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153 lines
16 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="0475C288DF1C5969996897D2534C4B40" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Rhaphaulus tonkinensis Pall-Gergely, Hunyadi & Maassen 2014" docType="treatment" docVersion="1" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661532978352" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0475C288DF1C5969996897D2534C4B40" httpUri="http://treatment.plazi.org/id/0475C288DF1C5969996897D2534C4B40" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName LSID="0475C288-DF1C-5969-9968-97D2534C4B40" authority="Pall-Gergely, Hunyadi & Maassen, 2014" authorityName="Pall-Gergely, Hunyadi & Maassen" authorityYear="2014" class="Gastropoda" domain="Unassigned" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus tonkinensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkinensis">
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Rhaphaulus tonkinensis
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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, Hunyadi & Maassen, 2014
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">Figs 13E, F</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">, 14A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pall-Gergely, Hunyadi & Maassen" authorityYear="2014" class="Gastropoda" domain="Unassigned" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus tonkinensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkinensis">Rhaphaulus tonkinensis</taxonomicName>
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2014
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</bibRefCitation>
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: 567, 569, fig. 1. Type locality: rocky wall, left side of the road nr. 6, 156 km towards Moc Chau, Ha Noi, Son La Province, Vietnam.
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<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
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: 128, fig. 7d, Son La Province, Vietnam.
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<bibRefCitation DOI="https://doi.org/10.1093/mollus/eyx006" author="Pall-Gergely, B" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="1" pagination="243 - 248" refId="B113" refString="Pall-Gergely, B, Gargominy, O, Fontaine, B, Asami, T, 2017. Breathing device of a new Streptaulus species from Vietnam extends understanding of the function and structure of respiratory tubes in cyclophoroids (Gastropoda: Caenogastropoda: Pupinidae). Journal of Molluscan Studies 83 (2): 243 - 248, DOI: https://doi.org/10.1093/mollus/eyx006" title="Breathing device of a new Streptaulus species from Vietnam extends understanding of the function and structure of respiratory tubes in cyclophoroids (Gastropoda: Caenogastropoda: Pupinidae)." url="https://doi.org/10.1093/mollus/eyx006" volume="83" year="2017">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2017
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</bibRefCitation>
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: fig. 1a-e.
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<bibRefCitation author="Raheem, DC" journalOrPublisher="The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" pageId="0" pageNumber="1" refId="B137" refString="Raheem, DC, Backeljau, T, Pearce-Kelly, P, Taylor, H, Fenn, J, Sutcharit, C, Panha, S, von, Oheimb KCM, von, Oheimb PV, Ikebe, C, Pall-Gergely, B, Gargominy, O, Hao, LV, Sang, PV, Tu, DV, Phong, DT, Naggs, M, Ablett, J, Dodds, JM, Wade, CM, Naggs, F, 2017. An Illustrated Guide to the Land Snails and Slugs of Vietnam. The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" title="An Illustrated Guide to the Land Snails and Slugs of Vietnam." year="2017">Raheem et al. 2017</bibRefCitation>
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: 6 (plate figure).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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HNHM 98757 from Ha Noi, Son La Province, Vietnam (Fig.
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">13E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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CUMZ 12163 (4 shells; Figs
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">13F</figureCitation>
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,
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">14A</figureCitation>
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) from Luang Cave, Mae Sai District, Chiang Rai Province,
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<date value="2015-10-23">23 Oct. 2015</date>
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. CUMZ 12164 (2 shells) from Pha Mee Cave, Mae Sai District, Chiang Rai Province,
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<date value="2015-10-23">23 Oct. 2015</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell elongated ovate; body whorls slightly bulging. Tube thick and flat, turning first straight upward then abruptly downward, highly widening and extending to nearly the entire last whorl height.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pall-Gergely, Hunyadi & Maassen" authorityYear="2014" class="Gastropoda" domain="Unassigned" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus tonkinensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkinensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhaphaulus tonkinensis</emphasis>
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</taxonomicName>
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can be distinguished from all other species from mainland Southeast Asia by a distinctive tube that is thick and flat, turning first straight upward then abruptly downward, greatly widening and extending to nearly the entire last whorl height.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Northern Vietnam (
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<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
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) and Chiang Rai Province, northern Thailand.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The tube of one specimen from Tham Luang, Mae Sai District, Chiang Rai Province when turning downward does not adhere to the apertural margin (Fig.
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">13F</figureCitation>
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). However, the tube of another specimen from the same locality adheres to the apertural margin (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A Rhaphaulus tonkinensis, specimen CUMZ 12163 / 2 from Luang Cave, Chiang Rai. B-D Rhaphaulus chrysalis B possible syntype NHMUK 2013.04.16 from Siam C specimen NHMUK 1871.9.23.52 from Burma, and D specimen NHMUK 1903.7.1.3073 from Molmein." figureDoi="10.3897/zookeys.1119.85400.figure14" httpUri="https://binary.pensoft.net/fig/735375" pageId="0" pageNumber="1">14A</figureCitation>
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), identical to the holotype (
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<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2014
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</bibRefCitation>
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). Thus, the extent of tube adherence to the apertural margin is treated as an infraspecific variation.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |