152 lines
13 KiB
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152 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.346.6114" ID-GBIF-Dataset="892cdf91-8f73-426f-9143-b21e3d4df75d" ID-PMC="PMC3821061" ID-Pensoft-Pub="1313-2970-346-1" ID-PubMed="24223483" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-346-1" ModsDocOrigin="ZooKeys 346" ModsDocTitle="Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)" checkinTime="1451246756994" checkinUser="pensoft" docAuthor="Perger, Robert & Grossi, Paschoal Coelho" docDate="2013" docId="07A4CA6B5A67BAF0B6C869F89EEFD56F" docLanguage="en" docName="ZooKeys 346: 1-16" docOrigin="ZooKeys 346" docSource="http://dx.doi.org/10.3897/zookeys.346.6114" docTitle="Oryctophileurus Kolbe 1878" docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="562FFFC4FFE7E0630407FF9EFF8CFFC5" masterDocTitle="Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="2" updateTime="1668156700007" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the rhinoceros beetle genus Oryctophileurus Kolbe with description of a new species, the male of O. varicosus Prell, and notes on biogeography (Scarabaeoidea, Dynastinae, Phileurini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perger, Robert</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grossi, Paschoal Coelho</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>346</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>16</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.346.6114</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.346.6114</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-346-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152049038" LSID="urn:lsid:plazi:treatment:07A4CA6B5A67BAF0B6C869F89EEFD56F" httpUri="http://treatment.plazi.org/id/07A4CA6B5A67BAF0B6C869F89EEFD56F" lastPageId="4" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection pageId="1" pageNumber="2" type="nomenclature">
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<paragraph pageId="1" pageNumber="2">
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<taxonomicName LSID="http://species-id.net/wiki/Oryctophileurus" authority="Kolbe, 1878" authorityName="Kolbe" authorityYear="1878" class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Oryctophileurus Kolbe, 1878</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="2" type="type species">
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<paragraph pageId="1" pageNumber="2">Type species.</paragraph>
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<paragraph pageId="1" pageNumber="2">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Phileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phileurus nasicornis" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="nasicornis">Phileurus nasicornis</taxonomicName>
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Burmeister, 1847 (original combination) (Fig. 3).
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</paragraph>
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<paragraph pageId="1" pageNumber="2">
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Species of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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are distinguished from other
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<taxonomicName lsidName="" pageId="1" pageNumber="2" rank="tribe" tribe="Phileurini">Phileurini</taxonomicName>
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by a combination of the following apomorphic characters: a single cephalic horn with lateral carina, pronotal cavity with ocellate punctures and two teeth or tubercles close to the anterior pronotal margin, teeth vertically positioned at about the same level as the outer eye margins.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="taxonomy and discussion">
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<paragraph pageId="2" pageNumber="3">
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<pageBreakToken pageId="2" pageNumber="3" start="start">Taxonomy</pageBreakToken>
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and discussion.
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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The following characters were cited by
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<bibRefCitation author="Endrodi, S" journalOrPublisher="Dr W. Junk, Dordrecht, Netherlands" pageId="13" pageNumber="14" title="The Dynastinae of the World." year="1985">
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<normalizedToken originalValue="Endrödi">Endroedi</normalizedToken>
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(1985)
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</bibRefCitation>
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for
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Dynastidae" genus="Microphileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microphileurus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Microphileurus</taxonomicName>
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Kolbe,
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<taxonomicName class="Insecta" family="Dynastidae" genus="Metaphileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metaphileurus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Metaphileurus</taxonomicName>
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Kolbe,
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<taxonomicName class="Insecta" family="Dynastidae" genus="Trioplus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trioplus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Trioplus</taxonomicName>
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Burmeister, and
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<taxonomicName class="Insecta" family="Dynastidae" genus="Goniophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goniophileurus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Goniophileurus</taxonomicName>
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Kolbe and are here not considered as generic: outer side of mandible tridentate; antenna 10-jointed; elytra with punctate rows; proleg sexually monomorphic, protarsus not thickened in male.
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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are distinguished from the other mentioned genera by large horns, a distinctly developed and posteriorly carinate pronotal cavity and four external teeth on the protibia (
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<bibRefCitation author="Endrodi, S" journalOrPublisher="Dr W. Junk, Dordrecht, Netherlands" pageId="13" pageNumber="14" title="The Dynastinae of the World." year="1985">
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<normalizedToken originalValue="Endrödi">Endroedi</normalizedToken>
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1985
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</bibRefCitation>
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;
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<bibRefCitation author="Ide, S" journalOrPublisher="Universidade de Sao Paulo, Brasil" pageId="13" pageNumber="14" title="Sistematica e evolucao dos generos neotropicais de Phileurini (Coleoptera: Scarabaeidae: Dynastinae)." year="1998">Ide 1998</bibRefCitation>
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). However, in
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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the two cephalic horns lack the lateral carina, the punctures in the pronotal cavity are predominately fused to short, transverse ridges, and the teeth close to the anterior pronotal margin are absent. Except for the horn number, these generic differences between
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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have not been recognized by previous workers.
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</paragraph>
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<paragraph pageId="3" pageNumber="4">
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<pageBreakToken pageId="3" pageNumber="4" start="start">Based</pageBreakToken>
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on the combination of apomorphic characters of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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that lacks in
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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we suggest retaining the genus
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="key to species of oryctophileurus">
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<paragraph pageId="3" pageNumber="4">
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Key to species of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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</paragraph>
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<paragraph lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<table lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<tr pageId="3" pageNumber="4">
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<td colspan="1" pageId="3" pageNumber="4" rowspan="1">Figs 3B, DFigs 3A, C</td>
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<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus nasicornis" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nasicornis">Oryctophileurus nasicornis</taxonomicName>
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</td>
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</tr>
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<tr pageId="3" pageNumber="4">
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<td colspan="1" pageId="3" pageNumber="4" rowspan="1">Figs 4A, C</td>
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<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus varicosus" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="varicosus">Oryctophileurus varicosus</taxonomicName>
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</td>
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</tr>
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<tr pageId="3" pageNumber="4">
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<td colspan="1" pageId="3" pageNumber="4" rowspan="1">Fig. 7AFig. 6CFig. 7C</td>
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<td colspan="1" pageId="3" pageNumber="4" rowspan="1">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus armicollis" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="armicollis">Oryctophileurus armicollis</taxonomicName>
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</td>
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</tr>
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<tr pageId="4" pageNumber="5">
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<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
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<pageBreakToken pageId="4" pageNumber="5" start="start">Fig</pageBreakToken>
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. 7BFig. 8CFig. 7D
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</td>
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<td colspan="1" pageId="4" pageNumber="5" rowspan="1">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus guerrai" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="guerrai">Oryctophileurus guerrai</taxonomicName>
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</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |