300 lines
26 KiB
XML
300 lines
26 KiB
XML
<document ID-DOI="10.5252/a2009n2a1" ID-ISSN="1639-4798" ID-Zenodo-Dep="4601217" approvalRequired="20" approvalRequired_for_textStreams="20" checkinTime="1615554471545" checkinUser="carolina" docAuthor="Rakotondrainibe, France" docDate="2009" docId="D53687E68F54FFEED08FFDEEFB655DAC" docLanguage="fr" docName="Adansonia.31.2.235-248.pdf" docOrigin="Adansonia (3) 31 (2)" docSource="http://www.bioone.org/doi/abs/10.5252/a2009n2a1" docStyle="DocumentStyle:4B218CE5E725ACA5B76DF48277ED11EE.1:Adansonia.2009-2010.journal_article" docStyleId="4B218CE5E725ACA5B76DF48277ED11EE" docStyleName="Adansonia.2009-2010.journal_article" docStyleVersion="1" docTitle="Triplophyllum varians subsp. varians T. Moore" docType="treatment" docVersion="2" lastPageId="12" lastPageNumber="247" masterDocId="290FFF9E8F5FFFE2D248FFFAFFDF5965" masterDocTitle="Le genre Triplophyllum Holttum (Pteridophyta, Tectariaceae) à Madagascar" masterLastPageNumber="248" masterPageNumber="235" pageId="11" pageNumber="246" updateTime="1615557924189" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Le genre Triplophyllum Holttum (Pteridophyta, Tectariaceae) à Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rakotondrainibe, France</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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<mods:title>Adansonia</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="series">
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<mods:title>3</mods:title>
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||
</mods:detail>
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||
<mods:detail type="pubDate">
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<mods:number>2009-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>31</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>2</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>235</mods:start>
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<mods:end>248</mods:end>
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</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://www.bioone.org/doi/abs/10.5252/a2009n2a1</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5252/a2009n2a1</mods:identifier>
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<mods:identifier type="ISSN">1639-4798</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4601217</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:D53687E68F54FFEED08FFDEEFB655DAC" httpUri="http://treatment.plazi.org/id/D53687E68F54FFEED08FFDEEFB655DAC" lastPageId="12" lastPageNumber="247" pageId="11" pageNumber="246">
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<subSubSection pageId="11" pageNumber="246" type="nomenclature">
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<paragraph blockId="11.[711,1175,532,591]" pageId="11" pageNumber="246">
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<heading centered="true" fontSize="11" level="2" pageId="11" pageNumber="246" reason="3">
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<taxonomicName authority="(T. Moore) Holttum subsp. varians" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="246" phylum="Tracheophyta" rank="subSpecies" species="varians" subSpecies="varians">
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<emphasis bold="true" box="[711,953,532,559]" italics="true" pageId="11" pageNumber="246">Triplophyllum varians</emphasis>
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(T.Moore) Holttum subsp.
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<emphasis bold="true" box="[940,1020,564,590]" italics="true" pageId="11" pageNumber="246">varians</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="246" type="distribution">
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<paragraph blockId="11.[677,1211,628,750]" pageId="11" pageNumber="246">
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DISTRIBUTION Afrique:
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<collectingCountry box="[778,860,659,686]" name="Guinea-Bissau" pageId="11" pageNumber="246">Guinée</collectingCountry>
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,
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<collectingCountry box="[873,1021,659,686]" name="Ivory Coast" pageId="11" pageNumber="246">Côte d’Ivoire</collectingCountry>
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,
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<collectingCountry box="[1034,1110,659,686]" name="Ghana" pageId="11" pageNumber="246">Ghana</collectingCountry>
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,
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<collectingCountry box="[1122,1205,659,686]" name="Nigeria" pageId="11" pageNumber="246">Nigeria</collectingCountry>
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,
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<collectingCountry box="[678,802,691,718]" name="Cameroon" pageId="11" pageNumber="246">Cameroun</collectingCountry>
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,
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<collectingCountry box="[817,898,691,718]" name="Gabon" pageId="11" pageNumber="246">Gabon</collectingCountry>
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,
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<collectingCountry box="[911,991,691,718]" name="Democratic Republic of the Congo" pageId="11" pageNumber="246">Congo</collectingCountry>
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,
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<collectingCountry box="[1006,1086,691,718]" name="Angola" pageId="11" pageNumber="246">Angola</collectingCountry>
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et
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<collectingCountry name="South Africa" pageId="11" pageNumber="246">Afrique du Sud</collectingCountry>
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.
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</paragraph>
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<paragraph blockId="11.[711,1175,817,908]" box="[711,1175,817,845]" pageId="11" pageNumber="246">
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<taxonomicName authorityName="Rakotondrainibe" authorityYear="2009" baseAuthorityName="Rakotondrainibe" baseAuthorityYear="2009" box="[711,1175,817,845]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="246" phylum="Tracheophyta" rank="species" species="varians" status="comb. et stat. nov." subSpecies="anjenabense">
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<emphasis bold="true" box="[711,953,817,844]" italics="true" pageId="11" pageNumber="246">Triplophyllum varians</emphasis>
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(T.Moore) Holttum
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="247" pageId="11" pageNumber="246" type="description">
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<paragraph blockId="11.[711,1175,817,908]" pageId="11" pageNumber="246">
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<taxonomicName authority="(Tardieu) Rakotondr." authorityName="Rakotondrainibe" authorityYear="2009" baseAuthorityName="Rakotondrainibe" baseAuthorityYear="2009" box="[718,1164,849,876]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="246" phylum="Tracheophyta" rank="subSpecies" species="varians" status="comb. et stat. nov." subSpecies="anjenabense">
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subsp.
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<emphasis bold="true" box="[793,924,849,876]" italics="true" pageId="11" pageNumber="246">anjenabense</emphasis>
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(Tardieu) Rakotondr.
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</taxonomicName>
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,
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<taxonomicNameLabel box="[844,1043,881,908]" pageId="11" pageNumber="246">comb. et stat. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph blockId="11.[674,1212,938,1354]" pageId="11" pageNumber="246">
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<taxonomicName authority="Tardieu, Notulae" authorityName="Tardieu, Notulae" box="[699,1084,938,962]" class="Polypodiopsida" family="Dryopteridaceae" genus="Ctenitis" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="246" phylum="Tracheophyta" rank="species" species="anjenabensis">
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<emphasis box="[699,906,938,962]" italics="true" pageId="11" pageNumber="246">Ctenitis anjenabensis</emphasis>
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Tardieu,
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<emphasis box="[1007,1084,938,962]" italics="true" pageId="11" pageNumber="246">Notulae</emphasis>
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</taxonomicName>
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<emphasis box="[1093,1209,939,962]" italics="true" pageId="11" pageNumber="246">systematicae</emphasis>
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16 (1-2): 204, fig. 1, 1-4 (1960). —
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<taxonomicName authority="Tardieu" authorityName="Tardieu" authorityYear="1965" baseAuthorityName="Tardieu" genus="Abacopteris" kingdom="Plantae" pageId="11" pageNumber="246" rank="species" species="anjenabensis">
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<emphasis italics="true" pageId="11" pageNumber="246">Abacopteris anjenabensis</emphasis>
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Tardieu
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</taxonomicName>
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,
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<taxonomicName box="[821,920,991,1015]" class="Magnoliopsida" family="Malvaceae" genus="Adansonia" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="246" phylum="Tracheophyta" rank="genus">
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<emphasis box="[821,920,991,1015]" italics="true" pageId="11" pageNumber="246">Adansonia</emphasis>
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</taxonomicName>
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5 (4): 494, fig. 2, 5-7 (1965),
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<emphasis box="[676,859,1017,1041]" italics="true" pageId="11" pageNumber="246">nom. illeg. superflu</emphasis>
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. —
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<typeStatus box="[906,954,1018,1042]" pageId="11" pageNumber="246">Type</typeStatus>
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:
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<collectingCountry box="[973,1090,1018,1042]" name="Madagascar" pageId="11" pageNumber="246">Madagascar</collectingCountry>
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, vallée inférieure de l’Androranga, affl uent de la Bemarivo (Nord- Est), aux environs d’Antongondriha, mont Anjenabe,
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<quantity box="[678,742,1097,1121]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="246" unit="m" value="600.0">600 m</quantity>
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,
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<date box="[751,873,1097,1121]" pageId="11" pageNumber="246" value="1950-11-03" valueMax="1950-11-07" valueMin="1950-11-03">3-7.XI.1950</date>
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,
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<emphasis box="[882,1141,1097,1121]" italics="true" pageId="11" pageNumber="246">Humbert et Capuron 24023</emphasis>
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(holo-, P00547065!, iso-, P00511814!, P00511815!, P00511816!, P00696321!, P0069622!).
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</paragraph>
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<paragraph blockId="11.[674,1212,938,1354]" pageId="11" pageNumber="246">
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||
<materialsCitation collectingDate="1949-02-10" collectingDateMax="2005-12-05" collectingDateMin="1949-02-10" collectionCode="P, G" collectorName="de l'Androranga & de la Bemarivo & de Binara" country="Madagascar" elevation="450" location="Humbert" municipality="Vallee" pageId="11" pageNumber="246" specimenCount="1" stateProvince="Antsiranana">
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MATÉRIEL ÉTUDIÉ. —
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<emphasis bold="true" box="[896,1019,1197,1221]" pageId="11" pageNumber="246">
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<collectingCountry box="[896,1015,1197,1221]" name="Madagascar" pageId="11" pageNumber="246">Madagascar</collectingCountry>
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.
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</emphasis>
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<collectingMunicipality box="[1026,1084,1197,1221]" pageId="11" pageNumber="246">Vallée</collectingMunicipality>
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moyenne
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<collectorName pageId="11" pageNumber="246">de l’Androranga</collectorName>
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, affl uent
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<collectorName box="[895,1036,1224,1248]" pageId="11" pageNumber="246">de la Bemarivo</collectorName>
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(Nord-Est),
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<locationDeviation firstLocation="Doany" pageId="11" pageNumber="246" secondLocation="Anketsahely">entre Doany et Anketsahely</locationDeviation>
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,
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<quantity box="[898,963,1250,1274]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="246" unit="m" value="450.0">
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||
<elevation box="[898,963,1250,1274]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="246" unit="m" value="450.0">450 m</elevation>
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</quantity>
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,
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<date box="[973,1114,1250,1274]" pageId="11" pageNumber="246" value="1949-02-10" valueMax="1949-02-12" valueMin="1949-02-10">
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<collectingDate box="[973,1114,1250,1274]" pageId="11" pageNumber="246" value="1949-02-10" valueMax="1949-02-12" valueMin="1949-02-10">10-12.II.1949</collectingDate>
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||
</date>
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||
,
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||
<emphasis italics="true" pageId="11" pageNumber="246">
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||
<location LSID="urn:lsid:plazi:treatment:D53687E68F54FFEED08FFDEEFB655DAC:5840602B8F54FFE9D62BFB18FB665D9F" box="[1123,1209,1250,1274]" country="Madagascar" municipality="Vallee" name="Humbert" pageId="11" pageNumber="246" stateProvince="Antsiranana">Humbert</location>
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23194
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</emphasis>
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(
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<collectionCode box="[758,775,1277,1301]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="11" pageNumber="246" type="Herbarium">P</collectionCode>
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). —
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<collectingRegion box="[829,952,1277,1301]" country="Madagascar" name="Antsiranana" pageId="11" pageNumber="246">Antsiranana</collectingRegion>
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, Vohémar, Daraina, forêt
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<collectorName box="[678,775,1303,1327]" pageId="11" pageNumber="246">de Binara</collectorName>
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,
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<quantity box="[786,854,1303,1327]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="11" pageNumber="246" unit="m" value="735.0">
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<elevation box="[786,854,1303,1327]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="11" pageNumber="246" unit="m" value="735.0">735 m</elevation>
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</quantity>
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,
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<date box="[866,978,1303,1327]" pageId="11" pageNumber="246" value="2005-12-05">
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<collectingDate box="[866,978,1303,1327]" pageId="11" pageNumber="246" value="2005-12-05">5.XII.2005</collectingDate>
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</date>
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,
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<emphasis box="[989,1212,1303,1327]" italics="true" pageId="11" pageNumber="246">Nusbaumer et al. 1733</emphasis>
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(
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<collectionCode box="[685,706,1330,1354]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="11" pageNumber="246" type="Herbarium">G</collectionCode>
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,
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<collectionCode box="[716,733,1330,1354]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="11" pageNumber="246" type="Herbarium">P</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="11.[675,1209,1389,1700]" box="[677,821,1389,1415]" pageId="11" pageNumber="246">DESCRIPTION</paragraph>
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<paragraph blockId="11.[675,1209,1389,1700]" pageId="11" pageNumber="246">
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Rhizome rampant, de
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<quantity box="[912,987,1420,1447]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="11" pageNumber="246" unit="cm" value="7.5" valueMax="8.0" valueMin="7.0">7-8 cm</quantity>
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de diamètre, portant des écailles triangulaires, jusqu’à 5,0 ×
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<quantity box="[1082,1167,1451,1478]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="246" unit="mm" value="0.7">0,7 mm</quantity>
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à la base, brun-ocre, à base tronquée, apex longuement atténué, marge munie de quelques prolongements fins et longs.
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</paragraph>
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<paragraph blockId="11.[675,1209,1389,1700]" lastBlockId="12.[111,649,216,940]" lastPageId="12" lastPageNumber="247" pageId="11" pageNumber="246">
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Frondes distantes de 0,5-1,0 cm; frondes stériles et fertiles plus ou moins dimorphes, limbe fertile plus étroit. Pétiole brun-ocre, canaliculé, long de (36)
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<quantity box="[721,829,1673,1700]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="11" pageNumber="246" unit="cm" value="57.5" valueMax="70.0" valueMin="45.0">45-70 cm</quantity>
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||
, portant le plus souvent sur toute sa longueur des écailles semblables à celles du rhizome.
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<collectingRegion box="[111,181,248,275]" country="Cameroon" name="West" pageId="12" pageNumber="247">Limbe</collectingRegion>
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||
subcoriace, triangulaire à lancéolé, de 31 × 23 à 57 ×
|
||
<quantity box="[213,281,279,306]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="247" unit="cm" value="60.0">60 cm</quantity>
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||
, imparipenné, 4-7 paires de pennes latérales; pennes proximales subopposées, courtement pétiolées, de 14,0-18,0(35,0) ×
|
||
<quantity box="[520,642,343,370]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="12" pageNumber="247" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2,5-3,5 cm</quantity>
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||
, à base le plus souvent dissymétrique, apex longuement acuminé, marge sinueuse à lobée; pennes médianes subopposées ou alternes, courtement pétiolées ou sessiles, libres ou légèrement adnées, de 12,0-14,0(25,0) ×
|
||
<quantity box="[339,456,501,528]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.2" metricValueMin="2.2" pageId="12" pageNumber="247" unit="cm" value="2.7" valueMax="3.2" valueMin="2.2">2,2-3,2 cm</quantity>
|
||
, à base le plus souvent symétrique, apex longuement acuminé, marge sinueuse à lobée; penne terminale semblable aux latérales, base le plus souvent dissymétrique.
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</paragraph>
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||
<paragraph blockId="12.[111,649,216,940]" pageId="12" pageNumber="247">
|
||
Face adaxiale du rachis et des costae planes ou canaliculée, portant un tomentum de poils courts dressés, brun-rougeâtres, formés de 2-4 cellules, longs de
|
||
<quantity box="[215,350,723,750]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="12" pageNumber="247" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0,1-0,2 mm</quantity>
|
||
, face abaxiale convexe, glabre. Surface et marge du limbe glabre. Nervation irrégulièrement anastomosée; présence d’au moins une rangée d’aréoles costales hautes avec chacune une nerville excurrente incluse.
|
||
</paragraph>
|
||
<paragraph blockId="12.[111,649,216,940]" pageId="12" pageNumber="247">Sores exindusiés, positionnés le plus souvent à l’extrémité des nervures.</paragraph>
|
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</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="247" type="distribution">
|
||
<paragraph blockId="12.[113,647,977,1098]" box="[113,416,977,1003]" pageId="12" pageNumber="247">
|
||
DISTRIBUTION
|
||
<collectingRegion box="[276,302,982,1001]" country="Cameroon" name="East" pageId="12" pageNumber="247">ET</collectingRegion>
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||
ÉCOLOGIE
|
||
</paragraph>
|
||
<paragraph blockId="12.[113,647,977,1098]" pageId="12" pageNumber="247">
|
||
Endémique de la région de Doany, province d’Antsiranana, Nord-Est de
|
||
<collectingCountry box="[376,505,1040,1067]" name="Madagascar" pageId="12" pageNumber="247">Madagascar</collectingCountry>
|
||
; forêt dense humide sur gneiss; rare (
|
||
<figureCitation box="[382,447,1071,1098]" captionStart="FIG" captionStartId="4.[677,688,1243,1260]" captionTargetBox="[677,1208,215,1200]" captionTargetId="figure-19@4.[679,1208,217,1200]" captionTargetPageId="4" captionText="FIG. 2. — Distribution de Triplophyllum fraternum (Mett. ex Kuhn) Holttum (●), T. pentagonum (Bonap.) Holttum (★) et T. varians (T.Moore) Holttum subsp. anjenabense (Tardieu) Rakotondr. (▲) sur la carte bioclimatique de Madagascar (Cornet 1974)." figureDoi="http://doi.org/10.5281/zenodo.4601223" httpUri="https://zenodo.org/record/4601223/files/figure.png" pageId="12" pageNumber="247">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="12.[112,649,1135,1700]" box="[113,243,1135,1161]" pageId="12" pageNumber="247">REMARQUES</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="247" type="description">
|
||
<paragraph blockId="12.[112,649,1135,1700]" pageId="12" pageNumber="247">
|
||
<taxonomicName authorityName="Holttum" authorityYear="1986" baseAuthorityName="T. Moore" box="[113,341,1166,1192]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="species" species="varians">
|
||
<emphasis box="[113,341,1166,1192]" italics="true" pageId="12" pageNumber="247">Triplophyllum varians</emphasis>
|
||
</taxonomicName>
|
||
et la sous-espèce
|
||
<emphasis box="[523,647,1166,1192]" italics="true" pageId="12" pageNumber="247">anjenabense</emphasis>
|
||
se distinguent de tous les autres
|
||
<taxonomicName box="[449,596,1198,1224]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[449,596,1198,1224]" italics="true" pageId="12" pageNumber="247">Triplophyllum</emphasis>
|
||
</taxonomicName>
|
||
africano-malgaches et américains par la morphologie générale de la fronde 1-pennée, imparipennée, avec toutes les pennes entières, y compris les proximales et la terminale, un rhizome rampant relativement épais (environ
|
||
<quantity box="[277,344,1356,1383]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="247" unit="mm" value="8.0">8 mm</quantity>
|
||
de diamètre) et une nervation réticulée avec des aréoles costales irrégulières et redressées à l’oblique.
|
||
</paragraph>
|
||
<paragraph blockId="12.[112,649,1135,1700]" lastBlockId="12.[673,1212,216,1225]" pageId="12" pageNumber="247">
|
||
La sous-espèce
|
||
<emphasis box="[306,432,1451,1477]" italics="true" pageId="12" pageNumber="247">anjenabense</emphasis>
|
||
se différencie de la sous-espèce
|
||
<typeStatus box="[251,298,1483,1510]" pageId="12" pageNumber="247">type</typeStatus>
|
||
africaine par des écailles sur le rhizome plus étroites, plus claires, brun-roux, des frondes non gemmifères, la présence sur la face adaxiale des costae d’un tomentum plus ou moins dense de poils courts, dressés, brun-rouges, 2-4-cellulaires et la présence, dans les aréoles foliaires, de quelques nervilles incluses excurrentes. La présence de ces nervilles incluses constitue une exception dans le genre.
|
||
</paragraph>
|
||
<paragraph blockId="12.[673,1212,216,1225]" pageId="12" pageNumber="247">
|
||
<collectingRegion box="[699,735,279,306]" country="Cameroon" name="East" pageId="12" pageNumber="247">Les</collectingRegion>
|
||
deux sous-espèces de
|
||
<taxonomicName authorityName="Holttum" authorityYear="1986" baseAuthorityName="T. Moore" box="[978,1209,279,305]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="species" species="varians">
|
||
<emphasis box="[978,1209,279,305]" italics="true" pageId="12" pageNumber="247">Triplophyllum varians</emphasis>
|
||
</taxonomicName>
|
||
se distinguent des
|
||
<taxonomicName box="[883,968,312,337]" class="Polypodiopsida" family="Tectariaceae" genus="Tectaria" kingdom="Animalia" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[883,968,312,337]" italics="true" pageId="12" pageNumber="247">Tectaria</emphasis>
|
||
</taxonomicName>
|
||
, genre dans lequel ils ont été placés (
|
||
<bibRefCitation author="CHRISTENSEN C." box="[837,1030,343,370]" pageId="12" pageNumber="247" refId="ref7274" refString="CHRISTENSEN C. 1934. - Index Filicum, Supplementum" type="book" year="1934">Christensen 1934</bibRefCitation>
|
||
; Tardieu-Blot
|
||
<emphasis italics="true" pageId="12" pageNumber="247">in sched.</emphasis>
|
||
), par la morphologie générale de leur fronde, leur limbe entièrement glabre même sur les marges, et leurs aréoles costales redressées et de forme irrégulière, non rectangulaires et non parallèles au costa. Ils se distinguent également des
|
||
<taxonomicName box="[1131,1212,501,527]" class="Polypodiopsida" family="Dryopteridaceae" genus="Ctenitis" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[1131,1212,501,527]" italics="true" pageId="12" pageNumber="247">Ctenitis</emphasis>
|
||
</taxonomicName>
|
||
par leur rhizome rampant portant des écailles non clathrées, leurs pennes entières, de grande taille, 12-16 ×
|
||
<quantity box="[767,887,596,623]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="12" pageNumber="247" unit="cm" value="3.0" valueMax="3.5" valueMin="2.5">2,5-3,5 cm</quantity>
|
||
, non écailleuses et entièrement glabres (mis à part le tomentum lâche présent sur la face adaxiale des costae de la sous-espèce
|
||
<emphasis italics="true" pageId="12" pageNumber="247">anjenabense</emphasis>
|
||
), la nervation réticulée et l’absence de glandes unicellulaires. Notons enfin que ces deux taxons présentent des ressemblances morphologiques avec les représentants du genre
|
||
<taxonomicName authority="Tardieu" authorityName="Tardieu" box="[947,1177,786,813]" class="Polypodiopsida" family="Dryopteridaceae" genus="Pseudotectaria" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[947,1092,786,812]" italics="true" pageId="12" pageNumber="247">Pseudotectaria</emphasis>
|
||
Tardieu
|
||
</taxonomicName>
|
||
: le limbe 1-penné à nervation réticulée et la base plus ou moins asymétrique des pennes proximales; mais ils s’en distinguent par leur rhizome rampant et la penne terminale entière, semblable aux latérales.
|
||
</paragraph>
|
||
<paragraph blockId="12.[673,1212,216,1225]" pageId="12" pageNumber="247">
|
||
Ainsi, malgré la forme particulière de leur limbe et le
|
||
<typeStatus box="[726,771,976,1003]" pageId="12" pageNumber="247">type</typeStatus>
|
||
de nervation observé, non conformes à la morphologie classique des
|
||
<taxonomicName box="[962,1106,1008,1034]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[962,1106,1008,1034]" italics="true" pageId="12" pageNumber="247">Triplophyllum</emphasis>
|
||
</taxonomicName>
|
||
connus à ce jour, avec quelques hésitations, nous avons suivi
|
||
<bibRefCitation author="HOLTTUM R. E." box="[677,852,1071,1098]" pageId="12" pageNumber="247" pagination="237 - 260" refId="ref7325" refString="HOLTTUM R. E. 1986. - Studies in the fern-genera allied to Tectaria Cav. V. Triplophyllum, a new genus of Africa and America. Kew Bulletin 41 (2): 237 - 260." type="journal article" year="1986">Holttum (1986)</bibRefCitation>
|
||
et
|
||
<bibRefCitation author="KRAMER K. U." box="[886,1047,1071,1098]" pageId="12" pageNumber="247" pagination="124" refId="ref7419" refString="KRAMER K. U. 1990. - Triplophyllum, in KUBITZKI K. (ed.), Families and Genera of Vascular Plants, 1: Pteridophytes and Gymnosperms. Springer, Berlin: 124." type="book chapter" year="1990">Kramer (1990)</bibRefCitation>
|
||
en incluant les deux sous-espèces,
|
||
<taxonomicName authorityName="Holttum" authorityYear="1986" baseAuthorityName="T. Moore" box="[884,959,1104,1129]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="species" species="varians">
|
||
<emphasis box="[884,959,1104,1129]" italics="true" pageId="12" pageNumber="247">varians</emphasis>
|
||
</taxonomicName>
|
||
et
|
||
<emphasis box="[996,1119,1103,1129]" italics="true" pageId="12" pageNumber="247">anjenabense</emphasis>
|
||
, dans le genre
|
||
<taxonomicName box="[740,884,1134,1160]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[740,884,1134,1160]" italics="true" pageId="12" pageNumber="247">Triplophyllum</emphasis>
|
||
</taxonomicName>
|
||
. Une étude phylogénétique est cependant nécessaire pour confirmer ou infirmer la monophylie du genre
|
||
<taxonomicName box="[910,1054,1198,1224]" class="Polypodiopsida" family="Tectariaceae" genus="Triplophyllum" kingdom="Plantae" order="Polypodiales" pageId="12" pageNumber="247" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[910,1054,1198,1224]" italics="true" pageId="12" pageNumber="247">Triplophyllum</emphasis>
|
||
</taxonomicName>
|
||
ainsi délimité.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |