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<document id="17E3A9F9A6D7A6DBD964E50F423D7D54" ID-DOI="10.11646/phytotaxa.601.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="8129605" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1688977395585" checkinUser="plazi" docAuthor="Londoño-Echeverri, Yeison, Trujillo-López, Ana María &amp; Pérez-Zabala, Jorge Andrés" docDate="2023" docId="D512878E2641FFE52EDDF897FE97C8EF" docLanguage="en" docName="phytotaxa.601.2.4.pdf" docOrigin="Phytotaxa 601 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.601.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Conchocarpus cardenasii Londono-Ech., A. M." docType="treatment" docVersion="4" lastPageNumber="181" masterDocId="292BFFF62640FFE22E55FFA3FFAACD03" masterDocTitle="A new species of Conchocarpus and first record of Euxylophora (Galipeinae, Zanthoxyloideae, Rutaceae) from Colombia" masterLastPageNumber="184" masterPageNumber="174" pageNumber="175" updateTime="1689073932988" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4B34E1FAE1848EFD8CFC5AF91EC16F46">A new species of Conchocarpus and first record of Euxylophora (Galipeinae, Zanthoxyloideae, Rutaceae) from Colombia</mods:title>
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<mods:namePart id="6543CF707D70C7DACE9E2C3CBC46E91E">Londoño-Echeverri, Yeison</mods:namePart>
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<mods:affiliation id="D9C7D5BFECE3FD7992FDFF0B38FBE0A6">Herbario Universidad de Antioquia (HUA) y Grupo de Estudios Botánicos (GEOBOTA), Instituto de Biología, Facultad de Ciencias Exactas y Naturales, Universidad de Antioquia, A. A, 1226, Medellín, Colombia &amp; Herbario Gabriel Gutiérrez Villegas (MEDEL) y Semillero en estudios taxonómicos de plantas de Colombia, Facultad de Ciencias, Universidad Nacional de Colombia, A. A, 3840, Medellín, Colombia &amp; ylondono @ unal. edu. co; https: // orcid. org / 0000 - 0001 - 5169 - 598 X</mods:affiliation>
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<mods:namePart id="72DF5BEE9EEFB41D86DE882FB0F65850">Trujillo-López, Ana María</mods:namePart>
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<mods:affiliation id="CD7FB9B30EA6DF32774BDFABBBC4A886">Herbario Universidad de Antioquia (HUA) y Grupo de Estudios Botánicos (GEOBOTA), Instituto de Biología, Facultad de Ciencias Exactas y Naturales, Universidad de Antioquia, A. A, 1226, Medellín, Colombia &amp; Herbario Gabriel Gutiérrez Villegas (MEDEL) y Semillero en estudios taxonómicos de plantas de Colombia, Facultad de Ciencias, Universidad Nacional de Colombia, A. A, 3840, Medellín, Colombia &amp; amtrujillol @ unal. edu. co; https: // orcid. org / 0000 - 0002 - 0898 - 1646</mods:affiliation>
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<mods:namePart id="BBD8E8323DCC78B2A31B7ADF9CB5CAC1">Pérez-Zabala, Jorge Andrés</mods:namePart>
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<mods:affiliation id="6BD7E81230951A7B5E2B8730F11BF2CC">Herbario Gabriel Gutiérrez Villegas (MEDEL) y Semillero en estudios taxonómicos de plantas de Colombia, Facultad de Ciencias, Universidad Nacional de Colombia, A. A, 3840, Medellín, Colombia &amp; jperez @ unal. edu. co; https: // orcid. org / 0000 - 0003 - 1388 - 7694</mods:affiliation>
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<mods:title id="1E694DE3C1679E297E566B92D517451E">Phytotaxa</mods:title>
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<mods:date id="BD59F763BE88D51DC827AA5C077F34D2">2023</mods:date>
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<taxonomicName id="9ABB4D1B2641FFE32EDDF897FD33CA4D" authority="Londono-Ech., A. M." authorityName="Londono-Ech., A. M." box="[136,665,1844,1871]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="175" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2641FFE32EDDF897FE05CA4C" bold="true" box="[136,431,1844,1871]" italics="true" pageId="1" pageNumber="175">Conchocarpus cardenasii</emphasis>
Londoño-Ech., A.M.
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<collectingRegion id="9F7FF87A2641FFE32CCDF897FD59CA4D" box="[664,755,1844,1870]" country="Venezuela" name="Trujillo" pageId="1" pageNumber="175">Trujillo</collectingRegion>
&amp; Pérez-Zab., 
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(
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&amp; 
<figureCitation id="C5802A1D2641FFE32A26F897FB28CA4D" box="[1139,1154,1844,1870]" captionStart="FIGURE 2" captionStartId="3.[136,229,1996,2018]" captionTargetBox="[169,1412,204,1963]" captionTargetId="figure-24@3.[155,1432,187,1974]" captionTargetPageId="3" captionText="FIGURE 2. Illustration of Conchocarpus cardenasii: A. Distal portion of flowering branch, B. Flower, C. Petal and two stamens in adaxial view, D. Disc surrounding the ovary, E. Mericarp, F. Embryo. (A–D from the holotype, E–F from D. Cárdenas 2090. Illustration by Ana Cristina Pareja, HUA illustrator.)" figureDoi="http://doi.org/10.5281/zenodo.8129609" httpUri="https://zenodo.org/record/8129609/files/figure.png" pageId="1" pageNumber="175">2</figureCitation>
)
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<emphasis id="6FCFEA8A2641FFE32EDDF8FFFF70CA72" bold="true" box="[136,218,1884,1905]" pageId="1" pageNumber="175">Type:—</emphasis>
<materialsCitation id="EDD33CC52641FFE32E8FF8FFFC4CCA95" collectingDate="1988-04-19" collectionCode="JAUM" collectorName="D. Cardenas" country="Colombia" county="La Eugenia" elevation="175" location="Uraba" municipality="Riosucio" pageId="1" pageNumber="175" specimenCode="JAUM-015131" specimenCount="1" stateProvince="Choco" typeStatus="holotype">
<collectingCountry id="25AC76082641FFE32E8FF8FFFEF7CA72" box="[218,349,1884,1905]" name="Colombia" pageId="1" pageNumber="175">COLOMBIA</collectingCountry>
. 
<collectingRegion id="9F7FF87A2641FFE32F3CF8FFFE01CA71" box="[361,427,1884,1906]" country="Colombia" name="Choco" pageId="1" pageNumber="175">Chocó</collectingRegion>
: Mun. 
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, zona de 
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, 
<location id="586460432641FFE32CA2F8FFFC6BCA71" LSID="urn:lsid:plazi:treatment:D512878E2641FFE52EDDF897FE97C8EF:586460432641FFE32CA2F8FFFC6BCA71" box="[759,961,1884,1906]" country="Colombia" county="La Eugenia" municipality="Riosucio" name="Cerros Del Cuchillo" pageId="1" pageNumber="175" stateProvince="Choco">Cerros Del Cuchillo</location>
, camino 
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(Nova) a la cumbre sureste, 
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, 
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(bud &amp; fl), 
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<collectorName id="F04E534E2641FFE32C76F823FD36CA96" box="[547,668,1920,1941]" pageId="1" pageNumber="175">D. Cárdenas</collectorName>
1735
</emphasis>
(
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: 
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!)
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.
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<subSubSection id="15A165132641FFE62EDDF867FCEACBA3" lastPageId="4" lastPageNumber="178" pageId="1" pageNumber="175" type="diagnosis">
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<emphasis id="6FCFEA8A2641FFE32EDDF867FE8BCADD" bold="true" box="[136,289,1988,2014]" pageId="1" pageNumber="175">Diagnosis:—</emphasis>
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<emphasis id="6FCFEA8A2641FFE32F74F867FDEBCADD" box="[289,577,1988,2014]" italics="true" pageId="1" pageNumber="175">Conchocarpus cardenasii</emphasis>
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is morphologically similar to 
<taxonomicName id="9ABB4D1B2641FFE32DCFF867FAAFCADD" authority="Kallunki" authorityName="Kallunki" box="[922,1285,1988,2014]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="175" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2641FFE32DCFF867FB31CADD" box="[922,1179,1988,2014]" italics="true" pageId="1" pageNumber="175">Conchocarpus grandis</emphasis>
Kallunki
</taxonomicName>
in 
<bibRefCitation id="392A4B692641FFE32B7EF867FEFCC501" author="Kallunki, J. A. &amp; Pirani, J. R." pageId="1" pageNumber="175" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998: 299)</bibRefCitation>
, 
<taxonomicName id="9ABB4D1B2641FFE32F34F84BFBA8C501" authority="Kallunki &amp; W. Meier (2019: 196)" authorityName="Kallunki &amp; W. Meier" authorityPageNumber="196" authorityYear="2019" box="[353,1026,2024,2051]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="175" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2641FFE32F34F84BFD22C501" box="[353,648,2024,2050]" italics="true" pageId="1" pageNumber="175">Conchocarpus jirajaranus</emphasis>
Kallunki &amp; W. Meier (2019: 196)
</taxonomicName>
and in minor degree to 
<taxonomicName id="9ABB4D1B2641FFE32B5DF84BFCE3C525" authority="(Pulle 1912: 142) Kallunki &amp; Pirani (1998: 300)" authorityName="Kallunki &amp; Pirani" authorityPageNumber="300" authorityYear="1998" baseAuthorityName="Pulle" baseAuthorityPageNumber="142" baseAuthorityYear="1912" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="175" phylum="Tracheophyta" rank="species" species="guyanensis">
<emphasis id="6FCFEA8A2641FFE32B5DF84BFEAEC525" italics="true" pageId="1" pageNumber="175">Conchocarpus guyanensis</emphasis>
(
<bibRefCitation id="392A4B692641FFE32F42F7AFFE78C525" author="Pulle, A." box="[279,466,2060,2086]" pageId="1" pageNumber="175" pagination="125 - 169" refId="ref6792" refString="Pulle, A. (1912) Neue Beitrage zur Flora Surinams III. Mit zwei Tafeln. Recuei des Travaux Botaniques Neerlandais 9 (2): 125 - 169." type="journal article" year="1912">Pulle 1912: 142</bibRefCitation>
) 
<bibRefCitation id="392A4B692641FFE32FB0F7AFFCE3C525" author="Kallunki, J. A. &amp; Pirani, J. R." box="[485,841,2060,2086]" pageId="1" pageNumber="175" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998: 300)</bibRefCitation>
</taxonomicName>
. 
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<emphasis id="6FCFEA8A2641FFE32D02F7AFFC5BC525" box="[855,1009,2060,2086]" italics="true" pageId="1" pageNumber="175">C. cardenasii</emphasis>
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can be distinguished from the former two species by its shorter petioles, up to 
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long (
<emphasis id="6FCFEA8A2644FFE62CAFFF3CFCE8CDBB" box="[762,834,159,184]" italics="true" pageId="4" pageNumber="178">versus</emphasis>
up to 
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long in 
<taxonomicName id="9ABB4D1B2644FFE62A6DFF3CFB1ACDBB" box="[1080,1200,158,185]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2644FFE62A6DFF3CFB1ACDBB" box="[1080,1200,158,185]" italics="true" pageId="4" pageNumber="178">C. grandis</emphasis>
</taxonomicName>
, up to 
<quantity id="9A439B7D2644FFE62B51FF3DFAF8CDBA" box="[1284,1362,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="178" unit="cm" value="7.5">7.5 cm</quantity>
long in 
<taxonomicName id="9ABB4D1B2644FFE62EDDFF60FE83CDDF" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[136,297,195,221]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2644FFE62EDDFF60FE83CDDF" box="[136,297,195,221]" italics="true" pageId="4" pageNumber="178">C. jirajaranus</emphasis>
</taxonomicName>
), its longer flowering pedicels 
<quantity id="9A439B7D2644FFE62CC4FF61FD51CDDE" box="[657,763,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="4" pageNumber="178" unit="cm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 cm</quantity>
long (
<emphasis id="6FCFEA8A2644FFE62D1CFF60FCCDCDDF" box="[841,871,195,220]" italics="true" pageId="4" pageNumber="178">vs.</emphasis>
<quantity id="9A439B7D2644FFE62D25FF61FC45CDDE" box="[880,1007,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="4" pageNumber="178" unit="cm" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.3–0.6 cm</quantity>
long in 
<emphasis id="6FCFEA8A2644FFE62A07FF60FB7BCDDF" box="[1106,1233,194,221]" italics="true" pageId="4" pageNumber="178">
<taxonomicName id="9ABB4D1B2644FFE62A07FF60FB67CDDF" box="[1106,1229,194,221]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="grandis">C. grandis</taxonomicName>
,
</emphasis>
ca. 
<quantity id="9A439B7D2644FFE62B56FF61FAFBCDDE" box="[1283,1361,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="178" unit="cm" value="0.2">0.2 cm</quantity>
long in 
<taxonomicName id="9ABB4D1B2644FFE62EDDFF44FE80CC03" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[136,298,231,257]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2644FFE62EDDFF44FE80CC03" box="[136,298,231,257]" italics="true" pageId="4" pageNumber="178">C. jirajaranus</emphasis>
</taxonomicName>
), its anthers basally and apically sterile (
<emphasis id="6FCFEA8A2644FFE62D5BFF44FC86CC03" box="[782,812,231,256]" italics="true" pageId="4" pageNumber="178">vs.</emphasis>
anthers only apically sterile), its smaller mericarps ca. 1.2 × 
<quantity id="9A439B7D2644FFE62E99FEA9FEE0CC26" box="[204,330,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="4" pageNumber="178" unit="cm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 cm</quantity>
(
<emphasis id="6FCFEA8A2644FFE62F09FEA8FED0CC27" box="[348,378,267,292]" italics="true" pageId="4" pageNumber="178">vs.</emphasis>
2–2.5 × 
<quantity id="9A439B7D2644FFE62FB7FEA9FDE1CC26" box="[482,587,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="4" pageNumber="178" unit="cm" value="1.85" valueMax="2.0" valueMin="1.7">1.7–2 cm</quantity>
in 
<taxonomicName id="9ABB4D1B2644FFE62C27FEA8FD40CC27" box="[626,746,266,293]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2644FFE62C27FEA8FD40CC27" box="[626,746,266,293]" italics="true" pageId="4" pageNumber="178">C. grandis</emphasis>
</taxonomicName>
, 1.5–2 × 
<quantity id="9A439B7D2644FFE62D0DFEA9FC7DCC26" box="[856,983,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="4" pageNumber="178" unit="cm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 cm</quantity>
in 
<taxonomicName id="9ABB4D1B2644FFE62DA8FEA8FB34CC27" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[1021,1182,267,293]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2644FFE62DA8FEA8FB34CC27" box="[1021,1182,267,293]" italics="true" pageId="4" pageNumber="178">C. jirajaranus</emphasis>
</taxonomicName>
) and its ovary glabrous (only different respect to 
<taxonomicName id="9ABB4D1B2644FFE62FE4FE8CFDF9CC4B" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[433,595,303,329]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2644FFE62FE4FE8CFDF9CC4B" box="[433,595,303,329]" italics="true" pageId="4" pageNumber="178">C. jirajaranus</emphasis>
</taxonomicName>
with ovary densely strigulose). From 
<taxonomicName id="9ABB4D1B2644FFE62A47FE8CFB18CC4B" authorityName="Kallunki &amp; Pirani" authorityYear="1998" baseAuthorityName="Pulle" baseAuthorityYear="1912" box="[1042,1202,303,329]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="guyanensis">
<emphasis id="6FCFEA8A2644FFE62A47FE8CFB18CC4B" box="[1042,1202,303,329]" italics="true" pageId="4" pageNumber="178">C. guyanensis</emphasis>
</taxonomicName>
, the new species can be distinguished by its racemiform thyrses, bearing 15–26 sessile cymules along rachis (
<emphasis id="6FCFEA8A2644FFE62ADBFEF0FB06CC6F" box="[1166,1196,339,364]" italics="true" pageId="4" pageNumber="178">vs.</emphasis>
long-pedunculate and corymbiform thyrses bearing 3–4 partial inflorescences near apex, each stalked, ca. 
<quantity id="9A439B7D2644FFE62A06FED5FB27CC92" box="[1107,1165,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="178" unit="cm" value="2.0">2 cm</quantity>
long) and its seeds with plano-convex cotyledons (
<emphasis id="6FCFEA8A2644FFE62FE4FE38FE7ACCB7" box="[433,464,411,436]" italics="true" pageId="4" pageNumber="178">vs.</emphasis>
plicate-conduplicate cotyledons).
</paragraph>
<caption id="09C466102642FFE02EDDF800FDD3CABA" ID-DOI="http://doi.org/10.5281/zenodo.8129607" ID-Zenodo-Dep="8129607" box="[136,633,1955,1977]" httpUri="https://zenodo.org/record/8129607/files/figure.png" pageId="2" pageNumber="176" startId="2.[136,229,1955,1977]" targetBox="[155,1432,187,1928]" targetPageId="2" targetType="figure">
<paragraph id="5D0436982642FFE02EDDF800FDD3CABA" blockId="2.[136,633,1955,1977]" box="[136,633,1955,1977]" pageId="2" pageNumber="176">
<emphasis id="6FCFEA8A2642FFE02EDDF800FF54CABB" bold="true" box="[136,254,1955,1977]" pageId="2" pageNumber="176">FIGURE 1.</emphasis>
Holotype of 
<taxonomicName id="9ABB4D1B2642FFE02F28F800FDD9CABB" box="[381,627,1955,1976]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="2" pageNumber="176" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2642FFE02F28F800FDD9CABB" box="[381,627,1955,1976]" italics="true" pageId="2" pageNumber="176">Conchocarpus cardenasii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="09C466102643FFE12EDDF86FFDA5C529" ID-DOI="http://doi.org/10.5281/zenodo.8129609" ID-Zenodo-Dep="8129609" httpUri="https://zenodo.org/record/8129609/files/figure.png" pageId="3" pageNumber="177" startId="3.[136,229,1996,2018]" targetBox="[169,1412,204,1963]" targetPageId="3" targetType="figure">
<paragraph id="5D0436982643FFE12EDDF86FFDA5C529" blockId="3.[136,1452,1996,2090]" pageId="3" pageNumber="177">
<emphasis id="6FCFEA8A2643FFE12EDDF86FFEAACAE2" bold="true" box="[136,256,1996,2018]" pageId="3" pageNumber="177">FIGURE 2.</emphasis>
Illustration of 
<taxonomicName id="9ABB4D1B2643FFE12FC2F86FFD3ACAE2" box="[407,656,1996,2017]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="3" pageNumber="177" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2643FFE12FC2F86FFD3ACAE2" box="[407,656,1996,2017]" italics="true" pageId="3" pageNumber="177">Conchocarpus cardenasii</emphasis>
</taxonomicName>
: 
<emphasis id="6FCFEA8A2643FFE12CF5F86FFD1DCAE2" bold="true" box="[672,695,1996,2017]" pageId="3" pageNumber="177">A.</emphasis>
Distal portion of flowering branch, 
<emphasis id="6FCFEA8A2643FFE12A77F86FFB92CAE2" bold="true" box="[1058,1080,1996,2017]" pageId="3" pageNumber="177">B.</emphasis>
Flower, 
<emphasis id="6FCFEA8A2643FFE12AC7F86FFB00CAE1" bold="true" box="[1170,1194,1996,2018]" pageId="3" pageNumber="177">C.</emphasis>
Petal and two stamens in adaxial view, 
<emphasis id="6FCFEA8A2643FFE12F58F853FE8EC506" bold="true" box="[269,292,2032,2053]" pageId="3" pageNumber="177">D.</emphasis>
Disc surrounding the ovary, 
<emphasis id="6FCFEA8A2643FFE12C6EF853FDF8C506" bold="true" box="[571,594,2032,2053]" pageId="3" pageNumber="177">E.</emphasis>
Mericarp, 
<emphasis id="6FCFEA8A2643FFE12CE9F853FD64C506" bold="true" box="[700,718,2032,2053]" pageId="3" pageNumber="177">F.</emphasis>
Embryo. (A–D from the holotype, E–F from 
<emphasis id="6FCFEA8A2643FFE12AD3F853FA97C506" box="[1158,1341,2032,2053]" italics="true" pageId="3" pageNumber="177">D. Cárdenas 2090.</emphasis>
Illustration by Ana 
<taxonomicName id="9ABB4D1B2643FFE12E86F7B7FE0AC52A" authority="Pareja, HUA" authorityName="Pareja, HUA" box="[211,416,2068,2090]" class="Arachnida" family="Phalangiidae" genus="Cristina" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="177" phylum="Arthropoda" rank="genus">Cristina Pareja, HUA</taxonomicName>
illustrator.)
</paragraph>
</caption>
<paragraph id="5D0436982644FFE62EE8FE1CFCEACBA3" blockId="4.[136,1452,158,2092]" pageId="4" pageNumber="178">
<emphasis id="6FCFEA8A2644FFE62EE8FE1CFF44CCDB" box="[189,238,447,472]" italics="true" pageId="4" pageNumber="178">Tree</emphasis>
<quantity id="9A439B7D2644FFE62EA1FE1DFEE3CCDA" box="[244,329,446,473]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="178" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">5–10 m</quantity>
tall, branchlets with internodes evident, velvety to densely short-strigose, glabrescent, longitudinally ridged. 
<emphasis id="6FCFEA8A2644FFE62EB2FE40FE9FCCFF" box="[231,309,483,508]" italics="true" pageId="4" pageNumber="178">Leaves</emphasis>
1-foliolate, alternate, the petiole (0.7–) 
<quantity id="9A439B7D2644FFE62D4FFE41FC0BCCFE" box="[794,929,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.3" metricValueMin="1.6" pageId="4" pageNumber="178" unit="cm" value="2.45" valueMax="3.3" valueMin="1.6">1.6–3.3 cm</quantity>
long, canaliculate adaxially, short-strigose, glabrescent, ridged longitudinally, transversely striate, swollen at the base and apex; blade (7–)15.5–26 × (3.4–) 
<quantity id="9A439B7D2644FFE62B2CFDA5FF4FCF46" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="11.4" metricValueMin="5.6" pageId="4" pageNumber="178" unit="cm" value="8.5" valueMax="11.4" valueMin="5.6">5.6– 11.4 cm</quantity>
, elliptic, basally obtuse and cuneate to convex; apically acute and slightly acuminate; sparsely short-strigose on both surfaces, glabrescent; surface pellucid-punctate, the glands visible as dark dots; the margin entire, slightly revolute; secondary veins 12–17 pairs, eucamptodromous becoming brochidodromous distally, raised on both surfaces, more distinct abaxially, intersecondary veins present; tertiary veins percurrent, raised on both surfaces, more distinct abaxially. 
<emphasis id="6FCFEA8A2644FFE62EA9FD19FE38CFD7" box="[252,402,698,724]" italics="true" pageId="4" pageNumber="178">Inflorescence</emphasis>
a racemiform thyrse, 
<quantity id="9A439B7D2644FFE62CDBFD19FCB0CFD6" box="[654,794,698,725]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" metricValueMax="1.91" metricValueMin="0.47" pageId="4" pageNumber="178" unit="cm" value="11.9" valueMax="19.1" valueMin="4.7">4.7–19.1 cm</quantity>
long (including peduncle), short-strigose and puberulous, slightly ridged longitudinally, apparently supra-axillary (originating from a terminal meristem but reoriented laterally by the sympodial growth), the axillar vegetative bud persisting below; the peduncle 
<quantity id="9A439B7D2644FFE62A1FFCA1FB63CE1E" box="[1098,1225,770,797]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="4.8" metricValueMin="0.9" pageId="4" pageNumber="178" unit="cm" value="2.85" valueMax="4.8" valueMin="0.9">0.9–4.8 cm</quantity>
long, the flowering portion 
<quantity id="9A439B7D2644FFE62EB7FC85FEC4CE42" box="[226,366,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" metricValueMax="14.3" metricValueMin="2.5" pageId="4" pageNumber="178" unit="cm" value="8.4" valueMax="14.3" valueMin="2.5">2.5–14.3 cm</quantity>
long, bearing 15–26 flowering nodes, each a dense and sessile cymule, 1–3-flowered, the axes indiscernible; the primary bracts 1–2 × 
<quantity id="9A439B7D2644FFE62C6FFCE9FDD6CE66" box="[570,636,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="178" unit="mm" value="1.0">1 mm</quantity>
, ovate, subtending each cymule, densely short-strigose on abaxial surface and on adaxial surface only towards the margin (glabrous toward the middle portion); the secondary bracts 0.2–0.5 × 
<quantity id="9A439B7D2644FFE62EDDFC31FEB8CEAE" box="[136,274,914,941]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="4" pageNumber="178" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
, ovate, subtending some pedicels, densely short-strigose on abaxial surface and on adaxial surface only towards the margin (glabrous towards the middle portion), pedicels 
<quantity id="9A439B7D2644FFE62DCEFC15FBACCED2" box="[923,1030,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="4" pageNumber="178" unit="cm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 cm</quantity>
long, short-strigose and puberulous, glabrescent. 
<emphasis id="6FCFEA8A2644FFE62F43FC79FED8CEF7" box="[278,370,986,1012]" italics="true" pageId="4" pageNumber="178">Flowers</emphasis>
5(–6)-merous, calyx cupular, 
<quantity id="9A439B7D2644FFE62C93FC79FCE4CEF6" box="[710,846,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="4" pageNumber="178" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.5–1.9 mm</quantity>
high, 5-dentate, the teeth glandular ca. 0.2 × 
<quantity id="9A439B7D2644FFE62B1BFC79FA0DCEF6" box="[1358,1447,986,1013]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="178" unit="mm" value="0.2">0.2 mm</quantity>
, densely to sparsely short-strigose abaxially, glabrous adaxially; corolla actinomorphic, cochlear, white, short-tubular by capillinection, the tube ca. 
<quantity id="9A439B7D2644FFE62F8CFB81FD85C93E" box="[473,559,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="178" unit="mm" value="4.5">4.5 mm</quantity>
high; petals 7–9.6 × 
<quantity id="9A439B7D2644FFE62D49FB81FC3AC93E" box="[796,912,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="4" pageNumber="178" unit="mm" value="1.3" valueMax="1.6" valueMin="1.0">1–1.6 mm</quantity>
, oblong, apically obtuse and rounded to slightly acuminate, coherent into a tube among them, tomentose abaxially, adaxially glabrous at the base and near to the portion adnate to the filaments, the rest of surface tomentose, on the mouth of tube mixed with little short-villous indumentum, pellucid-punctate more noticeably abaxially; androecium 5(–6)-merous, all stamens fertile, filaments 3–3.7 × 
<quantity id="9A439B7D2644FFE62EB1FB11FEC7C9CE" box="[228,365,1202,1229]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="4" pageNumber="178" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
, oblong to narrowly oblanceolate, coherent among them by capillinection, adherent to the corolla at the base and adnate to it only by a small dorsal portion at the tube throat (perhaps postgenital fusion, see discussion), abaxially tomentose, glabrous near to the portion adnate to the corolla, adaxially glabrous at the base, the rest of surface tomentose, mixed with short-villous indumentum at the throat; anthers 3.2–4 × 
<quantity id="9A439B7D2644FFE62A7DFABDFB1EC83A" box="[1064,1204,1310,1337]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="4" pageNumber="178" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.5–0.7 mm</quantity>
, narrowly lanceolate, broadly attached to the filaments, sterile at the base and at the apex, eglandular at the connective, not exserted along their whole length from the corolla tube; thecae with a basal sterile portion of 
<quantity id="9A439B7D2644FFE62A77FAC5FB06C882" box="[1058,1196,1382,1409]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="4" pageNumber="178" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
long, a middle fertile portion of 
<quantity id="9A439B7D2644FFE62EABFA29FE2FC8A6" box="[254,389,1418,1445]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="4" pageNumber="178" unit="mm" value="1.8" valueMax="2.1" valueMin="1.5">1.5–2.1 mm</quantity>
long, abaxially sparsely short-villous, adaxially short-villous; the apical sterile portion 
<quantity id="9A439B7D2644FFE62B02FA29FF1EC8CA" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="4" pageNumber="178" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
long, flattened, curved in anthesis, abaxially sparsely tomentose, adaxially sparsely tomentose mixed with little short-villous indumentum; disc ca. 
<quantity id="9A439B7D2644FFE62C41FA71FDC0C8EE" box="[532,618,1490,1517]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="178" unit="mm" value="0.7">0.7 mm</quantity>
high, glabrous, slightly shorter than the ovary, margin sinuate; gynoecium 5-merous, apocarpic, glabrous, not exserted from the throat at anthesis, ovary 
<quantity id="9A439B7D2644FFE62A5DFA55FB3BCB12" box="[1032,1169,1526,1553]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="4" pageNumber="178" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
high, ca. 
<quantity id="9A439B7D2644FFE62B50FA55FAF7CB12" box="[1285,1373,1526,1553]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="178" unit="mm" value="0.8">0.8 mm</quantity>
diam., umbilicate, style ca. 
<quantity id="9A439B7D2644FFE62F20F9B9FE67CB36" box="[373,461,1562,1589]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="178" unit="mm" value="0.3">0.3 mm</quantity>
long (measured over ovary apex), stigma 
<quantity id="9A439B7D2644FFE62DE7F9B9FB91CB36" box="[946,1083,1562,1589]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="178" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
long, capitate. 
<emphasis id="6FCFEA8A2644FFE62AB9F9B8FA9ACB37" box="[1260,1328,1563,1588]" italics="true" pageId="4" pageNumber="178">Fruits</emphasis>
composed by a single mericarp, ca. 1.2 × 
<quantity id="9A439B7D2644FFE62FB7F99DFDC8CB5A" box="[482,610,1598,1625]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="4" pageNumber="178" unit="cm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 cm</quantity>
, semi-orbicular laterally, ventrally straight, dorsally rounded, very sparsely short-strigose, pellucid-punctate, transversely ridged. 
<emphasis id="6FCFEA8A2644FFE62CBFF9C1FCB5CB7F" box="[746,799,1634,1660]" italics="true" pageId="4" pageNumber="178">Seed</emphasis>
1 per carpel, ca. 1 × 
<quantity id="9A439B7D2644FFE62A4CF9C1FBC3CB7E" box="[1049,1129,1634,1661]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="178" unit="cm" value="0.8">0.8 cm</quantity>
, glabrous, slightly rugulose, testa papery, embryo straight, cotyledons plano-convex, thick.
</paragraph>
</subSubSection>
<subSubSection id="15A165132644FFE62EE8F909FB8DCA57" pageId="4" pageNumber="178" type="distribution">
<paragraph id="5D0436982644FFE62EE8F909FB8DCA57" blockId="4.[136,1452,158,2092]" pageId="4" pageNumber="178">
<emphasis id="6FCFEA8A2644FFE62EE8F909FDA6CBC7" bold="true" box="[189,524,1706,1732]" pageId="4" pageNumber="178">Distribution and habitat:—</emphasis>
<taxonomicName id="9ABB4D1B2644FFE62C59F909FC85CBC7" box="[524,815,1706,1732]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="178" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2644FFE62C59F909FC85CBC7" box="[524,815,1706,1732]" italics="true" pageId="4" pageNumber="178">Conchocarpus cardenasii</emphasis>
</taxonomicName>
is endemic to 
<collectingCountry id="25AC76082644FFE62DB5F909FBF8CBC7" box="[992,1106,1706,1732]" name="Colombia" pageId="4" pageNumber="178">Colombia</collectingCountry>
. It has been collected only in a hill named as “Cerro del Cuchillo” on the Riosucio municipality of the 
<collectingRegion id="9F7FF87A2644FFE62D85F96DFB35CBEB" box="[976,1183,1742,1768]" country="Colombia" name="Choco" pageId="4" pageNumber="178">Chocó department</collectingRegion>
(
<figureCitation id="C5802A1D2644FFE62AE5F96DFB5CCBEB" box="[1200,1270,1742,1768]" captionStart="FIGURE 3" captionStartId="5.[136,229,1650,1672]" captionTargetBox="[161,1433,215,1623]" captionTargetId="figure-24@5.[151,1436,190,1625]" captionTargetPageId="5" captionText="FIGURE 3. Geographical distribution of Conchocarpus cardenasii and georeferenced locality of Euxylophora paraensis population in Colombia." figureDoi="http://doi.org/10.5281/zenodo.8129611" httpUri="https://zenodo.org/record/8129611/files/figure.png" pageId="4" pageNumber="178">Fig. 3</figureCitation>
). The Cerro del Cuchillo is a small and isolated hill on the Pacific Biogeographical Region (
<emphasis id="6FCFEA8A2644FFE62D83F950FBBECA0F" box="[982,1044,1779,1804]" italics="true" pageId="4" pageNumber="178">sensu</emphasis>
<bibRefCitation id="392A4B692644FFE62A4FF951FB0FCA0F" author="Bernal, R." box="[1050,1189,1778,1804]" pageId="4" pageNumber="178" pagination="19 - 32" refId="ref4866" refString="Bernal, R. (2016) Geografia de Colombia. In: Bernal, R., Gradstein, S. R. &amp; Celis, M. (Eds.) Catalogo de plantas y liquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, pp. 19 - 32." type="book chapter" year="2016">Bernal 2016</bibRefCitation>
) in a range of elevation of 
<quantity id="9A439B7D2644FFE62EF3F8B5FEB2CA32" box="[166,280,1814,1841]" metricMagnitude="2" metricUnit="m" metricValue="2.35" metricValueMax="4.5" metricValueMin="0.2" pageId="4" pageNumber="178" unit="m" value="235.0" valueMax="450.0" valueMin="20.0">20–450 m</quantity>
, with annual precipitation between 
<quantity id="9A439B7D2644FFE62CF1F8B5FCDDCA32" box="[676,887,1814,1841]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="4" pageNumber="178" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4.000 to 8.000 mm</quantity>
(
<bibRefCitation id="392A4B692644FFE62DD0F8B5FB28CA33" author="Cardenas-Lopez, D." box="[901,1154,1814,1840]" pageId="4" pageNumber="178" pagination="101 - 117" refId="ref5004" refString="Cardenas-Lopez, D. (2003) Inventario floristico en el Cerro del Cuchillo, Tapon del Darien colombiano. Caldasia 25 (1): 101 - 117." type="journal article" year="2003">Cárdenas-López 2003</bibRefCitation>
). This region is considered under the Equatorial Rainforest climate (Af climate 
<typeStatus id="8200883A2644FFE62C85F898FCAACA56" box="[720,768,1851,1877]" pageId="4" pageNumber="178">type</typeStatus>
<emphasis id="6FCFEA8A2644FFE62D52F898FCEFCA57" box="[775,837,1851,1876]" italics="true" pageId="4" pageNumber="178">sensu</emphasis>
<bibRefCitation id="392A4B692644FFE62D19F899FBB6CA57" author="Kottek, M. &amp; Grieser, J. &amp; Beck, C. &amp; Rudolf, B. &amp; Rubel, F." box="[844,1052,1850,1876]" pageId="4" pageNumber="178" pagination="259 - 263" refId="ref6354" refString="Kottek, M., Grieser, J., Beck, C., Rudolf, B. &amp; Rubel, F. (2006) World map of the Koppen Geiger climate classification updated. Meteorologische Zeitschrif 15 (3): 259 - 263. https: // doi. org / 10.1127 / 0941 - 2948 / 2006 / 0130" type="journal article" year="2006">
Kottek 
<emphasis id="6FCFEA8A2644FFE62DF5F898FC72CA57" box="[928,984,1850,1876]" italics="true" pageId="4" pageNumber="178">et al.</emphasis>
2006
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="15A165132644FFE62EE8F8FDFB65CA7A" box="[189,1231,1886,1913]" pageId="4" pageNumber="178" type="biology_ecology">
<paragraph id="5D0436982644FFE62EE8F8FDFB65CA7A" blockId="4.[136,1452,158,2092]" box="[189,1231,1886,1913]" pageId="4" pageNumber="178">
<emphasis id="6FCFEA8A2644FFE62EE8F8FDFEF5CA7B" bold="true" box="[189,351,1886,1912]" pageId="4" pageNumber="178">Phenology:—</emphasis>
Herbarium vouchers indicate blooming time in April and fruiting time in June.
</paragraph>
</subSubSection>
<subSubSection id="15A165132644FFE62EE8F821FDA1C52F" pageId="4" pageNumber="178" type="etymology">
<paragraph id="5D0436982644FFE62EE8F821FDA1C52F" blockId="4.[136,1452,158,2092]" pageId="4" pageNumber="178">
<emphasis id="6FCFEA8A2644FFE62EE8F821FECECA9F" bold="true" box="[189,356,1922,1948]" pageId="4" pageNumber="178">Etymology:—</emphasis>
The epithet honors to Dairon Cárdenas, a noteworthy Colombian botanist who dedicated most of his career to improving knowledge of the Colombian Amazonian flora. The only known specimens for the new species were collected by Dairon early in his career. This is a modest acknowledgment to Dairon, for his assiduous work that rendered so many contributions to the knowledge of the flora, also to his kindness and friendship with the authors when they visited the COAH herbarium.
</paragraph>
</subSubSection>
<caption id="09C466102645FFE72EDDF9D1FF44CBAF" ID-DOI="http://doi.org/10.5281/zenodo.8129611" ID-Zenodo-Dep="8129611" httpUri="https://zenodo.org/record/8129611/files/figure.png" pageId="5" pageNumber="179" startId="5.[136,229,1650,1672]" targetBox="[161,1433,215,1623]" targetPageId="5" targetType="figure">
<paragraph id="5D0436982645FFE72EDDF9D1FF44CBAF" blockId="5.[136,1452,1650,1708]" pageId="5" pageNumber="179">
<emphasis id="6FCFEA8A2645FFE72EDDF9D1FF55CB84" bold="true" box="[136,255,1650,1672]" pageId="5" pageNumber="179">FIGURE 3.</emphasis>
Geographical distribution of 
<taxonomicName id="9ABB4D1B2645FFE72C75F9D1FCBDCB84" box="[544,791,1650,1671]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="179" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2645FFE72C75F9D1FCBDCB84" box="[544,791,1650,1671]" italics="true" pageId="5" pageNumber="179">Conchocarpus cardenasii</emphasis>
</taxonomicName>
and georeferenced locality of 
<taxonomicName id="9ABB4D1B2645FFE72A16F9D1FA89CB84" authorityName="Huber" authorityYear="1909" box="[1091,1315,1650,1671]" class="Magnoliopsida" family="Rutaceae" genus="Euxylophora" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="179" phylum="Tracheophyta" rank="species" species="paraensis">
<emphasis id="6FCFEA8A2645FFE72A16F9D1FA89CB84" box="[1091,1315,1650,1671]" italics="true" pageId="5" pageNumber="179">Euxylophora paraensis</emphasis>
</taxonomicName>
population in Colombia.
</paragraph>
</caption>
<subSubSection id="15A165132645FFE72EE8F979FB6DCACF" pageId="5" pageNumber="179" type="conservation">
<paragraph id="5D0436982645FFE72EE8F979FB6DCACF" blockId="5.[136,1452,1754,2069]" pageId="5" pageNumber="179">
<emphasis id="6FCFEA8A2645FFE72EE8F979FDC8CBF7" bold="true" box="[189,610,1754,1780]" pageId="5" pageNumber="179">Preliminary conservation status:—</emphasis>
<taxonomicName id="9ABB4D1B2645FFE72C36F979FC2ECBF7" box="[611,900,1754,1780]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="179" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2645FFE72C36F979FC2ECBF7" box="[611,900,1754,1780]" italics="true" pageId="5" pageNumber="179">Conchocarpus cardenasii</emphasis>
</taxonomicName>
is only known by a subpopulation with AOO= 
<quantity id="9A439B7D2645FFE72BCBF979FF06CA1B" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="179" unit="km" value="4.0">4 km</quantity>
<superScript id="AACE9BD02645FFE72EF9F95DFF1ECA0F" attach="left" box="[172,180,1790,1804]" fontSize="6" pageId="5" pageNumber="179">2</superScript>
. The forest that inhabits is a small fragment &lt;
<quantity id="9A439B7D2645FFE72C8EF95DFC8FCA1B" box="[731,805,1790,1816]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="179" unit="km" value="70.0">70 km</quantity>
<superScript id="AACE9BD02645FFE72D70F95DFC87CA0F" attach="left" box="[805,813,1790,1804]" fontSize="6" pageId="5" pageNumber="179">2</superScript>
immersed in a matrix of severe deforestation with two important human settlements less than 
<quantity id="9A439B7D2645FFE72C6FF881FD2BCA3F" box="[570,641,1826,1852]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="179" unit="km" value="20.0">20 km</quantity>
away (
<emphasis id="6FCFEA8A2645FFE72C87F880FD5FCA3F" box="[722,757,1827,1852]" italics="true" pageId="5" pageNumber="179">i.e.</emphasis>
, the unique known population is severe fragmented and under continuous decrease of habitat quality). Although it can be classified as “Critically endangered” according to the criteria B: B2ab(iii) of 
<bibRefCitation id="392A4B692645FFE72FDBF8C9FD82CA87" author="IUCN" box="[398,552,1898,1924]" pageId="5" pageNumber="179" refId="ref5971" refString="IUCN. (2012) IUCN Red List Categories and Criteria: Version 3.1. Second edition. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, England, 32 pp." type="book" year="2012">IUCN (2012)</bibRefCitation>
, the Cerro del Cuchillo is currently a protected area, which decreases the effect from any threat. It should be rather considered as “Endangered” species, due to its small and restricted geographical distribution, surrounded by livestock pastures with null ecological connectivity with near forests.
</paragraph>
</subSubSection>
<subSubSection id="15A165132645FFE72EE8F875FA32C516" pageId="5" pageNumber="179" type="materials_examined">
<paragraph id="5D0436982645FFE72EE8F875FA32C516" blockId="5.[136,1452,1754,2069]" pageId="5" pageNumber="179">
<materialsCitation id="EDD33CC52645FFE72EE8F875FA3EC516" collectingDate="1988-06-23" collectionCode="JAUM" collectorName="D. Cardenas" country="Colombia" elevation="75" location="Uraba" municipality="Riosucio" pageId="5" pageNumber="179" specimenCode="JAUM-015132" specimenCount="1" stateProvince="Choco" typeStatus="paratype">
<emphasis id="6FCFEA8A2645FFE72EE8F875FD77CAF3" bold="true" box="[189,733,2006,2032]" pageId="5" pageNumber="179">
Additional specimens examined (
<typeStatus id="8200883A2645FFE72C17F875FD19CAF3" box="[578,691,2006,2032]" pageId="5" pageNumber="179">paratype</typeStatus>
):—
</emphasis>
<collectingCountry id="25AC76082645FFE72C88F875FCDCCAF3" box="[733,886,2006,2032]" name="Colombia" pageId="5" pageNumber="179">COLOMBIA</collectingCountry>
. 
<collectingRegion id="9F7FF87A2645FFE72D2AF875FC67CAF3" box="[895,973,2006,2032]" country="Colombia" name="Choco" pageId="5" pageNumber="179">Chocó</collectingRegion>
: Mun. 
<collectingMunicipality id="BD60ACE22645FFE72A42F875FBD7CAF3" box="[1047,1149,2006,2032]" pageId="5" pageNumber="179">Riosucio</collectingMunicipality>
, zona de 
<location id="586460432645FFE72AB7F875FA83CAF3" LSID="urn:lsid:plazi:treatment:D512878E2641FFE52EDDF897FE97C8EF:586460432645FFE72AB7F875FA83CAF3" box="[1250,1321,2006,2032]" country="Colombia" municipality="Riosucio" name="Uraba" pageId="5" pageNumber="179" stateProvince="Choco">Urabá</location>
, 
<location id="586460432645FFE72B66F875FF41C517" LSID="urn:lsid:plazi:treatment:D512878E2641FFE52EDDF897FE97C8EF:586460432645FFE72B66F875FF41C517" country="Colombia" municipality="Riosucio" name="Cerros Del Cuchillo" pageId="5" pageNumber="179" stateProvince="Choco">Cerros Del Cuchillo</location>
, camino de 
<location id="586460432645FFE72F24F859FE1FC517" LSID="urn:lsid:plazi:treatment:D512878E2641FFE52EDDF897FE97C8EF:586460432645FFE72F24F859FE1FC517" box="[369,437,2042,2068]" country="Colombia" municipality="Riosucio" name="Cidon" pageId="5" pageNumber="179" stateProvince="Choco">Cidon</location>
a la 
<location id="586460432645FFE72FBEF859FDEFC517" LSID="urn:lsid:plazi:treatment:D512878E2641FFE52EDDF897FE97C8EF:586460432645FFE72FBEF859FDEFC517" box="[491,581,2042,2068]" country="Colombia" municipality="Riosucio" name="Cumbre" pageId="5" pageNumber="179" stateProvince="Choco">Cumbre</location>
sureste, 
<quantity id="9A439B7D2645FFE72CFDF859FCB1C516" box="[680,795,2042,2069]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="179" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
<elevation id="D696D1AB2645FFE72CFDF859FCB1C516" box="[680,795,2042,2069]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="179" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">50–100 m</elevation>
</quantity>
, 
<date id="290510582645FFE72D72F859FC17C517" box="[807,957,2042,2069]" pageId="5" pageNumber="179" value="1988-06-23">
<collectingDate id="3941E9B02645FFE72D72F859FC17C517" box="[807,957,2042,2069]" pageId="5" pageNumber="179" value="1988-06-23">23 June 1988</collectingDate>
</date>
(fr), 
<emphasis id="6FCFEA8A2645FFE72DA2F858FB6EC517" box="[1015,1220,2042,2068]" italics="true" pageId="5" pageNumber="179">
<collectorName id="F04E534E2645FFE72DA2F858FB2FC517" box="[1015,1157,2042,2068]" pageId="5" pageNumber="179">D. Cárdenas</collectorName>
2090
</emphasis>
(
<specimenCode id="0D1D9EE32645FFE72A86F859FA23C516" box="[1235,1417,2042,2069]" pageId="5" pageNumber="179">JAUM-015132</specimenCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="15A165132646FFE52EE8FF3DFE97C8EF" lastPageId="7" lastPageNumber="181" pageId="6" pageNumber="180" type="discussion">
<paragraph id="5D0436982646FFE42EE8FF3DFF48CCDB" blockId="6.[136,1452,158,472]" pageId="6" pageNumber="180">
<emphasis id="6FCFEA8A2646FFE42EE8FF3DFE8CCDBB" bold="true" box="[189,294,158,184]" pageId="6" pageNumber="180">Notes:—</emphasis>
<taxonomicName id="9ABB4D1B2646FFE42F73FF3DFDEFCDBB" box="[294,581,158,184]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2646FFE42F73FF3DFDEFCDBB" box="[294,581,158,184]" italics="true" pageId="6" pageNumber="180">Conchocarpus cardenasii</emphasis>
</taxonomicName>
has a close resemblance to 
<taxonomicName id="9ABB4D1B2646FFE42D2FFF3CFC5BCDBB" box="[890,1009,158,185]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2646FFE42D2FFF3CFC5BCDBB" box="[890,1009,158,185]" italics="true" pageId="6" pageNumber="180">C. grandis</emphasis>
</taxonomicName>
and 
<emphasis id="6FCFEA8A2646FFE42A73FF3CFB66CDBB" box="[1062,1228,159,185]" italics="true" pageId="6" pageNumber="180">
<taxonomicName id="9ABB4D1B2646FFE42A73FF3CFB62CDBB" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[1062,1224,159,185]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="jirajaranus">C. jirajaranus</taxonomicName>
.
</emphasis>
These three species share the 1-foliolate leaves, the racemiform thyrses with partial inflorescences sessile (only stalked at the lowermost ones in 
<taxonomicName id="9ABB4D1B2646FFE42EB5FF44FEFDCC03" box="[224,343,230,257]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2646FFE42EB5FF44FEFDCC03" box="[224,343,230,257]" italics="true" pageId="6" pageNumber="180">C. grandis</emphasis>
</taxonomicName>
), the cupular calyx, the actinomorphic corolla (not verified for 
<taxonomicName id="9ABB4D1B2646FFE42A77FF44FB33CC03" box="[1058,1177,230,257]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2646FFE42A77FF44FB33CC03" box="[1058,1177,230,257]" italics="true" pageId="6" pageNumber="180">C. grandis</emphasis>
</taxonomicName>
), the all-fertile stamens, the lanceolate and apically sterile anthers, the umbilicate ovary (not verified for 
<taxonomicName id="9ABB4D1B2646FFE42A71FEA8FB6ECC27" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[1060,1220,267,293]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2646FFE42A71FEA8FB6ECC27" box="[1060,1220,267,293]" italics="true" pageId="6" pageNumber="180">C. jirajaranus</emphasis>
</taxonomicName>
) and the seeds with plano-convex cotyledons. On the other hand, 
<taxonomicName id="9ABB4D1B2646FFE42CC8FE8CFC94CC4B" authorityName="Kallunki &amp; Pirani" authorityYear="1998" baseAuthorityName="Pulle" baseAuthorityYear="1912" box="[669,830,303,329]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="guyanensis">
<emphasis id="6FCFEA8A2646FFE42CC8FE8CFC94CC4B" box="[669,830,303,329]" italics="true" pageId="6" pageNumber="180">C. guyanensis</emphasis>
</taxonomicName>
shares with the new species the unique in the genus presence of anthers with thecae basally sterile, but can be easily differentiated from it and the other two species above mentioned by its long-pedunculate and corymbiform thyrses and seeds with plicate-conduplicate cotyledons. Other features are relatively similar among the four species discussed (
<bibRefCitation id="392A4B692646FFE42D00FE39FBCACCB7" author="Kallunki, J. A. &amp; Pirani, J. R." box="[853,1120,410,436]" pageId="6" pageNumber="180" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani 1998</bibRefCitation>
, 
<bibRefCitation id="392A4B692646FFE42A3EFE39FADFCCB7" author="Kallunki, J. A. &amp; Meier, W." box="[1131,1397,410,436]" pageId="6" pageNumber="180" pagination="196 - 200" refId="ref6240" refString="Kallunki, J. A. &amp; Meier, W. (2019) Conchocarpus jirajaranus (Rutaceae), a new endemic species of the Coastal Cordillera of Venezuela. Brittonia 71 (2): 196 - 200. https: // doi. org / 10.1007 / s 12228 - 019 - 09572 - 1" type="journal article" year="2019">Kallunki &amp; Meier 2019</bibRefCitation>
), see 
<tableCitation id="103903232646FFE42EDDFE1DFF76CCDB" box="[136,220,446,472]" captionStart="TABLE 1" captionStartId="6.[136,216,522,543]" captionTargetPageId="6" captionText="TABLE 1. Comparison of the similar species to Conchocarpus cardenasii." httpUri="http://table.plazi.org/id/09C466102646FFE42EDDFDA9FCFDCF1C" pageId="6" pageNumber="180" tableUuid="09C466102646FFE42EDDFDA9FCFDCF1C">Table 1</tableCitation>
.
</paragraph>
<caption id="09C466102646FFE42EDDFDA9FCFDCF1C" ID-Table-UUID="09C466102646FFE42EDDFDA9FCFDCF1C" box="[136,855,522,544]" httpUri="http://table.plazi.org/id/09C466102646FFE42EDDFDA9FCFDCF1C" pageId="6" pageNumber="180" startId="6.[136,216,522,543]" targetBox="[159,1425,572,1735]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="5D0436982646FFE42EDDFDA9FCFDCF1C" blockId="6.[136,855,522,544]" box="[136,855,522,544]" pageId="6" pageNumber="180">
<emphasis id="6FCFEA8A2646FFE42EDDFDA9FF5ACF1C" bold="true" box="[136,240,522,543]" pageId="6" pageNumber="180">TABLE 1.</emphasis>
Comparison of the similar species to 
<taxonomicName id="9ABB4D1B2646FFE42C0EFDA9FCFBCF1C" box="[603,849,522,543]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2646FFE42C0EFDA9FCFBCF1C" box="[603,849,522,543]" italics="true" pageId="6" pageNumber="180">Conchocarpus cardenasii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="5D0436982646FFE42ECAFD9FFADDCB9C" pageId="6" pageNumber="180">
<table id="2FBBC4382646001D2ECAFD9FFA3BCBC4" box="[159,1425,572,1735]" gridcols="6" gridrows="12" pageId="6" pageNumber="180">
<tr id="E38B34DA2646001D2ECAFD9FFA3BCF51" box="[159,1425,572,594]" gridrow="0" pageId="6" pageNumber="180" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="A05A5DA62646001D2ECAFD9FFEF9CF51" box="[159,339,572,594]" gridcol="0" gridrow="0" pageId="6" pageNumber="180">
<emphasis id="6FCFEA8A2646FFE42ECAFD9FFEA2CF51" bold="true" box="[159,264,572,594]" pageId="6" pageNumber="180">Character</emphasis>
</th>
<th id="A05A5DA62646001D2D60FD9FFC75CF51" box="[821,991,572,594]" gridcol="3" gridrow="0" pageId="6" pageNumber="180">
<emphasis id="6FCFEA8A2646FFE42D33FD9FFC1BCF51" bold="true" box="[870,945,572,594]" pageId="6" pageNumber="180">Species</emphasis>
</th>
</tr>
<tr id="E38B34DA2646001D2ECAFD22FA3BCF94" box="[159,1425,641,663]" gridrow="1" pageId="6" pageNumber="180" rowspan-0="1">
<td id="A05A5DA62646001D2FD7FD22FD84CF94" box="[386,558,641,663]" gridcol="1" gridrow="1" pageId="6" pageNumber="180">
<taxonomicName id="9ABB4D1B2646FFE42FD7FD21FDAFCF94" box="[386,517,641,663]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2646FFE42FD7FD21FDAFCF94" bold="true" box="[386,517,641,663]" italics="true" pageId="6" pageNumber="180">C. cardenasii</emphasis>
</taxonomicName>
</td>
<td id="A05A5DA62646001D2C0EFD22FCACCF94" box="[603,774,641,663]" gridcol="2" gridrow="1" pageId="6" pageNumber="180">
<taxonomicName id="9ABB4D1B2646FFE42C0EFD21FD68CF94" box="[603,706,641,663]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2646FFE42C0EFD21FD68CF94" bold="true" box="[603,706,641,663]" italics="true" pageId="6" pageNumber="180">C. grandis</emphasis>
</taxonomicName>
</td>
<td id="A05A5DA62646001D2D60FD22FC75CF94" box="[821,991,641,663]" gridcol="3" gridrow="1" pageId="6" pageNumber="180">
<taxonomicName id="9ABB4D1B2646FFE42D60FD21FC6ACF94" authorityName="Kallunki &amp; Pirani" authorityYear="1998" baseAuthorityName="Pulle" baseAuthorityYear="1912" box="[821,960,642,663]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="guyanensis">
<emphasis id="6FCFEA8A2646FFE42D60FD21FC6ACF94" bold="true" box="[821,960,642,663]" italics="true" pageId="6" pageNumber="180">C. guyanensis</emphasis>
</taxonomicName>
<emphasis id="6FCFEA8A2646FFE42D93FD22FC67CF8E" bold="true" box="[966,973,641,653]" pageId="6" pageNumber="180">a</emphasis>
</td>
<td id="A05A5DA62646001D2A5AFD22FB11CF94" box="[1039,1211,641,663]" gridcol="4" gridrow="1" pageId="6" pageNumber="180">
<taxonomicName id="9ABB4D1B2646FFE42A5AFD21FB30CF94" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[1039,1178,642,663]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2646FFE42A5AFD21FB30CF94" bold="true" box="[1039,1178,642,663]" italics="true" pageId="6" pageNumber="180">C. jirajaranus</emphasis>
</taxonomicName>
</td>
<td id="A05A5DA62646001D2ABDFD22FA3BCF94" box="[1256,1425,641,663]" gridcol="5" gridrow="1" pageId="6" pageNumber="180">
<taxonomicName id="9ABB4D1B2646FFE42ABDFD21FACDCF94" box="[1256,1383,642,663]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="toxicarius">
<emphasis id="6FCFEA8A2646FFE42ABDFD21FACDCF94" bold="true" box="[1256,1383,642,663]" italics="true" pageId="6" pageNumber="180">C. toxicarius</emphasis>
</taxonomicName>
<emphasis id="6FCFEA8A2646FFE42B3EFD22FAD9CF8E" bold="true" box="[1387,1395,641,653]" pageId="6" pageNumber="180">b</emphasis>
</td>
</tr>
<tr id="E38B34DA2646001D2ECAFD68FA3BCFE2" box="[159,1425,715,737]" gridrow="2" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFD68FEF9CFE2" box="[159,339,715,737]" gridcol="0" gridrow="2" pageId="6" pageNumber="180">Leaves</th>
<td id="A05A5DA62646001D2FD7FD68FD84CFE2" box="[386,558,715,737]" gridcol="1" gridrow="2" pageId="6" pageNumber="180">1-foliolate</td>
<td id="A05A5DA62646001D2C0EFD68FCACCFE2" box="[603,774,715,737]" gridcol="2" gridrow="2" pageId="6" pageNumber="180">1-foliolate</td>
<td id="A05A5DA62646001D2D60FD68FC75CFE2" box="[821,991,715,737]" gridcol="3" gridrow="2" pageId="6" pageNumber="180">1-foliolate</td>
<td id="A05A5DA62646001D2A5AFD68FB11CFE2" box="[1039,1211,715,737]" gridcol="4" gridrow="2" pageId="6" pageNumber="180">1-foliolate</td>
<td id="A05A5DA62646001D2ABDFD68FA3BCFE2" box="[1256,1425,715,737]" gridcol="5" gridrow="2" pageId="6" pageNumber="180">5–7-foliolate</td>
</tr>
<tr id="E38B34DA2646001D2ECAFCAAFA3BCE38" box="[159,1425,777,827]" gridrow="3" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFCAAFEF9CE38" box="[159,339,777,827]" gridcol="0" gridrow="3" pageId="6" pageNumber="180">Petiole length (cm)</th>
<td id="A05A5DA62646001D2FD7FCAAFD84CE38" box="[386,558,777,827]" gridcol="1" gridrow="3" pageId="6" pageNumber="180">(0.7–)1.6–3.3</td>
<td id="A05A5DA62646001D2C0EFCAAFCACCE38" box="[603,774,777,827]" gridcol="2" gridrow="3" pageId="6" pageNumber="180">2.4–12</td>
<td id="A05A5DA62646001D2D60FCAAFC75CE38" box="[821,991,777,827]" gridcol="3" gridrow="3" pageId="6" pageNumber="180">0.5–6(–9.5)</td>
<td id="A05A5DA62646001D2A5AFCAAFB11CE38" box="[1039,1211,777,827]" gridcol="4" gridrow="3" pageId="6" pageNumber="180">(1.3–)2.5–7.5</td>
<td id="A05A5DA62646001D2ABDFCAAFA3BCE38" box="[1256,1425,777,827]" gridcol="5" gridrow="3" pageId="6" pageNumber="180">(28–)30–45</td>
</tr>
<tr id="E38B34DA2646001D2ECAFCC7FA3BCE9B" box="[159,1425,868,920]" gridrow="4" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFCC7FEF9CE9B" box="[159,339,868,920]" gridcol="0" gridrow="4" pageId="6" pageNumber="180">Thyrses morphology</th>
<td id="A05A5DA62646001D2FD7FCC7FD84CE9B" box="[386,558,868,920]" gridcol="1" gridrow="4" pageId="6" pageNumber="180">racemiform</td>
<td id="A05A5DA62646001D2C0EFCC7FCACCE9B" box="[603,774,868,920]" gridcol="2" gridrow="4" pageId="6" pageNumber="180">racemiform</td>
<td id="A05A5DA62646001D2D60FCC7FC75CE9B" box="[821,991,868,920]" gridcol="3" gridrow="4" pageId="6" pageNumber="180">corymbiform</td>
<td id="A05A5DA62646001D2A5AFCC7FB11CE9B" box="[1039,1211,868,920]" gridcol="4" gridrow="4" pageId="6" pageNumber="180">racemiform</td>
<td id="A05A5DA62646001D2ABDFCC7FA3BCE9B" box="[1256,1425,868,920]" gridcol="5" gridrow="4" pageId="6" pageNumber="180">racemiform</td>
</tr>
<tr id="E38B34DA2646001D2ECAFC63FA3BC90C" box="[159,1425,960,1039]" gridrow="5" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFC63FEF9C90C" box="[159,339,960,1039]" gridcol="0" gridrow="5" pageId="6" pageNumber="180">Flowering pedicels length (cm)</th>
<td id="A05A5DA62646001D2FD7FC63FD84C90C" box="[386,558,960,1039]" gridcol="1" gridrow="5" pageId="6" pageNumber="180">0.7–1</td>
<td id="A05A5DA62646001D2C0EFC63FCACC90C" box="[603,774,960,1039]" gridcol="2" gridrow="5" pageId="6" pageNumber="180">0.3–0.6</td>
<td id="A05A5DA62646001D2D60FC63FC75C90C" box="[821,991,960,1039]" gridcol="3" gridrow="5" pageId="6" pageNumber="180">(0.3–)0.5–0.6 (–0.7)</td>
<td id="A05A5DA62646001D2A5AFC63FB11C90C" box="[1039,1211,960,1039]" gridcol="4" gridrow="5" pageId="6" pageNumber="180">ca. 0.2</td>
<td id="A05A5DA62646001D2ABDFC63FA3BC90C" box="[1256,1425,960,1039]" gridcol="5" gridrow="5" pageId="6" pageNumber="180">0.1–0.2(–0.5)</td>
</tr>
<tr id="E38B34DA2646001D2ECAFB9BFA3BC98A" box="[159,1425,1080,1161]" gridrow="6" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFB9BFEF9C98A" box="[159,339,1080,1161]" gridcol="0" gridrow="6" pageId="6" pageNumber="180">Calyx morphology</th>
<td id="A05A5DA62646001D2FD7FB9BFD84C98A" box="[386,558,1080,1161]" gridcol="1" gridrow="6" pageId="6" pageNumber="180">cupular, 5-dentate</td>
<td id="A05A5DA62646001D2C0EFB9BFCACC98A" box="[603,774,1080,1161]" gridcol="2" gridrow="6" pageId="6" pageNumber="180">cupular, entire</td>
<td id="A05A5DA62646001D2D60FB9BFC75C98A" box="[821,991,1080,1161]" gridcol="3" gridrow="6" pageId="6" pageNumber="180">cupular, slightly 5-dentate</td>
<td id="A05A5DA62646001D2A5AFB9BFB11C98A" box="[1039,1211,1080,1161]" gridcol="4" gridrow="6" pageId="6" pageNumber="180">cupular, slightly 5-dentate</td>
<td id="A05A5DA62646001D2ABDFB9BFA3BC98A" box="[1256,1425,1080,1161]" gridcol="5" gridrow="6" pageId="6" pageNumber="180">cupular, obviously 5- lobed</td>
</tr>
<tr id="E38B34DA2646001D2ECAFB12FA3BC801" box="[159,1425,1201,1282]" gridrow="7" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFB12FEF9C801" box="[159,339,1201,1282]" gridcol="0" gridrow="7" pageId="6" pageNumber="180">Corolla symmetry</th>
<td id="A05A5DA62646001D2FD7FB12FD84C801" box="[386,558,1201,1282]" gridcol="1" gridrow="7" pageId="6" pageNumber="180">actinomorphic</td>
<td id="A05A5DA62646001D2C0EFB12FCACC801" box="[603,774,1201,1282]" gridcol="2" gridrow="7" pageId="6" pageNumber="180">probably actinomorphic, not verified here</td>
<td id="A05A5DA62646001D2D60FB12FC75C801" box="[821,991,1201,1282]" gridcol="3" gridrow="7" pageId="6" pageNumber="180">(actinomorphic)</td>
<td id="A05A5DA62646001D2A5AFB12FB11C801" box="[1039,1211,1201,1282]" gridcol="4" gridrow="7" pageId="6" pageNumber="180">actinomorphic</td>
<td id="A05A5DA62646001D2ABDFB12FA3BC801" box="[1256,1425,1201,1282]" gridcol="5" gridrow="7" pageId="6" pageNumber="180">actinomorphic (sub- actinomorphic)</td>
</tr>
<tr id="E38B34DA2646001D2ECAFA89FA3BC879" box="[159,1425,1322,1402]" gridrow="8" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFA89FEF9C879" box="[159,339,1322,1402]" gridcol="0" gridrow="8" pageId="6" pageNumber="180">Anthers morphology</th>
<td id="A05A5DA62646001D2FD7FA89FD84C879" box="[386,558,1322,1402]" gridcol="1" gridrow="8" pageId="6" pageNumber="180">lanceolate, thecae basally and apically sterile</td>
<td id="A05A5DA62646001D2C0EFA89FCACC879" box="[603,774,1322,1402]" gridcol="2" gridrow="8" pageId="6" pageNumber="180">lanceolate, thecae apically sterile</td>
<td id="A05A5DA62646001D2D60FA89FC75C879" box="[821,991,1322,1402]" gridcol="3" gridrow="8" pageId="6" pageNumber="180">lanceolate, thecae basally and apically sterile</td>
<td id="A05A5DA62646001D2A5AFA89FB11C879" box="[1039,1211,1322,1402]" gridcol="4" gridrow="8" pageId="6" pageNumber="180">lanceolate, thecae apically sterile</td>
<td id="A05A5DA62646001D2ABDFA89FA3BC879" box="[1256,1425,1322,1402]" gridcol="5" gridrow="8" pageId="6" pageNumber="180">lanceolate, sterile portions not verified here</td>
</tr>
<tr id="E38B34DA2646001D2ECAFA01FA3BCB13" box="[159,1425,1442,1552]" gridrow="9" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAFA01FEF9CB13" box="[159,339,1442,1552]" gridcol="0" gridrow="9" pageId="6" pageNumber="180">Ovary morphology and indumentum</th>
<td id="A05A5DA62646001D2FD7FA01FD84CB13" box="[386,558,1442,1552]" gridcol="1" gridrow="9" pageId="6" pageNumber="180">umbilicate, glabrous</td>
<td id="A05A5DA62646001D2C0EFA01FCACCB13" box="[603,774,1442,1552]" gridcol="2" gridrow="9" pageId="6" pageNumber="180">umbilicate, glabrous</td>
<td id="A05A5DA62646001D2D60FA01FC75CB13" box="[821,991,1442,1552]" gridcol="3" gridrow="9" pageId="6" pageNumber="180">umbilicate, glabrous (“cano- puberulous”)</td>
<td id="A05A5DA62646001D2A5AFA01FB11CB13" box="[1039,1211,1442,1552]" gridcol="4" gridrow="9" pageId="6" pageNumber="180">probably umbilicate, not verified here; densely strigulose</td>
<td id="A05A5DA62646001D2ABDFA01FA3BCB13" box="[1256,1425,1442,1552]" gridcol="5" gridrow="9" pageId="6" pageNumber="180">(umbilicate), glabrous</td>
</tr>
<tr id="E38B34DA2646001D2ECAF99BFA3BCB68" box="[159,1425,1592,1643]" gridrow="10" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAF99BFEF9CB68" box="[159,339,1592,1643]" gridcol="0" gridrow="10" pageId="6" pageNumber="180">Mericarps size (cm)</th>
<td id="A05A5DA62646001D2FD7F99BFD84CB68" box="[386,558,1592,1643]" gridcol="1" gridrow="10" pageId="6" pageNumber="180">ca. 1.2 × 0.9–1.1</td>
<td id="A05A5DA62646001D2C0EF99BFCACCB68" box="[603,774,1592,1643]" gridcol="2" gridrow="10" pageId="6" pageNumber="180">2–2.5 × 1.7–2</td>
<td id="A05A5DA62646001D2D60F99BFC75CB68" box="[821,991,1592,1643]" gridcol="3" gridrow="10" pageId="6" pageNumber="180">1.6(–2.2) × 1 (–1.5)</td>
<td id="A05A5DA62646001D2A5AF99BFB11CB68" box="[1039,1211,1592,1643]" gridcol="4" gridrow="10" pageId="6" pageNumber="180">1.5–2 × 1.5–1.8</td>
<td id="A05A5DA62646001D2ABDF99BFA3BCB68" box="[1256,1425,1592,1643]" gridcol="5" gridrow="10" pageId="6" pageNumber="180">1–1.3 × 1–1.1</td>
</tr>
<tr id="E38B34DA2646001D2ECAF930FA3BCBC4" box="[159,1425,1683,1735]" gridrow="11" pageId="6" pageNumber="180">
<th id="A05A5DA62646001D2ECAF930FEF9CBC4" box="[159,339,1683,1735]" gridcol="0" gridrow="11" pageId="6" pageNumber="180">Cotyledons morphology</th>
<td id="A05A5DA62646001D2FD7F930FD84CBC4" box="[386,558,1683,1735]" gridcol="1" gridrow="11" pageId="6" pageNumber="180">plano-convex</td>
<td id="A05A5DA62646001D2C0EF930FCACCBC4" box="[603,774,1683,1735]" gridcol="2" gridrow="11" pageId="6" pageNumber="180">plano-convex</td>
<td id="A05A5DA62646001D2D60F930FC75CBC4" box="[821,991,1683,1735]" gridcol="3" gridrow="11" pageId="6" pageNumber="180">plicate- conduplicate</td>
<td id="A05A5DA62646001D2A5AF930FB11CBC4" box="[1039,1211,1683,1735]" gridcol="4" gridrow="11" pageId="6" pageNumber="180">plano-convex</td>
<td id="A05A5DA62646001D2ABDF930FA3BCBC4" box="[1256,1425,1683,1735]" gridcol="5" gridrow="11" pageId="6" pageNumber="180">plano-convex c</td>
</tr>
</table>
</paragraph>
<tableNote id="A05D37162646FFE42EDDF942FD41CAAF" pageId="6" pageNumber="180" targetBox="[159,1425,572,1735]" targetPageId="6">
<paragraph id="5D0436982646FFE42EDDF942FE0CCA18" blockId="6.[136,1452,1761,1964]" pageId="6" pageNumber="180">
<superScript id="AACE9BD02646FFE42EDDF942FF40CBF4" attach="right" box="[136,234,1761,1783]" fontSize="5" pageId="6" pageNumber="180">aMeasures</superScript>
or features in parentheses were taken from synonyms protologues (
<bibRefCitation id="392A4B692646FFE42D3AF941FC4DCBF4" author="Ducke, W. A." box="[879,999,1762,1784]" pageId="6" pageNumber="180" pagination="3 - 269" refId="ref5119" refString="Ducke, W. A. (1922) Plantes nouvelles ou peu connues de la region amazonienne (II partie). Archivos do Jardim Botanico do Rio de Janeiro 3: 3 - 269." type="journal article" year="1922">Ducke 1922</bibRefCitation>
, 
<bibRefCitation id="392A4B692646FFE42DA4F941FB19CBF4" author="Porter, D. M. &amp; Elias, T. S." box="[1009,1203,1762,1784]" pageId="6" pageNumber="180" pagination="123 - 164" refId="ref6740" refString="Porter, D. M. &amp; Elias, T. S. (1979) Flora of Panama Part VI. Family 89. Rutaceae. Annals of the Missouri Botanical Garden 66 (2): 123 - 164. https: // doi. org / 10.2307 / 2398906" type="journal article" year="1979">Porter &amp; Elias 1979</bibRefCitation>
). For complete synonymy see 
<bibRefCitation id="392A4B692646FFE42EF8F8A5FE0BCA18" author="Kallunki, J. A. &amp; Pirani, J. R." box="[173,417,1798,1820]" pageId="6" pageNumber="180" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998)</bibRefCitation>
.
</paragraph>
<paragraph id="5D0436982646FFE42EDDF88AFB76CA3C" blockId="6.[136,1452,1761,1964]" box="[136,1244,1833,1856]" pageId="6" pageNumber="180">
<superScript id="AACE9BD02646FFE42EDDF88AFF41CA3C" attach="right" box="[136,235,1833,1855]" fontSize="5" pageId="6" pageNumber="180">bMeasures</superScript>
or features in parentheses were taken from synonyms protologues (
<bibRefCitation id="392A4B692646FFE42D21F889FB8ACA3C" author="Cuatrecasas, J." box="[884,1056,1833,1855]" pageId="6" pageNumber="180" pagination="464 - 488" refId="ref5034" refString="Cuatrecasas, J. (1952) Notas a la Flora de Colombia XII. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales 8 (32): 464 - 488." type="journal article" year="1952">Cuatrecasas 1952</bibRefCitation>
, 
<bibRefCitation id="392A4B692646FFE42A7FF889FB7BCA3C" author="McPherson, G." box="[1066,1233,1834,1856]" pageId="6" pageNumber="180" pagination="373 - 378" refId="ref6632" refString="McPherson, G. (1988) New and noteworthy taxa from Panama. Annals of the Missouri Botanical Garden 75 (1): 373 - 378." type="journal article" year="1988">McPherson 1988</bibRefCitation>
).
</paragraph>
<paragraph id="5D0436982646FFE42EDDF8EEFD41CAAF" blockId="6.[136,1452,1761,1964]" pageId="6" pageNumber="180">
<superScript id="AACE9BD02646FFE42EDDF8EEFF48CA67" attach="right" box="[136,226,1869,1892]" fontSize="5" pageId="6" pageNumber="180">cKallunki</superScript>
&amp; Pirani (1998) noted the cotyledons of 
<taxonomicName id="9ABB4D1B2646FFE42CD7F8EDFCABCA60" box="[642,769,1870,1891]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="6" pageNumber="180" phylum="Tracheophyta" rank="species" species="toxicarius">
<emphasis id="6FCFEA8A2646FFE42CD7F8EDFCABCA60" box="[642,769,1870,1891]" italics="true" pageId="6" pageNumber="180">C. toxicarius</emphasis>
</taxonomicName>
as “conduplicate, not plicate and thick”, clearly differencing it from plicate-conduplicate “conduplicate, plicate and thin” or plano-convex “plano-convex and thick”. Subsequently, 
<bibRefCitation id="392A4B692646FFE42AFBF8D1FADBCA84" author="Groppo, M. &amp; Bruniera, C. P. &amp; Ferreira, P. L. &amp; Ferreira, C. &amp; Pirani, J. R. &amp; Kallunki, J. A." box="[1198,1393,1906,1927]" pageId="6" pageNumber="180" refId="ref5499" refString="Groppo, M., Bruniera, C. P., Ferreira, P. L., Ferreira, C., Pirani, J. R. &amp; Kallunki, J. A. (2017) Phylogeny and delimitation of Galipeeae (Rutaceae, Sapindales) based on molecular data: insights on the evolution of zygomorphic flowers and staminodes. XIX International Botanical Congress, Shenzhen." type="book" year="2017">
Groppo 
<emphasis id="6FCFEA8A2646FFE42AA8F8D1FA87CA84" box="[1277,1325,1906,1927]" italics="true" pageId="6" pageNumber="180">et al.</emphasis>
(2017)
</bibRefCitation>
noted it under the character state “cotyledons plane or plane-convex”.
</paragraph>
</tableNote>
<paragraph id="5D0436982647FFE52EE8FF3DFD19CC4B" blockId="7.[136,1452,158,1517]" pageId="7" pageNumber="181">
The filaments of 
<taxonomicName id="9ABB4D1B2647FFE52FD2FF3DFD03CDBB" box="[391,681,158,184]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2647FFE52FD2FF3DFD03CDBB" box="[391,681,158,184]" italics="true" pageId="7" pageNumber="181">Conchocarpus cardenasii</emphasis>
</taxonomicName>
are here described as adnate to the corolla only by a small dorsal portion at the tube throat, because they are strongly united and hardly detachable without being torn apart. 
<bibRefCitation id="392A4B692647FFE52B05FF61FEBECC03" author="El Ottra, J. H. L. &amp; Demarco, D. &amp; Pirani, J. R." pageId="7" pageNumber="181" pagination="30 - 101" refId="ref5156" refString="El Ottra, J. H. L., Demarco, D. &amp; Pirani, J. R. (2019) Comparative floral structure and evolution in Galipeinae (Galipeeae: Rutaceae) and its implications at different systematic levels. Botanical Journal of the Linnean Society 191: 30 - 101. https: // doi. org / 10.1093 / botlinnean / boz 029" type="journal article" year="2019">
El Ottra 
<emphasis id="6FCFEA8A2647FFE52EDDFF44FF6BCC03" box="[136,193,230,256]" italics="true" pageId="7" pageNumber="181">et al.</emphasis>
(2019)
</bibRefCitation>
recently reported a kind of postgenital floral fusion in 
<taxonomicName id="9ABB4D1B2647FFE52DD1FF45FC57CC03" box="[900,1021,230,256]" class="Magnoliopsida" family="Rutaceae" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="183" phylum="Tracheophyta" rank="subTribe" subTribe="Galipeinae">Galipeinae</taxonomicName>
consisting in the union by papillae or cuticular projections; whether this is the situation responsible for the structure here observed in 
<taxonomicName id="9ABB4D1B2647FFE52AE3FEA8FAE6CC27" box="[1206,1356,266,293]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2647FFE52AE3FEA8FAE6CC27" box="[1206,1356,266,293]" italics="true" pageId="7" pageNumber="181">C. cardenasii</emphasis>
</taxonomicName>
remains a matter depending on further anatomical studies.
</paragraph>
<paragraph id="5D0436982647FFE52EE8FEF1FF7BCE43" blockId="7.[136,1452,158,1517]" pageId="7" pageNumber="181">
The cotyledons plano-convex in 
<taxonomicName id="9ABB4D1B2647FFE52C63FEF1FCF2CC6F" box="[566,856,338,364]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2647FFE52C63FEF1FCF2CC6F" box="[566,856,338,364]" italics="true" pageId="7" pageNumber="181">Conchocarpus cardenasii</emphasis>
</taxonomicName>
, 
<taxonomicName id="9ABB4D1B2647FFE52D3DFEF0FC4BCC6F" box="[872,993,338,365]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2647FFE52D3DFEF0FC4BCC6F" box="[872,993,338,365]" italics="true" pageId="7" pageNumber="181">C. grandis</emphasis>
</taxonomicName>
and 
<taxonomicName id="9ABB4D1B2647FFE52A49FEF0FB17CC6F" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[1052,1213,339,365]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2647FFE52A49FEF0FB17CC6F" box="[1052,1213,339,365]" italics="true" pageId="7" pageNumber="181">C. jirajaranus</emphasis>
</taxonomicName>
are morphologically atypical for 
<emphasis id="6FCFEA8A2647FFE52F44FED5FDE0CC93" box="[273,586,374,400]" italics="true" pageId="7" pageNumber="181">
<taxonomicName id="9ABB4D1B2647FFE52F44FED5FE1ECC93" authorityName="J. C. Mikan" authorityYear="1820" box="[273,436,374,400]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus" sensu="stricto">Conchocarpus</taxonomicName>
<taxonomicNameLabel id="74FC57F12647FFE52FE9FED4FDE0CC93" box="[444,586,375,400]" pageId="7" pageNumber="181" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
, and its taxonomic relevance could be tested in further phylogenetic studies. Similarly, other species currently under 
<taxonomicName id="9ABB4D1B2647FFE52C12FE39FD40CCB7" authorityName="J. C. Mikan" authorityYear="1820" box="[583,746,410,436]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus">
<emphasis id="6FCFEA8A2647FFE52C12FE39FD40CCB7" box="[583,746,410,436]" italics="true" pageId="7" pageNumber="181">Conchocarpus</emphasis>
</taxonomicName>
have uncommon features that could support new classification proposals. For example, 
<taxonomicName id="9ABB4D1B2647FFE52FFDFE1DFD5ECCDB" box="[424,756,446,472]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="nicaraguensis">
<emphasis id="6FCFEA8A2647FFE52FFDFE1DFD5ECCDB" box="[424,756,446,472]" italics="true" pageId="7" pageNumber="181">Conchocarpus nicaraguensis</emphasis>
</taxonomicName>
[
<bibRefCitation id="392A4B692647FFE52D50FE1DFB63CCDB" author="Standley, P. C. &amp; Williams, L. O." box="[773,1225,446,473]" pageId="7" pageNumber="181" pagination="187 - 220" refId="ref6876" refString="Standley, P. C. &amp; Williams, L. O. (1953) Plantae Centrali-Americanae V. Ceiba 3 (3): 187 - 220." type="journal article" year="1953">Standley &amp; Williams (1953: 206–207)</bibRefCitation>
] 
<bibRefCitation id="392A4B692647FFE52A8FFE1DFEA1CCFF" author="Kallunki, J. A. &amp; Pirani, J. R." pageId="7" pageNumber="181" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998: 314)</bibRefCitation>
and 
<taxonomicName id="9ABB4D1B2647FFE52F6AFE41FDC8CCFF" box="[319,610,482,508]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="ucayalinus">
<emphasis id="6FCFEA8A2647FFE52F6AFE41FDC8CCFF" box="[319,610,482,508]" italics="true" pageId="7" pageNumber="181">Conchocarpus ucayalinus</emphasis>
</taxonomicName>
[
<bibRefCitation id="392A4B692647FFE52C3BFE41FCD1CCFE" author="Huber, J." box="[622,891,482,509]" pageId="7" pageNumber="181" pagination="510 - 619" refId="ref5893" refString="Huber, J. (1906) Materiaes para a flora amazonica VI. Plantas vasculares colligidas e observadas no baixo Ucayali e no Pampa del Sacramento, nos mezes de outubro a dezembro de 1898. Boletim do Museo Goeldi de Historia Natural e Ethnographia 4: 510 - 619." type="journal article" year="1906">Huber (1906: 573–574)</bibRefCitation>
] 
<bibRefCitation id="392A4B692647FFE52DD2FE41FB7DCCFF" author="Kallunki, J. A. &amp; Pirani, J. R." box="[903,1239,482,508]" pageId="7" pageNumber="181" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998: 323)</bibRefCitation>
from northwestern South America, Central America and western Amazonia, resemble at first glance species of 
<taxonomicName id="9ABB4D1B2647FFE52AF8FDA5FAA6CF23" box="[1197,1292,518,544]" class="Magnoliopsida" family="Rutaceae" genus="Dryades" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus">
<emphasis id="6FCFEA8A2647FFE52AF8FDA5FAA6CF23" box="[1197,1292,518,544]" italics="true" pageId="7" pageNumber="181">Dryades</emphasis>
</taxonomicName>
because their calyx lobes imbricate and overlapping even after anthesis. However, not all diagnostic features of 
<taxonomicName id="9ABB4D1B2647FFE52A99FD89FA81CF47" box="[1228,1323,554,580]" class="Magnoliopsida" family="Rutaceae" genus="Dryades" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus">
<emphasis id="6FCFEA8A2647FFE52A99FD89FA81CF47" box="[1228,1323,554,580]" italics="true" pageId="7" pageNumber="181">Dryades</emphasis>
</taxonomicName>
are present in 
<taxonomicName id="9ABB4D1B2647FFE52EF3FDECFECFCF6B" box="[166,357,591,617]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="nicaraguensis">
<emphasis id="6FCFEA8A2647FFE52EF3FDECFECFCF6B" box="[166,357,591,617]" italics="true" pageId="7" pageNumber="181">C. nicaraguensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="9ABB4D1B2647FFE52FCAFDECFD90CF6B" box="[415,570,590,617]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="ucayalinus">
<emphasis id="6FCFEA8A2647FFE52FCAFDECFD90CF6B" box="[415,570,590,617]" italics="true" pageId="7" pageNumber="181">C. ucayalinus</emphasis>
</taxonomicName>
and these species have not been included in phylogenetic analyses. Added to the morphological relevance of the cotyledons in the classification of 
<emphasis id="6FCFEA8A2647FFE52DCCFDD1FB1CCF8F" box="[921,1206,626,652]" italics="true" pageId="7" pageNumber="181">
<taxonomicName id="9ABB4D1B2647FFE52DCCFDD1FB96CF8F" authorityName="J. C. Mikan" authorityYear="1820" box="[921,1084,626,652]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus" sensu="lato">Conchocarpus</taxonomicName>
<taxonomicNameLabel id="74FC57F12647FFE52A11FDD0FB1CCF8F" box="[1092,1206,626,652]" pageId="7" pageNumber="181" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
, 
<bibRefCitation id="392A4B692647FFE52A91FDD1FA01CF8F" author="Groppo, M. &amp; Bruniera, C. P. &amp; Ferreira, P. L. &amp; Ferreira, C. &amp; Pirani, J. R. &amp; Kallunki, J. A." box="[1220,1451,626,652]" pageId="7" pageNumber="181" refId="ref5499" refString="Groppo, M., Bruniera, C. P., Ferreira, P. L., Ferreira, C., Pirani, J. R. &amp; Kallunki, J. A. (2017) Phylogeny and delimitation of Galipeeae (Rutaceae, Sapindales) based on molecular data: insights on the evolution of zygomorphic flowers and staminodes. XIX International Botanical Congress, Shenzhen." type="book" year="2017">
Groppo 
<emphasis id="6FCFEA8A2647FFE52B74FDD0FAF0CF8F" box="[1313,1370,626,652]" italics="true" pageId="7" pageNumber="181">et al.</emphasis>
(2017)
</bibRefCitation>
found 
<taxonomicName id="9ABB4D1B2647FFE52E84FD35FE41CFB3" box="[209,491,662,688]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="toxicarius">
<emphasis id="6FCFEA8A2647FFE52E84FD35FE41CFB3" box="[209,491,662,688]" italics="true" pageId="7" pageNumber="181">Conchocarpus toxicarius</emphasis>
</taxonomicName>
[Spruce ex 
<bibRefCitation id="392A4B692647FFE52C27FD35FC2BCFB2" author="Engler, H. G. A." box="[626,897,662,689]" pageId="7" pageNumber="181" pagination="77 - 196" refId="ref5293" refString="Engler, H. G. A. (1874) Rutaceae. In: Martius, C. F. P. &amp; Eichler, A. G. (eds.) Flora Brasiliensis, enumeratio plantarum in Brasilia hactenus detectarum 12 (2). Lipsiae [Liepzig]: Apud Frid. Fleischer in Comm. pp. 77 - 196." type="book chapter" year="1874">Engler (1874: 114–115)</bibRefCitation>
] 
<bibRefCitation id="392A4B692647FFE52DDBFD35FB4FCFB3" author="Kallunki, J. A. &amp; Pirani, J. R." box="[910,1253,662,688]" pageId="7" pageNumber="181" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998: 322)</bibRefCitation>
, phylogenetically segregated from 
<emphasis id="6FCFEA8A2647FFE52F11FD19FDB8CFD7" box="[324,530,698,724]" italics="true" pageId="7" pageNumber="181">
<taxonomicName id="9ABB4D1B2647FFE52F11FD19FE4DCFD7" authorityName="J. C. Mikan" authorityYear="1820" box="[324,487,698,724]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus" sensu="stricto">Conchocarpus</taxonomicName>
<taxonomicNameLabel id="74FC57F12647FFE52FBAFD18FDB8CFD7" box="[495,530,699,724]" pageId="7" pageNumber="181" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="9ABB4D1B2647FFE52C1CFD19FD02CFD7" box="[585,680,698,724]" class="Magnoliopsida" family="Rutaceae" genus="Dryades" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus">
<emphasis id="6FCFEA8A2647FFE52C1CFD19FD02CFD7" box="[585,680,698,724]" italics="true" pageId="7" pageNumber="181">Dryades</emphasis>
</taxonomicName>
(this genus not segregated yet in 
<bibRefCitation id="392A4B692647FFE52A4AFD19FB5DCFD7" author="Groppo, M. &amp; Bruniera, C. P. &amp; Ferreira, P. L. &amp; Ferreira, C. &amp; Pirani, J. R. &amp; Kallunki, J. A." box="[1055,1271,698,724]" pageId="7" pageNumber="181" refId="ref5499" refString="Groppo, M., Bruniera, C. P., Ferreira, P. L., Ferreira, C., Pirani, J. R. &amp; Kallunki, J. A. (2017) Phylogeny and delimitation of Galipeeae (Rutaceae, Sapindales) based on molecular data: insights on the evolution of zygomorphic flowers and staminodes. XIX International Botanical Congress, Shenzhen." type="book" year="2017">
Groppo 
<emphasis id="6FCFEA8A2647FFE52A29FD18FB1ECFD7" box="[1148,1204,698,724]" italics="true" pageId="7" pageNumber="181">et al.</emphasis>
2017
</bibRefCitation>
), besides noting it under the character state “cotyledons plane or plane-convex” (see 
<tableCitation id="103903232647FFE52DDDFD7DFC70CFFB" box="[904,986,734,760]" captionStart="TABLE 1" captionStartId="6.[136,216,522,543]" captionTargetPageId="6" captionText="TABLE 1. Comparison of the similar species to Conchocarpus cardenasii." httpUri="http://table.plazi.org/id/09C466102646FFE42EDDFDA9FCFDCF1C" pageId="7" pageNumber="181" tableUuid="09C466102646FFE42EDDFDA9FCFDCF1C">Table 1</tableCitation>
), which suggest that this species perhaps might be recognized under another genus when a more complete sampling becomes available (Groppo, pers. comm. 2018).
</paragraph>
<paragraph id="5D0436982647FFE52EE8FCE9FB70C9F3" blockId="7.[136,1452,158,1517]" pageId="7" pageNumber="181">
Interestingly, although 
<taxonomicName id="9ABB4D1B2647FFE52F85FCE9FD58CE67" box="[464,754,842,868]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="toxicarius">
<emphasis id="6FCFEA8A2647FFE52F85FCE9FD58CE67" box="[464,754,842,868]" italics="true" pageId="7" pageNumber="181">Conchocarpus toxicarius</emphasis>
</taxonomicName>
can be distinguished by its leaves 5–7-foliolate from 
<taxonomicName id="9ABB4D1B2647FFE52BC7FCE8FF57CE8B" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2647FFE52BC7FCE8FF57CE8B" italics="true" pageId="7" pageNumber="181">C. cardenasii</emphasis>
</taxonomicName>
, 
<taxonomicName id="9ABB4D1B2647FFE52F59FCCCFE2ECE8B" box="[268,388,878,905]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="6FCFEA8A2647FFE52F59FCCCFE2ECE8B" box="[268,388,878,905]" italics="true" pageId="7" pageNumber="181">C. grandis</emphasis>
</taxonomicName>
and 
<taxonomicName id="9ABB4D1B2647FFE52FE9FCCCFDF1CE8B" authorityName="Kallunki &amp; W. Meier" authorityYear="2019" box="[444,603,879,905]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="jirajaranus">
<emphasis id="6FCFEA8A2647FFE52FE9FCCCFDF1CE8B" box="[444,603,879,905]" italics="true" pageId="7" pageNumber="181">C. jirajaranus</emphasis>
</taxonomicName>
, all four species share their androecium with all stamens fertile, lanceolate anthers and in a 
<emphasis id="6FCFEA8A2647FFE52F1EFC30FE15CEAF" box="[331,447,914,940]" italics="true" pageId="7" pageNumber="181">sensu lato</emphasis>
, their actinomorphic corolla, umbilicate ovary and plano-convex cotyledons (see 
<tableCitation id="103903232647FFE52B38FC31FF3FCED3" captionStart="TABLE 1" captionStartId="6.[136,216,522,543]" captionTargetPageId="6" captionText="TABLE 1. Comparison of the similar species to Conchocarpus cardenasii." httpUri="http://table.plazi.org/id/09C466102646FFE42EDDFDA9FCFDCF1C" pageId="7" pageNumber="181" tableUuid="09C466102646FFE42EDDFDA9FCFDCF1C">Table 1</tableCitation>
). Remarkably, one of the 
<specimenCount id="4BBDFD112647FFE52F9DFC14FDF4CED2" box="[456,606,951,977]" pageId="7" pageNumber="181" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
of 
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<emphasis id="6FCFEA8A2647FFE52CDCFC14FCB7CED3" box="[649,797,951,977]" italics="true" pageId="7" pageNumber="181">C. toxicarius</emphasis>
</taxonomicName>
belongs to 
<taxonomicName id="9ABB4D1B2647FFE52DFDFC15FB46CED3" box="[936,1260,950,976]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="6FCFEA8A2647FFE52DFDFC15FB46CED3" box="[936,1260,950,976]" italics="true" pageId="7" pageNumber="181">Conchocarpus heterophyllus</emphasis>
</taxonomicName>
[A. 
<bibRefCitation id="392A4B692647FFE52B41FC15FEA4CEF7" author="Saint-Hilaire, A. F. C. P." pageId="7" pageNumber="181" pagination="129 - 133" refId="ref6824" refString="Saint-Hilaire, A. F. C. P. (1823) Description de principaux genres nouveaux et de especes nouvelles de la Flore de Bresil, cites das le Memoire sur le Gynobase. Bulletin des Sciences, par la Societe Philomatique de Paris 1823: 129 - 133." type="journal article" year="1823">Saint-Hilaire (1823: 131)</bibRefCitation>
] 
<bibRefCitation id="392A4B692647FFE52F48FC79FDDFCEF7" author="Kallunki, J. A. &amp; Pirani, J. R." box="[285,629,986,1012]" pageId="7" pageNumber="181" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998: 300)</bibRefCitation>
, an inconsistence solved by 
<bibRefCitation id="392A4B692647FFE52DE3FC79FB7FCEF7" author="Kallunki, J. A. &amp; Pirani, J. R." box="[950,1237,986,1012]" pageId="7" pageNumber="181" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998)</bibRefCitation>
; thus, although the protologue of 
<taxonomicName id="9ABB4D1B2647FFE52F7CFC5CFE16C91B" box="[297,444,1023,1049]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="toxicarius">
<emphasis id="6FCFEA8A2647FFE52F7CFC5CFE16C91B" box="[297,444,1023,1049]" italics="true" pageId="7" pageNumber="181">C. toxicarius</emphasis>
</taxonomicName>
indicated only three fertile stamens, 
<bibRefCitation id="392A4B692647FFE52D37FC5DFB2BC91B" author="Kallunki, J. A. &amp; Pirani, J. R." box="[866,1153,1022,1048]" pageId="7" pageNumber="181" pagination="257 - 334" refId="ref6298" refString="Kallunki, J. A. &amp; Pirani, J. R. (1998) Synopses of Angostura Roem. &amp; Schult. and Conchocarpus J. C. Mikan (Rutaceae). Kew Bulletin 53 (2): 257 - 334. https: // doi. org / 10.2307 / 4114501" type="journal article" year="1998">Kallunki &amp; Pirani (1998)</bibRefCitation>
indicated that it have five stamens fertile, which is according to the original illustration at the complete flower drawings [see 
<bibRefCitation id="392A4B692647FFE52B58FB81FA02C93F" author="Engler, H. G. A." box="[1293,1448,1058,1084]" pageId="7" pageNumber="181" pagination="77 - 196" refId="ref5293" refString="Engler, H. G. A. (1874) Rutaceae. In: Martius, C. F. P. &amp; Eichler, A. G. (eds.) Flora Brasiliensis, enumeratio plantarum in Brasilia hactenus detectarum 12 (2). Lipsiae [Liepzig]: Apud Frid. Fleischer in Comm. pp. 77 - 196." type="book chapter" year="1874">Engler (1874</bibRefCitation>
: t.24, fig. 1) and the 
<typeStatus id="8200883A2647FFE52F2AFBE5FE4CC963" box="[383,486,1094,1120]" pageId="7" pageNumber="181">lectotype</typeStatus>
<emphasis id="6FCFEA8A2647FFE52FA5FBE4FD2AC963" box="[496,640,1094,1121]" italics="true" pageId="7" pageNumber="181">Spruce 2171</emphasis>
(K barcode 000531071 [digital image!]); note the original illustration at stamens details drawings showing those lanceolate anthers of 
<taxonomicName id="9ABB4D1B2647FFE52D3FFBC8FC57C987" box="[874,1021,1131,1157]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="toxicarius">
<emphasis id="6FCFEA8A2647FFE52D3FFBC8FC57C987" box="[874,1021,1131,1157]" italics="true" pageId="7" pageNumber="181">C. toxicarius</emphasis>
</taxonomicName>
as staminodes and probably, those of 
<taxonomicName id="9ABB4D1B2647FFE52EDDFB2CFEE8C9AB" box="[136,322,1166,1193]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="6FCFEA8A2647FFE52EDDFB2CFEE8C9AB" box="[136,322,1166,1193]" italics="true" pageId="7" pageNumber="181">C. heterophyllus</emphasis>
</taxonomicName>
as fertile stamens. Currently, the recognition of other segregate groups from 
<taxonomicName id="9ABB4D1B2647FFE52AEAFB2DFAC8C9AB" authorityName="J. C. Mikan" authorityYear="1820" box="[1215,1378,1166,1192]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus">
<emphasis id="6FCFEA8A2647FFE52AEAFB2DFAC8C9AB" box="[1215,1378,1166,1192]" italics="true" pageId="7" pageNumber="181">Conchocarpus</emphasis>
</taxonomicName>
is not definitive considering the lack of sampling species with deviating morphological combinations in molecular analyses (
<emphasis id="6FCFEA8A2647FFE52EC4FB74FF10C9F3" box="[145,186,1239,1264]" italics="true" pageId="7" pageNumber="181">e.g.</emphasis>
, actinomorphic corolla, five stamens fertile, lanceolate anthers and plano-convex cotyledons).
</paragraph>
<paragraph id="5D0436982647FFE52EE8FB59FE97C8EF" blockId="7.[136,1452,158,1517]" pageId="7" pageNumber="181">
Despite of the fact that 
<taxonomicName id="9ABB4D1B2647FFE52F86FB59FD5FC817" box="[467,757,1274,1300]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2647FFE52F86FB59FD5FC817" box="[467,757,1274,1300]" italics="true" pageId="7" pageNumber="181">Conchocarpus cardenasii</emphasis>
</taxonomicName>
could undergo nomenclatural and taxonomic changes if the splitting of 
<taxonomicName id="9ABB4D1B2647FFE52F5EFABDFE04C83B" authorityName="J. C. Mikan" authorityYear="1820" box="[267,430,1310,1336]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="genus">
<emphasis id="6FCFEA8A2647FFE52F5EFABDFE04C83B" box="[267,430,1310,1336]" italics="true" pageId="7" pageNumber="181">Conchocarpus</emphasis>
</taxonomicName>
is supported in future phylogenetic studies, its morphological features fits into the current circumscription of the genus. Considering the remarkably presence of cotyledons plano-convex and anthers basally sterile, 
<taxonomicName id="9ABB4D1B2647FFE52E89FAC4FEDEC883" box="[220,372,1382,1409]" class="Magnoliopsida" family="Rutaceae" genus="Conchocarpus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="181" phylum="Tracheophyta" rank="species" species="cardenasii">
<emphasis id="6FCFEA8A2647FFE52E89FAC4FEDEC883" box="[220,372,1382,1409]" italics="true" pageId="7" pageNumber="181">C. cardenasii</emphasis>
</taxonomicName>
may be a critical species to be included in future morphological and DNA systematics studies of the genus. Finally, given its condition of being a restricted endemic and endangered species, this formal taxonomic recognition is fundamental for supporting current and future conservation actions at the unique Cerros del Cuchillo remnant forests.
</paragraph>
</subSubSection>
</treatment>
</document>