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<mods:title id="68A7DB8B38CF5790FAD3B536DCAEAA11">Palaearctic osmiine bees of the genera Hofferia and Stenoheriades (Megachilidae, Osmiini): biology, taxonomy and key to species</mods:title>
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<mods:namePart id="E1E4631D3ECD281D05AF09E7A17A7783">Müller, Andreas</mods:namePart>
<mods:affiliation id="DC8DA6F8A2036FB38BD2C5CB027B26D4">ETH Zurich, Institute of Agricultural Sciences, Biocommunication and Entomology, Schmelzbergstrasse 9 / LFO, 8092 Zurich, Switzerland.</mods:affiliation>
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<mods:namePart id="AE0A2A7CC506B58E814B13E3E8959B3A">Trunz, Vincent</mods:namePart>
<mods:affiliation id="59C3D0A6CFC12B2D277D34AEC9A1B2F2">University of Neuchâtel, Institute of Biology, Laboratory of Evolutive Entomology, Rue Emile-Argand 11, 2000 Neuchâtel, Switzerland.</mods:affiliation>
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<treatment id="5F14F56C5E07FFC145DCF891FBA5C70E" ID-DOI="http://doi.org/10.5281/zenodo.6135418" ID-GBIF-Taxon="119414966" ID-Zenodo-Dep="6135418" LSID="urn:lsid:plazi:treatment:5F14F56C5E07FFC145DCF891FBA5C70E" httpUri="http://treatment.plazi.org/id/5F14F56C5E07FFC145DCF891FBA5C70E" lastPageId="6" lastPageNumber="181" pageId="5" pageNumber="180">
<subSubSection id="9FA717F15E07FFC245DCF891FDEEC30A" box="[151,576,1826,1852]" pageId="5" pageNumber="180" type="nomenclature">
<paragraph id="D702447A5E07FFC245DCF891FDEEC30A" blockId="5.[151,576,1826,1852]" box="[151,576,1826,1852]" pageId="5" pageNumber="180">
<heading id="8C4AF3165E07FFC245DCF891FDEEC30A" bold="true" box="[151,576,1826,1852]" fontSize="11" level="1" pageId="5" pageNumber="180" reason="1">
<emphasis id="E5C998685E07FFC245DCF891FDEEC30A" bold="true" box="[151,576,1826,1852]" pageId="5" pageNumber="180">
Genus
<taxonomicName id="10BD3FF95E07FFC245A5F891FDEEC30A" ID-CoL="7MN8" authority="Tkalcu, 1984" authorityName="Tkalcu" authorityYear="1984" box="[238,576,1826,1852]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="E5C998685E07FFC245A5F891FE36C30A" bold="true" box="[238,408,1826,1852]" italics="true" pageId="5" pageNumber="180">Stenoheriades</emphasis>
<bibRefCitation id="B32C398B5E07FFC244D4F891FDEEC30A" author="Tkalcu" box="[415,576,1826,1852]" pageId="5" pageNumber="180" refString="Tkalcu, B. (1984) Neue palaarktische Arten der Gattungen Pseudoheriades und Archeriades mit Beschreibung von Hofferia gen. n. (Hymenoptera, Apoidea, Megachilidae). Annotationes Zoologicae et Botanicae, 158, 1 - 22." type="journal article" year="1984">Tkalců, 1984</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9FA717F15E07FFC145DCF8DAFBA5C70E" lastPageId="6" lastPageNumber="181" pageId="5" pageNumber="180" type="discussion">
<paragraph id="D702447A5E07FFC245DCF8DAFC0DC3D8" blockId="5.[151,1436,1897,2030]" pageId="5" pageNumber="180">
Eleven species distributed in the Palaearctic and the Afrotropical regions (including
<collectingCountry id="AFAA04EA5E07FFC24102F8DAFB7CC3B4" box="[1097,1234,1897,1922]" name="Madagascar" pageId="5" pageNumber="180">Madagascar</collectingCountry>
) with five and six species, respectively. Six Afrotropical species are still undescribed (
<bibRefCitation id="B32C398B5E07FFC246D3F83EFBFEC390" author="Michener" box="[920,1104,1933,1958]" pageId="5" pageNumber="180" refString="Michener, C. D. (2007) The bees of the world, Second Edition. Johns Hopkins University Press, Baltimore and London, 953 pp." type="book" year="2007">Michener, 2007</bibRefCitation>
). The Palaearctic distribution range of
<taxonomicName id="10BD3FF95E07FFC2444FF802FE0CC3FC" box="[260,418,1969,1994]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="E5C998685E07FFC2444FF802FE0CC3FC" box="[260,418,1969,1994]" italics="true" pageId="5" pageNumber="180">Stenoheriades</emphasis>
</taxonomicName>
encompasses
<collectingCountry id="AFAA04EA5E07FFC24700F801FD1DC3FF" box="[587,691,1970,1993]" name="Morocco" pageId="5" pageNumber="180">Morocco</collectingCountry>
, southern
<collectingCountry id="AFAA04EA5E07FFC24664F802FCC1C3FC" box="[815,879,1969,1994]" name="Spain" pageId="5" pageNumber="180">Spain</collectingCountry>
and Sicily, southeastern Europe to easternmost
<collectingCountry id="AFAA04EA5E07FFC245DCF866FF49C3D8" box="[151,231,2005,2030]" name="Turkey" pageId="5" pageNumber="180">Turkey</collectingCountry>
and southern
<collectingCountry id="AFAA04EA5E07FFC244CFF866FE7AC3D8" box="[388,468,2005,2030]" name="Turkey" pageId="5" pageNumber="180">Turkey</collectingCountry>
to the Levant and the Arabian peninsula.
</paragraph>
<paragraph id="D702447A5E04FFC1458CFF24FECDC5B1" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="181">
<taxonomicName id="10BD3FF95E04FFC1458CFF24FD83C486" authority="Benoist, 1934" authorityName="Benoist" authorityYear="1934" box="[199,557,151,176]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="E5C998685E04FFC1458CFF24FE28C486" box="[199,390,151,176]" italics="true" pageId="6" pageNumber="181">Heriades integra</emphasis>
<bibRefCitation id="B32C398B5E04FFC144C5FF24FD83C486" author="Benoist" box="[398,557,151,176]" pageId="6" pageNumber="181" refString="Benoist, R. (1934) Descriptions d'especes nouvelles palearctiques d'hymenopteres melliferes. Bulletin de la Societe Entomologique de France, 39, 158 - 160." type="journal article" year="1934">Benoist, 1934</bibRefCitation>
</taxonomicName>
, is treated as a member of the genus
<taxonomicName id="10BD3FF95E04FFC1469AFF24FBC1C486" box="[977,1135,151,176]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="E5C998685E04FFC1469AFF24FBC1C486" box="[977,1135,151,176]" italics="true" pageId="6" pageNumber="181">Stenoheriades</emphasis>
</taxonomicName>
by
<bibRefCitation id="B32C398B5E04FFC141D1FF24FAE0C486" author="Griswold" box="[1178,1358,151,176]" pageId="6" pageNumber="181" refString="Griswold, T. L. (1985) A generic and subgeneric revision of the Heriades genus-group (Hymenoptera: Megachilidae). Dissertation, Utah State University, Logan, 207 pp." type="book" year="1985">Griswold (1985</bibRefCitation>
,
<bibRefCitation id="B32C398B5E04FFC14010FF2BFA3AC486" author="Griswold" box="[1371,1428,152,176]" pageId="6" pageNumber="181" refString="Griswold, T. L. (1994) Taxonomic notes on some heriadines with descriptions of three new species (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society, 67, 17 - 28." type="journal article" year="1994">1994</bibRefCitation>
) and
<bibRefCitation id="B32C398B5E04FFC1458CFF0FFE11C4E2" box="[199,447,188,213]" pageId="6" pageNumber="181" refString="Ungricht, S., Muller, A. &amp; Dorn, S. (2008) A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae). Zootaxa, 1865, 1 - 253." type="journal article">
Ungricht
<emphasis id="E5C998685E04FFC14479FF0EFEC5C4E3" box="[306,363,188,213]" italics="true" pageId="6" pageNumber="181">et al.</emphasis>
(2008)
</bibRefCitation>
. Unfortunately, the male
<typeStatus id="0806FAD85E04FFC14797FF0FFC90C4E3" box="[732,830,188,213]" pageId="6" pageNumber="181" type="holotype">holotype</typeStatus>
originating from northernmost
<collectingCountry id="AFAA04EA5E04FFC141E8FF0EFAA6C4E2" box="[1187,1288,189,212]" name="Morocco" pageId="6" pageNumber="181">Morocco</collectingCountry>
could not be located at the Muséum national dHistoire naturelle Paris. However, the original description lists several characters, such as spined axilla, lack of a transverse carina between vertical and horizontal part of tergum 1 and rounded tergum 7, which exclude its membership in
<taxonomicName id="10BD3FF95E04FFC147AFFE94FC2CC576" box="[740,898,295,320]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="E5C998685E04FFC147AFFE94FC2CC576" box="[740,898,295,320]" italics="true" pageId="6" pageNumber="181">Stenoheriades</emphasis>
</taxonomicName>
. Instead, these characters solely apply to
<taxonomicName id="10BD3FF95E04FFC14018FE9BFDC5C553" authority="Morawitz, 1868" authorityName="Morawitz" authorityYear="1868" class="Insecta" family="Megachilidae" genus="Osmia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="scutellaris" subGenus="Hoplosmia">
<emphasis id="E5C998685E04FFC14018FE9BFE0BC553" italics="true" pageId="6" pageNumber="181">Osmia (Hoplosmia) scutellaris</emphasis>
<bibRefCitation id="B32C398B5E04FFC144FBFEFFFDC5C553" author="Morawitz" box="[432,619,332,357]" pageId="6" pageNumber="181" refString="Morawitz, F. (1868) Ueber einige Faltenwespen und Bienen aus der Umgegend von Nizza. Horae Societatis Entomologicae Rossicae, 5, 145 - 156." type="journal article" year="1868">Morawitz, 1868</bibRefCitation>
</taxonomicName>
, rendering the name
<taxonomicName id="10BD3FF95E04FFC14623FEFFFB85C552" box="[872,1067,332,357]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="integra">
<emphasis id="E5C998685E04FFC14623FEFFFB85C552" box="[872,1067,332,357]" italics="true" pageId="6" pageNumber="181">Heriades integra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEFA25135E04FFC1417CFEFEFB0EC552" box="[1079,1184,333,357]" pageId="6" pageNumber="181">syn. nov.</taxonomicNameLabel>
a junior synonym of
<taxonomicName id="10BD3FF95E04FFC145DCFEC3FEF5C5BE" box="[151,347,367,392]" class="Insecta" family="Megachilidae" genus="Osmia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="scutellaris">
<emphasis id="E5C998685E04FFC145DCFEC3FEF5C5BE" box="[151,347,367,392]" italics="true" pageId="6" pageNumber="181">Osmia scutellaris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D702447A5E04FFC1458CFE26FBA5C70E" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="181">
No morphological characters are known so far to differentiate between the males of
<emphasis id="E5C998685E04FFC14137FE26FA32C59B" box="[1148,1436,404,429]" italics="true" pageId="6" pageNumber="181">
<taxonomicName id="10BD3FF95E04FFC14137FE26FB5AC59A" box="[1148,1268,405,428]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">S. asiatica</taxonomicName>
,
<taxonomicName id="10BD3FF95E04FFC1404BFE26FA32C59B" box="[1280,1436,404,429]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="coelostoma">S. coelostoma</taxonomicName>
</emphasis>
and
<taxonomicName id="10BD3FF95E04FFC1458DFE0BFEE6C5E6" box="[198,328,439,464]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="levantica">
<emphasis id="E5C998685E04FFC1458DFE0BFEE6C5E6" box="[198,328,439,464]" italics="true" pageId="6" pageNumber="181">S. levantica</emphasis>
</taxonomicName>
, raising the question whether the male
<typeStatus id="0806FAD85E04FFC147B5FE0BFC97C5E6" box="[766,825,440,464]" pageId="6" pageNumber="181">types</typeStatus>
of
<taxonomicName id="10BD3FF95E04FFC14615FE0BFC7DC5E6" box="[862,979,440,464]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC14615FE0BFC7DC5E6" box="[862,979,440,464]" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
might be conspecific with
<taxonomicName id="10BD3FF95E04FFC1404AFE0BFA32C5E6" box="[1281,1436,439,464]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="coelostoma">
<emphasis id="E5C998685E04FFC1404AFE0BFA32C5E6" box="[1281,1436,439,464]" italics="true" pageId="6" pageNumber="181">S. coelostoma</emphasis>
</taxonomicName>
or
<taxonomicName id="10BD3FF95E04FFC145FEFE6EFE96C5C3" box="[181,312,476,501]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="levantica">
<emphasis id="E5C998685E04FFC145FEFE6EFE96C5C3" box="[181,312,476,501]" italics="true" pageId="6" pageNumber="181">S. levantica</emphasis>
</taxonomicName>
.
<collectingCountry id="AFAA04EA5E04FFC1440EFE6FFECAC5C2" box="[325,356,476,500]" name="American Samoa" pageId="6" pageNumber="181">As</collectingCountry>
females of a
<taxonomicName id="10BD3FF95E04FFC144B0FE6FFD36C5C3" box="[507,664,476,501]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="E5C998685E04FFC144B0FE6FFD36C5C3" box="[507,664,476,501]" italics="true" pageId="6" pageNumber="181">Stenoheriades</emphasis>
</taxonomicName>
species have been collected near the
<typeStatus id="0806FAD85E04FFC14170FE6EFBC5C5C3" box="[1083,1131,477,501]" pageId="6" pageNumber="181">type</typeStatus>
locality of
<taxonomicName id="10BD3FF95E04FFC141A7FE6EFACEC5C2" box="[1260,1376,477,500]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC141A7FE6EFACEC5C2" box="[1260,1376,477,500]" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
(
<collectingCountry id="AFAA04EA5E04FFC14024FE6FFA32C5C2" box="[1391,1436,476,500]" name="Nauru" pageId="6" pageNumber="181">Nur</collectingCountry>
mountains,
<collectingCountry id="AFAA04EA5E04FFC14456FDB3FECEC62E" box="[285,352,512,536]" name="Turkey" pageId="6" pageNumber="181">Hatay</collectingCountry>
province,
<collectingCountry id="AFAA04EA5E04FFC14495FE4CFD9CC62E" box="[478,562,511,536]" name="Turkey" pageId="6" pageNumber="181">Turkey</collectingCountry>
;
<bibRefCitation id="B32C398B5E04FFC1470BFE4CFD7FC62E" author="Friese" box="[576,721,511,536]" pageId="6" pageNumber="181" refString="Friese, H. (1921) Apidae. In: Fahringer, J. &amp; Friese, H. (Eds.), Eine Hymenopteren-Ausbeute aus dem Amanusgebirge (Kleinasien und Nord-Syrien sudl. Armenien). Archiv fur Naturgeschichte, Abteilung A, 161 - 176." type="book chapter" year="1921">Friese, 1921</bibRefCitation>
), which neither belong to
<taxonomicName id="10BD3FF95E04FFC14143FDB3FB06C62E" box="[1032,1192,511,536]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="coelostoma">
<emphasis id="E5C998685E04FFC14143FDB3FB06C62E" box="[1032,1192,511,536]" italics="true" pageId="6" pageNumber="181">S. coelostoma</emphasis>
</taxonomicName>
nor
<taxonomicName id="10BD3FF95E04FFC141A9FDB3FAC6C62E" box="[1250,1384,511,536]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="levantica">
<emphasis id="E5C998685E04FFC141A9FDB3FAC6C62E" box="[1250,1384,511,536]" italics="true" pageId="6" pageNumber="181">S. levantica</emphasis>
</taxonomicName>
, the
<taxonomicName id="10BD3FF95E04FFC145DCFD97FE9BC60B" box="[151,309,548,573]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="E5C998685E04FFC145DCFD97FE9BC60B" box="[151,309,548,573]" italics="true" pageId="6" pageNumber="181">Stenoheriades</emphasis>
</taxonomicName>
species with mandibles of normal shape, a shallow impression at the clypeal margin and a basal tubercle on the labrum (see identification key) is most probably
<taxonomicName id="10BD3FF95E04FFC146CAFDFBFC57C656" box="[897,1017,584,608]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC146CAFDFBFC57C656" box="[897,1017,584,608]" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
. This assumption is in line with the currently known distribution range of
<taxonomicName id="10BD3FF95E04FFC1470FFDDEFD71C6B3" box="[580,735,620,645]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="coelostoma">
<emphasis id="E5C998685E04FFC1470FFDDEFD71C6B3" box="[580,735,620,645]" italics="true" pageId="6" pageNumber="181">S. coelostoma</emphasis>
</taxonomicName>
and
<taxonomicName id="10BD3FF95E04FFC1465DFDDEFC34C6B3" box="[790,922,620,645]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="levantica">
<emphasis id="E5C998685E04FFC1465DFDDEFC34C6B3" box="[790,922,620,645]" italics="true" pageId="6" pageNumber="181">S. levantica</emphasis>
</taxonomicName>
, which is—compared with
<taxonomicName id="10BD3FF95E04FFC1419AFDDEFAE9C6B2" box="[1233,1351,621,644]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC1419AFDDEFAE9C6B2" box="[1233,1351,621,644]" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
—more western and more southern, respectively (see species accounts). The clarification of the species identity of
<taxonomicName id="10BD3FF95E04FFC140CCFD23FF5EC6FA" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC140CCFD23FF5EC6FA" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
renders the information about the geographic range of
<taxonomicName id="10BD3FF95E04FFC146DEFD06FBBFC6FA" box="[917,1041,693,716]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC146DEFD06FBBFC6FA" box="[917,1041,693,716]" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
given by
<bibRefCitation id="B32C398B5E04FFC141DFFD07FA32C6FA" box="[1172,1436,692,717]" pageId="6" pageNumber="181" refString="Ungricht, S., Muller, A. &amp; Dorn, S. (2008) A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae). Zootaxa, 1865, 1 - 253." type="journal article">
Ungricht
<emphasis id="E5C998685E04FFC1404EFD06FAEAC6FB" box="[1285,1348,692,717]" italics="true" pageId="6" pageNumber="181">et al.</emphasis>
(2008)
</bibRefCitation>
erroneous:
<taxonomicName id="10BD3FF95E04FFC14453FD6BFE3CC6C6" box="[280,402,728,752]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC14453FD6BFE3CC6C6" box="[280,402,728,752]" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
neither occurs in Europe nor in western
<collectingCountry id="AFAA04EA5E04FFC14637FD64FC62C6C6" box="[892,972,727,752]" name="Turkey" pageId="6" pageNumber="181">Turkey</collectingCountry>
but ranges from central to easternmost
<collectingCountry id="AFAA04EA5E04FFC145DCFD4FFF47C723" box="[151,233,764,789]" name="Turkey" pageId="6" pageNumber="181">Turkey</collectingCountry>
. The
<taxonomicName id="10BD3FF95E04FFC1446DFD4FFE6AC723" box="[294,452,764,789]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="E5C998685E04FFC1446DFD4FFE6AC723" box="[294,452,764,789]" italics="true" pageId="6" pageNumber="181">Stenoheriades</emphasis>
</taxonomicName>
species that is distributed in southeastern Europe and western
<collectingCountry id="AFAA04EA5E04FFC141CCFD4FFB79C723" box="[1159,1239,764,789]" name="Turkey" pageId="6" pageNumber="181">Turkey</collectingCountry>
is
<taxonomicName id="10BD3FF95E04FFC141B2FD4EFA3AC723" box="[1273,1428,764,789]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="coelostoma">
<emphasis id="E5C998685E04FFC141B2FD4EFA3AC723" box="[1273,1428,764,789]" italics="true" pageId="6" pageNumber="181">S. coelostoma</emphasis>
</taxonomicName>
, which has been erroneously synonymized with
<taxonomicName id="10BD3FF95E04FFC147E0FC93FC8EC70E" box="[683,800,800,824]" class="Insecta" family="Megachilidae" genus="Stenoheriades" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis id="E5C998685E04FFC147E0FC93FC8EC70E" box="[683,800,800,824]" italics="true" pageId="6" pageNumber="181">S. asiatica</emphasis>
</taxonomicName>
by
<bibRefCitation id="B32C398B5E04FFC14602FCACFBA8C70E" author="Griswold" box="[841,1030,799,824]" pageId="6" pageNumber="181" refString="Griswold, T. L. (1994) Taxonomic notes on some heriadines with descriptions of three new species (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society, 67, 17 - 28." type="journal article" year="1994">Griswold (1994)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>