diversity by the
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<taxonomicName id="4C449AE0EA20FFF239217DF2C609FCF9" ID-CoL="BZS" authorityName="Rambur" authorityYear="1842" class="Insecta" family="Libellulidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="499" phylum="Arthropoda" rank="family">Libellulidae</taxonomicName>
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, which appear first in the Oligocene, and the Gomphidae, which has a great diversity in the late Jurassic through mid-Cretaceous, a single species in the Ypresian (early Eocene), and then is better known beginning in the Oligocene (
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<bibRefCitation id="EFD59C92EA20FFF23D427C6EC55DFC41" author="Archibald, S. B. & Cannings, R. A." box="[281,604,871,897]" pageId="3" pageNumber="499" pagination="783 - 816" refId="ref2515" refString="Archibald, S. B. & Cannings, R. A. (2019) Fossil dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands, western North America. The Canadian Entomologist, 151, 783 - 816. https: // doi. org / 10.4039 / tce. 2019.61" type="journal article" year="2019">Archibald & Cannings 2019</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BFBE163EA20FFF23C9C7C82C42AFBF5" blockId="3.[151,1437,727,1257]" pageId="3" pageNumber="499">
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In North America, fossil
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<taxonomicName id="4C449AE0EA20FFF23D837C82C54FFC65" ID-CoL="63Y" authorityName="Leach" authorityYear="1815" box="[472,590,907,933]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="499" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
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are found in the Paleocene of
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<collectingRegion id="49802F81EA20FFF23FC07C82C4EEFC65" box="[923,1007,907,933]" country="Canada" name="Alberta" pageId="3" pageNumber="499">Alberta</collectingRegion>
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and
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<collectingRegion id="49802F81EA20FFF238787C82C3BAFC65" box="[1059,1211,907,933]" country="United States of America" name="North Dakota" pageId="3" pageNumber="499">North Dakota</collectingRegion>
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(
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<bibRefCitation id="EFD59C92EA20FFF238937C82C7D2FC0A" author="Wighton, D. C. & Wilson, M. V. H." pageId="3" pageNumber="499" pagination="505 - 522" refId="ref3550" refString="Wighton, D. C. & Wilson, M. V. H. (1986) The Gomphaeschninae (Odonata: Aeshnidae): new fossil genus, reconstructed phylogeny, and geographical history. Systematic Entomology, 11, 505 - 522. https: // doi. org / 10.1111 / j. 1365 - 3113.1986. tb 00539. x" type="journal article" year="1986">Wighton & Wilson 1986</bibRefCitation>
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), the Paleocene or Eocene of
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<collectingRegion id="49802F81EA20FFF23E7F7CA6C573FC09" box="[548,626,943,969]" country="United States of America" name="Alaska" pageId="3" pageNumber="499">Alaska</collectingRegion>
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(
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<bibRefCitation id="EFD59C92EA20FFF23ED97CA6C484FC09" author="Garrouste, R. & Nel, A." box="[642,901,943,969]" pageId="3" pageNumber="499" pagination="1939 - 1946" refId="ref2894" refString="Garrouste, R. & Nel, A. (2019) Alaskan Palaeogene insects: a challenge for a better knowledge of the Beringian ' route' (Odonata: Aeshnidae, Dysagrionidae). Journal of Systematic Palaeontology, 17, 1939 - 1946. https: // doi. org / 10.1080 / 14772019.2019.1572235" type="journal article" year="2019">Garrouste & Nel 2019</bibRefCitation>
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), the Ypresian of the Okanagan Highlands (
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<bibRefCitation id="EFD59C92EA20FFF2392F7CA6C6B8FC2D" author="Archibald, S. B. & Cannings, R. A." pageId="3" pageNumber="499" pagination="783 - 816" refId="ref2515" refString="Archibald, S. B. & Cannings, R. A. (2019) Fossil dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands, western North America. The Canadian Entomologist, 151, 783 - 816. https: // doi. org / 10.4039 / tce. 2019.61" type="journal article" year="2019">Archibald & Cannings 2019</bibRefCitation>
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), and the Priabonian (late Eocene) at Florissant (summarised by
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<bibRefCitation id="EFD59C92EA20FFF238D37CDAC210FC2D" author="Meyer, W. H." box="[1160,1297,979,1005]" pageId="3" pageNumber="499" refId="ref3358" refString="Meyer, W. H. (2003) The Fossils of Florissant. Smithsonian Books, Washington, 258 pp." type="book" year="2003">Meyer 2003</bibRefCitation>
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) about 1200 kilometers to the southeast. The Gomphaeschninae is well represented in Paleocene and Eocene localities of Europe and North America (
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<bibRefCitation id="EFD59C92EA20FFF23D277B12C5BFFBF5" author="Archibald, S. B. & Cannings, R. A." box="[380,702,1051,1077]" pageId="3" pageNumber="499" pagination="783 - 816" refId="ref2515" refString="Archibald, S. B. & Cannings, R. A. (2019) Fossil dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands, western North America. The Canadian Entomologist, 151, 783 - 816. https: // doi. org / 10.4039 / tce. 2019.61" type="journal article" year="2019">Archibald & Cannings 2019</bibRefCitation>
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, Table 1).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35EB2E8EA20FFF23C9C7B36C647FB29" pageId="3" pageNumber="499" type="description">
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<paragraph id="8BFBE163EA20FFF23C9C7B36C647FB29" blockId="3.[151,1437,727,1257]" pageId="3" pageNumber="499">
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Lutetian dragonflies have only been reported from Eckfeld Maar (including
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<taxonomicName id="4C449AE0EA20FFF238427B36C3EDFB99" box="[1049,1260,1087,1113]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="497" phylum="Arthropoda" rank="subFamily" subFamily="Gomphaeschninae">Gomphaeschninae</taxonomicName>
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:
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<bibRefCitation id="EFD59C92EA20FFF238AD7B36C294FB99" author="Wappler, T." box="[1270,1429,1087,1113]" pageId="3" pageNumber="499" pagination="1 - 234" refId="ref3479" refString="Wappler, T. (2003) Die Insekten aus dem Mittel-Eozan des Eckfelder Maares, Vulkaneifel. Mainzer Naturwissenschaftliches Archiv Beiheft, 27, 1 - 234." type="journal article" year="2003">Wappler 2003</bibRefCitation>
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) (Grube Messel,
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<collectingCountry id="F353A1F3EA20FFF23D127B6AC6B3FBBD" box="[329,434,1123,1149]" name="Germany" pageId="3" pageNumber="499">Germany</collectingCountry>
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, referred to as Lutetian by
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<bibRefCitation id="EFD59C92EA20FFF23E817B6AC315FBBD" author="Archibald, S. B. & Cannings, R. A." box="[730,1044,1123,1149]" pageId="3" pageNumber="499" pagination="783 - 816" refId="ref2515" refString="Archibald, S. B. & Cannings, R. A. (2019) Fossil dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands, western North America. The Canadian Entomologist, 151, 783 - 816. https: // doi. org / 10.4039 / tce. 2019.61" type="journal article" year="2019">Archibald & Cannings 2019</bibRefCitation>
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is now dated as Ypresian including the Ypresian/Lutetian boundary by
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<bibRefCitation id="EFD59C92EA20FFF23E727B8EC5EBFB61" author="Lenz, O. K. & Wilde, V. & Mertz, D. F. & Riegel, W." box="[553,746,1158,1185]" pageId="3" pageNumber="499" pagination="873 - 889" refId="ref3278" refString="Lenz, O. K., Wilde, V., Mertz, D. F. & Riegel, W. (2015) New palynology-based astronomical and revised 40 Ar / 39 Ar ages for the Eocene maar lake of Messel (Germany). International Journal of Earth Sciences, Geologische Rundschau, 104, 873 - 889. https: // doi. org / 10.1007 / s 00531 - 014 - 1126 - 2" type="journal article" year="2015">
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Lenz
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<emphasis id="B9303D71EA20FFF23E317B8EC59CFB61" box="[618,669,1159,1185]" italics="true" pageId="3" pageNumber="499">et al</emphasis>
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. 2015
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</bibRefCitation>
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). Bartonian dragonflies are not known.
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<taxonomicName id="4C449AE0EA20FFF238EA7B8EC280FB60" authorityName="Archibald & Cannings & Greenwalt" authorityYear="2022" box="[1201,1409,1159,1185]" class="Insecta" family="Aeshnidae" genus="Kishenehna" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="499" phylum="Arthropoda" rank="species" species="prima">
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<emphasis id="B9303D71EA20FFF238EA7B8EC280FB60" box="[1201,1409,1159,1185]" italics="true" pageId="3" pageNumber="499">Kishenehna prima</emphasis>
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</taxonomicName>
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is the only dragonfly of this time known in the Western Hemisphere, increasing our scant knowledge of them in the middle Eocene.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |