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<document id="CC6C75149ACB731C1BB18E3CA3689312" ID-CLB-Dataset="32753" ID-DOI="10.11646/zootaxa.4286.2.3" ID-GBIF-Dataset="dfbcd01b-2ea6-4898-af4b-bb8ee425d26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="828403" ID-ZooBank="2B3C978D-C70B-4A31-9F2B-8E082FBBE51B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1500108776027" checkinUser="plazi" docAuthor="Nilsai, Areeruk, Jantarit, Sopark, Satasook, Chutamas &amp; Zhang, Feng" docDate="2017" docId="3E0D8E51FFFCFFC9AAD8FD23FEAE585E" docLanguage="en" docName="zootaxa.4286.2.3.pdf" docOrigin="Zootaxa 4286 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coecobrya cavicta" docType="treatment" docVersion="8" lastPageNumber="200" masterDocId="C234F629FFF7FFC4AA4FFFB1FFB05D75" masterDocTitle="Three new species of Coecobrya (Collembola: Entomobryidae) from caves in the Thai Peninsula" masterLastPageNumber="202" masterPageNumber="187" pageNumber="198" updateTime="1698465156621" updateUser="ExternalLinkService">
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<mods:title id="F45058B1C9569AB371B55C014FDAB659">Three new species of Coecobrya (Collembola: Entomobryidae) from caves in the Thai Peninsula</mods:title>
</mods:titleInfo>
<mods:name id="6077EE7A1CEB0339E9C10A8BB5417B61" type="personal">
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<mods:namePart id="8564FC48F17D0EAB109585D2A73F3563">Nilsai, Areeruk</mods:namePart>
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<mods:namePart id="A0CD280D8B6BC0D2FF447E757D3227FF">Jantarit, Sopark</mods:namePart>
</mods:name>
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<mods:namePart id="81C41B9FC0CE03E3B1FA1BBAF2DC48C0">Satasook, Chutamas</mods:namePart>
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<mods:namePart id="D3F58384BC0746086DA0553026F62C13">Zhang, Feng</mods:namePart>
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<mods:typeOfResource id="D878B6A8385A2A5C811248CED94E152B">text</mods:typeOfResource>
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<mods:title id="A4EBB2AE23EF0E789E614C526BA63D80">Zootaxa</mods:title>
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<mods:date id="3DDDFD53C8FC124A9E6080DD75AAE504">2017</mods:date>
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<mods:identifier id="BE890E23D9F666D3EBC73EA8FDE625BA" type="DOI">10.11646/zootaxa.4286.2.3</mods:identifier>
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<treatment id="3E0D8E51FFFCFFC9AAD8FD23FEAE585E" ID-DOI="http://doi.org/10.5281/zenodo.6045293" ID-GBIF-Taxon="132548722" ID-Zenodo-Dep="6045293" LSID="urn:lsid:plazi:treatment:3E0D8E51FFFCFFC9AAD8FD23FEAE585E" httpUri="http://treatment.plazi.org/id/3E0D8E51FFFCFFC9AAD8FD23FEAE585E" lastPageId="13" lastPageNumber="200" pageId="11" pageNumber="198">
<subSubSection id="FEBE6CCCFFFCFFCFAAD8FD23FE085FBB" pageId="11" pageNumber="198" type="nomenclature">
<paragraph id="B61B3F47FFFCFFCFAAD8FD23FD105FD8" blockId="11.[151,672,658,718]" box="[151,672,658,685]" pageId="11" pageNumber="198">
<heading id="ED53882BFFFCFFCFAAD8FD23FD105FD8" bold="true" box="[151,672,658,685]" fontSize="11" level="1" pageId="11" pageNumber="198" reason="1">
<emphasis id="84D0E355FFFCFFCFAAD8FD23FD105FD8" bold="true" box="[151,672,658,685]" pageId="11" pageNumber="198">
<taxonomicName id="71A444C4FFFCFFCFAAD8FD23FEC35FD9" box="[151,371,658,684]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="11" pageNumber="200" phylum="Arthropoda" rank="species" species="cavicta" status="sp. nov.">
<emphasis id="84D0E355FFFCFFCFAAD8FD23FEC35FD9" bold="true" box="[151,371,658,684]" italics="true" pageId="11" pageNumber="198">Coecobrya cavIcta</emphasis>
</taxonomicName>
Nilsai &amp; Zhang sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="B61B3F47FFFCFFCFAAD8FD04FE085FBB" blockId="11.[151,672,658,718]" box="[151,440,693,718]" pageId="11" pageNumber="198">
<figureCitation id="2E9F23C2FFFCFFCFAAD8FD04FF6D5FBB" box="[151,221,693,718]" captionStart="FIGURES 2 5" captionStartId="3.[151,264,1881,1904]" captionTargetBox="[151,1436,337,1860]" captionTargetId="figure@3.[151,1436,337,1860]" captionTargetPageId="3" captionText="FIGURES 2 5. Habitus: 2, Coecobrya chumphonensis sp. nov.: 3, Coecobrya polychaeta sp. nov.; 4, Coecobrya cf. polychaeta sp. nov.; 5, Coecobrya cavicta sp. nov. Scale bar: 1 mm." httpUri="https://zenodo.org/record/828407/files/figure.png" pageId="11" pageNumber="198">Figs 5</figureCitation>
,
<figureCitation id="2E9F23C2FFFCFFCFAAA5FD04FE825FBB" box="[234,306,693,718]" captionStart-0="FIGURES 43 54" captionStart-1="FIGURES 55 56" captionStartId-0="12.[151,264,1938,1961]" captionStartId-1="13.[151,264,1051,1074]" captionTargetBox-0="[196,1392,197,1888]" captionTargetBox-1="[177,1416,203,1023]" captionTargetId-0="figure@12.[190,1393,193,1894]" captionTargetId-1="figure@13.[151,1436,193,1030]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." captionText-1="FIGURES 55 56. Abdominal chaetotaxy in Coecobrya cavicta sp. nov. 55, Abd. I III; 56, Abd. IV." httpUri-0="https://zenodo.org/record/828419/files/figure.png" httpUri-1="https://zenodo.org/record/828421/files/figure.png" pageId="11" pageNumber="198">4355</figureCitation>
,
<tableCitation id="FB260AFCFFFCFFCFAB71FD04FE085FBB" box="[318,440,693,718]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="7.[129,206,1289,1310]" captionStartId-1="14.[151,239,152,175]" captionText-0="TABLE 2. The number variation of mac in sets of chaetae on thorax among three species. Character states in parentheses are rarely observeđ." captionText-1="TABLE 3. Comparison among C. chumphonensis sp. nov., C. polychaeta sp. nov., C. cavicta sp. nov., and C. annulata. s, smooth; c, ciliate. Character states in parentheses are rarely observed." pageId="11" pageNumber="198">Tables 23</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="FEBE6CCCFFFCFFCFAAD8FD4CFEAD5E2B" pageId="11" pageNumber="198" type="materials_examined">
<paragraph id="B61B3F47FFFCFFCFAAD8FD4CFEAD5E2B" blockId="11.[151,1437,765,1984]" pageId="11" pageNumber="198">
<materialsCitation id="06CC351AFFFCFFCFAAD8FD4CFEAD5E2B" ID-GBIF-Occurrence="1572325055" collectingDate="2016-09-08" collectorName="S. Jantarit &amp; A. Nilsai" country="Thailand" elevation="115" latitude="6.72636" location="Ton Din" longLatPrecision="1" longitude="100.16249" municipality="Khuan Don District" pageId="11" pageNumber="198" specimenCount="2" stateProvince="Satun Province" typeStatus="holotype">
<emphasis id="84D0E355FFFCFFCFAAD8FD4CFF685E63" bold="true" box="[151,216,765,790]" pageId="11" pageNumber="198">
<typeStatus id="691F81E5FFFCFFCFAAD8FD4CFF645E63" box="[151,212,765,790]" pageId="11" pageNumber="198">Type</typeStatus>
.
</emphasis>
<typeStatus id="691F81E5FFFCFFCFAAAAFD4CFEE25E63" box="[229,338,765,790]" pageId="11" pageNumber="198" type="holotype">Holotype</typeStatus>
: female on slide,
<collectingCountry id="CEB37FD7FFFCFFCFA863FD4CFD245E63" box="[556,660,765,790]" name="Thailand" pageId="11" pageNumber="198">Thailand</collectingCountry>
:
<collectingRegion id="7460F1A5FFFCFFCFA8EBFD4FFCE85E63" box="[676,856,765,790]" country="Thailand" name="Satun" pageId="11" pageNumber="198">Satun Province</collectingRegion>
:
<collectingMunicipality id="567FA53DFFFCFFCFA927FD4CFBE65E63" box="[872,1110,765,790]" pageId="11" pageNumber="198">Khuan Don District</collectingMunicipality>
:
<location id="B37B699CFFFCFFCFAE2AFD4FFB775E63" LSID="urn:lsid:plazi:treatment:3E0D8E51FFFCFFC9AAD8FD23FEAE585E:B37B699CFFFCFFCFAE2AFD4FFB775E63" box="[1125,1223,765,790]" country="Thailand" latitude="6.72636" longLatPrecision="1" longitude="100.16249" municipality="Khuan Don District" name="Ton Din" pageId="11" pageNumber="198" stateProvince="Satun Province">Ton Din</location>
cave,
<geoCoordinate id="D3905980FFFCFFCFAF54FD4CFA285E63" box="[1307,1432,765,790]" direction="north" orientation="latitude" pageId="11" pageNumber="198" precision="1" value="6.72636">6.72636°N</geoCoordinate>
,
<geoCoordinate id="D3905980FFFCFFCFAAD8FC90FE9D5E4F" box="[151,301,801,826]" direction="east" orientation="longitude" pageId="11" pageNumber="198" precision="1" value="100.16249">100.16249°E</geoCoordinate>
, altitude
<elevation id="3D89D874FFFCFFCFABDAFC90FE6E5E4C" box="[405,478,801,826]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="198" unit="m" value="115.0">
<quantity id="715C92A2FFFCFFCFABDAFC90FE6E5E4C" box="[405,478,801,826]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="198" unit="m" value="115.0">115 m</quantity>
</elevation>
,
<collectingDate id="D25EE06FFFFCFFCFABA4FC90FDE75E4F" box="[491,599,801,826]" pageId="11" pageNumber="198" value="2016-09-08">
<date id="C21A1987FFFCFFCFABA4FC90FDE75E4F" box="[491,599,801,826]" pageId="11" pageNumber="198" value="2016-09-08">8.ix.2016</date>
</collectingDate>
,
<collectorName id="1B515A91FFFCFFCFA82BFC93FD655E4F" box="[612,725,801,826]" pageId="11" pageNumber="198">S. Jantarit</collectorName>
and
<collectorName id="1B515A91FFFCFFCFA942FC93FCC35E4F" box="[781,883,801,826]" pageId="11" pageNumber="198">A. Nilsai</collectorName>
leg. (collection #THA_SJ_STN08), deposited in PSU-NHM.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FEBE6CCCFFFCFFCFAA88FCD8FB595AE9" pageId="11" pageNumber="198" type="description">
<paragraph id="B61B3F47FFFCFFCFAA88FCD8FC325EF7" blockId="11.[151,1437,765,1984]" box="[199,898,873,898]" pageId="11" pageNumber="198">
<emphasis id="84D0E355FFFCFFCFAA88FCD8FEE95EF7" bold="true" box="[199,345,873,898]" pageId="11" pageNumber="198">Description.</emphasis>
Body
<quantity id="715C92A2FFFCFFCFABECFCDBFDBA5EF4" box="[419,522,874,898]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="11" pageNumber="198" unit="mm" value="1.72">1.72 mm</quantity>
. Body whitish in alcohol (
<figureCitation id="2E9F23C2FFFCFFCFA97FFCD8FCC55EF7" box="[816,885,873,898]" captionStart="FIGURES 2 5" captionStartId="3.[151,264,1881,1904]" captionTargetBox="[151,1436,337,1860]" captionTargetId="figure@3.[151,1436,337,1860]" captionTargetPageId="3" captionText="FIGURES 2 5. Habitus: 2, Coecobrya chumphonensis sp. nov.: 3, Coecobrya polychaeta sp. nov.; 4, Coecobrya cf. polychaeta sp. nov.; 5, Coecobrya cavicta sp. nov. Scale bar: 1 mm." httpUri="https://zenodo.org/record/828407/files/figure.png" pageId="11" pageNumber="198">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="B61B3F47FFFCFFCFAA88FC3FFE075964" blockId="11.[151,1437,765,1984]" pageId="11" pageNumber="198">Antennae 2.67 times as long as cephalic diagonal. Antennal segment ratio as I: II: III: IV = 1: 1.48: 1.52: 2.39. Smooth spiny mic at base of antennae: three dorsal and three ventral on Ant. I, one internal, one external and one ventral on Ant. II. Long smooth straight chaetae present on the ventral side of Ant. I. Distal part of Ant. II and Ant. III organ not clearly seen.</paragraph>
<paragraph id="B61B3F47FFFCFFCFAA88FBAFFB825842" blockId="11.[151,1437,765,1984]" pageId="11" pageNumber="198">
Eyes absent. Prelabral and labral chaetae 4/5, 5, 4, all smooth; three median chaetae of the first and the second rows longer than lateral ones (
<figureCitation id="2E9F23C2FFFCFFCFABB5FBF0FDE2592F" box="[506,594,1089,1114]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 43</figureCitation>
). Clypeal chaetae smooth but their arrangement unclear. Dorsal cephalic chaetotaxy with four antennal (An), three median (M) and eight sutural (S) mac; Gr. II with four mac (
<figureCitation id="2E9F23C2FFFCFFCFAF76FBD4FA20590B" box="[1337,1424,1125,1150]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 44</figureCitation>
). Mandibles with 4+5 (4 left, 5 right) large apical teeth. Subapical chaeta of maxillary outer lobe thicker than apical one; three smooth sublobal hairs on maxillary outer lobe (
<figureCitation id="2E9F23C2FFFCFFCFA90BFB1CFC2D59B3" box="[836,925,1197,1222]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 45</figureCitation>
). Hypostomal chaetae h1 and h2 curved and subequal (
<figureCitation id="2E9F23C2FFFCFFCFAB45FB65FEEF5998" box="[266,351,1236,1261]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 46</figureCitation>
). Lateral process of labial palp as thick as normal chaetae, with tip beyond apex of labial papilla (
<figureCitation id="2E9F23C2FFFCFFCFAAD0FB49FF455865" box="[159,245,1272,1297]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 47</figureCitation>
). Labial chaetae as mrel1l2, all smooth; r, X,
<date id="C21A1987FFFCFFCFA8BEFB48FCBD5860" box="[753,781,1273,1301]" pageId="11" pageNumber="198">X2</date>
,
<date id="C21A1987FFFCFFCFA953FB48FC885860" box="[796,824,1273,1301]" pageId="11" pageNumber="198">X4</date>
, and a chaeta between H2 and H3 smooth and minute. Chaetae along cephalic groove 11, of which 34 smooth and two minute (
<figureCitation id="2E9F23C2FFFCFFCFA99EFAAEFB955842" box="[977,1061,1311,1336]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="B61B3F47FFFCFFCFAA88FAF2FE465B09" blockId="11.[151,1437,765,1984]" pageId="11" pageNumber="198">
Trochanteral organ with 1516 smooth spine-like chaetae; eight in arms and 78 between them (
<figureCitation id="2E9F23C2FFFCFFCFAF78FAF2FA20582E" box="[1335,1424,1347,1372]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 49</figureCitation>
). Partial inner differentiated tibiotarsal chaetae ciliate with ciliations not closely appressed to axis. Tibiotarsi distally with 10 chaetae in a whorl. Unguis with two paired inner teeth, unequal and 0.28 distance from base; unpaired tooth absent. Unguiculus lanceolate with outer edge serrate; the basal tooth clearly seen but the distal teeth extremely minute and often indistinct. Tenent hairs pointed (
<figureCitation id="2E9F23C2FFFCFFCFA971FA62FC235899" box="[830,915,1491,1516]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 50</figureCitation>
). Abd. IV 4.41 times as long as Abd. III along dorsal midline. Ventral tube not clearly seen. Furcula without smooth chaetae. Distal part of manubrium ventrally with 13+13 ciliate chaetae (
<figureCitation id="2E9F23C2FFFCFFCFAB98F9AAFD9D5B41" box="[471,557,1563,1588]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 51</figureCitation>
). Manubrial plaque with two pseudopores and four ciliate chaetae (
<figureCitation id="2E9F23C2FFFCFFCFAF76F9AAFA205B41" box="[1337,1424,1563,1588]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 52</figureCitation>
). Distal smooth part of dens 0.90 times as long as mucro. Mucro falcate with a long basal spine, nearly reaching the tip of the apical tooth (
<figureCitation id="2E9F23C2FFFCFFCFABD9F9D2FE595B09" box="[406,489,1635,1660]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 53</figureCitation>
).
</paragraph>
<paragraph id="B61B3F47FFFCFFCFAA88F936FE2D5A0D" blockId="11.[151,1437,765,1984]" pageId="11" pageNumber="198">
Th. II with three (m1, m2, m
<date id="C21A1987FFFCFFCFA862F936FDF25BD5" box="[557,578,1671,1696]" pageId="11" pageNumber="198">2i</date>
) medio-medial, four (m4, m4p, m
<date id="C21A1987FFFCFFCFA9AEF936FC475BD5" box="[993,1015,1671,1696]" pageId="11" pageNumber="198">4i</date>
, m
<date id="C21A1987FFFCFFCFAE6FF936FB855BD5" box="[1056,1077,1671,1696]" pageId="11" pageNumber="198">4i</date>
2) medio-sublateral and 3031 posterior mac; ms inner to sens; p5 as mac. Th. III with 35 mac; a
<date id="C21A1987FFFCFFCFA935F91AFC215BB1" box="[890,913,1707,1732]" pageId="11" pageNumber="198">4i</date>
2, m
<date id="C21A1987FFFCFFCFA98BF91AFC6A5BB1" box="[964,986,1707,1732]" pageId="11" pageNumber="198">5i</date>
, a
<date id="C21A1987FFFCFFCFA9BDF91AFBB85BB1" box="[1010,1032,1707,1732]" pageId="11" pageNumber="198">6i</date>
and m6ai as mac (
<figureCitation id="2E9F23C2FFFCFFCFAE95F91AFA9D5BB1" box="[1242,1325,1707,1732]" captionStart="FIGURES 43 54" captionStartId="12.[151,264,1938,1961]" captionTargetBox="[196,1392,197,1888]" captionTargetId="figure@12.[190,1393,193,1894]" captionTargetPageId="12" captionText="FIGURES 43 54. Coecobrya cavicta sp. nov. 43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy." httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="11" pageNumber="198">Fig. 54</figureCitation>
,
<tableCitation id="FB260AFCFFFCFFCFAF76F91AFA3F5BB6" box="[1337,1423,1707,1732]" captionStart="TABLE 2" captionStartId="7.[129,206,1289,1310]" captionText="TABLE 2. The number variation of mac in sets of chaetae on thorax among three species. Character states in parentheses are rarely observeđ." pageId="11" pageNumber="198">Table 2</tableCitation>
). Abd. I with seven (a2, a3, m24, m
<date id="C21A1987FFFCFFCFA864F97EFDF25B9D" box="[555,578,1743,1768]" pageId="11" pageNumber="198">2i</date>
, m4p) mac; sens posterior to ms. Abd. II with 4+5 (a2, a3, m3, m3e, m3ep) central and one (m5) lateral mac; a3 as mic on left side. Abd. III with one (m3) central and three (am6, pm6, p6) lateral mac (
<figureCitation id="2E9F23C2FFFCFFCFAB6EF8A6FEC55A45" box="[289,373,1815,1840]" captionStart="FIGURES 55 56" captionStartId="13.[151,264,1051,1074]" captionTargetBox="[177,1416,203,1023]" captionTargetId="figure@13.[151,1436,193,1030]" captionTargetPageId="13" captionText="FIGURES 55 56. Abdominal chaetotaxy in Coecobrya cavicta sp. nov. 55, Abd. I III; 56, Abd. IV." httpUri="https://zenodo.org/record/828421/files/figure.png" pageId="11" pageNumber="198">Fig. 55</figureCitation>
). Abd. IV with 7+9 central (I, M, A5, A5p, A6, B25) and 11 lateral mac (D3, E24, E2p, Ee8, Ee10, F13, F1p); B2 and B3 as mac on left side (
<figureCitation id="2E9F23C2FFFCFFCFA88AF88AFCA95A21" box="[709,793,1851,1876]" captionStart="FIGURES 55 56" captionStartId="13.[151,264,1051,1074]" captionTargetBox="[177,1416,203,1023]" captionTargetId="figure@13.[151,1436,193,1030]" captionTargetPageId="13" captionText="FIGURES 55 56. Abdominal chaetotaxy in Coecobrya cavicta sp. nov. 55, Abd. I III; 56, Abd. IV." httpUri="https://zenodo.org/record/828421/files/figure.png" pageId="11" pageNumber="198">Fig. 56</figureCitation>
). S-chaetae formula: 2+ms, 2/1+ms, 2, 2,?, 3; ps on Abd.
<date id="C21A1987FFFCFFCFAAD8F8D1FF755A02" box="[151,197,1887,1911]" pageId="11" pageNumber="198">IV 1</date>
/3 length of others.
</paragraph>
<paragraph id="B61B3F47FFFCFFCFAA88F832FB595AE9" blockId="11.[151,1437,765,1984]" box="[199,1257,1923,1948]" pageId="11" pageNumber="198">
<emphasis id="84D0E355FFFCFFCFAA88F832FE9B5AE9" bold="true" box="[199,299,1923,1948]" pageId="11" pageNumber="198">Ecology.</emphasis>
This species was found in a dark, humid cave near a stream bank, without bat guano.
</paragraph>
</subSubSection>
<subSubSection id="FEBE6CCCFFFCFFCFAA88F816FC4C5AB5" box="[199,1020,1959,1984]" pageId="11" pageNumber="198" type="etymology">
<paragraph id="B61B3F47FFFCFFCFAA88F816FC4C5AB5" blockId="11.[151,1437,765,1984]" box="[199,1020,1959,1984]" pageId="11" pageNumber="198">
<emphasis id="84D0E355FFFCFFCFAA88F816FEFE5AB5" bold="true" box="[199,334,1959,1984]" pageId="11" pageNumber="198">Etymology.</emphasis>
Named after the Latin word cavi- (&quot;cave&quot;) + victa (&quot;living&quot;).
</paragraph>
</subSubSection>
<caption id="E2DB6FCFFFFBFFC8AAD8F823FBDE5A93" httpUri="https://zenodo.org/record/828419/files/figure.png" pageId="12" pageNumber="199" targetBox="[196,1392,197,1888]" targetPageId="12">
<paragraph id="B61B3F47FFFBFFC8AAD8F823FBDE5A93" blockId="12.[151,1436,1938,2022]" pageId="12" pageNumber="199">
<emphasis id="84D0E355FFFBFFC8AAD8F823FEE85ADD" bold="true" box="[151,344,1938,1961]" pageId="12" pageNumber="199">FIGURES 4354.</emphasis>
<taxonomicName id="71A444C4FFFBFFC8AB2CF822FD965ADC" box="[355,550,1939,1961]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="12" pageNumber="199" phylum="Arthropoda" rank="species" species="cavicta" status="sp. nov.">
<emphasis id="84D0E355FFFBFFC8AB2CF822FD965ADC" box="[355,550,1939,1961]" italics="true" pageId="12" pageNumber="199">Coecobrya cavicta</emphasis>
</taxonomicName>
<emphasis id="84D0E355FFFBFFC8A87EF822FD365ADD" bold="true" box="[561,646,1939,1960]" pageId="12" pageNumber="199">
<taxonomicNameLabel id="9FE35E2EFFFBFFC8A87EF822FD365ADD" box="[561,646,1939,1960]" pageId="12" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
43, labrum; 44, dorsal cephalic chaetotaxy; 45, maxillary outer lobe; 46, hypostomal chaetae; 47, lateral process of labial palp; 48, labial and postlabial chaetae; 49, trochanteral organ; 50, hind claw; 51, distal part of manubrium ventrally; 52, manubrial plaque; 53, mucro; 54, thoracic chaetotaxy.
</paragraph>
</caption>
<caption id="E2DB6FCFFFFAFFC9AAD8FBAAFB235947" box="[151,1171,1051,1074]" httpUri="https://zenodo.org/record/828421/files/figure.png" pageId="13" pageNumber="200" targetBox="[177,1416,203,1023]" targetPageId="13">
<paragraph id="B61B3F47FFFAFFC9AAD8FBAAFB235947" blockId="13.[151,1171,1051,1074]" box="[151,1171,1051,1074]" pageId="13" pageNumber="200">
<emphasis id="84D0E355FFFAFFC9AAD8FBAAFEE45944" bold="true" box="[151,340,1051,1074]" pageId="13" pageNumber="200">FIGURES 5556.</emphasis>
Abdominal chaetotaxy in
<taxonomicName id="71A444C4FFFAFFC9A82FFBADFCAE5947" box="[608,798,1052,1074]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="cavicta" status="sp. nov.">
<emphasis id="84D0E355FFFAFFC9A82FFBADFCAE5947" box="[608,798,1052,1074]" italics="true" pageId="13" pageNumber="200">Coecobrya cavicta</emphasis>
</taxonomicName>
<emphasis id="84D0E355FFFAFFC9A96BFBADFCC55944" bold="true" box="[804,885,1052,1073]" pageId="13" pageNumber="200">
<taxonomicNameLabel id="9FE35E2EFFFAFFC9A96BFBADFCC55944" box="[804,885,1052,1073]" pageId="13" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
55, Abd. IIII; 56, Abd. IV.
</paragraph>
</caption>
<subSubSection id="FEBE6CCCFFFAFFC9AA88FBEFFEAE585E" pageId="13" pageNumber="200" type="discussion">
<paragraph id="B61B3F47FFFAFFC9AA88FBEFFEAE585E" blockId="13.[151,1437,1118,1323]" pageId="13" pageNumber="200">
<emphasis id="84D0E355FFFAFFC9AA88FBEFFE8B5902" bold="true" box="[199,315,1118,1143]" pageId="13" pageNumber="200">Remarks.</emphasis>
<taxonomicName id="71A444C4FFFAFFC9AB0CFBEEFDA75902" box="[323,535,1119,1143]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="species" species="cavicta" status="sp. nov.">
<emphasis id="84D0E355FFFAFFC9AB0CFBEEFDA75902" box="[323,535,1119,1143]" italics="true" pageId="13" pageNumber="200">Coecobrya cavicta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FE35E2EFFFAFFC9A850FBEFFDCA5902" box="[543,634,1118,1143]" pageId="13" pageNumber="200" rank="species">
<emphasis id="84D0E355FFFAFFC9A850FBEFFDCA5902" bold="true" box="[543,634,1118,1143]" pageId="13" pageNumber="200">sp. nov.</emphasis>
</taxonomicNameLabel>
can be separated from other
<taxonomicName id="71A444C4FFFAFFC9A989FBEEFBF15902" box="[966,1089,1119,1143]" class="Entognatha" family="Entomobryidae" genus="Coecobrya" kingdom="Animalia" order="Collembola" pageId="13" pageNumber="200" phylum="Arthropoda" rank="genus">
<emphasis id="84D0E355FFFAFFC9A989FBEEFBF15902" box="[966,1089,1119,1143]" italics="true" pageId="13" pageNumber="200">Coecobrya</emphasis>
</taxonomicName>
species by the combination of abundant posterior mac on dorsal head, labial chaetae r and six smooth and minute postlabial chaetae, two inner teeth on unguis, serrate outer edge on unguiculus, 13+13 chaetae on distal part of manubrium ventrally, four median-lateral mac on Th. II, five or four mac on Abd. II, and nine or seven central and 11 lateral mac on Abd. IV. It differs from closely related species in absence of unpaired tooth on unguis and four medio-sublateral mac on Th. II (
<tableCitation id="FB260AFCFFFAFFC9AAF7FAA3FEBE585E" box="[184,270,1298,1323]" captionStart="TABLE 3" captionStartId="14.[151,239,152,175]" captionText="TABLE 3. Comparison among C. chumphonensis sp. nov., C. polychaeta sp. nov., C. cavicta sp. nov., and C. annulata. s, smooth; c, ciliate. Character states in parentheses are rarely observed." pageId="13" pageNumber="200">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>