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<document id="DDDBCBF3731D1D09F5C5CE9F188AD3F1" ID-CLB-Dataset="31881" ID-DOI="10.5852/ejt.2016.255" ID-GBIF-Dataset="2719d968-2e75-4336-8e59-4c9563abdda2" ID-Zenodo-Dep="831094" ID-ZooBank="http://zoobank.org/urn:lsid:zoobank.org:pub:0E8DA4DC-FF4C-436E-94FB-CB89F6416C6E" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500286020512" checkinUser="jeremy" docAuthor="Nadine Duperre &amp; Elicio Tapia" docDate="2016" docId="F319EF090710FB14FD257B7C89F9C0DA" docLanguage="en" docName="Dupérré, N., _ Tapia, E. (2016). Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) (Done).pdf.imf" docOrigin="European Jornal of Taxonomy 255" docSource="http://sciencepress.mnhn.fr/en/periodiques/european-journal-taxonomy/2016/overview-anyphaenids-araneae-anyphaeninae-anyphaenidae-spider-fauna-choco-forest-ecuador-description-thirteen-new-species" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Katissa yaya Duperre &amp; Tapia, 2016, sp. nov." docType="treatment" docUuid="24956AEC-8F8D-4055-86B9-3253544FDE10" docUuidSource="ZooBank" docVersion="12" lastPageNumber="18" masterDocId="0F209771071EFB05FFBB7F778B3DC26A" masterDocTitle="Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Chocó forest of Ecuador, with the description of thirteen new species" masterLastPageNumber="50" masterPageNumber="1" pageNumber="15" updateTime="1698465609934" updateUser="ExternalLinkService">
<mods:mods id="567DE7162460217024BE0ED5964C9D51" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BDB447FFB71A5D766A4D93673AF42342">
<mods:title id="356A7464F71CF0CBA6310B02696670BD">Overview of the Anyphaenids (Araneae, Anyphaeninae, Anyphaenidae) spider fauna from the Chocó forest of Ecuador, with the description of thirteen new species</mods:title>
</mods:titleInfo>
<mods:name id="75ADD7BD1B40E2AFAFEB27181A94B5F8" type="personal">
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</mods:role>
<mods:namePart id="EA88D50C30F1F12CF0E4429979267B46">Nadine Duperre</mods:namePart>
</mods:name>
<mods:name id="B6CCA19303FBBCE4211F54EBF4675B0D" type="personal">
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<mods:namePart id="833B232AD0FF65E8F1A02DDB651F76FA">Elicio Tapia</mods:namePart>
</mods:name>
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<mods:title id="3F19CC829A25516724BCCCD1242BF3CD">European Jornal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="474DC9E321DC98A3BC56D87B33662C3D">
<mods:date id="151CDBADA3C58A5D2644609972C8EFA3">2016</mods:date>
<mods:detail id="60857E41C80F396EA907ADC0E162CA3E" type="volume">
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<mods:url id="77F66429E08243EA11BDFADBE0AA7C41">http://sciencepress.mnhn.fr/en/periodiques/european-journal-taxonomy/2016/overview-anyphaenids-araneae-anyphaeninae-anyphaenidae-spider-fauna-choco-forest-ecuador-description-thirteen-new-species</mods:url>
</mods:location>
<mods:classification id="F0D9A5A2687D752378DBB6D2EAB44F6D">journal article</mods:classification>
<mods:identifier id="09C4BB3D848A00D6EB4F12FB77EAF3DD" type="CLB-Dataset">31881</mods:identifier>
<mods:identifier id="468C097AA026974A4A556A80D6AC9B9D" type="DOI">10.5852/ejt.2016.255</mods:identifier>
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<mods:identifier id="594F55E81A6F62CFC71C3ADE5FC023E0" type="ZooBank">http://zoobank.org/urn:lsid:zoobank.org:pub:0E8DA4DC-FF4C-436E-94FB-CB89F6416C6E</mods:identifier>
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<treatment id="F319EF090710FB14FD257B7C89F9C0DA" ID-DOI="http://doi.org/10.5281/zenodo.6081741" ID-GBIF-Taxon="135129812" ID-Zenodo-Dep="6081741" LSID="urn:lsid:zoobank.org:act:24956AEC-8F8D-4055-86B9-3253544FDE10" httpUri="http://treatment.plazi.org/id/F319EF090710FB14FD257B7C89F9C0DA" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="33AA0D940710FB0BFD257B7C88ABC64F" box="[670,918,1035,1061]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="7B0F5E1F0710FB0BFD257B7C88ABC64F" blockId="14.[373,1214,1035,1131]" box="[670,918,1035,1061]" pageId="14" pageNumber="15">
<heading id="2047E9730710FB0BFD257B7C88ABC64F" box="[670,918,1035,1061]" centered="true" fontSize="11" level="5" pageId="14" pageNumber="15" reason="4">
<taxonomicName id="BCB0259C0710FB0BFD257B7C8809C64F" ID-CoL="6NJNN" box="[670,820,1035,1061]" class="Arachnida" family="Anyphaenidae" genus="Katissa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="yaya" status="sp. nov.">
<emphasis id="49C4820D0710FB0BFD257B7C8809C64F" bold="true" box="[670,820,1035,1061]" italics="true" pageId="14" pageNumber="15">Katissa yaya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52F73F760710FB0BFC877B7C88ABC64F" box="[828,918,1035,1061]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="33AA0D940710FB0BFECE7B598861C601" pageId="14" pageNumber="15" type="description">
<paragraph id="7B0F5E1F0710FB0BFECE7B598F83C623" blockId="14.[373,1214,1035,1131]" box="[373,1214,1070,1097]" pageId="14" pageNumber="15">urn:lsid:zoobank.org:act:24956AEC-8F8D-4055-86B9-3253544FDE10</paragraph>
<paragraph id="7B0F5E1F0710FB0BFD637B268861C601" blockId="14.[373,1214,1035,1131]" box="[728,860,1105,1131]" pageId="14" pageNumber="15">
<figureCitation id="E38B429A0710FB0BFD637B268861C601" box="[728,860,1105,1131]" captionStart="Figs 17 – 21" captionStartId="15.[189,241,1929,1955]" captionTargetBox="[217,1366,246,1881]" captionTargetId="figure@15.[217,1366,246,1881]" captionTargetPageId="15" captionText="Figs 17 – 21. Katissa yaya sp. nov. 17 .  Ƌ,   palp,   ventral  view.   18. Ƌ,   palp,   retrolateral  view. 19 .  ♀,   abdomen, dorsal view. 20 .  ♀,   epigynum,  ventral  view.   21 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars:   17 – 18, 20 – 21 = 0.1 mm; 19 = 1 mm." httpUri="https://zenodo.org/record/1038244/files/figure.png" pageId="14" pageNumber="15">Figs 17–21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="33AA0D940710FB0BFF067BE58AF6C72F" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="7B0F5E1F0710FB0BFF067BE58A0BC6C6" blockId="14.[189,310,1170,1196]" box="[189,310,1170,1196]" pageId="14" pageNumber="15">
<heading id="2047E9730710FB0BFF067BE58A0BC6C6" bold="true" box="[189,310,1170,1196]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="2">
<emphasis id="49C4820D0710FB0BFF067BE58A0BC6C6" bold="true" box="[189,310,1170,1196]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F0710FB0BFF067BB68AF6C72F" blockId="14.[189,1399,1217,1349]" pageId="14" pageNumber="15">
Males can be distinguished from all species by their elongated, serrated palpal retrolateral tibial apophysis with small dorsal spur, from
<taxonomicName id="BCB0259C0710FB0BFD2C7B92886CC694" box="[663,849,1253,1278]" class="Arachnida" family="Anyphaenidae" genus="Katissa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="guayasamini" status="sp. nov.">
<emphasis id="49C4820D0710FB0BFD2C7B92886CC694" box="[663,849,1253,1278]" italics="true" pageId="14" pageNumber="15">K. guayasamini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52F73F760710FB0BFCE17B92888AC695" box="[858,951,1253,1279]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
by the closely positioned dorsal spur (
<figureCitation id="E38B429A0710FB0BFF7D7A708A26C748" box="[198,283,1287,1314]" captionStart="Figs 17 – 21" captionStartId="15.[189,241,1929,1955]" captionTargetBox="[217,1366,246,1881]" captionTargetId="figure@15.[217,1366,246,1881]" captionTargetPageId="15" captionText="Figs 17 – 21. Katissa yaya sp. nov. 17 .  Ƌ,   palp,   ventral  view.   18. Ƌ,   palp,   retrolateral  view. 19 .  ♀,   abdomen, dorsal view. 20 .  ♀,   epigynum,  ventral  view.   21 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars:   17 – 18, 20 – 21 = 0.1 mm; 19 = 1 mm." httpUri="https://zenodo.org/record/1038244/files/figure.png" pageId="14" pageNumber="15">Fig. 18</figureCitation>
), remotely positioned in the latter species (
<figureCitation id="E38B429A0710FB0BFCB17A70885DC748" box="[778,864,1287,1314]" captionStart="Figs 22 – 26" captionStartId="18.[189,241,1930,1956]" captionTargetBox="[251,1339,265,1874]" captionTargetId="figure@18.[231,1359,248,1888]" captionTargetPageId="18" captionText="Figs 22 – 26. Katissa guayasamini sp. nov. 22 .  Ƌ,   palp,  ventral  view.   23. Ƌ,   palp,  retrolateral  view. 24. Ƌ,   habitus, dorsal view. 25. ♀,   epigynum,  ventral  view.   26 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars:   22 – 23, 25 – 26 = 0.1 mm; 24 = 1 mm." httpUri="https://zenodo.org/record/1038246/files/figure.png" pageId="14" pageNumber="15">Fig. 23</figureCitation>
). Females are distinguished by their knob-like epigynal Fap (Fig. 20).
</paragraph>
</subSubSection>
<subSubSection id="33AA0D940710FB0BFF067A1B8FC4C7DF" box="[189,1273,1388,1461]" pageId="14" pageNumber="15" type="etymology">
<paragraph id="7B0F5E1F0710FB0BFF067A1B8A79C7EC" blockId="14.[189,1273,1388,1461]" box="[189,324,1388,1414]" pageId="14" pageNumber="15">
<heading id="2047E9730710FB0BFF067A1B8A79C7EC" bold="true" box="[189,324,1388,1414]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="6">
<emphasis id="49C4820D0710FB0BFF067A1B8A79C7EC" bold="true" box="[189,324,1388,1414]" pageId="14" pageNumber="15">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F0710FB0BFF067AED8FC4C7DF" blockId="14.[189,1273,1388,1461]" box="[189,1273,1434,1461]" pageId="14" pageNumber="15">The speciFc name is a noun in apposition taken from the Kichwa language meaning ‘large’.</paragraph>
</subSubSection>
<subSubSection id="33AA0D940710FB0BFF067AAB8A61C44F" pageId="14" pageNumber="15" type="description">
<paragraph id="7B0F5E1F0710FB0BFF067AAB8A51C79C" blockId="14.[189,364,1500,1526]" box="[189,364,1500,1526]" pageId="14" pageNumber="15">
<heading id="2047E9730710FB0BFF067AAB8A51C79C" bold="true" box="[189,364,1500,1526]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="2">
<emphasis id="49C4820D0710FB0BFF067AAB8A51C79C" bold="true" box="[189,364,1500,1526]" pageId="14" pageNumber="15">
<typeStatus id="A40BE0BD0710FB0BFF067AAB8BC7C79C" box="[189,250,1500,1526]" pageId="14" pageNumber="15">Type</typeStatus>
material
</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F0710FB0BFF51797C8A61C44F" blockId="14.[189,1406,1547,1643]" box="[234,348,1547,1573]" pageId="14" pageNumber="15">
<heading id="2047E9730710FB0BFF51797C8A61C44F" bold="true" box="[234,348,1547,1573]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="6">
<emphasis id="49C4820D0710FB0BFF51797C8A61C44F" bold="true" box="[234,348,1547,1573]" pageId="14" pageNumber="15">
<typeStatus id="A40BE0BD0710FB0BFF51797C8A61C44F" box="[234,348,1547,1573]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="33AA0D940710FB0BFF0679598867C401" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="7B0F5E1F0710FB0BFF0679598867C401" blockId="14.[189,1406,1547,1643]" pageId="14" pageNumber="15">
<materialsCitation id="CBD854420710FB0BFF067959886BC401" ID-GBIF-Occurrence="1663864619" collectingDate="2014-09-04" collectingDateMax="2014-09-07" collectingDateMin="2014-09-04" collectingMethod=" hand  collected  in moss" collectionCode="QCAZ" collectorName="E. Tapia &amp; C. Tapia &amp; N. Duperre" country="Ecuador" elevation="2225" location=" Otonga  Biological  Reserve" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince=" Cotopaxi  Province" typeStatus="holotype">
<collectingCountry id="03A71E8F0710FB0BFF0679598A6DC422" box="[189,336,1582,1608]" name="Ecuador" pageId="14" pageNumber="15" value="Ecuador">ECUADOR</collectingCountry>
:
<specimenCount id="6DB695960710FB0BFEE179598A4CC422" box="[346,369,1582,1608]" pageId="14" pageNumber="15">Ƌ</specimenCount>
,
<collectingRegion id="B97490FD0710FB0BFECC79598966C422" box="[375,603,1582,1608]" pageId="14" pageNumber="15">Cotopaxi Province</collectingRegion>
,
<location id="7E6F08C40710FB0BFDDB79598898C422" LSID="urn:lsid:plazi:treatment:F319EF090710FB14FD257B7C89F9C0DA:7E6F08C40710FB0BFDDB79598898C422" box="[608,933,1582,1608]" country="Ecuador" name=" Otonga  Biological  Reserve" pageId="14" pageNumber="15" stateProvince=" Cotopaxi  Province">Otonga Biological Reserve</location>
,
<elevation id="F09DB92C0710FB0BFC0A79598F36C422" box="[945,1035,1582,1609]" pageId="14" pageNumber="15">2225 m</elevation>
,
<collectingDate id="1F4A81370710FB0BFBAC79598FFFC422" box="[1047,1218,1582,1608]" pageId="14" pageNumber="15">4–7 Sep. 2014</collectingDate>
,
<collectingMethod id="A2F126080710FB0BFB7D79598A27C401" pageId="14" pageNumber="15">hand collected in moss</collectingMethod>
,
<collectorName id="D6453BC90710FB0BFE9D79268AB7C401" box="[294,394,1617,1643]" pageId="14" pageNumber="15">E. Tapia</collectorName>
,
<collectorName id="D6453BC90710FB0BFE2D79268AC5C401" box="[406,504,1617,1643]" pageId="14" pageNumber="15">C. Tapia</collectorName>
and
<collectorName id="D6453BC90710FB0BFD8A79268988C401" box="[561,693,1617,1643]" pageId="14" pageNumber="15">N. Dupérré</collectorName>
leg. (
<collectionCode id="1DA1C6DA0710FB0BFD4C79268873C401" box="[759,846,1617,1643]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" pageId="14" pageNumber="15">QCAZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="33AA0D940710FB0BFF0679E58AC4C4C6" box="[189,505,1682,1708]" pageId="14" pageNumber="15" type="description">
<paragraph id="7B0F5E1F0710FB0BFF0679E58AC4C4C6" blockId="14.[189,505,1682,1708]" box="[189,505,1682,1708]" pageId="14" pageNumber="15">
<heading id="2047E9730710FB0BFF0679E58AC4C4C6" bold="true" box="[189,505,1682,1708]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="2">
<emphasis id="49C4820D0710FB0BFF0679E58AC4C4C6" bold="true" box="[189,505,1682,1708]" pageId="14" pageNumber="15">Other material examined</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="33AA0D940710FB0BFF0679B68AECC502" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="7B0F5E1F0710FB0BFF0679B68AECC502" blockId="14.[189,1406,1729,1896]" pageId="14" pageNumber="15">
<materialsCitation id="CBD854420710FB0BFF0679B68F7AC494" ID-GBIF-Occurrence="1663864613" collectingDate="2014-05-24" collectingDateMax="2014-05-30" collectingDateMin="2014-05-24" collectingMethod=" sifting  moss" collectionCode="QCAZ" collectorName="E. Tapia &amp; C. Tapia &amp; N. Duperre" country="Ecuador" elevation="1717" latitude="-0.41941" location=" Otonga  Biological  Reserve" longitude="-78.99607" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince=" Cotopaxi  Province">
<collectingCountry id="03A71E8F0710FB0BFF0679B68A6DC4B1" box="[189,336,1729,1755]" name="Ecuador" pageId="14" pageNumber="15" value="Ecuador">ECUADOR</collectingCountry>
:
<collectingRegion id="B97490FD0710FB0BFEE879B68909C4B1" box="[339,564,1729,1755]" pageId="14" pageNumber="15">Cotopaxi Province</collectingRegion>
,
<location id="7E6F08C40710FB0BFD8379B68846C4B1" LSID="urn:lsid:plazi:treatment:F319EF090710FB14FD257B7C89F9C0DA:7E6F08C40710FB0BFD8379B68846C4B1" box="[568,891,1729,1755]" country="Ecuador" name=" Otonga  Biological  Reserve" pageId="14" pageNumber="15" stateProvince=" Cotopaxi  Province">Otonga Biological Reserve</location>
: 1
<specimenCount id="6DB695960710FB0BFC2379B68893C4B1" box="[920,942,1729,1755]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
,
<collectingMethod id="A2F126080710FB0BFC0E79B68F74C4B1" box="[949,1097,1729,1755]" pageId="14" pageNumber="15">sifting moss</collectingMethod>
,
<geoCoordinate id="1E8438D80710FB0BFBE879B68FD6C4B1" box="[1107,1259,1729,1755]" pageId="14" pageNumber="15">-00.41941º S</geoCoordinate>
,
<geoCoordinate id="1E8438D80710FB0BFB4D79B68BE4C495" pageId="14" pageNumber="15">-78.99607º W</geoCoordinate>
,
<quantity id="BC48F3FA0710FB0BFF5F79938A03C495" box="[228,318,1764,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.7169999999999999" pageId="14" pageNumber="15" unit="m" value="1717.0">
<elevation id="F09DB92C0710FB0BFF5F79938A03C495" box="[228,318,1764,1791]" metricMagnitude="3" metricUnit="m" metricValue="1.7169999999999999" pageId="14" pageNumber="15" unit="m" value="1717.0">1717 m</elevation>
</quantity>
,
<date id="0F0E78DF0710FB0BFEF179938929C494" box="[330,532,1764,1791]" pageId="14" pageNumber="15" value="2014-05-24" valueMax="2014-05-30" valueMin="2014-05-24">
<collectingDate id="1F4A81370710FB0BFEF179938929C494" box="[330,532,1764,1791]" pageId="14" pageNumber="15" value="2014-05-24" valueMax="2014-05-30" valueMin="2014-05-24">24–30 May 2014</collectingDate>
</date>
,
<collectorName id="D6453BC90710FB0BFDA5799289BFC494" box="[542,642,1764,1791]" pageId="14" pageNumber="15">E. Tapia</collectorName>
,
<collectorName id="D6453BC90710FB0BFD37799389D3C494" box="[652,750,1764,1790]" pageId="14" pageNumber="15">C. Tapia</collectorName>
and
<collectorName id="D6453BC90710FB0BFC9E79928895C494" box="[805,936,1764,1791]" pageId="14" pageNumber="15">N. Dupérré</collectorName>
leg. (
<collectionCode id="1DA1C6DA0710FB0BFC5279938F02C494" box="[1001,1087,1764,1790]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" pageId="14" pageNumber="15">QCAZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="CBD854420710FB0BFBE879938F35C54B" ID-GBIF-Occurrence="1663864649" collectingDate="2014-09-04" collectingDateMax="2014-09-07" collectingDateMin="2014-09-04" collectingMethod=" beating  trees" collectionCode="DTC" collectorName="E. Tapia &amp; C. Tapia &amp; N. Duperre" country="Ecuador" elevation="1700" pageId="14" pageNumber="15" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince=" cotopaxi province">
<specimenCount id="6DB695960710FB0BFBE879938F43C494" box="[1107,1150,1762,1790]" pageId="14" pageNumber="15">1 Ƌ</specimenCount>
,
<specimenCount id="6DB695960710FB0BFB3779938FF3C495" box="[1164,1230,1764,1791]" pageId="14" pageNumber="15">2 ♀♀</specimenCount>
,
<collectingMethod id="A2F126080710FB0BFB6F79938E4FC495" box="[1236,1394,1764,1791]" pageId="14" pageNumber="15">beating trees</collectingMethod>
,
<date id="0F0E78DF0710FB0BFF0678708A51C54B" box="[189,364,1799,1825]" pageId="14" pageNumber="15" value="2014-09-04" valueMax="2014-09-07" valueMin="2014-09-04">
<collectingDate id="1F4A81370710FB0BFF0678708A51C54B" box="[189,364,1799,1825]" pageId="14" pageNumber="15" value="2014-09-04" valueMax="2014-09-07" valueMin="2014-09-04">4–7 Sep. 2014</collectingDate>
</date>
,
<quantity id="BC48F3FA0710FB0BFEC278708AE8C548" box="[377,469,1799,1826]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="15" unit="m" value="1700.0">
<elevation id="F09DB92C0710FB0BFEC278708AE8C548" box="[377,469,1799,1826]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="15" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<collectorName id="D6453BC90710FB0BFE58787F8975C54B" box="[483,584,1799,1826]" pageId="14" pageNumber="15">E. Tapia</collectorName>
,
<collectorName id="D6453BC90710FB0BFDEE78708984C54B" box="[597,697,1799,1825]" pageId="14" pageNumber="15">C. Tapia</collectorName>
and
<collectorName id="D6453BC90710FB0BFD4E787F8847C54B" box="[757,890,1799,1826]" pageId="14" pageNumber="15">N. Dupérré</collectorName>
leg. (
<collectionCode id="1DA1C6DA0710FB0BFC0478708F3DC54B" box="[959,1024,1799,1825]" collectionName="Dmitrijs Telnovs" pageId="14" pageNumber="15">DTC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="CBD854420710FB0BFBAD787089EAC52F" ID-GBIF-Occurrence="1663864628" collectingMethod=" sifting  moss" collectionCode="AMNH" collectorName="E.   Tapia &amp; C.  Tapia  &amp; N.  Duperre " country="Ecuador" pageId="14" pageNumber="15" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince=" cotopaxi province">
<specimenCount id="6DB695960710FB0BFBAD78708F64C54B" box="[1046,1113,1797,1825]" pageId="14" pageNumber="15">2 ƋƋ</specimenCount>
,
<specimenCount id="6DB695960710FB0BFBD178708F93C548" box="[1130,1198,1799,1826]" pageId="14" pageNumber="15">4 ♀♀</specimenCount>
,
<collectingMethod id="A2F126080710FB0BFB0F78708E6DC548" box="[1204,1360,1799,1826]" pageId="14" pageNumber="15">sifting moss</collectingMethod>
,
<collectorName id="D6453BC90710FB0BFAE6787F8A3CC52E" pageId="14" pageNumber="15">E. Tapia</collectorName>
,
<collectorName id="D6453BC90710FB0BFEB7785C8A4BC52E" box="[268,374,1834,1861]" pageId="14" pageNumber="15">C. Tapia</collectorName>
and
<collectorName id="D6453BC90710FB0BFE13785C8909C52F" box="[424,564,1835,1861]" pageId="14" pageNumber="15">N. Dupérré</collectorName>
leg. (
<collectionCode id="1DA1C6DA0710FB0BFDD5785C89F3C52F" box="[622,718,1835,1861]" collectionName="USA, New York, New York, American Museum of Natural History" pageId="14" pageNumber="15">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="CBD854420710FB0BFD60785C8AF0C502" ID-GBIF-Occurrence="1663864626" collectingDate="2014-09-04" collectingDateMax="2014-09-07" collectingDateMin="2014-09-04" collectingMethod=" sifting  moss" collectionCode="DTC" collectorName="E.   Tapia &amp; C.  Tapia  &amp; N. Duperre" country="Ecuador" elevation="1700" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince=" cotopaxi province">
<specimenCount id="6DB695960710FB0BFD58785C8832C52F" box="[739,783,1835,1861]" pageId="14" pageNumber="15">1 ♀</specimenCount>
,
<collectingMethod id="A2F126080710FB0BFCAD785D8892C52F" box="[790,943,1834,1861]" pageId="14" pageNumber="15">sifting moss</collectingMethod>
,
<collectingDate id="1F4A81370710FB0BFC00785C8F5AC52F" box="[955,1127,1835,1861]" pageId="14" pageNumber="15">4–7 Sep. 2014</collectingDate>
,
<collectorName id="D6453BC90710FB0BFBC8785C8FEAC52E" box="[1139,1239,1834,1861]" pageId="14" pageNumber="15">E. Tapia</collectorName>
,
<collectorName id="D6453BC90710FB0BFB59785C8E71C52E" box="[1250,1356,1834,1861]" pageId="14" pageNumber="15">C. Tapia</collectorName>
and
<collectorName id="D6453BC90710FB0BFF0678398A7CC502" box="[189,321,1870,1896]" pageId="14" pageNumber="15">N. Dupérré</collectorName>
leg. (
<collectionCode id="1DA1C6DA0710FB0BFE3878398AF9C502" box="[387,452,1870,1896]" collectionName="Dmitrijs Telnovs" pageId="14" pageNumber="15">DTC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="2FCF0E970711FB0AFF0678FE8952C580" httpUri="https://zenodo.org/record/1038244/files/figure.png" pageId="15" pageNumber="16" targetBox="[217,1366,246,1881]" targetPageId="15">
<paragraph id="7B0F5E1F0711FB0AFF0678FE8952C580" blockId="15.[189,1405,1929,2026]" pageId="15" pageNumber="16">
<emphasis id="49C4820D0711FB0AFF0678FE8A70C5C9" bold="true" box="[189,333,1929,1955]" pageId="15" pageNumber="16">Figs 17–21.</emphasis>
<taxonomicName id="BCB0259C0711FB0AFEE178FD8AC9C5C9" box="[346,500,1930,1955]" class="Arachnida" family="Anyphaenidae" genus="Katissa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="yaya" status="sp. nov.">
<emphasis id="49C4820D0711FB0AFEE178FD8AC9C5C9" box="[346,500,1930,1955]" italics="true" pageId="15" pageNumber="16">Katissa yaya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52F73F760711FB0AFDBA78FD895DC5CE" box="[513,608,1930,1956]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="49C4820D0711FB0AFDD078FE89B5C5C9" bold="true" box="[619,648,1929,1955]" pageId="15" pageNumber="16">17</emphasis>
. Ƌ, palp, ventral view.
<emphasis id="49C4820D0711FB0AFC0978FE88F2C5C9" bold="true" box="[946,975,1929,1955]" pageId="15" pageNumber="16">18</emphasis>
. Ƌ, palp, retrolateral view.
<emphasis id="49C4820D0711FB0AFA9278FE8E7BC5C9" bold="true" box="[1321,1350,1929,1955]" pageId="15" pageNumber="16">19</emphasis>
. ♀, abdomen, dorsal view.
<emphasis id="49C4820D0711FB0AFE7078DB8AD5C5AC" bold="true" box="[459,488,1964,1990]" pageId="15" pageNumber="16">20</emphasis>
. ♀, epigynum, ventral view.
<emphasis id="49C4820D0711FB0AFC8578DB8866C5AC" bold="true" box="[830,859,1964,1990]" pageId="15" pageNumber="16">21</emphasis>
. ♀, internal genitalia, dorsal view. Scale bars: 17–18, 20–21 = 0.1 mm; 19 = 1 mm.
</paragraph>
</caption>
<subSubSection id="33AA0D94070EFB14FF067E7C889CC02B" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="7B0F5E1F070EFB15FF067E7C8A73C34F" blockId="16.[189,334,267,291]" box="[189,334,267,293]" pageId="16" pageNumber="17">
<emphasis id="49C4820D070EFB15FF067E7C8A73C34F" bold="true" box="[189,334,267,293]" pageId="16" pageNumber="17">Description</emphasis>
</paragraph>
<paragraph id="7B0F5E1F070EFB15FF517E4D8A96C33E" blockId="16.[189,1074,313,375]" box="[234,427,313,340]" pageId="16" pageNumber="17">
<emphasis id="49C4820D070EFB15FF517E4D8A17C33E" bold="true" box="[234,298,314,340]" pageId="16" pageNumber="17">Male</emphasis>
(
<typeStatus id="A40BE0BD070EFB15FE827E4E8A9EC33E" box="[313,419,313,340]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067E2A8F0FC31D" blockId="16.[189,1074,313,375]" box="[189,1074,348,375]" pageId="16" pageNumber="17">MEASUREMENTS. Total length: 7.1; carapace length: 3.2; carapace width: 2.5.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067EE98F1EC394" blockId="16.[189,1399,414,511]" pageId="16" pageNumber="17">CEPHALOTHORAX. Pars cephalica light brown, with black mark near cephalic groove; pars thoracica yellowish, with a few dark marks dorsally along radiating lines; margin dark brown. Sternum light brown with two lateral dark brown bands, endites and labium dark brown.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067D518904C009" blockId="16.[189,1399,550,612]" pageId="16" pageNumber="17">CHELICERAE. Light brown basally, dark brown apically, excavated with antero-prolateral keel; promargin with 5, retromargin with 8 teeth.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067DFC89E8C158" blockId="16.[189,1399,650,818]" pageId="16" pageNumber="17">LEGS. Femora I–IV light yellow with brown mark at base of macrosetae. Tibiae light yellow with dark bands basally and apically. Metatarsi and tarsi dark brown. Claws unipectinate, I–II with 6, III with 5, IV with 6 teeth. Total length: I: 16.0; II: 15.1; III: 10.9; IV: 14.5; leg formula 1423; leg articles length (femur/patella/tibia/metatarsus/tarsus): leg I 4.5/1.2/4.6/3.9/1.8; leg II 4.1/1.2/4.3/3.9/1.6; leg III 3.0/1.1/2.5/3.2/1.1; leg IV 3.8/1.2/3.3/4.7/1.5.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067C2E8F7EC1B6" blockId="16.[189,1399,857,989]" pageId="16" pageNumber="17">
LEGS SPINATION. Femur I p1-1-1, r1-1-1; tibia I
<date id="0F0E78DF070EFB15FD607C2E8829C119" box="[731,788,857,883]" pageId="16" pageNumber="17">v2-2</date>
-2, p1-1-1, r1-1-0; metatarsus I
<date id="0F0E78DF070EFB15FB397C2E8F86C119" box="[1154,1211,857,883]" pageId="16" pageNumber="17">v2-2</date>
-0, p1-1-0, r1-1- 0. Femur II p1-1-1, r1-1-1; tibia II
<date id="0F0E78DF070EFB15FDF77C0B89B8C1FC" box="[588,645,892,918]" pageId="16" pageNumber="17">v2-2</date>
-2, p0-1-1, r1-1-0; metatarsus II
<date id="0F0E78DF070EFB15FC437C0B8F0CC1FC" box="[1016,1073,892,918]" pageId="16" pageNumber="17">v2-2</date>
-0, p0-1-0, r1-0-0. Femur III p1-1-1-, r1-1-1; tibia III d1-1-0,
<date id="0F0E78DF070EFB15FD8C7CE88952C1D3" box="[567,623,927,953]" pageId="16" pageNumber="17">v2-2</date>
-2, p1-1-0, r1-1-0; metatarsus III d0-1-0,
<date id="0F0E78DF070EFB15FBF77CE88FB8C1D3" box="[1100,1157,927,953]" pageId="16" pageNumber="17">v2-2</date>
-2. Femur IV p0-1-1, r0-0-1; tibia IV d1-1-0,
<date id="0F0E78DF070EFB15FE6F7CB58931C1B6" box="[468,524,962,988]" pageId="16" pageNumber="17">v2-2</date>
-2, p1-1-0, r1-1-0; metatarsus IV d0-1-0,
<date id="0F0E78DF070EFB15FC557CB58F1AC1B6" box="[1006,1063,962,988]" pageId="16" pageNumber="17">v2-2</date>
-2.
</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067B7389F8C62B" blockId="16.[189,1398,1028,1090]" pageId="16" pageNumber="17">ABDOMEN. Elongated oval. Light gray with whitish pale median bands with small branches laterally, with a few dark spots along midline and laterally.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067B1E8910C765" blockId="16.[189,1399,1128,1296]" pageId="16" pageNumber="17">
GENITALIA. Palpal tibia shorter than cymbium; elongated, serrated palpal retrolateral tibial apophysis with closely positioned basal spur (
<figureCitation id="E38B429A070EFB15FDDB7BFB8985C6CC" box="[608,696,1164,1190]" captionStart="Figs 17 – 21" captionStartId="15.[189,241,1929,1955]" captionTargetBox="[217,1366,246,1881]" captionTargetId="figure@15.[217,1366,246,1881]" captionTargetPageId="15" captionText="Figs 17 – 21. Katissa yaya sp. nov. 17 .  Ƌ,   palp,   ventral  view.   18. Ƌ,   palp,   retrolateral  view. 19 .  ♀,   abdomen, dorsal view. 20 .  ♀,   epigynum,  ventral  view.   21 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars:   17 – 18, 20 – 21 = 0.1 mm; 19 = 1 mm." httpUri="https://zenodo.org/record/1038244/files/figure.png" pageId="16" pageNumber="17">Fig. 18</figureCitation>
). Subtegulum pointed apically, with keel; tegulum rounded basally; ventral tegular process wide, rounded apically reaching median apophysis tip; median apophysis short, hook-shaped; embolic base protruding prolaterally; embolus ribbon-like, long with proximal part black, distal part pale (
<figureCitation id="E38B429A070EFB15FE737B828922C765" box="[456,543,1269,1295]" captionStart="Figs 17 – 21" captionStartId="15.[189,241,1929,1955]" captionTargetBox="[217,1366,246,1881]" captionTargetId="figure@15.[217,1366,246,1881]" captionTargetPageId="15" captionText="Figs 17 – 21. Katissa yaya sp. nov. 17 .  Ƌ,   palp,   ventral  view.   18. Ƌ,   palp,   retrolateral  view. 19 .  ♀,   abdomen, dorsal view. 20 .  ♀,   epigynum,  ventral  view.   21 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars:   17 – 18, 20 – 21 = 0.1 mm; 19 = 1 mm." httpUri="https://zenodo.org/record/1038244/files/figure.png" pageId="16" pageNumber="17">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="7B0F5E1F070EFB15FF517A408A78C73B" blockId="16.[189,1075,1335,1396]" box="[234,325,1335,1361]" pageId="16" pageNumber="17">
<heading id="2047E973070EFB15FF517A408A78C73B" bold="true" box="[234,325,1335,1361]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="6">
<emphasis id="49C4820D070EFB15FF517A408A78C73B" bold="true" box="[234,325,1335,1361]" pageId="16" pageNumber="17">Female</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067A2D8F0EC71E" blockId="16.[189,1075,1335,1396]" box="[189,1075,1370,1396]" pageId="16" pageNumber="17">MEASUREMENTS. Total length: 7.7; carapace length: 3.4; carapace width: 2.4.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067AEC8936C7DF" blockId="16.[189,523,1435,1462]" box="[189,523,1435,1462]" pageId="16" pageNumber="17">CEPHALOTHORAX. As in male.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF067AAA8A29C470" blockId="16.[189,1398,1500,1562]" pageId="16" pageNumber="17">CHELICERAE. Brown, not excavated without antero-prolateral keel; promargin with 5, retromargin with 7 teeth.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF0679358AD3C561" blockId="16.[189,1399,1601,1804]" pageId="16" pageNumber="17">LEGS. Femora I–II light yellow with brown mark at base of macrosetae. Femora III–IV light yellow with dark bands medially and apically. Tibiae light yellow with dark bands basally. Metatarsi I–II brown, metatarsi III–IV light yellow speckled with brown. Tarsi brown. Claws as in male. Palpal claws with 5 teeth. Total length: I: 12.2; II: 11.7; III: 9.3; IV: 12.3; leg formula 4123; leg articles length (femur/patella/ tibia/metatarsus/tarsus): leg I 3.5/1.2/3.2/2.8/1.5; leg II 3.2/1.2/3.2/2.7/1.4; leg III 2.5/1.1/2.1/2.7/0.9; leg IV 3.1/1.2/2.8/3.9/1.4.</paragraph>
<paragraph id="7B0F5E1F070EFB15FF0678448811C5DD" blockId="16.[189,1399,1843,1975]" pageId="16" pageNumber="17">
LEGS SPINATION. Femur I p1-1-1, r0-1-0; tibia I
<date id="0F0E78DF070EFB15FD4E78448810C527" box="[757,813,1843,1869]" pageId="16" pageNumber="17">v2-2</date>
-0, p0-1-1, r1-1-0; metatarsus I
<date id="0F0E78DF070EFB15FB1778448FD8C527" box="[1196,1253,1843,1869]" pageId="16" pageNumber="17">v2-2</date>
-0. Femur II p1-1-1, r0-1-0; tibia II
<date id="0F0E78DF070EFB15FE7678218938C51A" box="[461,517,1878,1904]" pageId="16" pageNumber="17">v2-2</date>
-0, p0-1-1, r0-1-0; metatarsus II
<date id="0F0E78DF070EFB15FC3878218899C51A" box="[899,932,1878,1904]" pageId="16" pageNumber="17">v2</date>
-0-0. Femur III p0-0-1-, r0-1-1; tibia III d1-1-0,
<date id="0F0E78DF070EFB15FEA1780E8A6FC5F9" box="[282,338,1913,1939]" pageId="16" pageNumber="17">v2-2</date>
-2, p1-1-0, r1-1-0; metatarsus III d0-1-0,
<date id="0F0E78DF070EFB15FC88780E8851C5F9" box="[819,876,1913,1939]" pageId="16" pageNumber="17">v2-2</date>
-2. Femur IV p0-0-1, r0-1-1; tibia IV d1-1-0,
<date id="0F0E78DF070EFB15FF0678EA8BCBC5DD" box="[189,246,1949,1975]" pageId="16" pageNumber="17">v2-2</date>
-2, p1-1-0, r1-1-0; metatarsus IV d0-0-1,
<date id="0F0E78DF070EFB15FD6C78EA882DC5DD" box="[727,784,1949,1975]" pageId="16" pageNumber="17">v2-2</date>
-2.
</paragraph>
<paragraph id="7B0F5E1F070FFB14FF067E7B898CC323" blockId="17.[189,1398,267,329]" pageId="17" pageNumber="18">
ABDOMEN. Oval. Light gray with whitish pale median band with small branches laterally, with a few dark spots along midline and laterally (
<figureCitation id="E38B429A070FFB14FDF77E59899EC323" box="[588,675,302,329]" captionStart="Figs 17 – 21" captionStartId="15.[189,241,1929,1955]" captionTargetBox="[217,1366,246,1881]" captionTargetId="figure@15.[217,1366,246,1881]" captionTargetPageId="15" captionText="Figs 17 – 21. Katissa yaya sp. nov. 17 .  Ƌ,   palp,   ventral  view.   18. Ƌ,   palp,   retrolateral  view. 19 .  ♀,   abdomen, dorsal view. 20 .  ♀,   epigynum,  ventral  view.   21 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars:   17 – 18, 20 – 21 = 0.1 mm; 19 = 1 mm." httpUri="https://zenodo.org/record/1038244/files/figure.png" pageId="17" pageNumber="18">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="7B0F5E1F070FFB14FF067E078F4CC3BA" blockId="17.[189,1405,368,465]" pageId="17" pageNumber="18">
GENITALIA. Epigynum with knob-like epigynal Fap; lateral epigynal grooves strongly curved (Fig. 20). Internal genitalia with copulatory ducts elongated and thin; seminal receptacles positioned at beginning of copulatory ducts course; spermathecae oval; fertilization ducts short (
<figureCitation id="E38B429A070FFB14FBB77EC18F59C3BA" box="[1036,1124,438,464]" captionStart="Figs 17 – 21" captionStartId="15.[189,241,1929,1955]" captionTargetBox="[217,1366,246,1881]" captionTargetId="figure@15.[217,1366,246,1881]" captionTargetPageId="15" captionText="Figs 17 – 21. Katissa yaya sp. nov. 17 .  Ƌ,   palp,   ventral  view.   18. Ƌ,   palp,   retrolateral  view. 19 .  ♀,   abdomen, dorsal view. 20 .  ♀,   epigynum,  ventral  view.   21 .  ♀,  internal  genitalia,  dorsal  view.  Scale  bars:   17 – 18, 20 – 21 = 0.1 mm; 19 = 1 mm." httpUri="https://zenodo.org/record/1038244/files/figure.png" pageId="17" pageNumber="18">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="7B0F5E1F070FFB14FF067E8F8A43C078" blockId="17.[189,929,504,577]" box="[189,382,504,530]" pageId="17" pageNumber="18">
<heading id="2047E973070FFB14FF067E8F8A43C078" bold="true" box="[189,382,504,530]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="6">
<emphasis id="49C4820D070FFB14FF067E8F8A43C078" bold="true" box="[189,382,504,530]" pageId="17" pageNumber="18">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070FFB14FF067D51889CC02B" blockId="17.[189,929,504,577]" box="[189,929,550,577]" pageId="17" pageNumber="18">Specimens where collected in moss from trees or beating trees.</paragraph>
</subSubSection>
<subSubSection id="33AA0D94070FFB14FF067D1F89F9C0DA" box="[189,708,616,689]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="7B0F5E1F070FFB14FF067D1F8A6BC0E8" blockId="17.[189,342,616,642]" box="[189,342,616,642]" pageId="17" pageNumber="18">
<heading id="2047E973070FFB14FF067D1F8A6BC0E8" bold="true" box="[189,342,616,642]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="2">
<emphasis id="49C4820D070FFB14FF067D1F8A6BC0E8" bold="true" box="[189,342,616,642]" pageId="17" pageNumber="18">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="7B0F5E1F070FFB14FF067DE189F9C0DA" blockId="17.[189,708,662,689]" box="[189,708,662,689]" pageId="17" pageNumber="18">
Ecuador: known only from the
<typeStatus id="A40BE0BD070FFB14FD977DE08962C0DB" box="[556,607,663,689]" pageId="17" pageNumber="18">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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