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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.603.8271" ID-GBIF-Dataset="07fddaf0-cd6d-460a-9078-d7c2339c64bc" ID-PMC="PMC4978003" ID-Pensoft-Pub="1313-2970-603-105" ID-PubMed="27551199" ID-ZBK="D44B76A79C0245D8B9F4996BFCB01579" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-603-105" ModsDocOrigin="ZooKeys 603" ModsDocTitle="Taxonomic revision of the Malagasy Nesomyrmexmadecassus species-group using a quantitative morphometric approach" checkinTime="1467898720993" checkinUser="pensoft" docAuthor="Csosz, Sandor &amp; Fisher, Brian L." docDate="2016" docId="A59C0AE686D5CF7B30F0EB81762673D6" docLanguage="en" docName="ZooKeys 603: 105-130" docOrigin="ZooKeys 603" docSource="http://dx.doi.org/10.3897/zookeys.603.8271" docTitle="Nesomyrmex nitidus Csosz &amp; Fisher, sp. n." docType="treatment" docUuid="F0E325CD-99C1-4CB0-817D-4D7EC060AD8F" docUuidSource="ZooBank" docVersion="5" lastPageNumber="125" masterDocId="F125FF8AFF8DFFF0FF8C31524066FFDF" masterDocTitle="Taxonomic revision of the Malagasy Nesomyrmexmadecassus species-group using a quantitative morphometric approach" masterLastPageNumber="130" masterPageNumber="105" pageNumber="120" updateTime="1668163297087" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic revision of the Malagasy Nesomyrmexmadecassus species-group using a quantitative morphometric approach</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Csosz, Sandor</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>603</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>105</mods:start>
<mods:end>130</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.603.8271</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.603.8271</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-603-105</mods:identifier>
<mods:identifier type="ZBK">D44B76A79C0245D8B9F4996BFCB01579</mods:identifier>
<mods:identifier type="ZooBank">D44B76A79C0245D8B9F4996BFCB01579</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127859328" LSID="urn:lsid:zoobank.org:act:F0E325CD-99C1-4CB0-817D-4D7EC060AD8F" httpUri="http://treatment.plazi.org/id/A59C0AE686D5CF7B30F0EB81762673D6" lastPageId="20" lastPageNumber="125" pageId="15" pageNumber="120">
<subSubSection pageId="15" pageNumber="120" type="multiple">
<paragraph pageId="15" pageNumber="120">Taxon classification Animalia Hymenoptera Formicidae</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="120" type="nomenclature">
<paragraph pageId="15" pageNumber="120">
<taxonomicName LSID="http://zoobank.org/F0E325CD-99C1-4CB0-817D-4D7EC060AD8F" authority="Csosz &amp; Fisher" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex nitidus" order="Hymenoptera" pageId="15" pageNumber="120" phylum="Arthropoda" rank="species" species="nitidus">
Nesomyrmex nitidus
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
&amp; Fisher
</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="120">sp. n.</taxonomicNameLabel>
Figs 13-15, Table 2
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="121" type="type material">
<paragraph pageId="16" pageNumber="121">
<pageBreakToken pageId="16" pageNumber="121" start="start">Type</pageBreakToken>
material investigated.
</paragraph>
<paragraph pageId="16" pageNumber="121">
Holotype: CASENT0163151, collection code: BLF24792: MADAGASCAR, Prov. Toamasina,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato river, N -16.81753,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.29498">E 49.29498</geoCoordinate>
, alt 360 m, B.L. Fisher et al. 2_25_2010, (1w, CAS);
</paragraph>
<paragraph pageId="16" pageNumber="121">
Paratypes: two workers and one gyne from the same locality under CASENT codes: CASENT0163112, collection code: BLF24794: MADAGASCAR, Prov. Toamasina,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato river, N -16.81753,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.29498">E 49.29498</geoCoordinate>
, alt 360 m, B.L. Fisher et al.2_25_2010, (1w, CAS); CASENT0162145, collection code: BLF24570: MADAGASCAR, Prov. Toamasina,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato river, N -16.7633,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.26692">E 49.26692</geoCoordinate>
, alt 520 m, B.L.Fisher et al.2_26_2010, (1w, CAS); CASENT0161445, collection code: BLF25001: MADAGASCAR, Prov. Toamasina,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato river, N -16.81209,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.29216">E 49.29216</geoCoordinate>
, alt 460 m, B.L. Fisher et al.2_22_2010, (1q, CAS);
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="124" pageId="16" pageNumber="121" type="materials_examined">
<paragraph pageId="16" pageNumber="121">Material examined.</paragraph>
<paragraph lastPageId="19" lastPageNumber="124" pageId="16" pageNumber="121">
MADAGASCAR: BLF1888(24) (collection code: BLF01888, 1w, CAS): Prov. Antsiranana, R.S. Manongarivo, 12.8 km 228° SW Antanambao, N -13.97667,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.42333">E 48.42333</geoCoordinate>
, alt 780 m, B.L. Fisher, 10_12_1998; CASENT0059931 (collection code: BLF12392, 1w, CAS): Prov. Fianarantsoa, 7.6 km 122
<normalizedToken originalValue="º">°</normalizedToken>
Kianjavato,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="Classée">Classee</normalizedToken>
Vatovavy, N -21.4,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="47.94">E 47.94</geoCoordinate>
, alt 175 m, B.L. Fisher et al., 6_6_2005; CASENT0067500 (collection code: BLF12687, 1w, CAS): Prov. Toamasina, Parc National Mananara-Nord, 7.1 km 261° Antanambe, N -16.455,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="49.7875">E 49.7875</geoCoordinate>
, alt 225 m, B.L. Fisher et al., 11_16_2005; CASENT0068002 (collection code: BLF12780, 2w, CAS): Prov. Toamasina, Res. Ambodiriana, 4.8 km 306° Manompana, along Manom
<pageBreakToken pageId="17" pageNumber="122" start="start">pana</pageBreakToken>
river, N -16.67233,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.70117">E 49.70117</geoCoordinate>
, alt 125 m, B.L.Fisher et al., 11_18_2005; CASENT0076214 (collection code: BLF09620, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
de Binara, 7.5 km 230° SW Daraina, N -13.255,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.61667">E 49.61667</geoCoordinate>
, alt 375 m, B.L. Fisher, 12_2_2003; CASENT0077523 (collection code: BLF09713, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
de Binara, 9.1km 233° SW Daraina, N -13.26333,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.60333">E 49.60333</geoCoordinate>
, alt 650-800 m, B.L. Fisher, 12_4_2003; CASENT0107046 (collection code: BLF11562, 1w, CAS), CASENT0107052 (collection code: BLF11562, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
Ambato, 26.6 km 33° Ambanja, N -13.4645,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.55167">E 48.55167</geoCoordinate>
, alt 150 m, B.L. Fisher, 12_9_2004; CASENT0107060 (collection code: BLF11610, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
Ambato, 26.6 km 33° Ambanja, N -13.4645,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.55167">E 48.55167</geoCoordinate>
, alt 150 m, B.L. Fisher, 12_10_2004; CASENT0110675 (collection code: BLF11220, 1w, CAS): Prov. Antsiranana, Ambondrobe, 41.1km 175° Vohemar, N -13.71533,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="50.10167">E 50.10167</geoCoordinate>
, alt 10 m, B.L. Fisher, 11_30_2004; CASENT0129913 (collection code: BLF15100, 1w, CAS): Prov. Toliara, Parc National Andohahela, Col de Tanatana, 33.3km NW Tolagnaro, N -24.7585,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.85367">E 46.85367</geoCoordinate>
, alt 275 m, B.L. Fisher et al., 11_22_2006; CASENT0136588 (collection code: BLF18628, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAmpombofofo">d'Ampombofofo</normalizedToken>
, N -12.09949,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.33874">E 49.33874</geoCoordinate>
, alt 25 m, B.L. Fisher et al., 11_21_2007; CASENT0151045 (collection code: BLF22399, 1w, CAS): Prov. Toamasina, Parc National de Zahamena, Sahavorondrano River, N -17.75257,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.85725">E 48.85725</geoCoordinate>
, alt 765 m, B.L. Fisher et al., 2_23_2009; CASENT0151511 (collection code: BLF23080, 1w, CAS): Prov. Mahajanga,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Beanka, 50.2 km E Maintirano, N -17.88756,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="44.47265">E 44.47265</geoCoordinate>
, alt 153 m, B.L.Fisher et al., 10_31_2009; CASENT0151914 (collection code: BLF22603, 1w, CAS, CASENT0151914): Prov. Antsiranana, Betaolana Forest, along Bekona River, N -14.52996,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.44039">E 49.44039</geoCoordinate>
, alt 880 m, B.L. Fisher et al., 3_5_2009; CASENT0152470 (collection code: BLF22141, 1w, CAS, CASENT0152470): Prov.
<pageBreakToken pageId="18" pageNumber="123" start="start">Toamasina</pageBreakToken>
, Parc National de Zahamena, Tetezambatana forest, near junction of Nosivola and Manakambahiny Rivers, N -17.74298,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.72936">E 48.72936</geoCoordinate>
, alt 860 m, B.L. Fisher et al., 2_19_2009; CASENT0155948 (collection code: BLF22797, 1w, CAS): Prov. Mahajanga,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Beanka, 50.2 km E Maintirano, N -18.02649,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="44.05051">E 44.05051</geoCoordinate>
, alt 250 m, B.L. Fisher et al., 10_19_2009; CASENT0156676 (collection code: BLF22969, 1w, CAS): Prov. Mahajanga,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="forestière">forestiere</normalizedToken>
Beanka, 53.6 km E Maintirano, N -18.04014,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="44.53394">E 44.53394</geoCoordinate>
, alt 272 m, B.L. Fisher et al., 10_25_2009; CASENT0162145 (collection code: BLF24570, 2w, CAS): Prov. Toamasina,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato river, N -16.7633,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.26692">E 49.26692</geoCoordinate>
, alt 520 m, B.L. Fisher et al., 2_22_2010; CASENT0162819 (collection code: BLF24484, 1w, CAS): Prov. Toamasina,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato river, N -16.76912,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.26704">E 49.26704</geoCoordinate>
, alt 475 m, B.L. Fisher et al., 2_21_2010; CASENT0163112 (collection code: BLF24794, 1w, CAS), CASENT0163151 (collection code: BLF24792, 1w, CAS): Prov. Toamasina,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato river, N -16.81753,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.29498">E 49.29498</geoCoordinate>
, alt 360 m, B.L. Fisher et al., 2_25_2010; CASENT0205731 (collection code: BLF25790, 1w, CAS): Prov. Toliara, Makay Mts., N -21.25864,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.16412">E 45.16412</geoCoordinate>
, alt 500 m, B.L. Fisher et al., 12_8_2010; CASENT0208609 (collection code: BLF25261, 1w, CAS): Prov. Toliara, Makay Mts., N -21.21985,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.32396">E 45.32396</geoCoordinate>
, alt 500 m, B.L. Fisher et al., 11_25_2010; CASENT0245134 (collection code: BLF26356, 1w, CAS): Prov. Antananarivo, Ankalalahana, N -19.00659,
<geoCoordinate direction="east" orientation="longitude" precision="5" value="47.1122">E 47.1122</geoCoordinate>
, alt 1375 m, B.L. Fisher et al., 3_29_2011; CASENT0261123 (collection code: BLF27634, 2w, CAS): Prov. Fianarantsoa, Andrambovato along river Tatamaly, N -21.51082,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="47.40992">E 47.40992</geoCoordinate>
, alt 1063 m, B.L. Fisher et al., 10_24_2011; CASENT0419627 (collection code: BLF04344, 1w, CAS): Prov. Mahajanga, Parc National Tsingy de Bemaraha, 2.5 km 62° ENE Bekopaka, Ankidrodroa River, N -19.13222,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="44.81467">E 44.81467</geoCoordinate>
, alt 100 m, Fisher-Griswold Arthropod Team, 11_11_2001; CASENT0419848 (collection code: BLF04434, 1w, CAS), CASENT0419849 (collection code: BLF04434, 1w, CAS): Prov. Mahajanga, Parc National Tsingy de Bemaraha, 10.6 km ESE 123° Antsalova, N -18.70944,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="44.71817">E 44.71817</geoCoordinate>
, alt 150 m, Fisher-Griswold Arthropod Team, 11_16_2001; CASENT0422571 (collection code: BLF03132, 1w, CAS): Prov. Antsiranana, Montagne des
<normalizedToken originalValue="Français">Francais</normalizedToken>
, 7.2 km 142° SE Antsiranana (=Diego Suarez), N -12.32278,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.33817">E 49.33817</geoCoordinate>
, alt 180 m, Fisher, Griswold et al., 2_22_2001; CASENT0422585 (collection code: BLF03426, 1w, CAS), CASENT0422593 (collection code: BLF03426, 1w, CAS), CASENT0422597 (collection code: BLF03426, 2w, CAS): Prov. Antsiranana, Nosy Be,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
Naturelle
<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
de Lokobe, 6.3 km 112° ESE Hellville, N -13.41933,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.33117">E 48.33117</geoCoordinate>
, alt 30 m, Fisher, Griswold et al., 3_19_2001; CASENT0422629 (collection code: BLF02859, 2w, CAS), CASENT0422651 (collection code: BLF02859, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
de
<normalizedToken originalValue="lAnkarana">l'Ankarana</normalizedToken>
, 22.9 km 224° SW Anivorano Nord, N -12.90889,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.10983">E 49.10983</geoCoordinate>
, alt 80 m, Fisher, Griswold et al., 2_10_2001; CASENT0422673 (collection code: BLF02660, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
<normalizedToken originalValue="dAmbre">d'Ambre</normalizedToken>
, 3.5 km 235° SW Sakaramy, N -12.46889,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="49.24217">E 49.24217</geoCoordinate>
, alt 325 m, Fisher, Griswold et al., 1_26_2001; CASENT0422690 (collection code: BLF03426, 1w, CAS), CASENT0422691 (collection code: BLF03426, 2w, CAS):
<pageBreakToken pageId="19" pageNumber="124" start="start">Prov</pageBreakToken>
. Antsiranana, Nosy Be,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
Naturelle
<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
de Lokobe, 6.3 km 112° ESE Hellville, N -13.41933,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="48.33117">E 48.33117</geoCoordinate>
, alt 30 m, Fisher, Griswold et al., 3_19_2001; CASENT0443282 (collection code: BLF04234, 2w, CAS), CASENT0443283 (collection code: BLF04234, 2w, CAS), CASENT0443284 (collection code: BLF04234, 2w, CAS): Prov. Mahajanga, Parc National Tsingy de Bemaraha, 3.4 km 93° E Bekopaka, Tombeau Vazimba, N -19.14194,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="44.828">E 44.828</geoCoordinate>
, alt 50 m, Fisher-Griswold Arthropod Team, 11_6_2001; CASENT0474737 (collection code: BLF06448, 1w, CAS): Prov. Mahajanga, Parc National de Namoroka, 9.8 km 300° WNW Vilanandro, N -16.46667,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="45.35">E 45.35</geoCoordinate>
, alt 140 m, Fisher, Griswold et al., 11_4_2002; CASENT0484804 (collection code: BLF07511, 2w, CAS), CASENT0484870 (collection code: BLF07511, 2w, CAS), CASENT0484900 (collection code: BLF07511, 1w, CAS): Prov. Toliara, Parc National de Zombitse, 19.8 km 84° E Sakaraha, N -22.84333,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="44.71">E 44.71</geoCoordinate>
, alt 770 m, Fisher, Griswold et al., 2_5_2003; CASENT0490345 (collection code: BLF07384, 1w, CAS), CASENT0490346 (collection code: BLF07384, 2w, CAS): Prov. Fianarantsoa,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAnalalava">d'Analalava</normalizedToken>
, 29.6 km 280° W Ranohira, N -22.59167,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.12833">E 45.12833</geoCoordinate>
, alt 700 m, Fisher, Griswold et al., 2_1_2003; CASENT0490719 (collection code: BLF07703, 2w, CAS), CASENT0491357 (collection code: BLF07762, 2w, CAS), CASENT0491554 (collection code: BLF07293, 1w, CAS): Prov. Fianarantsoa,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAtsirakambiaty">d'Atsirakambiaty</normalizedToken>
, 7.6 km 285° WNW Itremo, N -20.59333,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.56333">E 46.56333</geoCoordinate>
, alt 1550 m, Fisher, Griswold et al., 1_22_2003; CASENT0491364 (collection code: BLF07761, 2w, CAS), CASENT0492591 (collection code: BLF07652, 2w, CAS), CASENT0492611 (collection code: BLF07652, 2w, CAS), CASENT0492612 (collection code: BLF07652, 2w, CAS): Prov. Fianarantsoa, Parc National
<normalizedToken originalValue="dIsalo">d'Isalo</normalizedToken>
, Sahanafa River, 29.2 km 351° N Ranohira, N -22.31333,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="45.29167">E 45.29167</geoCoordinate>
, alt 500 m, Fisher, Griswold et al., 2_10_2003; CASENT0494326 (collection code: BLF09951, 1w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
de Bekaraoka, 6.8 km 60° ENE Daraina, N -13.16667,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="49.71">E 49.71</geoCoordinate>
, alt 150 m, B.L. Fisher, 12_8_2003; CASENT0495109 (collection code: BLF08147, 2w, CAS): Prov. Toamasina, Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
, 18.0 km 21° NNE Ambinanitelo, N -15.18833,
<geoCoordinate direction="east" orientation="longitude" precision="55" value="49.615">E 49.615</geoCoordinate>
, alt 470 m, Fisher, Griswold et al., 3_8_2003; CASENT0498718 (collection code: BLF10016, 2w, CAS), CASENT0498721 (collection code: BLF10016, 2w, CAS): Prov. Antsiranana,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAmpondrabe">d'Ampondrabe</normalizedToken>
, 26.3 km 10° NNE Daraina, N -12.97,
<geoCoordinate direction="east" orientation="longitude" precision="5555" value="49.7">E 49.7</geoCoordinate>
, alt 175 m, B.L. Fisher, 12_10_2003.
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="125" pageId="19" pageNumber="124" type="description">
<paragraph pageId="19" pageNumber="124">Description of workers.</paragraph>
<paragraph lastPageId="20" lastPageNumber="125" pageId="19" pageNumber="124">
Body color: yellow. Body color pattern: Body concolorous. Absolute cephalic size: 496 [460, 574]. Cephalic length vs. maximum width of head capsule (CL/CWb): 1.23 [1.16, 1.35]. Postocular distance vs. cephalic length (PoOc/CL): 0.48 [0.46, 0.50]. Postocular sides of cranium contour frontal view orientation: converging posteriorly. Postocular sides of cranium contour frontal view shape: convex. Vertex contour line in frontal view shape: straight; slightly concave. Vertex sculpture: main sculpture inconspuous, ground sculpture smooth. Gena contour line in frontal view shape: convex. Genae contour from anterior view orientation: converging. Gena sculpture: rugoso-reticulate with feeble areolate ground sculpture. Concentric carinae laterally surrounding antennal foramen: present. Eye length vs. absolute cephalic size (EL/CS): 0.26 [0.23, 0.27]. Frontal carina distance vs. absolute cephalic
<pageBreakToken pageId="20" pageNumber="125" start="start">size</pageBreakToken>
(FRS/CS): 0.31 [0.29, 0.33]. Longitudinal carinae on median region of frons: absent. Smooth median region on frons: present. Antennomere count: 12. Scape length vs. absolute cephalic size (SL/CS): 0.74 [0.69, 0.78]. Median clypeal notch: variable. Ground sculpture of submedian area of clypeus: smooth; present. Median carina of clypeus: variable. Lateral carinae of clypeus count: present. Metanotal depression: variable. Dorsal region of mesosoma sculpture: areolate ground sculpture, superimposed by dispersed rugae. Lateral region of pronotum sculpture: ground sculpture areolate, main sculpture absent. Mesopleuron sculpture: ground sculpture areolate, main sculpture absent. Metapleuron sculpture: ground sculpture areolate, main sculpture absent. Petiole width vs. absolute cephalic size (PEW/CS): 0.22 [0.19, 0.24]. Anterior profile of petiolar node contour line in lateral view shape: concave. Dorso-caudal petiolar profile contour line in lateral view shape: convex. Dorsal region of petiole sculpture: ground sculpture smooth, main sculpture absent. Postpetiole width vs. absolute cephalic size (PPW/CS): 0.33 [0.30, 0.36]. Dorsal region of postpetiole sculpture: ground sculpture smooth, main sculpture dispersed rugose.
</paragraph>
<caption pageId="20" pageNumber="125">
<paragraph pageId="20" pageNumber="125">
Figures 13-15.
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex nitidus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="nitidus">Nesomyrmex nitidus</taxonomicName>
sp. n. holotype worker (CASENT0163151). Head in full-face view (13), dorsal view of the body (14), lateral view of the body (15).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="20" pageNumber="125" type="diagnosis">
<paragraph pageId="20" pageNumber="125">Diagnosis.</paragraph>
<paragraph pageId="20" pageNumber="125">
Workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex nitidus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="nitidus">Nesomyrmex nitidus</taxonomicName>
cannot be confused with
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex gibber" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="gibber">Nesomyrmex gibber</taxonomicName>
because the conspicuous mesothoracic hump that is a diagnostic character of the latter species is absent in
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex nitidus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="nitidus">Nesomyrmex nitidus</taxonomicName>
workers. This species also can be easily separated from dark phenotypes of
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex madecassus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="madecassus">Nesomyrmex madecassus</taxonomicName>
based on color: the dark
<taxonomicName lsidName="madecassus" pageId="20" pageNumber="125" rank="species" species="madecassus">madecassus</taxonomicName>
phenotypes are dark brown but the workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex nitidus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="nitidus">Nesomyrmex nitidus</taxonomicName>
are light yellow. Morphometric ratio (PoOC/CW) and discriminant D4 function helps to separate
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex nitidus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="nitidus">Nesomyrmex nitidus</taxonomicName>
from ocher
<taxonomicName lsidName="madecassus" pageId="20" pageNumber="125" rank="species" species="madecassus">madecassus</taxonomicName>
phenotypes; further details are given in Diagnosis under
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex madecassus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="madecassus">Nesomyrmex madecassus</taxonomicName>
.
</paragraph>
<paragraph pageId="20" pageNumber="125">
The workers of this species are the most similar to that of
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex flavus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="flavus">Nesomyrmex flavus</taxonomicName>
. The broadly overlapping elevational distribution as well as qualitative and quantitative traits of
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex flavus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="flavus">Nesomyrmex flavus</taxonomicName>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex nitidus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="nitidus">Nesomyrmex nitidus</taxonomicName>
workers hamper easy separation. A simplified discriminant D2 function with a greatly reduced character set for safe separation is provided in the diagnosis section of
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex flavus" order="Hymenoptera" pageId="20" pageNumber="125" phylum="Arthropoda" rank="species" species="flavus">Nesomyrmex flavus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="125" type="biology_ecology">
<paragraph pageId="20" pageNumber="125">Biology and distribution.</paragraph>
<paragraph pageId="20" pageNumber="125">
This species typically occurs in
<normalizedToken originalValue="Madagascars">Madagascar's</normalizedToken>
rain forests at lower altitudes between 10 and 1550 meter, mean: 383 m (Fig. 3). This species is known to forage in low vegetation, nests can often be found in dead twigs, stems above ground or rarely in rotten logs at higher elevations.
</paragraph>
</subSubSection>
</treatment>
</document>