155 lines
15 KiB
XML
155 lines
15 KiB
XML
<document id="6ECC4CCF033514B58BBD89A49918F858" ID-CLB-Dataset="27116" ID-DOI="http://dx.doi.org/10.3897/zookeys.840.33326" ID-GBIF-Dataset="363f2c7c-0635-46c8-b598-9e9aa014b6d2" ID-PMC="PMC6482118" ID-Pensoft-Pub="1313-2970-840-35" ID-PubMed="31065227" ID-ZooBank="C607C2FDC17644FABBC582D561C6434C" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-840-35" ModsDocOrigin="ZooKeys 840" ModsDocTitle="Conchological and molecular analysis of the “non-scaly” Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" checkinTime="1555526082661" checkinUser="pensoft" docAuthor="Zacaery Khalik, Mohd, P. Hendriks, Kasper, J. Vermeulen, Jaap & Menno Schilthuizen," docDate="2019" docId="1EC52054B87E4CB118EB7FC4F8E86A96" docLanguage="en" docName="ZooKeys 840: 35-86" docOrigin="ZooKeys 840" docSource="http://dx.doi.org/10.3897/zookeys.840.33326" docTitle="Georissa everetti Smith 1895" docType="treatment" docVersion="6" lastPageNumber="47" masterDocId="647C1D35F570C105FFFCFFCDB169212D" masterDocTitle="Conchological and molecular analysis of the " non-scaly " Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" masterLastPageNumber="86" masterPageNumber="35" pageNumber="47" updateTime="1701381596384" updateUser="ExternalLinkService">
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<mods:title id="C8042EA3DC1DD0F72AF7F08208EEBD65">Conchological and molecular analysis of the " non-scaly " Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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<mods:namePart id="83F15D004320D0250FE3B6908DE4938F">Zacaery Khalik, Mohd</mods:namePart>
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<mods:namePart id="DC7617FFC086435C6F4FEED9EE02ACC6">P. Hendriks, Kasper</mods:namePart>
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<mods:namePart id="FD431FF8D9F37D09FF754A026C63EAD1">J. Vermeulen, Jaap</mods:namePart>
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<mods:namePart id="9BA2AE7BCD7F417D4EA2980639B3D309">Menno Schilthuizen,</mods:namePart>
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<mods:typeOfResource id="CD3CF54E9F3EAABFB99501697FFE17DF">text</mods:typeOfResource>
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<mods:title id="49E069DD3F800E899BCA2A5DFC127590">ZooKeys</mods:title>
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<mods:date id="C5E46187CE74A16D337F1CDA52D31920">2019</mods:date>
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<mods:detail id="331147A1915B804659414D29381BA70D" type="volume">
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<mods:number id="6BC1E5BEF14338156019932AEFF6798F">840</mods:number>
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<mods:url id="8B8F3EF5BFA796FF5FF806FA7A8B6066">http://dx.doi.org/10.3897/zookeys.840.33326</mods:url>
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<mods:classification id="DF3624CF5F79E80419BF8864B787F3AB">journal article</mods:classification>
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<mods:identifier id="8E5178E873D16B35858FF3A01ECD162B" type="DOI">http://dx.doi.org/10.3897/zookeys.840.33326</mods:identifier>
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<treatment id="1EC52054B87E4CB118EB7FC4F8E86A96" ID-GBIF-Taxon="156787948" LSID="urn:lsid:plazi:treatment:1EC52054B87E4CB118EB7FC4F8E86A96" httpUri="http://treatment.plazi.org/id/1EC52054B87E4CB118EB7FC4F8E86A96" lastPageNumber="47" pageId="12" pageNumber="47">
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<subSubSection id="196A406808E9724375DF7149761DBA2F" pageId="12" pageNumber="47" type="nomenclature">
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<paragraph id="E6F478795AC3940C6D7E11E017B2D715" pageId="12" pageNumber="47">
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<taxonomicName id="4AE5BAAB51F9BAD1D438C2FCBB99BB69" ID-CoL="7RDTZ" authority="Smith, 1895" authorityName="Smith" authorityYear="1895" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa everetti" order="Cycloneritida" pageId="12" pageNumber="47" phylum="Mollusca" rank="species" species="everetti">Georissa everetti Smith, 1895</taxonomicName>
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Figures 1B, 7
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<normalizedToken id="85391FCC57E089326A09822D14DE95D1" originalValue="A–K">A-K</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection id="73DDFB43F68BB4E9ABAB69A5B3580745" pageId="12" pageNumber="47" type="reference_group">
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<paragraph id="408454EC456A4E027CE609321BC6BF48" pageId="12" pageNumber="47">
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<taxonomicName id="B756E15333D787FD46AF697F78C2DEB5" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa everetti" order="Cycloneritida" pageId="12" pageNumber="47" phylum="Mollusca" rank="species" species="everetti">Georissa everetti</taxonomicName>
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Smith, 1895: 125, plate IV fig. 15;
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<bibRefCitation id="6372A0E59C04EE5E6E56AD89A05492A8" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
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: 120, figs 55-57.
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</paragraph>
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</subSubSection>
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<subSubSection id="BDFF19B12CBDB82ED3FFEB2590204266" pageId="12" pageNumber="47" type="type locality">
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<paragraph id="8780FBBC9CEF350ADE980F78F8ED8D04" pageId="12" pageNumber="47">Type locality.</paragraph>
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<paragraph id="6ADC5FC3C7531505888FE86D6FC76A86" pageId="12" pageNumber="47">Rumbang, W. Sarawak.</paragraph>
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</subSubSection>
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<subSubSection id="B7F4F55460E07E3CE898A9D0111A7938" pageId="12" pageNumber="47" type="type material">
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<paragraph id="F6AB7D4C2D851638CAB2F2A5D9862CF8" pageId="12" pageNumber="47">Type material.</paragraph>
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<paragraph id="842C223D1F6E636BD7F208D835CFB735" pageId="12" pageNumber="47">
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Holotype (Holotype by original monotypy). Rumbang, Sarawak: NHMUK 1893.6.7.69 (glued on paper) (Fig. 1B) (
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<bibRefCitation id="5375EFB009C9D7DE8613AC5892BA1882" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="2A2EE955D00B15FC47C62293F5FF0CD8" pageId="12" pageNumber="47" type="materials_examined">
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<paragraph id="F2B5830F3AC54080B6CD5B5BE935C376" pageId="12" pageNumber="47">Other material.</paragraph>
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<paragraph id="FC7F7AE7DAEE668CB0CF50243FC79369" pageId="12" pageNumber="47">
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Kampung Giam, Lower Penrissen valley, Sarawak: JJV 12546. Kampung Benuk, Lower Penrissen valley, Sarawak (
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<geoCoordinate id="CFD3541F65B8EB042F9624AEE6175AA2" direction="north" orientation="latitude" precision="9" value="1.3078333">01°18.47'N</geoCoordinate>
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,
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<geoCoordinate id="1202A582B45D0F49EB98C7A3437AA612" direction="east" orientation="longitude" precision="9" value="110.28817">110°17.29'E</geoCoordinate>
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): JJV 12548. Kampung Temurang, Upper Penrissen valley, Sarawak (
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<geoCoordinate id="72599E42F739E36759EFEA357CCA4407" direction="north" orientation="latitude" precision="9" value="1.2025">01°12.15'N</geoCoordinate>
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,
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<geoCoordinate id="468C1385F82821A853CD0BB6A6BDADF9" direction="east" orientation="longitude" precision="9" value="110.26967">110°16.18'E</geoCoordinate>
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): JJV 12547. Great Cave, Niah National Park, Sarawak: RMNH/MOL 336264, JJV 10185 (Fig. 7), JJV 13119. Batu Punggul, Sepulut valley, Interior Province, Sabah: JJV 1906. Bukit Tinahas, Sepulut valley, Interior Province, Sabah (
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<geoCoordinate id="2535859E7408DCF9D60DE9F62121B5FA" direction="north" orientation="latitude" precision="9" value="4.638">04°38.28'N</geoCoordinate>
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,
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<geoCoordinate id="46C872040C6668BE3627DC005AACF278" direction="east" orientation="longitude" precision="9" value="116.6175">116°37.05'E</geoCoordinate>
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): JJV 7622.
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</paragraph>
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<caption id="F07D87A021046AA990D5023CBAAEA9AD" pageId="12" pageNumber="47">
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<paragraph id="1995B4B3A0AE885CA9B6EECC4BECEA6B" pageId="12" pageNumber="47">
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Figure 7.
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<taxonomicName id="200620BAE7191AC979E8CB16B8B024A6" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa everetti" order="Cycloneritida" pageId="12" pageNumber="47" phylum="Mollusca" rank="species" species="everetti">Georissa everetti</taxonomicName>
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Smith, 1895.
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<normalizedToken id="46C8ACA2E95F964262771F2E57031461" originalValue="A–KJJV">A-KJJV</normalizedToken>
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10185 A, D shell apertural view B shell side view C shell rear view E, F shell cross-section from 3D model G, H operculum frontal and ventral view from 3D model I shell top view J protoconch side view K close up of protoconch from top at 1000
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<normalizedToken id="A1897E2B8178F21E191137A9593D9FEA" originalValue="×">x</normalizedToken>
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magnification. Scale bars: 500
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<normalizedToken id="FB2B223A3184F4C91E4CDD906BB82A54" originalValue="µm">µm</normalizedToken>
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(
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<normalizedToken id="3D7A106FE93EFCD9E142153A1E7730F5" originalValue="A–I">A-I</normalizedToken>
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); 200
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<normalizedToken id="496EE85A4E32E61FDFE5763968F1CBE7" originalValue="µm">µm</normalizedToken>
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(J); 10
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<normalizedToken id="B531B8A447C785EC47896DC87EB548C7" originalValue="µm">µm</normalizedToken>
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(K).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="11E04096C66B1254710B532255ECC016" pageId="12" pageNumber="47" type="description">
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<paragraph id="225E434C1B5FBD7F1069680885F75F29" pageId="12" pageNumber="47">Description.</paragraph>
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<paragraph id="FAC7E9B2F5F92EF2A2974900D3666DFB" pageId="12" pageNumber="47">
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Protoconch. Colour: orange to red. Sculpture: rounded to ellipsoidal mesh pattern, mixed with irregular sculptural shapes (whenever two or more rounded or ellipsoidal meshes are connected or combined). Mesh width: 4-30
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<normalizedToken id="22E3FC6EE9368878F6A2A567DBC5EF06" originalValue="µm">µm</normalizedToken>
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. Teleoconch. Colour: orange to red. First whorl: convex. Subsequent whorls: convex, with relatively wide penultimate and final whorls. Suture: clearly impressed. Shoulder: narrow. Number of whorls: 2
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<normalizedToken id="6D30CF5DFC5159B94D3D2FE746A57C79" originalValue="¾–">3/4-</normalizedToken>
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3
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<normalizedToken id="4BF4D90DDC2A00063393C77DCE59D5F0" originalValue="¼">1/4</normalizedToken>
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. SH: 1.82-2.23 mm. SW: 1.52-1.75 mm. SI: 1.16-1.30. Shell sculpture. Radial sculpture: present, thin, forming small nodules when intersecting with spiral sculpture; these nodules are also present on the shoulder close to the suture. Spiral sculpture: present, thin, regularly spaced, oblique, appearing immediately after the protoconch, distorted/discontinuous by radial ribs. Aperture. Shape: rounded to ovoid, straight to concave parietal side, palatal edge contiguous with the body whorl, basal side convex. AH: 1.05-0.92 mm. AW: 1.09-0.96 mm. AI: 0.89-0.99.
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</paragraph>
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</subSubSection>
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<subSubSection id="181D58E60FB020DADA6D6584BE78A040" pageId="12" pageNumber="47" type="diagnosis">
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<paragraph id="0220A4903D76EE9F007375C3780FE18E" pageId="12" pageNumber="47">Diagnosis.</paragraph>
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<paragraph id="945F6C4E2F4083E77772945AD2D3F3E6" pageId="12" pageNumber="47">
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The strong and thin oblique spiral sculpture on its shell is diagnostic for
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<taxonomicName id="EC8DEA05651366CDCAC8817B1668E9CD" lsidName="G. everetti" pageId="12" pageNumber="47" rank="species" species="everetti">G. everetti</taxonomicName>
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.
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<taxonomicName id="061CDD52C8909CC943F542348C59C0D0" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa similis" order="Cycloneritida" pageId="12" pageNumber="47" phylum="Mollusca" rank="species" species="similis">Georissa similis</taxonomicName>
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has a somewhat similar knitted sculpture pattern resulting from the intersection of radial and spiral ribbing, but the shell shape is entirely distinct, with broad penultimate and final whorls. Based on the shell shape and habitus,
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<taxonomicName id="3AAB0A9F0C22CA601E242914616BFBC6" lsidName="G. everetti" pageId="12" pageNumber="47" rank="species" species="everetti">G. everetti</taxonomicName>
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resembles
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<taxonomicName id="127D01F6E9D0C16BB33870D23195279B" lsidName="G. gomantonensis" pageId="12" pageNumber="47" rank="species" species="gomantonensis">G. gomantonensis</taxonomicName>
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and
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<taxonomicName id="A053A491C6A498D97F44D444DF5B77B4" lsidName="G. williamsi" pageId="12" pageNumber="47" rank="species" species="williamsi">G. williamsi</taxonomicName>
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, which, however, have clear, regular, spiral shell ribs.
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</paragraph>
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</subSubSection>
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<subSubSection id="F12B5824637EA20A292B73FEC5F17CFA" pageId="12" pageNumber="47" type="distribution">
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<paragraph id="D6833F4D175A551C6B4335D9C683D471" pageId="12" pageNumber="47">Distribution.</paragraph>
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<paragraph id="DCD51918926E11203A780C7724858260" pageId="12" pageNumber="47">
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<taxonomicName id="669BDF62FED593A8A66D45B214B9BEEE" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa everetti" order="Cycloneritida" pageId="12" pageNumber="47" phylum="Mollusca" rank="species" species="everetti">Georissa everetti</taxonomicName>
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is widely distributed in Sabah and Sarawak, but is found in low abundances. The species known to occur from Padawan/Penrissen, Sarawak in the South (where Rumbang, the type locality is located), to further north, Niah, Sarawak, and Sepulut valley, Sabah.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |