treatments-xml/data/03/87/47/038747324C34C6111EB72C8FFDC9260E.xml
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<document id="D6997962777EA3CA49A845D4863A8C01" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C34C6111EB72C8FFDC9260E" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteropsis barbarae Lees &amp; Kremen, sp. nov." docType="treatment" docVersion="9" lastPageNumber="70" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="67" updateTime="1698662715872" updateUser="ExternalLinkService">
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<mods:title id="4FBB69259C28476E6F1BA2E0E0626753">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:namePart id="31090D394ECFEB386474B5406247019C">C, Lees David</mods:namePart>
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<mods:typeOfResource id="6D9285F7B3D6CA59C667F59D7043022C">text</mods:typeOfResource>
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<mods:date id="32FA70A4CD9E23268EE51E443D50C8E4">2016</mods:date>
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<mods:number id="FBC302F5B53B44403048FD941220A595">4118</mods:number>
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<mods:identifier id="2B3657E26E252C5E0F06FC096723987B" type="DOI">10.11646/zootaxa.4118.1.1</mods:identifier>
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<treatment id="038747324C34C6111EB72C8FFDC9260E" ID-DOI="http://doi.org/10.5281/zenodo.6086448" ID-GBIF-Taxon="120817147" ID-Zenodo-Dep="6086448" LSID="urn:lsid:plazi:treatment:038747324C34C6111EB72C8FFDC9260E" httpUri="http://treatment.plazi.org/id/038747324C34C6111EB72C8FFDC9260E" lastPageId="70" lastPageNumber="70" pageId="66" pageNumber="67">
<subSubSection id="C334A5AF4C34C6151EB72C8FFBC120AA" pageId="66" pageNumber="67" type="nomenclature">
<paragraph id="8B91F6244C34C6151EB72C8FFD1620C9" blockId="66.[151,1042,418,478]" box="[151,709,418,445]" pageId="66" pageNumber="67">
<heading id="D0D941484C34C6151EB72C8FFD1620C9" bold="true" box="[151,709,418,445]" fontSize="11" level="1" pageId="66" pageNumber="67" reason="1">
<emphasis id="B95A2A364C34C6151EB72C8FFD1620C9" bold="true" box="[151,709,418,445]" pageId="66" pageNumber="67">
<taxonomicName id="4C2E8DA74C34C6151EB72C8FFD8820C9" authority="Lees &amp; Kremen" authorityName="Lees &amp; Kremen" box="[151,603,418,445]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
<emphasis id="B95A2A364C34C6151EB72C8FFE5F20C8" bold="true" box="[151,396,418,444]" italics="true" pageId="66" pageNumber="67">Heteropsis barbarae</emphasis>
Lees &amp; Kremen
</taxonomicName>
,
<taxonomicNameLabel id="A269974D4C34C6151C482C8EFD1620C9" box="[616,709,419,445]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B91F6244C34C6151EB72CEBFBC120AA" blockId="66.[151,1042,418,478]" box="[151,1042,453,478]" pageId="66" pageNumber="67">LSID: urn:lsid:zoobank.org:act:DC87986A-3822-43C9-B1B9-83A2714D088B</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C34C6151EB72F27FC812357" box="[151,850,522,547]" pageId="66" pageNumber="67" type="description">
<paragraph id="8B91F6244C34C6151EB72F27FC812357" blockId="66.[151,1437,522,2035]" box="[151,850,522,547]" pageId="66" pageNumber="67">
<emphasis id="B95A2A364C34C6151EB72F27FE842357" bold="true" box="[151,343,522,547]" pageId="66" pageNumber="67">Prior references</emphasis>
: sp. 16 (
<bibRefCitation id="EFBF8BD54C34C6151F982F26FDE32357" author="Lees" box="[440,560,523,547]" pageId="66" pageNumber="67" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
;
<bibRefCitation id="EFBF8BD54C34C6151C1C2F26FCDF2357" author="Torres" box="[572,780,522,547]" pageId="66" pageNumber="67" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres
<emphasis id="B95A2A364C34C6151CAA2F21FD1A2357" box="[650,713,522,547]" italics="true" pageId="66" pageNumber="67">et al.,</emphasis>
2001
</bibRefCitation>
: 462).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C34C6151EE72F01FE6122BA" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="8B91F6244C34C6151EE72F01FC2F23F9" blockId="66.[151,1437,522,2035]" pageId="66" pageNumber="67">
<emphasis id="B95A2A364C34C6151EE72F01FD202331" bold="true" box="[199,755,556,581]" pageId="66" pageNumber="67">
<typeStatus id="549548864C34C6151EE72F01FED12331" box="[199,258,556,581]" pageId="66" pageNumber="67">Type</typeStatus>
material., Deposition
<collectionCode id="ED3F6EE14C34C6151C382F01FDA62331" box="[536,629,556,581]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="66" pageNumber="67">MNHN</collectionCode>
:
<typeStatus id="549548864C34C6151CA62F01FD202331" box="[646,755,556,581]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
: ♂ (
<figureCitation id="1315EAA14C34C6151D102F01FC552331" box="[816,902,556,581]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
A),
<collectingCountry id="F339B6B44C34C6151D962F01FBEE2331" box="[950,1085,556,581]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
NE, Maitsoarongana, Sikiory River, near camp,
<geoCoordinate id="EE1A90E34C34C6151F4E2F7FFE32231E" box="[366,481,590,619]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="5" value="-15.1885">15.1885o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C34C6151FD12F7FFDB6231E" box="[497,613,590,619]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="5" value="49.5163">49.5163o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C34C6151CBA2F7FFD2F231F" box="[666,764,594,619]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="66" pageNumber="67" unit="km" value="0.02">0.02 km</quantity>
,
<quantity id="4CD65BC14C34C6151D2B2F7FFC7B231E" box="[779,936,594,619]" metricMagnitude="2" metricUnit="m" metricValue="6.85" metricValueMax="7.08" metricValueMin="6.62" pageId="66" pageNumber="67" unit="m" value="685.0" valueMax="708.0" valueMin="662.0">685 +/- 23 m</quantity>
,
<date id="FF90D0E44C34C6151D972F7FFBEF231E" box="[951,1084,594,618]" pageId="66" pageNumber="67" value="2001-12-12">12/12/2001</date>
: 15:29, D.C. Lees:
<collectionCode id="ED3F6EE14C34C6151B002F7FFA9A231E" box="[1312,1353,594,618]" pageId="66" pageNumber="67">DL</collectionCode>
01-227,
<collectionCode id="ED3F6EE14C34C6151EB72F59FF3D23F8" box="[151,238,628,652]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E)
<date id="FF90D0E44C34C6151F352F59FEA023F9" box="[277,371,628,653]" pageId="66" pageNumber="67" value="2008" valueMax="2069">2008-69</date>
[accession number],
<collectionCode id="ED3F6EE14C34C6151C422F59FD6A23F8" box="[610,697,628,652]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E) 1717096 [
<collectingCountry id="F339B6B44C34C6151D702F59FC5823F8" box="[848,907,628,652]" name="Qatar" pageId="66" pageNumber="67">QTR</collectingCountry>
barcode].
</paragraph>
<paragraph id="8B91F6244C34C6151EE72FBBFE6122BA" blockId="66.[151,1437,522,2035]" pageId="66" pageNumber="67">
<emphasis id="B95A2A364C34C6151EE72FBBFD6F23DB" bold="true" box="[199,700,662,687]" pageId="66" pageNumber="67">
<typeStatus id="549548864C34C6151EE72FBBFEEC23DB" box="[199,319,662,687]" pageId="66" pageNumber="67" type="paratype">Paratypes</typeStatus>
(accession
<collectionCode id="ED3F6EE14C34C6151FED2FBBFDFB23DB" box="[461,552,662,687]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E)
<date id="FF90D0E44C34C6151C752FBBFD6023DB" box="[597,691,662,687]" pageId="66" pageNumber="67" value="2008" valueMax="2069">2008-69</date>
)
</emphasis>
: ♂,
<collectingCountry id="F339B6B44C34C6151CD62FBBFCAE23DB" box="[758,893,662,687]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
NE, Maitsoarongana, Sikiory R.,
<quantity id="4CD65BC14C34C6151B312FBBFA4B23DA" box="[1297,1432,662,687]" metricMagnitude="2" metricUnit="m" metricValue="7.05" metricValueMax="7.5" metricValueMin="6.6" pageId="66" pageNumber="67" unit="m" value="705.0" valueMax="750.0" valueMin="660.0">660750 m</quantity>
,
<geoCoordinate id="EE1A90E34C34C6151EB72F96FED923A7" box="[151,266,695,724]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="5" value="-15.1907">15.1907o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C34C6151F3C2F96FE4323A7" box="[284,400,695,724]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="5" value="49.5309">49.5309o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C34C6151FE92F96FDDA23A0" box="[457,521,699,724]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="66" pageNumber="67" unit="km" value="2.0">2 km</quantity>
,
<quantity id="4CD65BC14C34C6151C392F96FD6923A7" box="[537,698,699,724]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="7.6" metricValueMin="7.4" pageId="66" pageNumber="67" unit="m" value="750.0" valueMax="760.0" valueMin="740.0">750 +/- 10 m</quantity>
<date id="FF90D0E44C34C6151CE52F96FC9B23A7" box="[709,840,699,723]" pageId="66" pageNumber="67" value="2001-12-12">12/12/2001</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C34C6151DC12F96FBD923A7" box="[993,1034,699,723]" pageId="66" pageNumber="67">DL</collectionCode>
01-241,
<collectionCode id="ED3F6EE14C34C6151A492F96FB1323A7" box="[1129,1216,699,723]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E) 1717097 [
<collectingCountry id="F339B6B44C34C6151B412F96FA4F23A7" box="[1377,1436,699,723]" name="Qatar" pageId="66" pageNumber="67">QTR</collectingCountry>
barcode]; ♂, NE, Maitsoarongana, c.
<quantity id="4CD65BC14C34C6151C602FCDFD59238C" box="[576,650,736,760]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="66" pageNumber="67" unit="m" value="710.0">710 m</quantity>
,
<geoCoordinate id="EE1A90E34C34C6151CB72FCDFCD8238C" box="[663,779,733,761]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="5" value="-15.1885">15.1885o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C34C6151D392FCDFC5F238C" box="[793,908,733,761]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="5" value="49.5163">49.5163o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C34C6151D9D2FCDFBCD238D" box="[957,1054,736,761]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="66" pageNumber="67" unit="km" value="0.02">0.02 km</quantity>
,
<quantity id="4CD65BC14C34C6151A0A2FCDFB6A238C" box="[1066,1209,736,760]" metricMagnitude="2" metricUnit="m" metricValue="7.1" metricValueMax="7.33" metricValueMin="6.87" pageId="66" pageNumber="67" unit="m" value="710.0" valueMax="733.0" valueMin="687.0">710 +/- 23m</quantity>
,
<date id="FF90D0E44C34C6151AE62FCDFA9A238C" box="[1222,1353,736,760]" pageId="66" pageNumber="67" value="2001-12-13">13/12/2001</date>
: 06:00, D.C. Lees:
<collectionCode id="ED3F6EE14C34C6151F032E2FFE9F226E" box="[291,332,770,794]" pageId="66" pageNumber="67">DL</collectionCode>
01-247,
<collectionCode id="ED3F6EE14C34C6151F8C2E2FFDD0226E" box="[428,515,770,794]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E) 1717098 [
<collectingCountry id="F339B6B44C34C6151C872E2FFD31226E" box="[679,738,770,794]" name="Qatar" pageId="66" pageNumber="67">QTR</collectingCountry>
barcode]; ♂, NE, Maitsoarongana, ridge just S. of camp,
<geoCoordinate id="EE1A90E34C34C6151EB72E0AFED9224B" box="[151,266,804,832]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="5" value="-15.1907">15.1907o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C34C6151F372E0AFE58224B" box="[279,395,804,832]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="5" value="49.5309">49.5309o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C34C6151F9B2E0AFE2B2234" box="[443,504,807,832]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="66" pageNumber="67" unit="km" value="2.0">2 km</quantity>
,
<quantity id="4CD65BC14C34C6151C242E0AFD44224B" box="[516,663,807,832]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="7.6" metricValueMin="7.4" pageId="66" pageNumber="67" unit="m" value="750.0" valueMax="760.0" valueMin="740.0">750 +/- 10 m</quantity>
<date id="FF90D0E44C34C6151CBE2E0AFCC6224B" box="[670,789,807,831]" pageId="66" pageNumber="67" value="2001-12-07">7/12/2001</date>
: 16:39, D.C. Lees:
<collectionCode id="ED3F6EE14C34C6151DCF2E0AFBCB224B" box="[1007,1048,807,831]" pageId="66" pageNumber="67">DL</collectionCode>
01-132,
<collectionCode id="ED3F6EE14C34C6151A522E0AFB1A224B" box="[1138,1225,807,831]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E) 1717099 [
<collectingCountry id="F339B6B44C34C6151B412E0AFA4F224B" box="[1377,1436,807,831]" name="Qatar" pageId="66" pageNumber="67">QTR</collectingCountry>
barcode]; ♀ (
<figureCitation id="1315EAA14C34C6151F0C2E61FE532210" box="[300,384,844,869]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
B),
<collectingCountry id="F339B6B44C34C6151F872E61FDFD2211" box="[423,558,844,869]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
NE, Sikiory R., riparian,
<geoCoordinate id="EE1A90E34C34C6151D702E61FC172210" box="[848,964,840,869]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="5" value="-15.1848">15.1848o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C34C6151DF12E61FB942210" box="[977,1095,840,869]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="5" value="49.4659">49.4659o E</geoCoordinate>
, +/-
<quantity id="4CD65BC14C34C6151A5B2E61FB092211" box="[1147,1242,844,869]" metricMagnitude="3" metricUnit="m" metricValue="5.42" pageId="66" pageNumber="67" unit="km" value="5.42">5.42 km</quantity>
,
<quantity id="4CD65BC14C34C6151AC52E61FABF2210" box="[1253,1388,844,869]" metricMagnitude="2" metricUnit="m" metricValue="7.1" metricValueMax="7.15" metricValueMin="7.05" pageId="66" pageNumber="67" unit="m" value="710.0" valueMax="715.0" valueMin="705.0">710 +/- 5 m</quantity>
,
<date id="FF90D0E44C34C6151B582E61FF2422F3" pageId="66" pageNumber="67" value="2001-12-12">12/ 12/2001</date>
: 17:12, D.C. Lees:
<collectionCode id="ED3F6EE14C34C6151FF22E42FE2822F3" box="[466,507,879,903]" pageId="66" pageNumber="67">DL</collectionCode>
01-231,
<collectionCode id="ED3F6EE14C34C6151C752E42FD7F22F3" box="[597,684,879,903]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E) 1717100 [
<collectingCountry id="F339B6B44C34C6151D652E42FC5322F3" box="[837,896,879,903]" name="Qatar" pageId="66" pageNumber="67">QTR</collectingCountry>
barcode]; ♂,
<collectingCountry id="F339B6B44C34C6151A3E2E43FB7622F3" box="[1054,1189,878,903]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
NE, Maitsoarongana, near path,
<geoCoordinate id="EE1A90E34C34C6151F372EBEFE5922DF" box="[279,394,911,940]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="5" value="-15.1907">
15.1907
<emphasis id="B95A2A364C34C6151F522EA2FEA922E9" bold="true" box="[370,378,911,925]" pageId="66" pageNumber="67">o</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C34C6151FBC2EBEFDC122DF" box="[412,530,911,940]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="5" value="49.5309">49.5309o E</geoCoordinate>
, +/-
<quantity id="4CD65BC14C34C6151C712EBEFD4622D8" box="[593,661,915,940]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="66" pageNumber="67" unit="km" value="2.0">2 km</quantity>
.,
<quantity id="4CD65BC14C34C6151C892EBEFC8D22DF" box="[681,862,915,940]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="66" pageNumber="67" unit="m" value="750.0" valueMax="850.0" valueMin="650.0">
750 +/-
<specimenCount id="9D283DAD4C34C6151D2C2EBEFC8D22DF" box="[780,862,915,939]" pageId="66" pageNumber="67" type="generic">100 m</specimenCount>
</quantity>
.,
<date id="FF90D0E44C34C6151D532EBEFC2522DF" box="[883,1014,915,939]" pageId="66" pageNumber="67" value="2001-12-13">13/12/2001</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C34C6151AB02EBEFB6A22DF" box="[1168,1209,915,939]" pageId="66" pageNumber="67">DL</collectionCode>
01-248,
<collectionCode id="ED3F6EE14C34C6151B042EBEFAA822DF" box="[1316,1403,915,939]" pageId="66" pageNumber="67">BMNH</collectionCode>
(E) 1717095 [
<collectingCountry id="F339B6B44C34C6151F262E9BFE9222BA" box="[262,321,950,974]" name="Qatar" pageId="66" pageNumber="67">QTR</collectingCountry>
barcode].
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C34C6151EE72EFAFC8B2284" box="[199,856,983,1008]" pageId="66" pageNumber="67" type="description">
<paragraph id="8B91F6244C34C6151EE72EFAFC8B2284" blockId="66.[151,1437,522,2035]" box="[199,856,983,1008]" pageId="66" pageNumber="67">
<emphasis id="B95A2A364C34C6151EE72EFAFE6D2284" bold="true" box="[199,446,983,1008]" pageId="66" pageNumber="67">Deposition summary</emphasis>
: BMNH (HT ♂, 4 PT ♂♂, 1 PT ♀).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C34C6151EE72ED1FF672542" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="8B91F6244C34C6151EE72ED1FF672542" blockId="66.[151,1437,522,2035]" pageId="66" pageNumber="67">
<emphasis id="B95A2A364C34C6151EE72ED1FEB52561" bold="true" box="[199,358,1020,1045]" pageId="66" pageNumber="67">
<typeStatus id="549548864C34C6151EE72ED1FED12561" box="[199,258,1020,1045]" pageId="66" pageNumber="67">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F339B6B44C34C6151F4D2ED1FE272561" box="[365,500,1020,1045]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
NE, Maitsoarongana, Sikiory River,
<geoCoordinate id="EE1A90E34C34C6151DB72ED1FBD82561" box="[919,1035,1017,1045]" direction="south" orientation="latitude" pageId="66" pageNumber="67" precision="5" value="-15.1885">15.1885o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C34C6151A382ED1FB5F2560" box="[1048,1164,1017,1045]" direction="east" orientation="longitude" pageId="66" pageNumber="67" precision="5" value="49.5163">49.5163o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C34C6151A9B2ED0FAC82561" box="[1211,1307,1020,1045]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="66" pageNumber="67" unit="km" value="0.02">0.02 km</quantity>
,
<quantity id="4CD65BC14C34C6151B062ED1FF632542" metricMagnitude="2" metricUnit="m" metricValue="6.85" metricValueMax="7.08" metricValueMin="6.62" pageId="66" pageNumber="67" unit="m" value="685.0" valueMax="708.0" valueMin="662.0">
685 +/-
<specimenCount id="9D283DAD4C34C6151BA02ED0FF632542" pageId="66" pageNumber="67" type="generic">23 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C34C6151EE7296DFC632743" pageId="66" pageNumber="67" type="diagnosis">
<paragraph id="8B91F6244C34C6151EE7296DFC632743" blockId="66.[151,1437,522,2035]" pageId="66" pageNumber="67">
<emphasis id="B95A2A364C34C6151EE7296DFE92252D" bold="true" box="[199,321,1088,1113]" pageId="66" pageNumber="67">Diagnosis.</emphasis>
The ventral wings of
<taxonomicName id="4C2E8DA74C34C6151C17296CFD1A252D" box="[567,713,1088,1113]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C34C6151C17296CFD1A252D" box="[567,713,1088,1113]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
</taxonomicName>
are similar to some other members of the
<taxonomicName id="4C2E8DA74C34C6151A82296CFAF0252D" box="[1186,1315,1088,1113]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
<emphasis id="B95A2A364C34C6151A82296CFAF0252D" box="[1186,1315,1088,1113]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
</taxonomicName>
group that exhibit a crenulate HW margin, notably
<taxonomicName id="4C2E8DA74C34C6151C582949FC9E250F" box="[632,845,1122,1147]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="menamenoides" status="sp. nov.">
<emphasis id="B95A2A364C34C6151C582949FC9E250F" box="[632,845,1122,1147]" italics="true" pageId="66" pageNumber="67">Ht. menamenoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269974D4C34C6151D79294FFC64250F" box="[857,951,1122,1147]" pageId="66" pageNumber="67" rank="species">
<emphasis id="B95A2A364C34C6151D79294FFC64250F" bold="true" box="[857,951,1122,1147]" pageId="66" pageNumber="67">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C2E8DA74C34C6151DD72949FBAE250F" box="[1015,1149,1122,1147]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
<emphasis id="B95A2A364C34C6151DD72949FBAE250F" box="[1015,1149,1122,1147]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
</taxonomicName>
. Upperside dark brown, lacking light orange wash and smooth HW margin of
<taxonomicName id="4C2E8DA74C34C6151CD429ABFC1625E9" box="[756,965,1156,1181]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="menamenoides">
<emphasis id="B95A2A364C34C6151CD429ABFC1625E9" box="[756,965,1156,1181]" italics="true" pageId="66" pageNumber="67">Ht. menamenoides</emphasis>
</taxonomicName>
(
<figureCitation id="1315EAA14C34C6151DF529A9FBF625E9" box="[981,1061,1156,1181]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, ( Antakares; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 ( Antakares; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 ( MadagascarNW; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
A) and
<taxonomicName id="4C2E8DA74C34C6151A5B29ABFB3D25E9" box="[1147,1262,1158,1181]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="maeva">
<emphasis id="B95A2A364C34C6151A5B29ABFB3D25E9" box="[1147,1262,1158,1181]" italics="true" pageId="66" pageNumber="67">Ht. maeva</emphasis>
</taxonomicName>
(
<figureCitation id="1315EAA14C34C6151ADE29A9FA9C25E9" box="[1278,1359,1156,1181]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, ( Antakares; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 ( Antakares; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 ( MadagascarNW; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
D) and the HWD orange crescent (Mf) typical of
<taxonomicName id="4C2E8DA74C34C6151CA22985FCD625CB" box="[642,773,1190,1215]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
<emphasis id="B95A2A364C34C6151CA22985FCD625CB" box="[642,773,1190,1215]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
</taxonomicName>
. Fresh specimens of
<taxonomicName id="4C2E8DA74C34C6151DDF2985FB4725CB" box="[1023,1172,1190,1215]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C34C6151DDF2985FB4725CB" box="[1023,1172,1190,1215]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
</taxonomicName>
are distinctive in their strong slightly dingy orange ventral cast (
<figureCitation id="1315EAA14C34C6151C4629E4FD6A2595" box="[614,697,1225,1250]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
A,E), and the ventral scales are easily displaced/rubbed. Worn ♀♀ can easily be confused with those of often sympatric
<taxonomicName id="4C2E8DA74C34C6151CCB29C1FCB72477" box="[747,868,1260,1283]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="B95A2A364C34C6151CCB29C1FCB72477" box="[747,868,1260,1283]" italics="true" pageId="66" pageNumber="67">Ht. pauper</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2E8DA74C34C6151DBB29C1FBFF2470" box="[923,1068,1259,1284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C34C6151DBB29C1FBFF2470" box="[923,1068,1259,1284]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
</taxonomicName>
. Indeed the ♀ of
<taxonomicName id="4C2E8DA74C34C6151AD129C1FA522470" box="[1265,1409,1259,1284]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C34C6151AD129C1FA522470" box="[1265,1409,1259,1284]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
</taxonomicName>
is sometimes almost indistinguishable (
<figureCitation id="1315EAA14C34C6151C172820FD5A2452" box="[567,649,1293,1318]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
D) to that of
<taxonomicName id="4C2E8DA74C34C6151D392823FC7F2452" box="[793,940,1293,1318]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C34C6151D392823FC7F2452" box="[793,940,1293,1318]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
</taxonomicName>
(
<figureCitation id="1315EAA14C34C6151D9B2820FBDC2452" box="[955,1039,1293,1318]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
B), but
<taxonomicName id="4C2E8DA74C34C6151A412823FB202452" box="[1121,1267,1293,1318]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C34C6151A412823FB202452" box="[1121,1267,1293,1318]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
</taxonomicName>
♀♀ often tend to be more flushed with reddish scales on the upperside, while
<taxonomicName id="4C2E8DA74C34C6151D87281DFB93243C" box="[935,1088,1327,1352]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C34C6151D87281DFB93243C" box="[935,1088,1327,1352]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
</taxonomicName>
can be distinguished by the combination of rich and dense slightly dingy orange scales on the ventral surface which lack a yellow cast (such as in
<taxonomicName id="4C2E8DA74C34C6151E942858FEE724F9" box="[180,308,1396,1421]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="strigula">
<emphasis id="B95A2A364C34C6151E942858FEE724F9" box="[180,308,1396,1421]" italics="true" pageId="66" pageNumber="67">Ht. strigula</emphasis>
</taxonomicName>
), combined with a less contrasting, pale orange band between Pml and Sml (
<figureCitation id="1315EAA14C34C6151AB22859FB3624F8" box="[1170,1253,1396,1421]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="66" pageNumber="67">Fig. 20</figureCitation>
C,D,F), which is pale ochreous in both sexes of some populations of
<taxonomicName id="4C2E8DA74C34C6151CF528BAFCB524DA" box="[725,870,1429,1454]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C34C6151CF528BAFCB524DA" box="[725,870,1429,1454]" italics="true" pageId="66" pageNumber="67">Ht. undulans</emphasis>
</taxonomicName>
(however that is a very variable species), but there will always be a few especially more worn specimens of
<taxonomicName id="4C2E8DA74C34C6151D302894FC7124A5" box="[784,930,1464,1489]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C34C6151D302894FC7124A5" box="[784,930,1464,1489]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
</taxonomicName>
that are hard to place without dissection. The similarly lowland
<taxonomicName id="4C2E8DA74C34C6151F4228F6FE062486" box="[354,469,1499,1522]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="maeva">
<emphasis id="B95A2A364C34C6151F4228F6FE062486" box="[354,469,1499,1522]" italics="true" pageId="66" pageNumber="67">Ht. maeva</emphasis>
</taxonomicName>
has a brighter orange underside (
<figureCitation id="1315EAA14C34C6151D6928F7FC492487" box="[841,922,1498,1523]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, ( Antakares; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 ( Antakares; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 ( MadagascarNW; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="66" pageNumber="67">Fig. 21</figureCitation>
D), denser black ventral brush in the ♂, and is distinctive in that its upperside is orange too, except for HWs of western specimens which are usually suffused with brown (but
<taxonomicName id="4C2E8DA74C34C6151F392B32FE7F2743" box="[281,428,1566,1591]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C34C6151F392B32FE7F2743" box="[281,428,1566,1591]" italics="true" pageId="66" pageNumber="67">Ht. barbarae</emphasis>
</taxonomicName>
is anyway absent in the west of
<collectingCountry id="F339B6B44C34C6151D372B33FC772743" box="[791,932,1566,1591]" name="Madagascar" pageId="66" pageNumber="67">Madagascar</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C34C6131EE72B6DFD912229" lastPageId="68" lastPageNumber="69" pageId="66" pageNumber="67" type="description">
<paragraph id="8B91F6244C34C6131EE72B6DFEC020A4" blockId="66.[151,1437,522,2035]" lastBlockId="68.[151,1437,151,2013]" lastPageId="68" lastPageNumber="69" pageId="66" pageNumber="67">
<emphasis id="B95A2A364C34C6151EE72B6DFE7C272D" bold="true" box="[199,431,1600,1625]" pageId="66" pageNumber="67">Description. Wings</emphasis>
: Upperside mid-brown with a slight russet brown cast. Space-CuA1 ocellus expressed strongly in FWD; orange ring sub-hexagonal., M1 ocellus very small black white pupilled spot surrounded by a narrow orange ring. HWD ocellus CuA1 fairly small and sub-elliptic, surrounded by a narrow orange ring. Underside has a lighter dingy orange cast, only slightly lighter distad of the Mb. Mb darker orange-brown, gently convex and slightly irregular (relatively straight in the HT HWV) with paler orange highlights distal to it in space- M2 to CuA1. FWV space-CuA1 ocellus similar size to FWD (in the HT), as is FWV M1 ocellus, which is reduced to a very small white point surrounded by a narrow black ring and orange ring that slightly contrasts with background. FWV space-CuA1 ocellus follows concave shape of the FWV darker orange-brown Mb before it bends back toward mid costa, with a ring that is orange near the black iris (which is significantly smaller than on FWD) and is yellowy-orange towards the greatest inflection of the bend in the Mb, forming a spiral arm (Ore). HWV space-CuA1 ocellus similar size and shape to that on dorsal surface, surrounded by pale orange ring not much lighter than background; no other ocelli conspicuously expressed on the HWV nor HWD. The HWV basal area is sparsely irrorated with brown strigulae, the area between PMb and Mb more evenly and finely irrorated. A wavy-zigzag darker diffuse brownish line inset from margin follows a double narrow Sml divided by the ground colour, not very ochreous as usual in
<taxonomicName id="4C2E8DA74C32C6131C1D2D90FD1D21A1" box="[573,718,188,213]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C32C6131C1D2D90FD1D21A1" box="[573,718,188,213]" italics="true" pageId="68" pageNumber="69">Ht. undulans</emphasis>
</taxonomicName>
) that closely tracks the margin of both wings, the margin being relatively smooth in the FW and distinctly crenulate in the HW. The reddish/dark orange-brown HWV PMb is slightly more convex than the Mb and both lines terminate abruptly at 1A. In FWV in the cell area are four dark orange-brown Cbs meeting almost perpendicular at the costa, the outermost pair delineating relatively paler areas, the outermost of which is more convex than the others, with its borders diverging towards the costa (in the HT).
<emphasis id="B95A2A364C32C6131EB72C42FED820FC" bold="true" box="[151,267,367,392]" pageId="68" pageNumber="69">Variation.</emphasis>
♀♀ similar but larger and slightly paler (
<figureCitation id="1315EAA14C32C6131CFB2C42FCFC20FC" box="[731,815,367,392]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="68" pageNumber="69">Fig. 20</figureCitation>
B). Space-CuA1 ocellus FWD is a little wider proximad in some specimens, and FWD M1 ocellus sometimes not visible. HW space-CuA1 ocellus is more circular in some specimens.
</paragraph>
<caption id="DF51A6AC4C35C6141EB72A69FE7E26A1" httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="67" pageNumber="68" targetBox="[278,1309,211,1838]" targetPageId="67">
<paragraph id="8B91F6244C35C6141EB72A69FE7E26A1" blockId="67.[151,1436,1860,2005]" pageId="67" pageNumber="68">
<emphasis id="B95A2A364C35C6141EB72A69FEF1262D" bold="true" box="[151,290,1860,1882]" pageId="67" pageNumber="68">FIGURE 20.</emphasis>
Adults in DV (left), VV (right) for mounted specimens and adults in natural posture.
<emphasis id="B95A2A364C35C6141A892A69FB68262D" bold="true" box="[1193,1211,1860,1881]" pageId="67" pageNumber="68">A</emphasis>
: HT ♂ of
<taxonomicName id="4C2E8DA74C35C6141B0F2A68FE7B260D" authority="Lees &amp; Kremen" authorityName="Lees &amp; Kremen" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae" status="sp. nov.">
<emphasis id="B95A2A364C35C6141B0F2A68FA4F262E" box="[1327,1436,1861,1882]" italics="true" pageId="67" pageNumber="68">Heteropsis</emphasis>
<emphasis id="B95A2A364C35C6141EB72A4EFF26260D" box="[151,245,1891,1913]" italics="true" pageId="67" pageNumber="68">barbarae</emphasis>
Lees &amp; Kremen
</taxonomicName>
,
<taxonomicNameLabel id="A269974D4C35C6141F962A4EFDDA260C" box="[438,521,1891,1912]" pageId="67" pageNumber="68" rank="species">
<emphasis id="B95A2A364C35C6141F962A4EFDDA260C" bold="true" box="[438,521,1891,1912]" pageId="67" pageNumber="68">sp. nov.</emphasis>
</taxonomicNameLabel>
(Maitsoarongana, Makira; BMNH(E) 1717096).
<emphasis id="B95A2A364C35C6141A2E2A4EFBCC260C" bold="true" box="[1038,1055,1891,1912]" pageId="67" pageNumber="68">B</emphasis>
: PT ♀ of
<taxonomicName id="4C2E8DA74C35C6141AB12A49FACB260D" box="[1169,1304,1891,1913]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C35C6141AB12A49FACB260D" box="[1169,1304,1891,1913]" italics="true" pageId="67" pageNumber="68">Ht. barbarae</emphasis>
</taxonomicName>
(Sikiory R., Makira; BMNH(E) 1717100).
<emphasis id="B95A2A364C35C6141FF02AACFE3126E3" bold="true" box="[464,482,1921,1943]" pageId="67" pageNumber="68">C</emphasis>
: LT ♂ of
<taxonomicName id="4C2E8DA74C35C6141C6A2AACFC4926E3" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[586,922,1921,1943]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C35C6141C6A2AACFD2A26E3" box="[586,761,1921,1943]" italics="true" pageId="67" pageNumber="68">Culapa undulans</emphasis>
Oberthür, 1916
</taxonomicName>
(Fito; BMNH(E) #674752).
<emphasis id="B95A2A364C35C6141AE22AAFFB0726E3" bold="true" box="[1218,1236,1922,1943]" pageId="67" pageNumber="68">D</emphasis>
: ♀ of
<taxonomicName id="4C2E8DA74C35C6141B382AAFFA4F26E3" box="[1304,1436,1921,1943]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C35C6141B382AAFFA4F26E3" box="[1304,1436,1921,1943]" italics="true" pageId="67" pageNumber="68">Ht. undulans</emphasis>
</taxonomicName>
(Antanandava, PN Andohahela).
<emphasis id="B95A2A364C35C6141FC62A8DFE2426C1" bold="true" box="[486,503,1952,1973]" pageId="67" pageNumber="68">E</emphasis>
: ♀ of
<taxonomicName id="4C2E8DA74C35C6141C182A8CFD6826C2" box="[568,699,1952,1974]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C35C6141C182A8CFD6826C2" box="[568,699,1952,1974]" italics="true" pageId="67" pageNumber="68">Ht. barbarae</emphasis>
</taxonomicName>
(Makira; 27/01/2003).
<emphasis id="B95A2A364C35C6141D882A8DFC6426C1" bold="true" box="[936,951,1952,1973]" pageId="67" pageNumber="68">F</emphasis>
: ♀ of
<taxonomicName id="4C2E8DA74C35C6141DD82A8CFBA926C2" box="[1016,1146,1952,1974]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C35C6141DD82A8CFBA926C2" box="[1016,1146,1952,1974]" italics="true" pageId="67" pageNumber="68">Ht. undulans</emphasis>
</taxonomicName>
(Ankirindro, Makira; 17/01/ 2003) (photos: D. C. Lees).
</paragraph>
</caption>
<paragraph id="8B91F6244C32C6131EE72CF1FDAB2349" blockId="68.[151,1437,151,2013]" pageId="68" pageNumber="69">
<emphasis id="B95A2A364C32C6131EE72CF1FEBF2081" bold="true" box="[199,364,476,501]" pageId="68" pageNumber="69">Wingspan/fwl</emphasis>
: range 33.538.5/
<quantity id="4CD65BC14C32C6131C192CF1FD082080" box="[569,731,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.9149999999999998" metricValueMax="2.05" metricValueMin="1.78" pageId="68" pageNumber="69" unit="mm" value="19.15" valueMax="20.5" valueMin="17.8">17.820.5 mm</quantity>
(n=14 ♂♂), including HT ♂, 36.3/
<quantity id="4CD65BC14C32C6131A442CF1FB182080" box="[1124,1227,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="68" pageNumber="69" unit="mm" value="20.7">20.7 mm</quantity>
; mean 35.7 +/- 1.1 SD/19.2 +/- 0.97 SD mm (n=10 ♂♂, including HT). Range 36.740.4/
<quantity id="4CD65BC14C32C6131D992CD2FB8D2363" box="[953,1118,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" metricValueMax="2.15" metricValueMin="2.01" pageId="68" pageNumber="69" unit="mm" value="20.8" valueMax="21.5" valueMin="20.1">20.121.5 mm</quantity>
; mean 37.0+/- 2.6 SD/20+/- 1.5 SD mm (n=4 ♀♀; referred specimens).
</paragraph>
<paragraph id="8B91F6244C32C6131EE72F6AFC192261" blockId="68.[151,1437,151,2013]" pageId="68" pageNumber="69">
<emphasis id="B95A2A364C32C6131EE72F6AFE872314" bold="true" box="[199,340,583,608]" pageId="68" pageNumber="69">Androconia</emphasis>
: (observations on ♂ specimen MAD182 from Marojejy, referred specimen). Discocellular brush HWD dark grey at base, rufous/reddish-brown in distal ½; underlying patch light brown to reddish-brown, long, very narrow lenticular, surrounded by silvery scales or yellowish ones distal to brush. HWD Cub (cubital brush) above 1A+2A base, mid brown, dispersed, juxtaposed potentially under abdominal sternal segments A2A5, approximately where there is a conspicuous black androconial patch, which is not divided by lighter overlying scales as it is in
<taxonomicName id="4C2E8DA74C32C6131F6C2FD0FE0F2261" box="[332,476,764,789]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C32C6131F6C2FD0FE0F2261" box="[332,476,764,789]" italics="true" pageId="68" pageNumber="69">Ht. undulans</emphasis>
</taxonomicName>
. Inflated vein 2A swollen within basal half.
</paragraph>
<paragraph id="8B91F6244C32C6131EE72E32FD912229" blockId="68.[151,1437,151,2013]" pageId="68" pageNumber="69">
<emphasis id="B95A2A364C32C6131EE72E32FEDB224C" bold="true" box="[199,264,799,824]" pageId="68" pageNumber="69">Palps</emphasis>
: Ochreous, with intermingled brown scales on outside face away from compound eye, dark brown scales above where potentially brushing eye.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C32C6131EE72E4AFC9B220B" box="[199,840,871,896]" pageId="68" pageNumber="69" type="etymology">
<paragraph id="8B91F6244C32C6131EE72E4AFC9B220B" blockId="68.[151,1437,151,2013]" box="[199,840,871,896]" pageId="68" pageNumber="69">
<emphasis id="B95A2A364C32C6131EE72E4AFE9D22F4" bold="true" box="[199,334,871,896]" pageId="68" pageNumber="69">Etymology.</emphasis>
After Barbara, the mother of Claire Kremen.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C32C6121EE72EA1FB472384" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69" type="discussion">
<paragraph id="8B91F6244C32C6121EE72EA1FE692010" blockId="68.[151,1437,151,2013]" lastBlockId="69.[151,1436,151,2013]" lastPageId="69" lastPageNumber="70" pageId="68" pageNumber="69">
<emphasis id="B95A2A364C32C6131EE72EA1FE9F22D1" bold="true" box="[199,332,908,933]" pageId="68" pageNumber="69">Discussion.</emphasis>
This species was first recognized in the field in
<date id="FF90D0E44C32C6131DA52EA1FBD022D0" box="[901,1027,908,932]" pageId="68" pageNumber="69" value="1991" valueMax="1993">19911993</date>
by Claire Kremen in the survey of what is now Masoala National Park (
<bibRefCitation id="EFBF8BD54C32C6131C1E2E9DFD3522BC" author="Lees" box="[574,742,943,968]" pageId="68" pageNumber="69" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997: 64</bibRefCitation>
). It had been collected once before on the expedition of M.I. Evans to RS Ambatovaky in 1988 (specimen in BMNH) but surprisingly there were no (other) historical specimens in BMNH nor MNHN. It is a quite striking and common lowland riparian species in large parts of the Northeast of
<collectingCountry id="F339B6B44C32C6131EB72931FEF32541" box="[151,288,1052,1077]" name="Madagascar" pageId="68" pageNumber="69">Madagascar</collectingCountry>
, so obviously completely overlooked in historic collecting in the region. All available
<typeStatus id="549548864C32C6131ACF2930FAF92541" box="[1263,1322,1053,1077]" pageId="68" pageNumber="69">types</typeStatus>
in the
<taxonomicName id="4C2E8DA74C32C6131B592930FED4252C" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C32C6131B592930FED4252C" italics="true" pageId="68" pageNumber="69">Ht. subsimilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2E8DA74C32C6131F65296CFE19252C" box="[325,458,1087,1112]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="strigula">
<emphasis id="B95A2A364C32C6131F65296CFE19252C" box="[325,458,1087,1112]" italics="true" pageId="68" pageNumber="69">Ht. strigula</emphasis>
</taxonomicName>
groups (all in BMNH) were examined. In particular, among the orange underside species, the following specimens are typified. These include two of up to three (although in his original description, Mabille does not state the number of specimens; but see also Mabille [1887]: 77) possible ♂♂ STs in BMNH of
<taxonomicName id="4C2E8DA74C32C6131EB72981FD8125B0" authority="Mabille, 1878" authorityName="Mabille" authorityYear="1878" box="[151,594,1196,1221]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="maeva">
<emphasis id="B95A2A364C32C6131EB72981FEA025B0" box="[151,371,1196,1221]" italics="true" pageId="68" pageNumber="69">Mycalesis maeva</emphasis>
<bibRefCitation id="EFBF8BD54C32C6131FB62981FD8125B0" author="Mabille" box="[406,594,1196,1221]" pageId="68" pageNumber="69" refString="Mabille, P. (1878) Lepidoptera Africana. Bulletin de la Societe Zoologique de France, 3, 81 - 95." type="journal article" year="1878">Mabille, 1878</bibRefCitation>
</taxonomicName>
; LT ♂, here designated (
<figureCitation id="1315EAA14C32C6131DD12981FB8C25B0" box="[1009,1119,1196,1221]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, ( Antakares; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 ( Antakares; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 ( MadagascarNW; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
D): BMNH(E) 673788;
<typeStatus id="549548864C32C6131EB729E2FED9259C" box="[151,266,1231,1256]" pageId="68" pageNumber="69" type="lectotype">Lectotype</typeStatus>
|MadagascarNo10 [?sic; or MadagascarNW?]|Ex Coll. CHRIS WARD|Ex Oberthür Coll. Brit. Mus. 1927-3.|
<taxonomicName id="4C2E8DA74C32C6131ECB29D9FD922479" authority="Mabille" authorityName="Mabille" box="[235,577,1268,1293]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="maeva">Culapa [sic] maeva Mabille</taxonomicName>
|P.E.L. Viette de. 1968
<taxonomicName id="4C2E8DA74C32C6131D7029D9FBC42478" box="[848,1047,1268,1293]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="maeva">Mycalesis maeva</taxonomicName>
<typeStatus id="549548864C32C6131A6429D9FB342478" box="[1092,1255,1268,1292]" pageId="68" pageNumber="69" type="lectotype">LECTOTYPE</typeStatus>
. For
<taxonomicName id="4C2E8DA74C32C6131B0A29D9FF30245B" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="maeva">Mycalesis maeva</taxonomicName>
, the potential ST male BMNH(E) 673789, “NW Madgsr” from Joicey Bequest (presumably from the Grose- Smith collection) is here designated PLT. Regarding
<taxonomicName id="4C2E8DA74C32C6131CD52811FBB62421" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[757,1125,1340,1365]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laetifica">
<emphasis id="B95A2A364C32C6131CD52811FC7E2421" box="[757,941,1340,1365]" italics="true" pageId="68" pageNumber="69">Culapa laetifica</emphasis>
<bibRefCitation id="EFBF8BD54C32C6131D952811FBB62421" author="Oberthur" box="[949,1125,1340,1365]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
</taxonomicName>
, a LT ♂ is here designated (
<figureCitation id="1315EAA14C32C6131EBF2872FF20240C" box="[159,243,1375,1400]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, ( Antakares; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 ( Antakares; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 ( MadagascarNW; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
B): the specimen bearing labels “BMNH(E) #674741;
<typeStatus id="549548864C32C6131D7E2872FC1C240C" box="[862,975,1375,1400]" pageId="68" pageNumber="69" type="lectotype">Lectotype</typeStatus>
|Nord-Madagascar (Antakares) Isokitra a Diego Suarez
<date id="FF90D0E44C32C6131F1C28A9FDF524E8" box="[316,550,1412,1437]" pageId="68" pageNumber="69" value="1891-05" valueMax="1891-10-31" valueMin="1891-05-00">Mai a Octobre 1891</date>
E.&amp;B. Perrot; illustrated by
<bibRefCitation id="EFBF8BD54C32C6131D5828A9FB2C24E8" author="Oberthur" box="[888,1279,1412,1437]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916" yearSuffix="f">Oberthür (1916: Pl. 368, f. 3072)</bibRefCitation>
”. For the last species, the text by
<bibRefCitation id="EFBF8BD54C32C6131F5D288AFDE224B4" author="d'Abrera" box="[381,561,1447,1472]" pageId="68" pageNumber="69" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">dAbrera (1980</bibRefCitation>
: 185) “♀ (?
<typeStatus id="549548864C32C6131CE6288AFCF924B4" box="[710,810,1447,1472]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
) as illustrated”, is here considered ambiguous, but the corresponding PLT specimen (also illustrated by
<bibRefCitation id="EFBF8BD54C32C6131CE828E1FB9D2490" author="Oberthur" box="[712,1102,1484,1509]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916" yearSuffix="f">Oberthür (1916: Pl. 368, f. 3073))</bibRefCitation>
is now numbered BMNH(E) #674742, while four additional PLT ♂♂ of
<taxonomicName id="4C2E8DA74C32C6131CB028C2FC9B277C" box="[656,840,1519,1544]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laetifica">
<emphasis id="B95A2A364C32C6131CB028C2FC9B277C" box="[656,840,1519,1544]" italics="true" pageId="68" pageNumber="69">Culapa laetifica</emphasis>
</taxonomicName>
are numbered BMNH(E) #674743-674746. Among STs of
<taxonomicName id="4C2E8DA74C32C6131EC72B39FDFC2759" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[231,559,1556,1581]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laeta">
<emphasis id="B95A2A364C32C6131EC72B39FEA92759" box="[231,378,1556,1581]" italics="true" pageId="68" pageNumber="69">Culapa laeta</emphasis>
<bibRefCitation id="EFBF8BD54C32C6131FA12B39FDFC2759" author="Oberthur" box="[385,559,1556,1581]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
</taxonomicName>
, a LT ♂ in BMNH is here designated (
<figureCitation id="1315EAA14C32C6131DC72B39FBE92758" box="[999,1082,1556,1581]" captionStart="FIGURE 21" captionStartId="72.[151,250,1772,1794]" captionTargetBox="[290,1282,340,1732]" captionTargetId="figure@72.[270,1317,337,1750]" captionTargetPageId="72" captionText="FIGURE 21. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; NHMUK 010289153). B: LT ♂ of Culapa laetifica Oberthür, 1916, ( Antakares; BMNH (E) # 674741). C: LT ♂ of C. laeta Oberthür, 1916 ( Antakares; BMNH (E) # 675416). D: LT ♂ of Mycalesis maeva Mabille, 1877 ( MadagascarNW; BMNH (E) # 673788). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264618/files/figure.png" pageId="68" pageNumber="69">Fig. 21</figureCitation>
C), the specimen bearing labels “LT|Nord-Madagascar (Antakares) Isokitra à Diego-Suarez
<date id="FF90D0E44C32C6131D1E2B1AFBF02724" box="[830,1059,1591,1616]" pageId="68" pageNumber="69" value="1891-05" valueMax="1891-10-31" valueMin="1891-05-00">Mai à Octobre 1891</date>
E.&amp;B. Perrot|
<taxonomicName id="4C2E8DA74C32C6131AE52B1AFA4F2724" authority="Obthr" authorityName="Obthr" box="[1221,1436,1591,1616]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laeta">Culapa laeta Obthr</taxonomicName>
<typeStatus id="549548864C32C6131E932B71FF3A2700" box="[179,233,1628,1652]" pageId="68" pageNumber="69">type</typeStatus>
|P.E.L. Viette det. 1968
<taxonomicName id="4C2E8DA74C32C6131FCC2B71FD3E2701" authority="Ch. Obthr" authorityName="Ch. Obthr" box="[492,749,1628,1653]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laeta">
Culapa laeta
<collectingCountry id="F339B6B44C32C6131C5E2B71FD712701" box="[638,674,1628,1653]" name="Switzerland" pageId="68" pageNumber="69">Ch</collectingCountry>
. Obthr
</taxonomicName>
<typeStatus id="549548864C32C6131D312B71FC672701" box="[785,948,1628,1653]" pageId="68" pageNumber="69" type="lectotype">LECTOTYPE</typeStatus>
|Supposed “
<typeStatus id="549548864C32C6131A142B71FBA32700" box="[1076,1136,1628,1652]" pageId="68" pageNumber="69">Type</typeStatus>
” (
<typeStatus id="549548864C32C6131AA92B71FB302700" box="[1161,1251,1628,1652]" pageId="68" pageNumber="69" type="syntype">syntype</typeStatus>
) of
<taxonomicName id="4C2E8DA74C32C6131B302B71FED127EC" authority="Oberthur" authorityName="Oberthur" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laeta">Culapa laeta Oberthür</taxonomicName>
)|[Copy of f. 3075]|Ex Oberthür Coll. Brit. Mus. 1927-3.|
<typeStatus id="549548864C32C6131D4D2BADFC3427EC" box="[877,999,1664,1688]" pageId="68" pageNumber="69" type="lectotype">Lectotype</typeStatus>
♂ of
<taxonomicName id="4C2E8DA74C32C6131A072B52FA4B27E3" authority="Oberthur D.C. Lees" authorityName="Oberthur D.C. Lees" box="[1063,1432,1663,1688]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laeta">Culapa laeta Oberthür D.C. Lees</taxonomicName>
, det.
<date id="FF90D0E44C32C6131EE92B89FE9527C8" box="[201,326,1700,1724]" pageId="68" pageNumber="69" value="2000-09">Sept. 2000</date>
|BMNH(E) 675416”; a surviving further 5 of
<specimenCount id="9D283DAD4C32C6131D712B88FCAE27C8" box="[849,893,1701,1724]" pageId="68" pageNumber="69" type="generic">7 ♂</specimenCount>
STs and all 4 ♀ PLTs of
<taxonomicName id="4C2E8DA74C32C6131A882B89FAE827C9" box="[1192,1339,1700,1725]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="laeta">
<emphasis id="B95A2A364C32C6131A882B89FAE827C9" box="[1192,1339,1700,1725]" italics="true" pageId="68" pageNumber="69">Culapa laeta</emphasis>
</taxonomicName>
become PLTs. The STs of
<taxonomicName id="4C2E8DA74C32C6131F4F2BEAFD3F2794" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" box="[367,748,1735,1760]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C32C6131F4F2BEAFDE02794" box="[367,563,1735,1760]" italics="true" pageId="68" pageNumber="69">Culapa undulans</emphasis>
<bibRefCitation id="EFBF8BD54C32C6131C1E2BEAFD3F2794" author="Oberthur" box="[574,748,1735,1760]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
</taxonomicName>
were also examined; the LT ♂ is here considered to have designated by
<bibRefCitation id="EFBF8BD54C32C6131F1A2BC1FE3A2670" author="d'Abrera" box="[314,489,1772,1797]" pageId="68" pageNumber="69" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">dAbrera, 1980</bibRefCitation>
: 184 by the text “♂ (
<typeStatus id="549548864C32C6131CF72BC1FCEF2671" box="[727,828,1772,1797]" pageId="68" pageNumber="69" type="holotype">holotype</typeStatus>
) and ♀ as illustrated”, as illustrated on p. 185; the LT (
<figureCitation id="1315EAA14C32C6131EBF2A22FF22265C" box="[159,241,1807,1832]" captionStart="FIGURE 20" captionStartId="67.[151,250,1860,1882]" captionTargetBox="[278,1309,211,1838]" captionTargetId="figure@67.[266,1320,193,1839]" captionTargetPageId="67" captionText="FIGURE 20. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture. A: HT ♂ of Heteropsis barbarae Lees &amp; Kremen, sp. nov. (Maitsoarongana, Makira; BMNH (E) 1717096). B: PT ♀ of Ht. barbarae (Sikiory R., Makira; BMNH (E) 1717100). C: LT ♂ of Culapa undulans Oberthür, 1916 (Fito; BMNH (E) # 674752). D: ♀ of Ht. undulans (Antanandava, PN Andohahela). E: ♀ of Ht. barbarae (Makira; 27 / 01 / 2003). F: ♀ of Ht. undulans (Ankirindro, Makira; 17 / 01 / 2003) (photos: D. C. Lees)." httpUri="https://zenodo.org/record/264617/files/figure.png" pageId="68" pageNumber="69">Fig. 20</figureCitation>
C) bears the labels “
<collectingCountry id="F339B6B44C32C6131FF32A22FD8E265C" box="[467,605,1807,1832]" name="Madagascar" pageId="68" pageNumber="69">Madagascar</collectingCountry>
Fito
<date id="FF90D0E44C32C6131CB92A22FC89265C" box="[665,858,1807,1832]" pageId="68" pageNumber="69" value="1897-05" valueMax="1897-08-31" valueMin="1897-05-00">Mai a Aout 1897</date>
Perrot Freres|BMNH(E) #674752|
<taxonomicName id="4C2E8DA74C32C6131AFF2A22FA4F265C" box="[1247,1436,1807,1832]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">Culapa undulans</taxonomicName>
<typeStatus id="549548864C32C6131E942A18FF382639" box="[180,235,1845,1869]" pageId="68" pageNumber="69">Type</typeStatus>
Oberthür [handwriting]|[label bearing copy of f. 3062]” and is indeed the specimen previously illustrated by
<bibRefCitation id="EFBF8BD54C32C6131E9E2A7AFDEC2604" author="Oberthur" box="[190,575,1879,1904]" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916" yearSuffix="f">Oberthür (1916: Pl.368, f. 3062)</bibRefCitation>
; the corresponding PLT ♂ of
<taxonomicName id="4C2E8DA74C32C6131D852A7AFBB92604" box="[933,1130,1879,1904]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C32C6131D852A7AFBB92604" box="[933,1130,1879,1904]" italics="true" pageId="68" pageNumber="69">Culapa undulans</emphasis>
</taxonomicName>
from the same locality is numbered BMNN(E) #674753. The specimen illustrated with no abdomen by
<bibRefCitation id="EFBF8BD54C32C6131A052A51FB0926E0" author="d'Abrera" box="[1061,1242,1916,1941]" pageId="68" pageNumber="69" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">dAbrera (1980</bibRefCitation>
: 185; BMNH(E) #674891) represents a PLT of
<taxonomicName id="4C2E8DA74C32C6131FD32A8DFD6626CD" box="[499,693,1952,1977]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C32C6131FD32A8DFD6626CD" box="[499,693,1952,1977]" italics="true" pageId="68" pageNumber="69">Culapa undulans</emphasis>
</taxonomicName>
as treated syntypically by
<bibRefCitation id="EFBF8BD54C32C6131DCA2A8DFEBC26A9" author="Oberthur" pageId="68" pageNumber="69" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916" yearSuffix="f">Oberthür (1916: 220, pl. 367, f. 3063, from Antsianaka)</bibRefCitation>
but is misidentified and in fact represents a ♀ of
<taxonomicName id="4C2E8DA74C32C6131DBD2AE8FBC526A8" box="[925,1046,1989,2012]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="B95A2A364C32C6131DBD2AE8FBC526A8" box="[925,1046,1989,2012]" italics="true" pageId="68" pageNumber="69">Ht. pauper</emphasis>
</taxonomicName>
, to which “
<taxonomicName id="4C2E8DA74C32C6131ABA2AE9FB3E26A9" box="[1178,1261,1988,2013]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B95A2A364C32C6131ABA2AE9FB3E26A9" box="[1178,1261,1988,2013]" italics="true" pageId="68" pageNumber="69">Culapa</emphasis>
</taxonomicName>
(var. ou espèce séparée)
<taxonomicName id="4C2E8DA74C33C6121EDA2DBAFE6121C4" authority="Oberthur" authorityName="Oberthur" box="[250,434,151,176]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseudonarcissus">
<emphasis id="B95A2A364C33C6121EDA2DBAFE6121C4" box="[250,434,151,176]" italics="true" pageId="69" pageNumber="70">pseudonarcissus</emphasis>
</taxonomicName>
<bibRefCitation id="EFBF8BD54C33C6121FE52DBAFC9321C4" author="Oberthur" box="[453,832,151,176]" pageId="69" pageNumber="70" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür (1916: 223, fasc. 11: 41)</bibRefCitation>
(that
<bibRefCitation id="EFBF8BD54C33C6121DA02DBAFBFC21C4" author="d'Abrera" box="[896,1071,151,176]" pageId="69" pageNumber="70" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">dAbrera, 1980</bibRefCitation>
: 184 treated under
<taxonomicName id="4C2E8DA74C33C6121B232DB4FA4021C4" box="[1283,1427,151,176]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="undulans">
<emphasis id="B95A2A364C33C6121B232DB4FA4021C4" box="[1283,1427,151,176]" italics="true" pageId="69" pageNumber="70">Ht. undulans</emphasis>
</taxonomicName>
) also would belong. The last form (represented by a ♀ without abdomen;
<figureCitation id="1315EAA14C33C6121DE52D91FBC721A0" box="[965,1044,188,213]" captionStart="FIGURE 14" captionStartId="48.[151,250,1802,1824]" captionTargetBox="[299,1270,199,1771]" captionTargetId="figure@48.[287,1299,193,1781]" captionTargetPageId="48" captionText="FIGURE 14. Adults in DV (left), VV (right). A: HT ♂ of Pseudonympha subsimilis Butler, 1879 ( Antananarivo; BMNH (E) # 674882). B: PLT ♀ of Culapa undulata Oberthür, 1916 ( Antsianaka; BMNH (E) # 674884). C: LT ♂ of Culapa undulata Oberthür, 1916 ( Antsianaka; BMNH (E) # 674883). D: LT ♂ of Culapa pauper Oberthür, 1916 ( Antsianaka; BMNH (E) # 674888). E: LT ♀ of Culapa pauper sp. or var. pseudonarcissus Oberthür, 1916 ( Antakares; BMNH (E) # 674890). Scale bar: 30 mm." httpUri="https://zenodo.org/record/264611/files/figure.png" pageId="69" pageNumber="70">Fig. 14</figureCitation>
E) is here designated LT of
<taxonomicName id="4C2E8DA74C33C6121B692D91FE9C218C" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseudonarcissus">
<emphasis id="B95A2A364C33C6121B692D91FE9C218C" italics="true" pageId="69" pageNumber="70">Culapa pseudonarcissus</emphasis>
</taxonomicName>
, bearing labels “Nord
<collectingCountry id="F339B6B44C33C6121C6E2DF2FD06218C" box="[590,725,223,248]" name="Madagascar" pageId="69" pageNumber="70">Madagascar</collectingCountry>
Antakares Isokitra a Diego Suarez
<date id="FF90D0E44C33C6121A462DF2FA95218C" box="[1126,1350,223,248]" pageId="69" pageNumber="70" value="1891-05" valueMax="1891-10-31" valueMin="1891-05-00">Mai a Octobre 1891</date>
|
<taxonomicName id="4C2E8DA74C33C6121B682DF2FF302068" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pauper">Culapa pauper</taxonomicName>
var?
<taxonomicName id="4C2E8DA74C33C6121F152C29FE382069" box="[309,491,260,285]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseudonarcissus">pseudonarcissus</taxonomicName>
(Bdv. in ms) Obthr.
<typeStatus id="549548864C33C6121CC82C28FCCC2069" box="[744,799,261,285]" pageId="69" pageNumber="70">Type</typeStatus>
♀)|BMNH(E) #674890” and this is presumably a dry season individual; although this has no consequence,
<bibRefCitation id="EFBF8BD54C33C6121CD92C0AFBA22034" author="Oberthur" box="[761,1137,295,320]" pageId="69" pageNumber="70" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916" yearSuffix="f">Oberthür (1916, pl. 368, f. 3068)</bibRefCitation>
also captioned it “
<emphasis id="B95A2A364C33C6121B692C0AFE742011" italics="true" pageId="69" pageNumber="70">
<taxonomicName id="4C2E8DA74C33C6121B692C0AFF352010" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pauper">Culapa pauper</taxonomicName>
-
<taxonomicName id="4C2E8DA74C33C6121ECF2C61FE742011" box="[239,423,332,357]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pseudonarcissus">pseudonarcissus</taxonomicName>
</emphasis>
”.
</paragraph>
<paragraph id="8B91F6244C33C6121EE72C42FB472384" blockId="69.[151,1436,151,2013]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72C42FDA220FC" bold="true" box="[199,625,367,392]" pageId="69" pageNumber="70">Additional information. ♂ genitalia</emphasis>
: 141DL (
<figureCitation id="1315EAA14C33C6121CC12C42FCE120FC" box="[737,818,367,392]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui &amp; Aduse-Poku, sp. nov., ♂ genitalia ( Mahassabe; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="69" pageNumber="70">Fig. 18</figureCitation>
C, referred specimen): the basic configuration is fairly typical for
<taxonomicName id="4C2E8DA74C33C6121F382CB8FE4F20D9" box="[280,412,404,429]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="strigula">
<emphasis id="B95A2A364C33C6121F382CB8FE4F20D9" box="[280,412,404,429]" italics="true" pageId="69" pageNumber="70">Ht. strigula</emphasis>
</taxonomicName>
group. From LV, the approximately parallel-sided uncus is very slightly longer than the tegumen, the dorsal margin approximately in line with that of the tegumen and slightly upraised after its brow, with a smoothly round head before the ventral hook (inflated from DV before tip), which projects forward rather than down. The gnathos originates on an elliptic base and is gently tapered and straight to an approx. 35-degree angle to uncus in the specimen examined; from DV it features a typical bulls horn sinuate appearance with pointed tips inrecurved. Only a slight constriction is found at hinge with vinculum. The valve has a fairly parallel-sided, flattened arm which is bent over at tip (inwards from DV) exposing a dense coverage of spinoid setae and the uncus tip when not downflexed is about the same extension as, not proud of the valve tips. The saccus is of normal length for the
<taxonomicName id="4C2E8DA74C33C6121EC72F98FEB423B9" box="[231,359,692,717]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="strigula">
<emphasis id="B95A2A364C33C6121EC72F98FEB423B9" box="[231,359,692,717]" italics="true" pageId="69" pageNumber="70">Ht. strigula</emphasis>
</taxonomicName>
group and the aedeagus is about the same length as the valve and strongly recurved distad of the (also proximally uprecurved) ostium. The juxta is narrow and not sharply lipped proximad.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C33C6121EE72FD1FC1125B1" pageId="69" pageNumber="70" type="description">
<paragraph id="8B91F6244C33C6121EE72FD1FBC822D1" blockId="69.[151,1436,151,2013]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72FD1FE7A2261" bold="true" box="[199,425,764,789]" pageId="69" pageNumber="70">DNA divergences:</emphasis>
COI-5P cluster number BOLD:ACW4995 (exemplar BMAD248-15, DL14M-0033, Marojejy) is 4.86% divergent to
<taxonomicName id="4C2E8DA74C33C6121C6C2E0CFD18224C" box="[588,715,801,824]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="tianae">
<emphasis id="B95A2A364C33C6121C6C2E0CFD18224C" box="[588,715,801,824]" italics="true" pageId="69" pageNumber="70">Ht. tianae</emphasis>
</taxonomicName>
(BOLD:AAE4112), and 5.63% divergent to
<taxonomicName id="4C2E8DA74C33C6121B3A2E0CFA4F224C" box="[1306,1436,801,824]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="maeva">
<emphasis id="B95A2A364C33C6121B3A2E0CFA4F224C" box="[1306,1436,801,824]" italics="true" pageId="69" pageNumber="70">Ht. maeva</emphasis>
</taxonomicName>
(BOLD:AAE5488), its sister species in the study of Aduse-Poku
<emphasis id="B95A2A364C33C6121D532E68FC602229" box="[883,947,836,861]" italics="true" pageId="69" pageNumber="70">et al.,</emphasis>
(2015). In the COII dataset of
<bibRefCitation id="EFBF8BD54C33C6121B2E2E68FF3222F4" pageId="69" pageNumber="70" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article">
Torres
<emphasis id="B95A2A364C33C6121B7D2E68FA4F2229" box="[1373,1436,836,861]" italics="true" pageId="69" pageNumber="70">et al.,</emphasis>
(2001)
</bibRefCitation>
,
<taxonomicName id="4C2E8DA74C33C6121ECF2E44FE5222F4" box="[239,385,871,896]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="barbarae">
<emphasis id="B95A2A364C33C6121ECF2E44FE5222F4" box="[239,385,871,896]" italics="true" pageId="69" pageNumber="70">Ht. barbarae</emphasis>
</taxonomicName>
(their “
<emphasis id="B95A2A364C33C6121FF72E44FDD522F4" box="[471,518,873,896]" italics="true" pageId="69" pageNumber="70">Hen</emphasis>
. sp. 16”;
<accessionNumber id="947D6BC74C33C6121C512E45FD3822F4" box="[625,747,872,896]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY040128" pageId="69" pageNumber="70">
AY
<date id="FF90D0E44C33C6121CB82E45FD3822F4" box="[664,747,872,896]" pageId="69" pageNumber="70" value="28-04-01">040128</date>
</accessionNumber>
based on 1 ♀ from Ivontaka Sud) was 6.54% divergent to
<taxonomicName id="4C2E8DA74C33C6121B592E44FF3322D0" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="maeva">
<emphasis id="B95A2A364C33C6121B592E44FF3322D0" italics="true" pageId="69" pageNumber="70">Ht. maeva</emphasis>
</taxonomicName>
(
<accessionNumber id="947D6BC74C33C6121ECE2EA1FEBA22D0" box="[238,361,908,932]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY040132" pageId="69" pageNumber="70">
AY
<date id="FF90D0E44C33C6121F352EA1FEBA22D0" box="[277,361,908,932]" pageId="69" pageNumber="70" value="32-04-01">040132</date>
</accessionNumber>
based on 3 ♂♂, and 1 ♀ from Masoala; 390 bp comparable).
</paragraph>
<paragraph id="8B91F6244C33C6121EE72E82FCBF2564" blockId="69.[151,1436,151,2013]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72E82FE3422BC" bold="true" box="[199,487,943,968]" pageId="69" pageNumber="70">Phylogeny/sister species</emphasis>
: possibly
<taxonomicName id="4C2E8DA74C33C6121C7E2E9CFCE322BC" box="[606,816,943,968]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="menamenoides">
<emphasis id="B95A2A364C33C6121C7E2E9CFCE322BC" box="[606,816,943,968]" italics="true" pageId="69" pageNumber="70">Ht. menamenoides</emphasis>
</taxonomicName>
(combined tree in
<bibRefCitation id="EFBF8BD54C33C6121A2D2E82FAF522BC" author="Aduse-Poku" box="[1037,1318,943,968]" pageId="69" pageNumber="70" refString="Aduse-Poku, K., Lees, D. C., Brattstrom, O., Kodandaramaiah, U., Collins, S. &amp; Brakefield, P. M., (2016) Molecular phylogeny and generic-level taxonomy of the widespread palaeotropical ' Heteropsis clade' (Nymphalidae: Satyrinae: Mycalesina). Systematic Entomology, in press. http: // dx. doi. org / syen. 12183" type="book" year="2016">
Aduse-Poku
<emphasis id="B95A2A364C33C6121A802E9CFB0822BC" box="[1184,1243,943,968]" italics="true" pageId="69" pageNumber="70">et al.</emphasis>
, 2016
</bibRefCitation>
, in press).
<bibRefCitation id="EFBF8BD54C33C6121EB72EF8FEA42298" author="Torres" box="[151,375,980,1005]" pageId="69" pageNumber="70" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres
<emphasis id="B95A2A364C33C6121EC72EF8FEFB2299" box="[231,296,980,1005]" italics="true" pageId="69" pageNumber="70">et al.,</emphasis>
(2001
</bibRefCitation>
: 467) suggested a topological relationship with
<taxonomicName id="4C2E8DA74C33C6121DBD2EF8FBC02298" box="[925,1043,981,1004]" class="Insecta" family="Nymphalidae" genus="Mycalesis" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="maeva">
<emphasis id="B95A2A364C33C6121DBD2EF8FBC02298" box="[925,1043,981,1004]" italics="true" pageId="69" pageNumber="70">Ht. maeva</emphasis>
</taxonomicName>
in their cladistic analysis of COII sequences, but this was not supported (
<bibRefCitation id="EFBF8BD54C33C6121C6B2ED5FCF52564" author="Torres" box="[587,806,1015,1040]" pageId="69" pageNumber="70" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres
<emphasis id="B95A2A364C33C6121CBB2ED4FD092564" box="[667,730,1015,1040]" italics="true" pageId="69" pageNumber="70">et al.,</emphasis>
(2001
</bibRefCitation>
: 466).
</paragraph>
<paragraph id="8B91F6244C33C6121EE72931FE252541" blockId="69.[151,1436,151,2013]" box="[199,502,1052,1077]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72931FE252541" bold="true" box="[199,502,1052,1077]" pageId="69" pageNumber="70">Ecology and distribution.</emphasis>
</paragraph>
<paragraph id="8B91F6244C33C6121EE72912FDB1252C" blockId="69.[151,1436,151,2013]" box="[199,610,1087,1112]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72912FEF0252C" bold="true" box="[199,291,1087,1112]" pageId="69" pageNumber="70">Habitat</emphasis>
: riparian primary rainforest.
</paragraph>
<paragraph id="8B91F6244C33C6121EE72949FCAB2509" blockId="69.[151,1436,151,2013]" box="[199,888,1124,1149]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72949FE902509" bold="true" box="[199,323,1124,1149]" pageId="69" pageNumber="70">Behaviour</emphasis>
: flies low and both sexes come easily to fruit bait.
</paragraph>
<paragraph id="8B91F6244C33C6121EE729AAFCF925D4" blockId="69.[151,1436,151,2013]" box="[199,810,1159,1184]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE729AAFEEE25D4" bold="true" box="[199,317,1159,1184]" pageId="69" pageNumber="70">Hostplant</emphasis>
: unknown but almost certainly low grasses.
</paragraph>
<paragraph id="8B91F6244C33C6121EE72981FC1125B1" blockId="69.[151,1436,151,2013]" box="[199,962,1196,1221]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72981FE8B25B1" bold="true" box="[199,344,1196,1221]" pageId="69" pageNumber="70">Early stages</emphasis>
: unknown but for expressed eggs, which were whitish.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C33C6121EE729E2FB4A2444" pageId="69" pageNumber="70" type="distribution">
<paragraph id="8B91F6244C33C6121EE729E2FDB42479" blockId="69.[151,1436,151,2013]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE729E2FE8A259C" bold="true" box="[199,345,1231,1256]" pageId="69" pageNumber="70">Distribution</emphasis>
: endemic to the northeastern rainforests of
<collectingCountry id="F339B6B44C33C6121D7929E2FC31259C" box="[857,994,1231,1256]" name="Madagascar" pageId="69" pageNumber="70">Madagascar</collectingCountry>
, from Marojejy and Masoala south to Ambatovaky (
<figureCitation id="1315EAA14C33C6121F1729D9FE5A2478" box="[311,393,1268,1293]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees &amp; Kremen, sp. nov. Light blue: Ht. hazovola Lees &amp; Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees &amp; Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees &amp; Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees &amp; Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees &amp; Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees &amp; Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui &amp; Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees &amp; Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="69" pageNumber="70">Fig. 30</figureCitation>
C, light green dots).
</paragraph>
<paragraph id="8B91F6244C33C6121EE7283AFB4A2444" blockId="69.[151,1436,151,2013]" box="[199,1177,1303,1328]" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE7283AFE4A2444" bold="true" box="[199,409,1303,1328]" pageId="69" pageNumber="70">Elevational range</emphasis>
:
<quantity id="4CD65BC14C33C6121F882835FDDD245B" box="[424,526,1304,1328]" metricMagnitude="2" metricUnit="m" metricValue="4.45" metricValueMax="8.9" metricValueMin="0.0" pageId="69" pageNumber="70" unit="m" value="445.0" valueMax="890.0" valueMin="0.0">
0
<specimenCount id="9D283DAD4C33C6121FE42835FDDD245B" box="[452,526,1304,1328]" pageId="69" pageNumber="70" type="generic">890 m</specimenCount>
</quantity>
. (n= 153, including referred specimens and observations).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C33C6111EE72813FDC9260E" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="materials_examined">
<paragraph id="8B91F6244C33C6111EE72813FDC9260E" blockId="69.[151,1436,151,2013]" lastBlockId="70.[151,1437,151,1914]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
<emphasis id="B95A2A364C33C6121EE72813FE692423" bold="true" box="[199,442,1342,1367]" pageId="69" pageNumber="70">Referred specimens.</emphasis>
♀,
<collectingCountry id="F339B6B44C33C6121FCF2813FDA52423" box="[495,630,1342,1367]" name="Madagascar" pageId="69" pageNumber="70">Madagascar</collectingCountry>
NE, Marojejy PN, Camp II [above],
<geoCoordinate id="EE1A90E34C33C6121A3F2812FB402423" box="[1055,1171,1339,1367]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-14.4344">14.4344o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121A812813FAD52423" box="[1185,1286,1339,1367]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="55" value="49.759">49.759o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121B182813FA4B2423" box="[1336,1432,1342,1367]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="69" pageNumber="70" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C33C6121EB72848FEFC2409" box="[151,303,1381,1406]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="69" pageNumber="70" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">850 +/- 50 m</quantity>
,
<date id="FF90D0E44C33C6121F1D2848FE13240A" box="[317,448,1381,1406]" pageId="69" pageNumber="70" value="2006-11-12">12/11/2006</date>
: 10:00, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121CBE2848FD15240A" box="[670,710,1381,1406]" pageId="69" pageNumber="70">DL</collectionCode>
06-298; ♂, NE, Marojejy PN,
<geoCoordinate id="EE1A90E34C33C6121A3E284BFB41240A" box="[1054,1170,1378,1406]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-14.4344">14.4344o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121A802848FAC72409" box="[1184,1300,1378,1406]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.7606">49.7606o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121B652848FA4B240A" box="[1349,1432,1381,1406]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="69" pageNumber="70" unit="km" value="0.5">0.5 km</quantity>
,
<quantity id="4CD65BC14C33C6121EB728A7FEF924D5" box="[151,298,1417,1442]" metricMagnitude="2" metricUnit="m" metricValue="7.3" metricValueMax="7.8" metricValueMin="6.8" pageId="69" pageNumber="70" unit="m" value="730.0" valueMax="780.0" valueMin="680.0">730 +/- 50 m</quantity>
,
<date id="FF90D0E44C33C6121F1628A4FE6424D6" box="[310,439,1417,1442]" pageId="69" pageNumber="70" value="2006-11-11">11/11/2006</date>
: 15:48, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121CAF28A4FD6424D6" box="[655,695,1417,1442]" pageId="69" pageNumber="70">DL</collectionCode>
06-269,
<collectionCode id="ED3F6EE14C33C6121D3128A4FC9424D6" box="[785,839,1417,1442]" country="USA" httpUri="http://grbio.org/cool/s9x3-awht" name="Dicty Stock Center" pageId="69" pageNumber="70">DSC</collectionCode>
_0826 [photo]; ♂, data as above but:
<date id="FF90D0E44C33C6121AC828A4FA8E24D6" box="[1256,1373,1417,1442]" pageId="69" pageNumber="70" value="2006-12-03">3/12/2006</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121EFA289DFEC324BD" box="[218,272,1456,1481]" country="USA" httpUri="http://grbio.org/cool/s9x3-awht" name="Dicty Stock Center" pageId="69" pageNumber="70">DSC</collectionCode>
_0826 [photo]; ♀, NE, Manantenina camp 2, Marojejy PN,
<geoCoordinate id="EE1A90E34C33C6121D8B289CFBCD24BD" box="[939,1054,1453,1481]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-14.4347">14.4347o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121A0B289DFB4D24BC" box="[1067,1182,1453,1481]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.7601">49.7601o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121AEC289DFACE24BD" box="[1228,1309,1456,1481]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="69" pageNumber="70" unit="km" value="0.5">0.5 km</quantity>
,
<quantity id="4CD65BC14C33C6121B08289CFF7C249B" metricMagnitude="2" metricUnit="m" metricValue="7.3" metricValueMax="7.8" metricValueMin="6.8" pageId="69" pageNumber="70" unit="m" value="730.0" valueMax="780.0" valueMin="680.0">730 +/- 50 m</quantity>
,
<date id="FF90D0E44C33C6121E9F28FAFEED2484" box="[191,318,1495,1520]" pageId="69" pageNumber="70" value="2006-11-10">10/11/2006</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121FF828FAFDD22484" box="[472,513,1495,1520]" pageId="69" pageNumber="70">DL</collectionCode>
06-169; specimen, NE, Marojejy PN [between camp 1 and camp2],
<geoCoordinate id="EE1A90E34C33C6121B0228FAFA45249B" box="[1314,1430,1492,1520]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-14.4362">14.4362o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121EB728D0FED92762" box="[151,266,1530,1558]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.7679">49.7679o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121F1928D0FE4B2762" box="[313,408,1533,1558]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="69" pageNumber="70" unit="km" value="0.85">0.85 km</quantity>
,
<quantity id="4CD65BC14C33C6121F8328D0FD972762" box="[419,580,1533,1558]" metricMagnitude="2" metricUnit="m" metricValue="6.13" metricValueMax="7.38" metricValueMin="4.88" pageId="69" pageNumber="70" unit="m" value="613.0" valueMax="738.0" valueMin="488.0">613 +/- 125 m</quantity>
,
<date id="FF90D0E44C33C6121C7028D0FD022762" box="[592,721,1533,1558]" pageId="69" pageNumber="70" value="2006-11-10">10/11/2006</date>
, 15:58, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121D8A28D0FC002762" box="[938,979,1533,1558]" pageId="69" pageNumber="70">DL</collectionCode>
06-170; ♀, NE, Marojejy PN,
<geoCoordinate id="EE1A90E34C33C6121B0228D3FA452762" box="[1314,1430,1530,1558]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-14.4377">14.4377o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121EB72B09FED92748" box="[151,266,1569,1597]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.7756">49.7756o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121F1B2B09FE492749" box="[315,410,1572,1597]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="69" pageNumber="70" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C33C6121F862B08FDEF2748" box="[422,572,1572,1597]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="69" pageNumber="70" unit="m" value="400.0" valueMax="450.0" valueMin="350.0">400 +/- 50 m</quantity>
,
<date id="FF90D0E44C33C6121C682B09FD182749" box="[584,715,1572,1597]" pageId="69" pageNumber="70" value="2006-11-20">20/11/2006</date>
: 15:07, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121D862B09FC1D2749" box="[934,974,1572,1597]" pageId="69" pageNumber="70">DL</collectionCode>
06-456; ♂, NE, Marojejy PN,
<geoCoordinate id="EE1A90E34C33C6121B022B09FA452749" box="[1314,1430,1569,1597]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-14.4425">14.4425o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121EB72B61FED92717" box="[151,266,1608,1636]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.7818">49.7818o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121F192B66FE592710" box="[313,394,1611,1636]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="69" pageNumber="70" unit="km" value="0.5">0.5 km</quantity>
,
<quantity id="4CD65BC14C33C6121FB52B66FDE52717" box="[405,566,1611,1636]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="69" pageNumber="70" unit="m" value="450.0" valueMax="600.0" valueMin="300.0">450 +/- 150 m</quantity>
,
<date id="FF90D0E44C33C6121C622B66FD162710" box="[578,709,1611,1636]" pageId="69" pageNumber="70" value="2006-11-10">10/11/2006</date>
: 10:11, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121DBC2B66FC172710" box="[924,964,1611,1636]" pageId="69" pageNumber="70">DL</collectionCode>
06-205; ♂, NE, Marojejy PN,
<quantity id="4CD65BC14C33C6121B352B61FA9D2710" box="[1301,1358,1611,1636]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="69" pageNumber="70" unit="km" value="1.0">1 km</quantity>
before [after?] entrée,
<geoCoordinate id="EE1A90E34C33C6121F672B5FFE1A27FF" box="[327,457,1647,1675]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="-14.45015">14.45015o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121FF72B5FFDEE27FE" box="[471,573,1647,1675]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="55" value="49.786">49.786o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121C512B5EFD0127FF" box="[625,722,1650,1675]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="69" pageNumber="70" unit="km" value="1.78">1.78 km</quantity>
,
<quantity id="4CD65BC14C33C6121CFF2B5FFC5B27FE" box="[735,904,1650,1675]" metricMagnitude="2" metricUnit="m" metricValue="3.26" metricValueMax="5.01" metricValueMin="1.51" pageId="69" pageNumber="70" unit="m" value="326.0" valueMax="501.0" valueMin="151.0">326 +/- 175 m</quantity>
,
<date id="FF90D0E44C33C6121DB62B5FFBCA27FF" box="[918,1049,1650,1675]" pageId="69" pageNumber="70" value="2006-11-10">10/11/2006</date>
: 13:16, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121ADA2B5FFAF127FF" box="[1274,1314,1650,1675]" pageId="69" pageNumber="70">DL</collectionCode>
06-204; ♂, NE, Marojejy PN [around entrée],
<geoCoordinate id="EE1A90E34C33C6121C3C2BB4FD4327C5" box="[540,656,1686,1714]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-14.4626">14.4626o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121CBC2BB4FCC327C5" box="[668,784,1686,1714]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.7964">49.7964o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121D1F2BB4FC4327C6" box="[831,912,1689,1714]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="69" pageNumber="70" unit="km" value="0.5">0.5 km</quantity>
,
<quantity id="4CD65BC14C33C6121DBB2BB4FBEF27C5" box="[923,1084,1689,1714]" metricMagnitude="2" metricUnit="m" metricValue="1.56" metricValueMax="2.56" metricValueMin="0.5599999999999999" pageId="69" pageNumber="70" unit="m" value="156.0" valueMax="256.0" valueMin="56.0">156 +/- 100 m</quantity>
,
<date id="FF90D0E44C33C6121A682BB4FB1F27C6" box="[1096,1228,1689,1714]" pageId="69" pageNumber="70" value="2006-11-10">10/11/2006</date>
: 12:48, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121EB72B90FF1327A2" box="[151,192,1725,1750]" pageId="69" pageNumber="70">DL</collectionCode>
06-168; ♂, NE, Marojejy,
<quantity id="4CD65BC14C33C6121FC22B90FDF827A1" box="[482,555,1725,1750]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="69" pageNumber="70" unit="m" value="650.0">650 m</quantity>
,
<date id="FF90D0E44C33C6121C162B93FD7F27A2" box="[566,684,1726,1750]" pageId="69" pageNumber="70" value="2014-01-30">30/1/2014</date>
: 12:57, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121DA22B93FC1627A2" box="[898,965,1726,1750]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="69" pageNumber="70">MAD</collectionCode>
182 [pheromone voucher], IA387 [isotope voucher]; ♂ [
<collectionCode id="ED3F6EE14C33C6121F0D2BC8FE5A2788" box="[301,393,1765,1788]" pageId="69" pageNumber="70">BMNH</collectionCode>
], NE, Andranobe, Masoala Peninsula,
<geoCoordinate id="EE1A90E34C33C6121D1B2BC9FC7C2788" box="[827,943,1760,1789]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-15.6816">15.6816o S</geoCoordinate>
, 49.9573oE +/-
<quantity id="4CD65BC14C33C6121A462BC9FB162789" box="[1126,1221,1764,1789]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="69" pageNumber="70" unit="km" value="1.31">1.31 km</quantity>
, 0 m,
<date id="FF90D0E44C33C6121B2A2BC9FAA42788" box="[1290,1399,1764,1788]" pageId="69" pageNumber="70" value="1991-12-03">3-Dec-91</date>
, C. Kremen: CK826,
<collectionCode id="ED3F6EE14C33C6121F7E2A25FE662654" box="[350,437,1800,1824]" pageId="69" pageNumber="70">BMNH</collectionCode>
(E)
<date id="FF90D0E44C33C6121FFD2A2AFDEE2654" box="[477,573,1799,1824]" pageId="69" pageNumber="70" value="2008" valueMax="2069">2008-69</date>
; ♂, NE, Masoala, Anaovandrano,
<date id="FF90D0E44C33C6121D9E2A25FBE12654" box="[958,1074,1800,1824]" pageId="69" pageNumber="70" value="1994-01-14">14/1/1994</date>
, D.C. Lees: WG5 [wing image], IA38 [isotope voucher]; ♂, NE, Masoala, Ambavaony,
<quantity id="4CD65BC14C33C6121D342A01FC8C2630" box="[788,863,1836,1861]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="69" pageNumber="70" unit="m" value="125.0">125 m</quantity>
,
<date id="FF90D0E44C33C6121D4D2A01FC3D2630" box="[877,1006,1836,1860]" pageId="69" pageNumber="70" value="1994-11-27">27/11/1994</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C33C6121AA02A01FB682630" box="[1152,1211,1836,1860]" pageId="69" pageNumber="70">DCL</collectionCode>
47a, IA108 [isotope voucher]; ♂, NE, Masoala, Ambanivony,
<date id="FF90D0E44C33C6121C5C2A62FD2E261C" box="[636,765,1871,1896]" pageId="69" pageNumber="70" value="1993-11-25">25/11/1993</date>
, C. Kremen: CK26, IA109 [pheromone voucher]; ♀, NE, Makira, Vohitaly,
<geoCoordinate id="EE1A90E34C33C6121F472A5BFE0926FB" box="[359,474,1907,1935]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-15.4414">15.4414o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121FCA2A5BFD8D26FB" box="[490,606,1907,1935]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.5309">49.5309o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121CB32A5BFD2526FB" box="[659,758,1910,1935]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="69" pageNumber="70" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C33C6121D242A5BFC8326FB" box="[772,848,1910,1935]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="69" pageNumber="70" unit="m" value="645.0">645 m</quantity>
,
<date id="FF90D0E44C33C6121D7F2A5AFC3226FB" box="[863,993,1911,1935]" pageId="69" pageNumber="70" value="2002-12-30">30/12/2002</date>
, D.C. Lees: DL-SF00, IA112 [isotope voucher]; ♀, NE, Makira, Ankirindro,
<geoCoordinate id="EE1A90E34C33C6121C672AB0FD6926C2" box="[583,698,1946,1974]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="5" value="-15.2931">15.2931o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121CE62AB0FCE926C1" box="[710,826,1946,1974]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="5" value="49.5472">49.5472o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C33C6121D482AB0FC1526C2" box="[872,966,1949,1974]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="69" pageNumber="70" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C33C6121DF12AB0FBB726C1" box="[977,1124,1949,1974]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="69" pageNumber="70" unit="m" value="675.0" valueMax="700.0" valueMin="650.0">675 +/- 25 m</quantity>
,
<date id="FF90D0E44C33C6121A4F2AB0FB3026C2" box="[1135,1251,1949,1974]" pageId="69" pageNumber="70" value="2003-01-15">15/1/2003</date>
, D.C. Lees: DL- SE00, IA114 [isotope voucher]; specimen, NE, Ankiatomboka forest, c.
<quantity id="4CD65BC14C33C6121DE62AE8FBC326A8" box="[966,1040,1989,2013]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="69" pageNumber="70" unit="m" value="710.0">710 m</quantity>
,
<geoCoordinate id="EE1A90E34C33C6121A3C2AE9FB5226A9" box="[1052,1153,1985,2013]" direction="south" orientation="latitude" pageId="69" pageNumber="70" precision="55" value="-15.179">15.179o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C33C6121AAE2AE8FB2526A8" box="[1166,1270,1985,2013]" direction="east" orientation="longitude" pageId="69" pageNumber="70" precision="55" value="49.412">49.412o E</geoCoordinate>
,
<quantity id="4CD65BC14C33C6121B222AE8FA4B26A8" box="[1282,1432,1988,2013]" metricMagnitude="2" metricUnit="m" metricValue="7.1" metricValueMax="7.6" metricValueMin="6.6" pageId="69" pageNumber="70" unit="m" value="710.0" valueMax="760.0" valueMin="660.0">710 +/- 50 m</quantity>
,
<date id="FF90D0E44C30C6111EB72DB6FEDB21C7" box="[151,264,155,179]" pageId="70" pageNumber="71" value="2001-12-09">9/12/2001</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C30C6111FB42DB7FE6D21C7" box="[404,446,154,179]" pageId="70" pageNumber="71">DL</collectionCode>
01-169; ♂, NE, Bivontro, Makira,
<geoCoordinate id="EE1A90E34C30C6111D1B2DB7FC7C21C7" box="[827,943,151,179]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.3984">15.3984o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111D9B2DB6FBFD21C6" box="[955,1070,151,179]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.4477">49.4477o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111A7C2DB6FB7F21C7" box="[1116,1196,154,179]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="km" value="0.1">0.1 km</quantity>
,
<quantity id="4CD65BC14C30C6111A972DB7FA9A21C6" box="[1207,1353,154,179]" metricMagnitude="2" metricUnit="m" metricValue="7.62" metricValueMax="7.87" metricValueMin="7.37" pageId="70" pageNumber="71" unit="m" value="762.0" valueMax="787.0" valueMin="737.0">762 +/- 25 m</quantity>
,
<date id="FF90D0E44C30C6111B742DB6FF1C21AE" pageId="70" pageNumber="71" value="2002-12-21">21/12/ 2002</date>
, D.C. Lees: DL-SG77; ♂, NE, Vohitaly-Anjiahely,
<geoCoordinate id="EE1A90E34C30C6111D3C2DECFC4321AE" box="[796,912,190,218]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.3989">15.3989o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111DBE2DECFBC321AD" box="[926,1040,190,218]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.5211">49.5211o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111A622DEFFB7121AE" box="[1090,1186,193,218]" metricMagnitude="3" metricUnit="m" metricValue="1.82" pageId="70" pageNumber="71" unit="km" value="1.82">1.82 km</quantity>
,
<quantity id="4CD65BC14C30C6111A8F2DECFA9421AD" box="[1199,1351,193,218]" metricMagnitude="2" metricUnit="m" metricValue="4.35" metricValueMax="4.85" metricValueMin="3.85" pageId="70" pageNumber="71" unit="m" value="435.0" valueMax="485.0" valueMin="385.0">435 +/- 50 m</quantity>
,
<date id="FF90D0E44C30C6111B742DEFFF1C2075" pageId="70" pageNumber="71" value="2002-12-31">31/12/ 2002</date>
,
<collectionCode id="ED3F6EE14C30C6111EFD2DC4FEE72075" box="[221,308,233,257]" pageId="70" pageNumber="71">BMNH</collectionCode>
(E) #672331 [
<collectionCode id="ED3F6EE14C30C6111FEC2DC4FDDF2075" box="[460,524,233,257]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="70" pageNumber="71">DNA</collectionCode>
voucher]; ♀, NE, Anjiahely,
<quantity id="4CD65BC14C30C6111D792DC4FC662075" box="[857,949,232,257]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="70" pageNumber="71" unit="km" value="0.37">0.37 km</quantity>
E.,
<geoCoordinate id="EE1A90E34C30C6111DC32DC5FB852075" box="[995,1110,229,257]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.412">15.4120o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111A442DC5FB042074" box="[1124,1239,229,257]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.5022">49.5022o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111B272DC5FAB42075" box="[1287,1383,232,257]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="70" pageNumber="71" unit="km" value="0.25">0.25 km</quantity>
,
<quantity id="4CD65BC14C30C6111B522DC4FF262050" metricMagnitude="2" metricUnit="m" metricValue="3.8" metricValueMax="3.85" metricValueMin="3.75" pageId="70" pageNumber="71" unit="m" value="380.0" valueMax="385.0" valueMin="375.0">380 +/- 5 m</quantity>
<date id="FF90D0E44C30C6111F212C20FE572051" box="[257,388,269,293]" pageId="70" pageNumber="71" value="2002-12-12">12/12/2002</date>
, D.C. Lees: DL-SG1,
<collectionCode id="ED3F6EE14C30C6111CB32C20FD392051" box="[659,746,269,293]" pageId="70" pageNumber="71">BMNH</collectionCode>
(E)
<date id="FF90D0E44C30C6111D382C21FCAB2051" box="[792,888,268,293]" pageId="70" pageNumber="71" value="2008" valueMax="2069">2008-69</date>
,
<collectionCode id="ED3F6EE14C30C6111DA92C20FC332051" box="[905,992,269,293]" pageId="70" pageNumber="71">BMNH</collectionCode>
(E) 1717101; ♂, NE, Sahantaha, near fanihy roost area,
<geoCoordinate id="EE1A90E34C30C6111F552C1EFE3A203F" box="[373,489,303,332]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.1963">15.1963o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111FD62C1EFDBA203F" box="[502,617,303,332]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.5564">49.5564o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111CBA2C1EFD2A2038" box="[666,761,307,332]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="70" pageNumber="71" unit="km" value="0.98">0.98 km</quantity>
,
<quantity id="4CD65BC14C30C6111D242C1EFC41203F" box="[772,914,307,332]" metricMagnitude="2" metricUnit="m" metricValue="5.17" metricValueMax="5.71" metricValueMin="4.63" pageId="70" pageNumber="71" unit="m" value="517.0" valueMax="571.0" valueMin="463.0">517 +/-54 m</quantity>
,
<date id="FF90D0E44C30C6111DBE2C1EFBF1203F" box="[926,1058,307,331]" pageId="70" pageNumber="71" value="2001-12-13">13/12/2001</date>
: 10:46, D.C. Lees:
<collectionCode id="ED3F6EE14C30C6111ADC2C1EFAF62038" box="[1276,1317,307,332]" pageId="70" pageNumber="71">DL</collectionCode>
01-257; ♂, NE, Anjanaharibe Mt.,
<geoCoordinate id="EE1A90E34C30C6111F842C77FDDA2006" box="[420,521,342,371]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="55" value="-15.188">15.188o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111C382C77FD5F2006" box="[536,652,342,371]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.6139">49.6139o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111C9F2C77FCC22007" box="[703,785,346,371]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="km" value="0.1">0.1 km</quantity>
,
<quantity id="4CD65BC14C30C6111D3E2C77FC782006" box="[798,939,346,371]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.7" pageId="70" pageNumber="71" unit="m" value="475.0" valueMax="480.0" valueMin="470.0">475 +/- 5 m</quantity>
,
<date id="FF90D0E44C30C6111D982C77FBCF2007" box="[952,1052,346,371]" pageId="70" pageNumber="71" value="2003-02-06">6/2/2003</date>
, D.C. Lees; ♀, NE, Tampolo, W. Masoala, ca.
<quantity id="4CD65BC14C30C6111F0F2CADFEA820EC" box="[303,379,384,409]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="m" value="150.0">150 m</quantity>
, primary rainforest,
<geoCoordinate id="EE1A90E34C30C6111C462CADFD0920ED" box="[614,730,381,409]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.7301">15.7301o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111CC82CADFC8F20EC" box="[744,860,381,409]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.9604">49.9604o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111DB02CACFC1D20ED" box="[912,974,384,409]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="70" pageNumber="71" unit="km" value="2.0">2 km</quantity>
,
<date id="FF90D0E44C30C6111DFB2CADFB9F20ED" box="[987,1100,384,409]" pageId="70" pageNumber="71" value="2001-11-06">6/11/2001</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C30C6111AC12CACFADE20ED" box="[1249,1293,385,409]" pageId="70" pageNumber="71">DL</collectionCode>
01-1; ♀, NE, Tampolo, W. Masoala, course of old railroad
<quantity id="4CD65BC14C30C6111CAF2C85FD3720B4" box="[655,740,423,448]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="70" pageNumber="71" unit="km" value="1.5" valueMax="2.0" valueMin="1.0">12 km</quantity>
SE of camp,
<geoCoordinate id="EE1A90E34C30C6111D592C8AFC3E20B4" box="[889,1005,420,448]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.7279">15.7279o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111DD92C8AFBBE20CB" box="[1017,1133,420,448]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.9751">49.9751o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111ABB2C85FB3820B4" box="[1179,1259,423,448]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="70" pageNumber="71" unit="km" value="1.8">1.8 km</quantity>
,
<quantity id="4CD65BC14C30C6111AD62C85FAA820CB" box="[1270,1403,423,448]" metricMagnitude="1" metricUnit="m" metricValue="3.2" metricValueMax="5.7" metricValueMin="0.7" pageId="70" pageNumber="71" unit="m" value="32.0" valueMax="57.0" valueMin="7.0">32 +/- 25 m</quantity>
,
<date id="FF90D0E44C30C6111BA62C85FF222090" pageId="70" pageNumber="71" value="2001-11-09">9/ 11/2001</date>
, D.C. Lees,
<collectionCode id="ED3F6EE14C30C6111FA32CE1FE7F2090" box="[387,428,460,484]" pageId="70" pageNumber="71">DL</collectionCode>
01-27,
<collectionCode id="ED3F6EE14C30C6111FDA2CE1FD822090" box="[506,593,460,484]" pageId="70" pageNumber="71">BMNH</collectionCode>
(E) #697188 [
<collectionCode id="ED3F6EE14C30C6111CCB2CE1FCF82090" box="[747,811,460,484]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="70" pageNumber="71">DNA</collectionCode>
voucher], KA556 [=KA-P556;
<collectionCode id="ED3F6EE14C30C6111AB72CE1FB072090" box="[1175,1236,460,484]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="70" pageNumber="71">DNA</collectionCode>
extract voucher]; ♀, NE, Andranobe, W. Masoala,
<quantity id="4CD65BC14C30C6111C0A2CDEFDAA237E" box="[554,633,499,523]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="70" pageNumber="71" unit="m" value="240.0">240 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111CAB2CDFFD06237F" box="[651,725,495,523]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5555" value="-15.4">15.4o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111CC72CDFFCEC237E" box="[743,831,495,523]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="555" value="49.58">49.58o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111D5A2CDEFC0C237F" box="[890,991,498,523]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="70" pageNumber="71" unit="km" value="1.78">1.78 km</quantity>
,
<specimenCount id="9D283DAD4C30C6111DD02CDEFB96237E" box="[1008,1093,499,523]" pageId="70" pageNumber="71" type="generic">
<quantity id="4CD65BC14C30C6111DD02CDEFB92237E" box="[1008,1089,499,523]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="70" pageNumber="71" unit="m" value="240.0">240 m</quantity>
.
</specimenCount>
<date id="FF90D0E44C30C6111A722CDEFB66237F" box="[1106,1205,499,523]" pageId="70" pageNumber="71" value="1991-02-03">3/2/1991</date>
, C. Kremen et al., Genitalia157 [genitalia voucher]; ♀, NE, Andranobe, W. Masoala,
<quantity id="4CD65BC14C30C6111DA72F34FC112345" box="[903,962,537,561]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="70" pageNumber="71" unit="m" value="90.0">90 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111DEE2F34FB912345" box="[974,1090,534,562]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.6816">15.6816o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111A6E2F34FB112345" box="[1102,1218,534,562]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.9573">49.9573o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111AD12F34FA832346" box="[1265,1360,537,562]" metricMagnitude="3" metricUnit="m" metricValue="1.79" pageId="70" pageNumber="71" unit="km" value="1.79">1.79 km</quantity>
,
<quantity id="4CD65BC14C30C6111B7C2F34FA4B2345" box="[1372,1432,537,561]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="70" pageNumber="71" unit="m" value="90.0">90 m</quantity>
,
<date id="FF90D0E44C30C6111EB72F13FEC62322" box="[151,277,574,598]" pageId="70" pageNumber="71" value="1991-11-14">14/11/1991</date>
, C. Kremen et al.: Biodiversity voucher; ♀, NE, W. Masoala, Andranobe field station, T2S2 [site],
<quantity id="4CD65BC14C30C6111BA02F13FF7C2308" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="70" pageNumber="71" unit="m" value="90.0">90 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111E9C2F49FEE32308" box="[188,304,608,637]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.6816">15.6816o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111F1E2F49FE612308" box="[318,434,608,637]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.9573">49.9573o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111FC52F49FD952309" box="[485,582,612,637]" metricMagnitude="3" metricUnit="m" metricValue="1.79" pageId="70" pageNumber="71" unit="km" value="1.79">1.79 km</quantity>
,
<quantity id="4CD65BC14C30C6111C732F49FD432308" box="[595,656,612,636]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="70" pageNumber="71" unit="m" value="90.0">90 m</quantity>
,
<date id="FF90D0E44C30C6111CBE2F49FCCF2308" box="[670,796,612,636]" pageId="70" pageNumber="71" value="1991-11-14">14/11/1991</date>
, C. Kremen
<emphasis id="B95A2A364C30C6111D922F48FC3E2309" box="[946,1005,612,637]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
, Biodiversity survey voucher|Gen158 [genitalia], IA594 [isotope voucher]; ♂, data as above but:
<date id="FF90D0E44C30C6111D192FA5FC7B23D4" box="[825,936,648,672]" pageId="70" pageNumber="71" value="1991-11-02">2/11/1991</date>
, C. Kremen et al., Biodiversity voucher; ♂, NE, Andranobe, W. Masoala,
<quantity id="4CD65BC14C30C6111FDD2F83FD9F23B3" box="[509,588,686,711]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="70" pageNumber="71" unit="m" value="520.0">520 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111C7D2F83FD6723B3" box="[605,692,683,711]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="555" value="-15.65">15.65o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111CE62F82FCFF23B3" box="[710,812,683,711]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="55" value="49.983">49.983o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111D462F82FC1A23B3" box="[870,969,686,711]" metricMagnitude="3" metricUnit="m" metricValue="1.79" pageId="70" pageNumber="71" unit="km" value="1.79">1.79 km</quantity>
,
<quantity id="4CD65BC14C30C6111DFA2F83FBFA23B3" box="[986,1065,686,711]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="70" pageNumber="71" unit="m" value="520.0">520 m</quantity>
,
<date id="FF90D0E44C30C6111A1A2F82FB6B23B3" box="[1082,1208,687,711]" pageId="70" pageNumber="71" value="1991-11-14">14/11/1991</date>
, C. Kremen et al., Biodiversity voucher; ♂, data as above but:
<quantity id="4CD65BC14C30C6111CA62FFFFD1C239E" box="[646,719,722,747]" metricMagnitude="2" metricUnit="m" metricValue="5.99" pageId="70" pageNumber="71" unit="m" value="599.0">599 m</quantity>
,
<date id="FF90D0E44C30C6111CFB2FFEFC89239F" box="[731,858,723,747]" pageId="70" pageNumber="71" value="1991-10-20">20/10/1991</date>
, C. Kremen
<emphasis id="B95A2A364C30C6111DCB2FF9FBF0239F" box="[1003,1059,722,747]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
, Biodiversity voucher; ♂, data as above but:
<date id="FF90D0E44C30C6111F322FDAFE51227B" box="[274,386,759,783]" pageId="70" pageNumber="71" value="1991-11-08">8/11/1991</date>
, C. Kremen
<emphasis id="B95A2A364C30C6111C312FD5FD9A227B" box="[529,585,758,783]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
: Biodiversity voucher; ♀, data as above but:
<quantity id="4CD65BC14C30C6111A612FDAFB5A227B" box="[1089,1161,759,783]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="m" value="100.0">100 m</quantity>
,
<date id="FF90D0E44C30C6111AB52FDBFAC7227B" box="[1173,1300,758,783]" pageId="70" pageNumber="71" value="1991-10-26">26/10/1991</date>
, C. Kremen
<emphasis id="B95A2A364C30C6111EB72E31FF1D2247" box="[151,206,794,819]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
: Bio|Gen156 [genitalia]; ♀, data as above but:
<date id="FF90D0E44C30C6111CFB2E36FC892247" box="[731,858,795,819]" pageId="70" pageNumber="71" value="1991-10-29">29/10/1991</date>
, C. Kremen
<emphasis id="B95A2A364C30C6111DC92E31FBF32247" box="[1001,1056,794,819]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
: Bio|Gen159; ♀, NE, Andranobe, W. Masoala,
<quantity id="4CD65BC14C30C6111F0B2E6CFEA7222D" box="[299,372,833,858]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="70" pageNumber="71" unit="m" value="520.0">520 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111FA02E6CFE0B222E" box="[384,472,830,858]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="555" value="-15.65">15.65o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111FC52E6FFD98222D" box="[485,587,830,858]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="55" value="49.983">49.983o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111C5B2E6FFD09222E" box="[635,730,833,858]" metricMagnitude="3" metricUnit="m" metricValue="1.79" pageId="70" pageNumber="71" unit="km" value="1.79">1.79 km</quantity>
,
<quantity id="4CD65BC14C30C6111CC52E6CFCFC222D" box="[741,815,833,858]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="70" pageNumber="71" unit="m" value="520.0">520 m</quantity>
,
<date id="FF90D0E44C30C6111D1B2E6FFC7F222E" box="[827,940,834,858]" pageId="70" pageNumber="71" value="1991-11-08">8/11/1991</date>
, C. Kremen
<emphasis id="B95A2A364C30C6111A1D2E6FFBA6222E" box="[1085,1141,833,858]" italics="true" pageId="70" pageNumber="71">et al.</emphasis>
: Biodiversity voucher; ♂, NE, Be Dinta, upper Anaovandrano R., W. Masoala,
<quantity id="4CD65BC14C30C6111CC62E45FCFC22F4" box="[742,815,872,897]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">500 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111D1A2E45FC7E22F5" box="[826,941,869,897]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.6733">15.6733o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111D9A2E45FBFE22F4" box="[954,1069,869,897]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.9885">49.9885o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111A7B2E45FB6922F5" box="[1115,1210,872,897]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C30C6111AE52E45FA8422F4" box="[1221,1367,872,897]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="70" pageNumber="71" unit="m" value="500.0" valueMax="550.0" valueMin="450.0">500 +/- 50 m</quantity>
,
<date id="FF90D0E44C30C6111B422E44FF1C22D1" pageId="70" pageNumber="71" value="1993-02-18">18/2/ 1993</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C30C6111F442EA0FE5F22D1" box="[356,396,909,933]" pageId="70" pageNumber="71">DL</collectionCode>
93-0012, IA593 [isotope voucher]; ♂, NE, Masoala E, Be Dinta,
<quantity id="4CD65BC14C30C6111A5D2EA1FB1A22D0" box="[1149,1225,908,933]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="70" pageNumber="71" unit="m" value="550.0">550 m</quantity>
,
<date id="FF90D0E44C30C6111AF72EA0FA8A22D1" box="[1239,1369,909,933]" pageId="70" pageNumber="71" value="1993-02-17">17/02/1993</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C30C6111EFF2E9CFED422BD" box="[223,263,945,969]" pageId="70" pageNumber="71">DL</collectionCode>
93-0029, IA592 [isotope voucher]; ♂, NE, Be Dinta, upper Anaovandrano R., W. Masoala,
<quantity id="4CD65BC14C30C6111B692E9DFA4B22BC" box="[1353,1432,944,969]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">500 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111EB72EFAFED9229B" box="[151,266,980,1008]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.6733">15.6733o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111F3A2EFAFE5E229B" box="[282,397,980,1008]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.9885">49.9885o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111FE22EFAFDF02284" box="[450,547,983,1008]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C30C6111C112EFAFD1E229B" box="[561,717,983,1008]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="70" pageNumber="71" unit="m" value="500.0" valueMax="550.0" valueMin="450.0">500 +/- 50 m</quantity>
, swamp forest,
<date id="FF90D0E44C30C6111DA62EFAFC24229B" box="[902,1015,983,1007]" pageId="70" pageNumber="71" value="1993-02-18">18/2/1993</date>
, D.C. Lees:
<collectionCode id="ED3F6EE14C30C6111AAD2EFAFB66229B" box="[1165,1205,983,1007]" pageId="70" pageNumber="71">DL</collectionCode>
93-0030; ♂, NE, Be Dinta, upper Anaovandrano R., W. Masoala,
<quantity id="4CD65BC14C30C6111CB62ED0FD332562" box="[662,736,1021,1046]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">500 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111CCD2ED0FCB22562" box="[749,865,1018,1046]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.6733">15.6733o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111D4E2ED0FC312562" box="[878,994,1018,1046]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.9885">49.9885o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111A332ED0FBA02562" box="[1043,1139,1021,1046]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C30C6111A5F2ED0FAC52562" box="[1151,1302,1021,1046]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="70" pageNumber="71" unit="m" value="500.0" valueMax="550.0" valueMin="450.0">500 +/- 50 m</quantity>
,
<date id="FF90D0E44C30C6111B032ED3FA4B2562" box="[1315,1432,1022,1046]" pageId="70" pageNumber="71" value="1993-02-17">17/2/1993</date>
: 16:30, D.C. Lees:
<collectionCode id="ED3F6EE14C30C6111F48290FFE43254E" box="[360,400,1058,1082]" pageId="70" pageNumber="71">DL</collectionCode>
93-0031; ♂ [
<collectionCode id="ED3F6EE14C30C6111C3F290EFDA8254E" box="[543,635,1059,1082]" pageId="70" pageNumber="71">BMNH</collectionCode>
],
<collectingCountry id="F339B6B44C30C6111CAE290FFCC6254F" box="[654,789,1058,1083]" name="Madagascar" pageId="70" pageNumber="71">Madagascar</collectingCountry>
NE, Masoala, Be Dinta, R. Anaovandrano,
<quantity id="4CD65BC14C30C6111B2B290FFA86254E" box="[1291,1365,1058,1083]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="500.0">500 m</quantity>
,
<date id="FF90D0E44C30C6111B43290FFF012515" pageId="70" pageNumber="71" value="1993-02-20">20/2/ 1993</date>
, D.C. Lees; ♀, NE, Iketry [Iketra], lower Anaovandrano R., W7-L2, Masoala E.,
<geoCoordinate id="EE1A90E34C30C6111A5C2965FB232515" box="[1148,1264,1093,1121]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.7216">15.7216o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111ADE2965FAA12514" box="[1278,1394,1093,1121]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.1753">50.1753o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111EB72942FF2625FC" box="[151,245,1135,1160]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C30C6111F22295DFE9825F3" box="[258,331,1136,1160]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="70" pageNumber="71" unit="m" value="270.0">270 m</quantity>
,
<date id="FF90D0E44C30C6111F77295DFE1A25FC" box="[343,457,1136,1160]" pageId="70" pageNumber="71" value="1994-01-18">18/1/1994</date>
, D.C. Lees; ♀, NE, Manosona, Anaovandrano R., E Masoala,
<geoCoordinate id="EE1A90E34C30C6111AAD2942FB2025FC" box="[1165,1267,1132,1160]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="55" value="-15.784">15.784o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111B202942FAA025F3" box="[1280,1395,1132,1160]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.2164">50.2164o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111EB729BBFF2425DB" box="[151,247,1174,1199]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C30C6111F2329BBFEEC25DA" box="[259,319,1174,1199]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="70" pageNumber="71" unit="m" value="45.0">45 m</quantity>
,
<date id="FF90D0E44C30C6111F6D29BAFE6325DB" box="[333,432,1175,1199]" pageId="70" pageNumber="71" value="1994-02-01">1/2/1994</date>
, D.C. Lees; ♂, NE, Masoala E., Antsamanarana R.,
<quantity id="4CD65BC14C30C6111A2929BBFB8025DA" box="[1033,1107,1174,1199]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="70" pageNumber="71" unit="m" value="275.0">275 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111A4029BBFB1625DB" box="[1120,1221,1171,1199]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="55" value="-15.295">15.295o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111AF329BBFAEA25DA" box="[1235,1337,1171,1199]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="55" value="50.227">50.227o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111B4B29BBFF6D25A7" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C30C6111EF22997FE5225A6" box="[210,385,1210,1235]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="3.25" metricValueMin="2.25" pageId="70" pageNumber="71" unit="m" value="275.0" valueMax="325.0" valueMin="225.0">275 +/- 50 m</quantity>
, site W2L2-275M 5 LO,
<date id="FF90D0E44C30C6111CEC2997FCED25A7" box="[716,830,1210,1235]" pageId="70" pageNumber="71" value="1993-12-05">5/12/1993</date>
, H. Raharitsimba:
<collectionCode id="ED3F6EE14C30C6111A0E2996FB8525A7" box="[1070,1110,1211,1235]" pageId="70" pageNumber="71">DL</collectionCode>
93-0028; ♂ [developmental abnormality LHS CuA1 HWV], NE, E Masoala, Ambavony R. [Ambanivony], W2-L1-125M [site], disturbed forest,
<geoCoordinate id="EE1A90E34C30C6111EC52828FE8A246A" box="[229,345,1282,1310]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.2813">15.2813o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111F462828FE092469" box="[358,474,1282,1310]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.2877">50.2877o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111C2B282BFDB9246A" box="[523,618,1285,1310]" metricMagnitude="2" metricUnit="m" metricValue="7.199999999999999" pageId="70" pageNumber="71" unit="km" value="0.72">0.72 km</quantity>
,
<quantity id="4CD65BC14C30C6111C56282BFCDF2469" box="[630,780,1285,1310]" metricMagnitude="2" metricUnit="m" metricValue="2.23" metricValueMax="2.73" metricValueMin="1.73" pageId="70" pageNumber="71" unit="m" value="223.0" valueMax="273.0" valueMin="173.0">223 +/- 50 m</quantity>
,
<date id="FF90D0E44C30C6111D39282BFC4B246A" box="[793,920,1286,1310]" pageId="70" pageNumber="71" value="1993-11-29">29/11/1993</date>
, C. Kremen: CK93-0009; ♀, NE, E Masoala, [Ambavaony R.], bifurcation to
<quantity id="4CD65BC14C30C6111C242801FD9D2430" box="[516,590,1324,1348]" metricMagnitude="2" metricUnit="m" metricValue="3.7" pageId="70" pageNumber="71" unit="m" value="370.0">370 m</quantity>
, riparian, W2L1 Saharand. [Sarahandrano]
<geoCoordinate id="EE1A90E34C30C6111A1C2801FB472430" box="[1084,1172,1320,1349]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="555" value="-15.28">15.28o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111A812801FAC62430" box="[1185,1301,1320,1349]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.286">50.2860o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111B662801FA4B2431" box="[1350,1432,1324,1349]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="70" pageNumber="71" unit="km" value="1.4">1.4 km</quantity>
,
<quantity id="4CD65BC14C30C6111EB7287DFEEA2413" box="[151,313,1360,1384]" metricMagnitude="2" metricUnit="m" metricValue="2.4" metricValueMax="4.3" metricValueMin="0.5" pageId="70" pageNumber="71" unit="m" value="240.0" valueMax="430.0" valueMin="50.0">240 +/- 190 m</quantity>
,
<date id="FF90D0E44C30C6111F642862FE11241C" box="[324,450,1359,1384]" pageId="70" pageNumber="71" value="1993-11-25">25/11/1993</date>
, H. Raharitsimba:
<collectingCountry id="F339B6B44C30C6111CB72862FD6E241C" box="[663,701,1359,1384]" name="Chile" pageId="70" pageNumber="71">TR</collectingCountry>
46A; ♂, NE, Masoala E., Antsamanarana R., W2L2 [site],
<quantity id="4CD65BC14C30C6111B6E2862FA442413" box="[1358,1431,1359,1384]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="70" pageNumber="71" unit="m" value="275.0">275 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111EB7285BFED924FB" box="[151,266,1395,1423]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.2813">15.2813o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111F37285BFE5824FB" box="[279,395,1395,1423]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.2877">50.2877o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111F9A285AFDCB24FB" box="[442,536,1398,1423]" metricMagnitude="2" metricUnit="m" metricValue="7.199999999999999" pageId="70" pageNumber="71" unit="km" value="0.72">0.72 km</quantity>
, 5LO [fruit trap], riparian,
<date id="FF90D0E44C30C6111D67285AFC1524FB" box="[839,966,1399,1423]" pageId="70" pageNumber="71" value="1993-12-11">11/12/1993</date>
, H. Raharitsimba:
<collectionCode id="ED3F6EE14C30C6111ABA285BFB1124FB" box="[1178,1218,1398,1423]" pageId="70" pageNumber="71">DL</collectionCode>
93-0026; ♀, data as above but:
<quantity id="4CD65BC14C30C6111F3A28B7FEB524C6" box="[282,358,1434,1459]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="70" pageNumber="71" unit="m" value="275.0">275 m</quantity>
,
<date id="FF90D0E44C30C6111F5528B7FE3524C7" box="[373,486,1434,1459]" pageId="70" pageNumber="71" value="1993-12-05">5/12/1993</date>
, H. Raharitsimba, GEN 117
<collectionCode id="ED3F6EE14C30C6111D0E28B6FC8624C7" box="[814,853,1435,1459]" pageId="70" pageNumber="71">DL</collectionCode>
[genitalia]; ♂ (
<figureCitation id="1315EAA14C30C6111A2C28B7FBB324C7" box="[1036,1120,1434,1459]" captionStart="FIGURE 18" captionStartId="59.[151,250,1877,1899]" captionTargetBox="[234,1357,184,1854]" captionTargetId="figure@59.[220,1368,169,1871]" captionTargetPageId="59" captionText="FIGURE 18. ♂ genitalia in LV (left), DV (right). A: Heteropsis tornado Lees, Allaoui &amp; Aduse-Poku, sp. nov., ♂ genitalia ( Mahassabe; 362 DL) (detail of valve inset; drawing modified after Lees, 1997: 106); not to scale. B: ♂ genitalia of Ht. lanyvary. (left: DL- 05 - 761; right: DL- 05 - 845). C: ♂ genitalia of Heteropsis barbarae sp. nov. (Antsamanarana R., Masoala; CK 30, 141 DL). D: ♂ genitalia of Heteropsis menamenoides Lees, sp. nov. (Mt. Bekolosy; DLBEK 94 _ 180; 231 DL). Scale bar: 3 mm." httpUri="https://zenodo.org/record/264615/files/figure.png" pageId="70" pageNumber="71">Fig. 18</figureCitation>
C), NE, Antsamanarana R., Masoala E., [
<quantity id="4CD65BC14C30C6111F1128ECFE7F24AD" box="[305,428,1473,1498]" metricMagnitude="2" metricUnit="m" metricValue="2.85" metricValueMax="5.2" metricValueMin="0.5" pageId="70" pageNumber="71" unit="m" value="285.0" valueMax="520.0" valueMin="50.0">50520 m</quantity>
],
<geoCoordinate id="EE1A90E34C30C6111FE328ECFDFB24AE" box="[451,552,1470,1498]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="55" value="-15.307">15.307o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111C1828ECFD4D24AD" box="[568,670,1470,1498]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="55" value="50.233">50.233o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111CF428EFFCFB24AE" box="[724,808,1473,1498]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="km" value="0.1">0.1 km</quantity>
,
<quantity id="4CD65BC14C30C6111D1728EFFC5724AD" box="[823,900,1474,1498]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="70" pageNumber="71" unit="m" value="100.0">100 m</quantity>
,
<date id="FF90D0E44C30C6111DB328EFFBC124AE" box="[915,1042,1474,1498]" pageId="70" pageNumber="71" value="1993-11-29">29/11/1993</date>
, C. Kremen: CK30, GEN 141
<collectionCode id="ED3F6EE14C30C6111B5428EFFA4F24AE" box="[1396,1436,1474,1498]" pageId="70" pageNumber="71">DL</collectionCode>
[genitalia]; ♀, data as above but:
<geoCoordinate id="EE1A90E34C30C6111C3528C5FD5A2775" box="[533,649,1509,1537]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.2968">15.2968o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111CB728C5FCD82774" box="[663,779,1509,1537]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.2271">50.2271o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111D1C28C4FC4E2775" box="[828,925,1512,1537]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="70" pageNumber="71" unit="km" value="1.31">1.31 km</quantity>
,
<quantity id="4CD65BC14C30C6111D8928C4FB832774" box="[937,1104,1512,1537]" metricMagnitude="2" metricUnit="m" metricValue="2.94" metricValueMax="5.1899999999999995" metricValueMin="0.69" pageId="70" pageNumber="71" unit="m" value="294.0" valueMax="519.0" valueMin="69.0">294 +/- 225 m</quantity>
,
<date id="FF90D0E44C30C6111A7D28C4FB0E2775" box="[1117,1245,1513,1537]" pageId="70" pageNumber="71" value="1993-12-10">10/12/1993</date>
, D.C. Lees: E64 (egg voucher); ♂, NE, Antsamanarana R., Masoala E.,
<quantity id="4CD65BC14C30C6111D222B22FC982753" box="[770,843,1551,1576]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="70" pageNumber="71" unit="m" value="250.0">250 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111D772B22FC732753" box="[855,928,1548,1576]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5555" value="-15.3">15.3o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111D8D2B22FBD62753" box="[941,1029,1548,1576]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="555" value="50.23">50.23o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111A142B22FB40275C" box="[1076,1171,1551,1576]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="km" value="0.15">0.15 km</quantity>
,
<quantity id="4CD65BC14C30C6111ABE2B22FB342753" box="[1182,1255,1551,1576]" metricMagnitude="2" metricUnit="m" metricValue="2.69" pageId="70" pageNumber="71" unit="m" value="269.0">269 m</quantity>
,
<date id="FF90D0E44C30C6111AD32B22FAA12753" box="[1267,1394,1551,1575]" pageId="70" pageNumber="71" value="1993-12-12">12/12/1993</date>
, H. Raharitsimba, GEN 142
<collectionCode id="ED3F6EE14C30C6111F892B1BFE03273A" box="[425,464,1590,1614]" pageId="70" pageNumber="71">DL</collectionCode>
[genitalia]; ♀, NE, Ambavaony R., E Masoala, [
<quantity id="4CD65BC14C30C6111A272B18FB52273A" box="[1031,1153,1589,1614]" metricMagnitude="2" metricUnit="m" metricValue="2.15" metricValueMax="3.8" metricValueMin="0.5" pageId="70" pageNumber="71" unit="m" value="215.0" valueMax="380.0" valueMin="50.0">50380 m</quantity>
],
<geoCoordinate id="EE1A90E34C30C6111AB72B18FB2F273A" box="[1175,1276,1586,1614]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="55" value="-15.279">15.279o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111B2C2B18FAA2273A" box="[1292,1393,1586,1614]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="55" value="50.288">50.288o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111EB72B77FF242707" box="[151,247,1626,1651]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="70" pageNumber="71" unit="km" value="0.12">0.12 km</quantity>
,
<quantity id="4CD65BC14C30C6111F232B77FE9E2706" box="[259,333,1626,1651]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="70" pageNumber="71" unit="m" value="150.0">150 m</quantity>
,
<date id="FF90D0E44C30C6111F792B77FE182706" box="[345,459,1626,1650]" pageId="70" pageNumber="71" value="1993-12-01">1/12/1993</date>
, C. Kremen: CK37, EGG [egg voucher]; ♀, data as above but:
<quantity id="4CD65BC14C30C6111ABE2B77FB3B2706" box="[1182,1256,1626,1650]" metricMagnitude="2" metricUnit="m" metricValue="2.23" pageId="70" pageNumber="71" unit="m" value="223.0">223 m</quantity>
,
<date id="FF90D0E44C30C6111AD52B77FAA02706" box="[1269,1395,1626,1650]" pageId="70" pageNumber="71" value="1993-11-29">29/11/1993</date>
, C. Kremen; ♀, NE, E Masoala, Ambavaony R., riparian, W2-L1-50M [
<collectionCode id="ED3F6EE14C30C6111DBB2BACFC0E27ED" box="[923,989,1665,1689]" pageId="70" pageNumber="71">SITE</collectionCode>
],
<geoCoordinate id="EE1A90E34C30C6111DD02BADFBB727ED" box="[1008,1124,1661,1689]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.2813">15.2813o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111A512BADFB3627EC" box="[1137,1253,1661,1689]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.2877">50.2877o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111B352BACFAA727ED" box="[1301,1396,1664,1689]" metricMagnitude="2" metricUnit="m" metricValue="7.199999999999999" pageId="70" pageNumber="71" unit="km" value="0.72">0.72 km</quantity>
,
<quantity id="4CD65BC14C30C6111BA02BADFF7C27C8" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="m" value="50.0">50 m</quantity>
, 2LO [fruit trap]
<date id="FF90D0E44C30C6111F552B88FE2027C9" box="[373,499,1701,1725]" pageId="70" pageNumber="71" value="1993-11-28">28/11/1993</date>
, D.C. Lees: E3 [egg voucher; “pearly white, smallish, like
<taxonomicName id="4C2E8DA74C30C6111AB32B8BFADE27C9" box="[1171,1293,1702,1725]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="B95A2A364C30C6111AB32B8BFADE27C9" box="[1171,1293,1702,1725]" italics="true" pageId="70" pageNumber="71">Ht. pauper</emphasis>
</taxonomicName>
”]; ♂, NE, E Masoala, Ambavaony [Ambanivony],
<quantity id="4CD65BC14C30C6111C782BE1FD742797" box="[600,679,1740,1764]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="70" pageNumber="71" unit="m" value="380.0">380 m</quantity>
, W2L1 [
<collectionCode id="ED3F6EE14C30C6111D342BE1FC872790" box="[788,852,1740,1764]" pageId="70" pageNumber="71">SITE</collectionCode>
]
<geoCoordinate id="EE1A90E34C30C6111D472BE6FC082790" box="[871,987,1736,1764]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="-15.2813">15.2813o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111DCD2BE6FBB22797" box="[1005,1121,1736,1764]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="50.2877">50.2877o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111ABC2BE6FB0D2790" box="[1180,1246,1739,1764]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="70" pageNumber="71" unit="km" value="5.0">5 km</quantity>
,
<date id="FF90D0E44C30C6111ACE2BE1FABE2790" box="[1262,1389,1740,1764]" pageId="70" pageNumber="71" value="1993-11-30">30/11/1993</date>
, H. Raharitsimba:
<collectingCountry id="F339B6B44C30C6111F1D2BDFFEB6267F" box="[317,357,1778,1803]" name="Chile" pageId="70" pageNumber="71">TR</collectingCountry>
61; ♂, NE, Ambavaony R., E Masoala,
<quantity id="4CD65BC14C30C6111D0B2BDFFCA5267E" box="[811,886,1778,1803]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="70" pageNumber="71" unit="m" value="125.0">125 m</quantity>
,
<geoCoordinate id="EE1A90E34C30C6111DA32BDFFC3A267F" box="[899,1001,1775,1803]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="55" value="-15.279">15.279o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111DD72BDFFB8E267E" box="[1015,1117,1775,1803]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="55" value="50.288">50.288o E</geoCoordinate>
+/-
<quantity id="4CD65BC14C30C6111AB02BDEFB23267F" box="[1168,1264,1778,1803]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="70" pageNumber="71" unit="km" value="0.12">0.12 km</quantity>
,
<quantity id="4CD65BC14C30C6111ADD2BDFFA9B267E" box="[1277,1352,1778,1803]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="70" pageNumber="71" unit="m" value="125.0">125 m</quantity>
,
<date id="FF90D0E44C30C6111B762BDEFF1C265B" pageId="70" pageNumber="71" value="1993-11-27">27/11/ 1993</date>
, C. Kremen, GEN 124
<collectionCode id="ED3F6EE14C30C6111FC62A3AFDDE265B" box="[486,525,1815,1839]" pageId="70" pageNumber="71">DL</collectionCode>
[genitalia]; ♂, NE, Masoala, 1993, C. Kremen: CK93-0005, IA595 [isotope voucher]; ♂ [
<collectionCode id="ED3F6EE14C30C6111F102A13FE5F2621" box="[304,396,1854,1877]" pageId="70" pageNumber="71">BMNH</collectionCode>
], NE, Reserve Speciale d'Ambatovaky,
<quantity id="4CD65BC14C30C6111D722A13FC482621" box="[850,923,1854,1878]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="70" pageNumber="71" unit="m" value="700.0">700 m</quantity>
[
<geoCoordinate id="EE1A90E34C30C6111D8B2A10FC242622" box="[939,1015,1850,1878]" direction="south" orientation="latitude" pageId="70" pageNumber="71" precision="5555" value="-16.8">16.8o S</geoCoordinate>
,
<geoCoordinate id="EE1A90E34C30C6111A252A13FBA82621" box="[1029,1147,1850,1878]" direction="east" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="49.1222">49.1222o E</geoCoordinate>
,
<quantity id="4CD65BC14C30C6111AA92A10FB052621" box="[1161,1238,1853,1878]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="70" pageNumber="71" unit="m" value="580.0">580 m</quantity>
],
<date id="FF90D0E44C30C6111ACB2A13FA8F2622" box="[1259,1372,1854,1878]" pageId="70" pageNumber="71" value="1990-02-11">11/2/1990</date>
, M.I. Evans [#1.16], 254
<collectionCode id="ED3F6EE14C30C6111F532A4FFE4B260D" box="[371,408,1890,1913]" pageId="70" pageNumber="71">DL</collectionCode>
[genitalia].
</paragraph>
</subSubSection>
</treatment>
</document>