163 lines
16 KiB
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163 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo" docDate="2022" docId="3E7DF42329B45868B4C3729423361A02" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Huancabamba Benedetti & Pinto-da-Rocha 2022, gen. nov." docType="treatment" docUuid="4A0804FA-3092-4585-A75A-51146B4ECB5B" docUuidSource="ZooBank" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>309</mods:start>
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<mods:end>388</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
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<mods:identifier type="ZooBank">D5C0468B99A14EF39237D9BC51A8BDA3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:4A0804FA-3092-4585-A75A-51146B4ECB5B" httpUri="http://treatment.plazi.org/id/3E7DF42329B45868B4C3729423361A02" lastPageNumber="309" pageId="0" pageNumber="309">
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<subSubSection pageId="0" pageNumber="309" type="nomenclature">
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<paragraph pageId="0" pageNumber="309">
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3.28.
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<taxonomicName LSID="http://zoobank.org/4A0804FA-3092-4585-A75A-51146B4ECB5B" authority="Benedetti & Pinto-da-Rocha, 2022" authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Insecta" family="Metasarcidae" genus="Huancabamba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huancabamba" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus" status="gen. nov.">Huancabamba</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="309">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="description">
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<paragraph pageId="0" pageNumber="309">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury & Maury, 1998; C I. dianae Kury & Maury, 1998; D I. ochoai Kury & Maury, 1998; E I. pictus Kury & Maury, 1998; F I. viracocha Kury & Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">Figs 4A</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male trochanter-patella / tibia IV of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha, dorsal and ventral view, respectively. A - B H. kubricki gen. et sp. nov.; C - D L. antonionii gen. et sp. nov.; E - F L. woodyalleni gen. et sp. nov.; G - H T. chaplini sp. nov.; I - J T. joseochoai sp. nov.; K - L T. scorsesei sp. nov.; M - N T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure11" httpUri="https://binary.pensoft.net/fig/729885" pageId="0" pageNumber="309">, 11A, B</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Penis of Huancabamba gen. nov. and Incasarcus, dorsal, lateral and ventral views, respectively. A - C H. kubricki gen. et sp. nov.; D - F I. ochoai Kury & Maury, 1998; G - I I. viracocha Kury & Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure18" httpUri="https://binary.pensoft.net/fig/729892" pageId="0" pageNumber="309">, 18A-C</figureCitation>
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Live specimens of Huancabamba gen. nov. and Lumieria gen. nov. A H. kubricki gen. nov. et sp. nov., male; B female; C L. antonionii gen. nov. sp. nov., male; D female; E L. woodyalleni gen. nov. sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure24" httpUri="https://binary.pensoft.net/fig/729898" pageId="0" pageNumber="309">, 24A, B</figureCitation>
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha in Peru. A H. kubricki gen. et sp. nov., L. antonionii gen. et sp. nov., T. chaplini sp. nov., T. scorsesei sp. nov., T. weyrauchi; B L. woodyalleni gen. et sp. nov., T. joseochoai sp. nov." figureDoi="10.3897/asp.80.e73829.figure29" httpUri="https://binary.pensoft.net/fig/729903" pageId="0" pageNumber="309">, 29A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="type species">
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<paragraph pageId="0" pageNumber="309">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName class="Insecta" family="Metasarcidae" genus="Huancabamba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huancabamba kubricki" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kubricki">
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<emphasis italics="true" pageId="0" pageNumber="309">Huancabamba kubricki</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">gen. et. sp. nov.</emphasis>
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by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="diagnosis">
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<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Insecta" family="Metasarcidae" genus="Huancabamba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huancabamba" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="309">Huancabamba</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">gen. nov.</emphasis>
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can be distinguished from all other
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<taxonomicName authorityName="Kury" authorityYear="1994" family="Metasarcidae" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="309" rank="family">Metasarcidae</taxonomicName>
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genera by the combination of following: Kappa-type DSS; male femur IV at least 1.6 longer than DS length and with low tubercles; ocularium with two low tubercles; area III with spines; coxa IV apex reaching area III; penis with more than 13 MS C; a dry-mark in depression of ocularium, carapace, and lateral region to ocularium.
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</paragraph>
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<caption doi="10.3897/asp.80.e73829.figure21" httpUri="https://binary.pensoft.net/fig/729895" pageId="0" pageNumber="309" start="Figure 21" startId="F21">
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<paragraph pageId="0" pageNumber="309">
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<emphasis bold="true" pageId="0" pageNumber="309">Figure 21.</emphasis>
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Penis of
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<taxonomicName authorityName="Roewer" authorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Tschaidicancha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tschaidicancha" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="309">Tschaidicancha</emphasis>
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</taxonomicName>
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, dorsal, lateral and ventral views, respectively.
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<emphasis bold="true" pageId="0" pageNumber="309">A</emphasis>
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-
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<emphasis bold="true" pageId="0" pageNumber="309">C</emphasis>
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<taxonomicName family="Metasarcidae" genus="T." kingdom="Animalia" lsidName="T. chaplini" order="Opiliones" pageId="0" pageNumber="309" rank="species" species="chaplini">
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<emphasis italics="true" pageId="0" pageNumber="309">T. chaplini</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
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;
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<emphasis bold="true" pageId="0" pageNumber="309">D</emphasis>
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-
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<emphasis bold="true" pageId="0" pageNumber="309">F</emphasis>
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<taxonomicName family="Metasarcidae" genus="T." kingdom="Animalia" lsidName="T. scorsesei" order="Opiliones" pageId="0" pageNumber="309" rank="species" species="scorsesei">
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<emphasis italics="true" pageId="0" pageNumber="309">T. scorsesei</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
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; Legend bars = 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="description">
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<paragraph pageId="0" pageNumber="309">Description.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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Kappa-type DSS, with carapace very wide, constriction I weakly marked and coda undefined, coalescing with mid-bulge. Ocularium low, medially depressed. Ocularium with two low tubercles. Areas of dorsal scutum moderately tuberculate. Area I undivided. Area III armed with two high spines. Posterior margin of DS armed with a pair of high tubercles. Coda short, without constriction. Coxa IV reaching area III. Coxa IV unarmed (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus, dorsal of males of Huancabamba gen. nov. and Incasarcus. A H. kubricki gen. et sp. nov.; B I. argenteus Kury & Maury, 1998; C I. dianae Kury & Maury, 1998; D I. ochoai Kury & Maury, 1998; E I. pictus Kury & Maury, 1998; F I. viracocha Kury & Maury, 1998; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure4" httpUri="https://binary.pensoft.net/fig/729878" pageId="0" pageNumber="309">4A</figureCitation>
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). Femur IV about same size as dorsal scutum length (Figs
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male trochanter-patella / tibia IV of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha, dorsal and ventral view, respectively. A - B H. kubricki gen. et sp. nov.; C - D L. antonionii gen. et sp. nov.; E - F L. woodyalleni gen. et sp. nov.; G - H T. chaplini sp. nov.; I - J T. joseochoai sp. nov.; K - L T. scorsesei sp. nov.; M - N T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure11" httpUri="https://binary.pensoft.net/fig/729885" pageId="0" pageNumber="309">11A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Live specimens of Huancabamba gen. nov. and Lumieria gen. nov. A H. kubricki gen. nov. et sp. nov., male; B female; C L. antonionii gen. nov. sp. nov., male; D female; E L. woodyalleni gen. nov. sp. nov., male; F female." figureDoi="10.3897/asp.80.e73829.figure24" httpUri="https://binary.pensoft.net/fig/729898" pageId="0" pageNumber="309">24A</figureCitation>
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). More than 13 MS C. Penis stylus thin thickness. Penis VP thin thickness (Fig.
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Penis of Huancabamba gen. nov. and Incasarcus, dorsal, lateral and ventral views, respectively. A - C H. kubricki gen. et sp. nov.; D - F I. ochoai Kury & Maury, 1998; G - I I. viracocha Kury & Maury, 1998; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure18" httpUri="https://binary.pensoft.net/fig/729892" pageId="0" pageNumber="309">18A-C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="etymology">
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<paragraph pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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The genus name, a noun in the nominative singular, from Quechua,
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<emphasis italics="true" pageId="0" pageNumber="309">huanca</emphasis>
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(stone) +
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<emphasis italics="true" pageId="0" pageNumber="309">bamba</emphasis>
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(plain). It refers to
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<taxonomicName authorityName="Benedetti & Pinto-da-Rocha" authorityYear="2022" class="Insecta" family="Metasarcidae" genus="Huancabamba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huancabamba" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Huancabamba</taxonomicName>
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depression, an interruption of the Andean Mountains, located between southern Ecuador and northern Peru. This depression constitutes a biogeographic barrier between the northern Andes and the central Andes. The name is a reference to this
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<taxonomicName authorityName="Kury" authorityYear="1994" family="Metasarcidae" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="309" rank="family">Metasarcidae</taxonomicName>
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genus occurred fartherst north. Gender: feminine.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="distribution">
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<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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(Fig.
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha in Peru. A H. kubricki gen. et sp. nov., L. antonionii gen. et sp. nov., T. chaplini sp. nov., T. scorsesei sp. nov., T. weyrauchi; B L. woodyalleni gen. et sp. nov., T. joseochoai sp. nov." figureDoi="10.3897/asp.80.e73829.figure29" httpUri="https://binary.pensoft.net/fig/729903" pageId="0" pageNumber="309">29A</figureCitation>
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) PERU.
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<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="309" type="species composition">
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<paragraph pageId="0" pageNumber="309">Species composition.</paragraph>
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<paragraph pageId="0" pageNumber="309">
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<taxonomicName class="Insecta" family="Metasarcidae" genus="Huancabamba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Huancabamba kubricki" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kubricki">
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<emphasis italics="true" pageId="0" pageNumber="309">Huancabamba kubricki</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="309">gen. et. sp. nov.</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |